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<mods:title id="3B05B9F8B061026FE29ADADED5424FB4">Two new species and two new records of the genus Tenuibaetis Kang &amp; Yang, 1994 (Ephemeroptera: Baetidae) from Indian Himalaya</mods:title>
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Kubendran, Vasanth &amp; Subramanian
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(
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<figureCitation id="13437DE8FFCEFFA81E03F9D6FEC9FB58" box="[297,357,1159,1185]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="1" pageNumber="512">2A–I</figureCitation>
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<figureCitation id="13437DE8FFCEFFA81E5AF9D6FE07FB58" box="[368,427,1159,1185]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="1" pageNumber="512">3A–I</figureCitation>
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(
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<specimenCount id="9D7EAAE4FFCEFFA81AA4F9A2FF72FAC8" pageId="1" pageNumber="512" type="larva">3 larvae</specimenCount>
, same data as holotype (
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</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFCEFFAA1FEDF86BFE3BFB58" lastPageId="3" lastPageNumber="514" pageId="1" pageNumber="512" type="description">
<paragraph id="8BC7616DFFCEFFA81FEDF86BFD41FA80" blockId="1.[151,1437,1230,2013]" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDF86BFEF5FAAD" bold="true" box="[199,345,1338,1364]" pageId="1" pageNumber="512">Description.</emphasis>
Mature larva (
<figureCitation id="13437DE8FFCEFFA81ED7F86AFDD0FAAC" box="[509,636,1339,1365]" captionStart="FIGURES 1" captionStartId="2.[151,264,1813,1837]" captionTargetBox="[152,1436,381,1790]" captionTargetId="figure-105@2.[151,1436,381,1790]" captionTargetPageId="2" captionText="FIGURES 1. Larvae: A–B, Tenuibaetis himani sp. nov. A, color pattern, dorsal view; B, habitus, lateral view; C, Tenuibaetis kangi sp. nov., color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7235704" httpUri="https://zenodo.org/record/7235704/files/figure.png" pageId="1" pageNumber="512">Figs 1A–B</figureCitation>
). Body length: 
<quantity id="4C80CC88FFCEFFA81C0CF86AFC1CFAAC" box="[806,944,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.4" metricValueMin="4.2" pageId="1" pageNumber="512" unit="mm" value="4.3" valueMax="4.4" valueMin="4.2">4.2–4.4 mm</quantity>
; cerci length 2.5–3.0 mm; paracercus length 
<quantity id="4C80CC88FFCEFFA81FBDF80FFF5CFA80" box="[151,240,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="512" unit="mm" value="1.5">1.5 mm</quantity>
; antenna 1.2× as long as head capsule length.
</paragraph>
<paragraph id="8BC7616DFFCEFFA81FEDF8D3FC16F9A8" blockId="1.[151,1437,1230,2013]" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDF8D3FEE3FA65" bold="true" box="[199,335,1410,1436]" pageId="1" pageNumber="512">Coloration.</emphasis>
Head uniformly brown with a pair of medially pale white; brown with light yellow transverse band on the posterior half of mesonotum; base of clypeus pale yellow (
<figureCitation id="13437DE8FFCEFFA81C5DF8F6FC7CFA38" box="[887,976,1447,1473]" captionStart="FIGURES 1" captionStartId="2.[151,264,1813,1837]" captionTargetBox="[152,1436,381,1790]" captionTargetId="figure-105@2.[151,1436,381,1790]" captionTargetPageId="2" captionText="FIGURES 1. Larvae: A–B, Tenuibaetis himani sp. nov. A, color pattern, dorsal view; B, habitus, lateral view; C, Tenuibaetis kangi sp. nov., color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7235704" httpUri="https://zenodo.org/record/7235704/files/figure.png" pageId="1" pageNumber="512">Fig. 1A</figureCitation>
); dark or light brown tergal color pattern typical of the genus, with a pale-yellow transverse band on the posterior half of mesonotum; two paired pale-yellow maculae on abdominal terga IV–V and terga IX–X pale yellow; cerci light brown without a brown band at ca. ½ of cerci; thorax and abdomen ventrally brownish white, brighter than dorsally; legs light brown; femur dorsally pale and light brown apically, distomedially brown; claws distally brownish.
</paragraph>
<paragraph id="8BC7616DFFCEFFA81FEDFB0AFE3DF960" blockId="1.[151,1437,1230,2013]" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDFB0AFECDF98C" bold="true" box="[199,353,1627,1653]" pageId="1" pageNumber="512">Morphology.</emphasis>
<emphasis id="B90CBD7FFFCEFFA81E43FB0AFE6AF98C" box="[361,454,1627,1653]" italics="true" pageId="1" pageNumber="512">Antenna</emphasis>
(
<figureCitation id="13437DE8FFCEFFA81EF2FB0AFD9EF98C" box="[472,562,1627,1653]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="1" pageNumber="512">Fig. 3C</figureCitation>
). Scape and pedicel sub-cylindrical, flagellum with thin and simple setae on apex of each segment.
</paragraph>
<paragraph id="8BC7616DFFCEFFA81FEDFBF2FE1CF8B4" blockId="1.[151,1437,1230,2013]" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDFBF2FE8CF944" box="[199,288,1699,1725]" italics="true" pageId="1" pageNumber="512">Labrum</emphasis>
(
<figureCitation id="13437DE8FFCEFFA81E18FBF2FE3CF944" box="[306,400,1699,1725]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="1" pageNumber="512">Fig. 2A</figureCitation>
). Subrectangular with length 0.7× maximum width. Medial emargination of distal margin with a small, apically pointed process. Dorsally with many medium, fine, simple setae; submarginal arc of setae composed of 1+4 simple setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with two short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph id="8BC7616DFFCEFFA81FEDFA06FE83F840" blockId="1.[151,1437,1230,2013]" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDFA06FEDEF888" box="[199,370,1879,1905]" italics="true" pageId="1" pageNumber="512">Right mandible</emphasis>
(
<figureCitation id="13437DE8FFCEFFA81E55FA06FE56F888" box="[383,506,1879,1905]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="1" pageNumber="512">Figs 2B, C</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 3+3 denticles each. Prostheca slender, distolaterally denticulate. Margin between prostheca and mola without tubercle. Tuft of setae at apex of mola present.
</paragraph>
<paragraph id="8BC7616DFFCEFFAB1FEDFA92FD4DFEE4" blockId="1.[151,1437,1230,2013]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="513" pageId="1" pageNumber="512">
<emphasis id="B90CBD7FFFCEFFA81FEDFA92FECDF824" box="[199,353,1987,2013]" italics="true" pageId="1" pageNumber="512">Left mandible</emphasis>
(
<figureCitation id="13437DE8FFCEFFA81E5AFA92FE42F824" box="[368,494,1987,2013]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="1" pageNumber="512">Figs 2D, E</figureCitation>
). Incisors fused. Outer and inner sets of denticles with 3+4 denticles each. Prostheca robust, apically with small denticles and comb-shape structure. Margin between prostheca and mola straight. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present. Both mandibles with lateral margins almost straight; basal half with fine, simple setae scattered over dorsal surface.
</paragraph>
<paragraph id="8BC7616DFFCDFFAB1FEDFC76FBB2FE9C" blockId="2.[151,1437,151,357]" pageId="2" pageNumber="513">
<emphasis id="B90CBD7FFFCDFFAB1FEDFC76FEF3FEB8" box="[199,351,295,321]" italics="true" pageId="2" pageNumber="513">Hypopharynx</emphasis>
(
<figureCitation id="13437DE8FFCDFFAB1E44FC76FE6EFEB8" box="[366,450,295,321]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="2" pageNumber="513">Fig. 2F</figureCitation>
). Lingua approx. as long superlingua. Lingua longer than broad, distal half not expanded. Superlingua rounded, lateral margin rounded with thin setae along distal margin.
</paragraph>
<caption id="DF0731E5FFCDFFAB1FBDFA44FD90F8A8" ID-DOI="http://doi.org/10.5281/zenodo.7235704" ID-Zenodo-Dep="7235704" httpUri="https://zenodo.org/record/7235704/files/figure.png" pageId="2" pageNumber="513" startId="2.[151,264,1813,1837]" targetBox="[152,1436,381,1790]" targetPageId="2">
<paragraph id="8BC7616DFFCDFFAB1FBDFA44FD90F8A8" blockId="2.[151,1436,1813,1874]" pageId="2" pageNumber="513">
<emphasis id="B90CBD7FFFCDFFAB1FBDFA44FE8EF8D4" bold="true" box="[151,290,1813,1837]" pageId="2" pageNumber="513">FIGURES 1.</emphasis>
Larvae: A–B, 
<taxonomicName id="4C781AEEFFCDFFAB1E91FA44FDD0F8D4" authority="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha, 2022" authorityName="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha" authorityYear="2022" box="[443,636,1813,1837]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="species" species="himani" status="sp. nov.">
<emphasis id="B90CBD7FFFCDFFAB1E91FA44FDD0F8D4" box="[443,636,1813,1837]" italics="true" pageId="2" pageNumber="513">Tenuibaetis himani</emphasis>
</taxonomicName>
<emphasis id="B90CBD7FFFCDFFAB1DA9FA47FD79F8D7" bold="true" box="[643,725,1814,1838]" pageId="2" pageNumber="513">
<taxonomicNameLabel id="A23F0004FFCDFFAB1DA9FA47FD79F8D7" box="[643,725,1814,1838]" pageId="2" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, color pattern, dorsal view; B, habitus, lateral view; C, 
<taxonomicName id="4C781AEEFFCDFFAB1A02FA44FF63F8A8" authority="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha, 2022" authorityName="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha" authorityYear="2022" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="513" phylum="Arthropoda" rank="species" species="kangi" status="sp. nov.">
<emphasis id="B90CBD7FFFCDFFAB1A02FA44FF63F8A8" italics="true" pageId="2" pageNumber="513">Tenuibaetis kangi</emphasis>
</taxonomicName>
<emphasis id="B90CBD7FFFCDFFAB1FFCFA6BFE81F8AB" bold="true" box="[214,301,1850,1874]" pageId="2" pageNumber="513">
<taxonomicNameLabel id="A23F0004FFCDFFAB1FFCFA6BFE84F8AB" box="[214,296,1850,1874]" pageId="2" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
color pattern, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC7616DFFCDFFAA1FEDFAD1FA9AFEE4" blockId="2.[151,1436,1920,2018]" lastBlockId="3.[151,1437,151,1905]" lastPageId="3" lastPageNumber="514" pageId="2" pageNumber="513">
<emphasis id="B90CBD7FFFCDFFAB1FEDFAD1FEB2F863" box="[199,286,1920,1946]" italics="true" pageId="2" pageNumber="513">Maxilla</emphasis>
(
<figureCitation id="13437DE8FFCDFFAB1E04FAD1FE03F863" box="[302,431,1920,1946]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="2" pageNumber="513">Figs 2G, H</figureCitation>
). Maxilla apically with three canines and three dentisetae, lacinia with two rows of setae, one row with abundant thin setae ending with stout and long setae, second row with fine, long stout setae. Inner dorsal row of setae with three dentisetae, distal dentiseta tooth-like; middle dentiseta slender, bifid and serrate; proximal dentiseta slender, biserrate and with proximal furcation strongly developed and abducted. Medially with one feathered spine-like seta and 4–5 long, simple setae. Maxillary palp slightly longer than galea–lacinia; two segmented. Segment I equal in length to segment II. Apex of segment II rounded without nipple and without excavation at inner distolateral margin. Setae on maxillary palp fine and simple over surface of segments.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFC76FE97FD98" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFC76FEB1FEB8" box="[199,285,295,321]" italics="true" pageId="3" pageNumber="514">Labium</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1E06FC76FED0FEB8" box="[300,380,295,321]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Fig. 2I</figureCitation>
). Glossa shorter than paraglossa; inner margin with 6 spine-like setae. Most apical setae much longer than other setae; apex with one long, robust, spatulate setae and one medium, robust seta; outer margin with 4–5 spine-like setae, base with simple setae. Paraglossa subrectangular slightly curved inward; apex rounded with three rows of long robust distally pectinate setae in apical area and few simple setae in antero-medial area. Dorsally with a row of long spine-like setae near inner margin. Labial palp 3-segmented; segment I shorter than segments II and III combined. Segment II slightly produced inward to form a moderately expanded lobe at distal corner; dorsally with a row of fine medium simple setae; segment III conical, slightly asymmetrical with a small concavity at inner apex; covered with short stout spine-like setae. Segments II and III with fine simple setae scattered over dorsal surface.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFF3AFCA4FD7C" blockId="3.[151,1437,151,1905]" box="[199,776,619,645]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFF3AFED5FD7C" box="[199,377,619,645]" italics="true" pageId="3" pageNumber="514">Hind wing pads</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1EA2FF3AFE4EFD7C" box="[392,482,619,645]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3D</figureCitation>
). Present, well developed.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFFDEFB53FC78" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFFDEFEB3FD50" box="[199,287,655,681]" italics="true" pageId="3" pageNumber="514">Foreleg</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1E18FFDEFE26FD50" box="[306,394,655,681]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3F</figureCitation>
). Ratio of foreleg segments 1.1:1.0:0.5:0.2. Femur length ca. 3× maximum width. Dorsal margin with a row of ca 16–17 long, stout, apically rounded setae; length of setae 0.25× maximum width of femur. Apex rounded with a row of short, stout, pointed setae, many stouts lanceolate setae on lateral surface. Villopore well developed. Dorsal margin of tibia with a row of fine simple setae, stout setae absent; ventral margin with a row of 6–7 spine-like setae and a tuft of long, fine, simple setae; anterior surface scattered with stout, short, spatulate setae. Dorsal margin of tarsus with hair–like setae; ventral margin with 9–10 stout setae increasing in length towards apex. Claw (
<figureCitation id="13437DE8FFCCFFAA1E0EFE36FED0FC78" box="[292,380,871,897]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3G</figureCitation>
) with a row of 11 denticles; distally pointed, 6–7 stripes, subapical setae absent.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFEDAFD08FC30" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFEDAFEAAFC5D" box="[199,262,907,932]" italics="true" pageId="3" pageNumber="514">Terga</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1E3DFEDAFE3FFC5C" box="[279,403,907,933]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3A–B</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae, with stripes and tergum IX irregular spines apically.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFE82FD1AFBCC" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFE82FF51FC14" box="[199,253,979,1005]" italics="true" pageId="3" pageNumber="514">Gills</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1E24FE82FE2DFC14" box="[270,385,979,1005]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3H, I</figureCitation>
). Single lamellate gills present on segments I–VII. Margin with small denticles intercalating fine, simple setae. Abdominal gills I, II, VI and VII without visible trachea and III–V with poorly visible trachea. Tracheae limited to proximal part of main trunk.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDF96EFAA0FB84" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDF96EFE95FBA1" box="[199,313,1087,1112]" italics="true" pageId="3" pageNumber="514">Paraproct</emphasis>
(
<figureCitation id="13437DE8FFCCFFAA1E6DF96EFE30FBA0" box="[327,412,1087,1113]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3E</figureCitation>
). Distally not expanded, with 12–13 marginal stout spines. Surface scattered with micropores and fine simple setae, and with a patch of notched scales. Cercotractor with medium, marginal spines.
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDF9D6FE3BFB58" blockId="3.[151,1437,151,1905]" box="[199,407,1159,1185]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDF9D6FEB6FB58" bold="true" box="[199,282,1159,1185]" pageId="3" pageNumber="514">Imago.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFCCFFAA1FEDF9FAFD05FB10" pageId="3" pageNumber="514" type="etymology">
<paragraph id="8BC7616DFFCCFFAA1FEDF9FAFD05FB10" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDF9FAFEE2FB3C" bold="true" box="[199,334,1195,1221]" pageId="3" pageNumber="514">Etymology.</emphasis>
The new species is named after the 
<typeStatus id="54C3DFCFFFCCFFAA1DD6F9FAFC80FB3C" box="[764,812,1195,1221]" pageId="3" pageNumber="514">type</typeStatus>
locality located near snow (Himani meaning snow in Hindi), in the state of 
<collectingRegion id="49BCAF8FFFCCFFAA1EA7F99EFDF1FB10" box="[397,605,1231,1257]" country="India" name="Himachal Pradesh" pageId="3" pageNumber="514">Himachal Pradesh</collectingRegion>
, 
<collectingCountry id="F36F21FDFFCCFFAA1D4DF99EFD08FB10" box="[615,676,1231,1257]" name="India" pageId="3" pageNumber="514">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFCCFFAA1FEDF9A3FC1FFAF4" box="[199,947,1266,1293]" pageId="3" pageNumber="514" type="distribution">
<paragraph id="8BC7616DFFCCFFAA1FEDF9A3FC1FFAF4" blockId="3.[151,1437,151,1905]" box="[199,947,1266,1293]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDF9A3FECDFAF5" bold="true" box="[199,353,1266,1292]" pageId="3" pageNumber="514">Distribution.</emphasis>
North Western Himalaya (
<collectingRegion id="49BCAF8FFFCCFFAA1DA7F9A2FCF3FAF4" box="[653,863,1267,1293]" country="India" name="Himachal Pradesh" pageId="3" pageNumber="514">Himachal Pradesh</collectingRegion>
), 
<collectingCountry id="F36F21FDFFCCFFAA1C58F9A2FC03FAF4" box="[882,943,1267,1293]" name="India" pageId="3" pageNumber="514">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFCCFFAA1FEDF847FA3FF888" pageId="3" pageNumber="514" type="diagnosis">
<paragraph id="8BC7616DFFCCFFAA1FEDF847FD1FF8FC" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDF847FE34FAC8" bold="true" box="[199,408,1302,1329]" pageId="3" pageNumber="514">Diagnosis. Larva.</emphasis>
<taxonomicName id="4C781AEEFFCCFFAA1EB5F846FBB8FAC8" authority="Kubendran, Vasanth &amp; Subramanian" authorityName="Kubendran, Vasanth &amp; Subramanian" authorityYear="2022" box="[415,1044,1303,1329]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="himani" status="sp. nov.">
<emphasis id="B90CBD7FFFCCFFAA1EB5F846FDD8FAC8" box="[415,628,1303,1329]" italics="true" pageId="3" pageNumber="514">Tenuibaetis himani</emphasis>
Kubendran, Vasanth &amp; Subramanian
</taxonomicName>
<emphasis id="B90CBD7FFFCCFFAA1B31F846FBD8FAC8" bold="true" box="[1051,1140,1303,1329]" pageId="3" pageNumber="514">
<taxonomicNameLabel id="A23F0004FFCCFFAA1B31F846FBD8FAC8" box="[1051,1140,1303,1329]" pageId="3" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all known species of 
<taxonomicName id="4C781AEEFFCCFFAA1EA9F86AFDAFFAAC" authority="Kang &amp; Yang, 1994" box="[387,515,1339,1365]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B90CBD7FFFCCFFAA1EA9F86AFDAFFAAC" box="[387,515,1339,1365]" italics="true" pageId="3" pageNumber="514">Tenuibaetis</emphasis>
</taxonomicName>
by the following combination of characters: (i) dark and light brown tergal colour pattern typical of the genus with a pale yellow transverse band on the posterior half of mesonotum, two paired pale yellow maculae on abdominal terga IV–V and terga IX–X pale yellow (
<figureCitation id="13437DE8FFCCFFAA1C9FF8D2FB98FA64" box="[949,1076,1411,1437]" captionStart="FIGURES 1" captionStartId="2.[151,264,1813,1837]" captionTargetBox="[152,1436,381,1790]" captionTargetId="figure-105@2.[151,1436,381,1790]" captionTargetPageId="2" captionText="FIGURES 1. Larvae: A–B, Tenuibaetis himani sp. nov. A, color pattern, dorsal view; B, habitus, lateral view; C, Tenuibaetis kangi sp. nov., color pattern, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7235704" httpUri="https://zenodo.org/record/7235704/files/figure.png" pageId="3" pageNumber="514">Figs 1A–B</figureCitation>
); (ii) labrum dorsal submarginal arc of setae composed of one plus four simple setae (
<figureCitation id="13437DE8FFCCFFAA1C23F8F6FCCBFA38" box="[777,871,1447,1473]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Fig. 2A</figureCitation>
); (iii) right mandible: canine with 3+3 denticles, prostheca slender, apically denticulate (
<figureCitation id="13437DE8FFCCFFAA1D71F89AFD71FA1C" box="[603,733,1483,1509]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Figs 2B–C</figureCitation>
); (iv) left mandible: canine with 3+4 denticles (
<figureCitation id="13437DE8FFCCFFAA1A2DF89AFAC9FA1C" box="[1287,1381,1483,1509]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Fig. 2D</figureCitation>
); (v) maxilla: distal dentiseta slender, trifid and pectinate; middle dentiseta slender, bifid and serrate; proximal dentiseta slender, biserrate and with proximal furcation strongly developed and abducted (
<figureCitation id="13437DE8FFCCFFAA1B08FB42FBD1F9D4" box="[1058,1149,1555,1581]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Fig. 2H</figureCitation>
); (vi) labial palp segment II slightly produced inward to form a moderately expanded lobe at distal corner, segment III conical, slightly asymmetrical with a small concavity at inner apex (
<figureCitation id="13437DE8FFCCFFAA1DF1FB0AFC87F98C" box="[731,811,1627,1653]" captionStart="FIGURES 2" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[201,1392,189,1911]" captionTargetId="figure-21@4.[184,1404,181,1919]" captionTargetPageId="4" captionText="FIGURES 2. Larva: Tenuibaetis himani sp. nov., details of mouthparts: A, labrum (right part-dorsal view, left part-ventral view); B–G dorsal view; right mandible; C, right prostheca; D, left mandible; E, left prostheca; F, hypopharynx; G, maxilla; H, maxilla (arrow indicated dentiseta); I, labium (right part-dorsal, left part-ventral)." figureDoi="http://doi.org/10.5281/zenodo.7235706" httpUri="https://zenodo.org/record/7235706/files/figure.png" pageId="3" pageNumber="514">Fig. 2I</figureCitation>
); (vii) fore femur dorsal margin with 16–17 long, stout, apically rounded setae, anterior surface with a row of robust setae with median spine-like setae (
<figureCitation id="13437DE8FFCCFFAA1BEBFB2EFAB9F960" box="[1217,1301,1663,1689]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3F</figureCitation>
); (viii) claw with a row of 11 denticles, subapical seta absent (
<figureCitation id="13437DE8FFCCFFAA1DFBFBF2FC81F944" box="[721,813,1699,1725]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3G</figureCitation>
); (ix) abdominal gills I, II, VI and VII without trachea and III–V with poorly developed trachea (
<figureCitation id="13437DE8FFCCFFAA1D58FB96FD4BF918" box="[626,743,1735,1761]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Figs 3H, I</figureCitation>
); (x) paraproct distally not expanded, 12–13 marginal spines, surface with a patch of notched scales (
<figureCitation id="13437DE8FFCCFFAA1D65FBBAFD0AF8FC" box="[591,678,1771,1797]" captionStart="FIGURES 3" captionStartId="5.[151,264,1882,1906]" captionTargetBox="[162,1425,190,1847]" captionTargetId="figure-17@5.[151,1436,181,1858]" captionTargetPageId="5" captionText="FIGURES 3. Larva: Tenuibaetis himani sp. nov., dorsal view: A, posterior margin of tergum IV; B, posterior margin of tergum IX; C, antenna; D, hind wing pads; E, paraproct; F, foreleg; G, claw; H, gill IV; I, gill VII." figureDoi="http://doi.org/10.5281/zenodo.7235708" httpUri="https://zenodo.org/record/7235708/files/figure.png" pageId="3" pageNumber="514">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8BC7616DFFCCFFAA1FEDFA5FFA3FF888" blockId="3.[151,1437,151,1905]" pageId="3" pageNumber="514">
<emphasis id="B90CBD7FFFCCFFAA1FEDFA5FFE86F8D1" bold="true" box="[199,298,1806,1832]" pageId="3" pageNumber="514">Habitat.</emphasis>
<taxonomicName id="4C781AEEFFCCFFAA1E18FA5EFDABF8D0" authority="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha, 2022" authorityName="Kubendran &amp; Vasanth &amp; Subramanian &amp; Gattolliat &amp; Selvakumar &amp; Jabeen &amp; Sinha" authorityYear="2022" box="[306,519,1807,1833]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="514" phylum="Arthropoda" rank="species" species="himani" status="sp. nov.">
<emphasis id="B90CBD7FFFCCFFAA1E18FA5EFDABF8D0" box="[306,519,1807,1833]" italics="true" pageId="3" pageNumber="514">Tenuibaetis himani</emphasis>
</taxonomicName>
<emphasis id="B90CBD7FFFCCFFAA1D24FA5EFDC5F8D0" bold="true" box="[526,617,1807,1833]" pageId="3" pageNumber="514">
<taxonomicNameLabel id="A23F0004FFCCFFAA1D24FA5EFDC5F8D0" box="[526,617,1807,1833]" pageId="3" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected in Baspa River (
<figureCitation id="13437DE8FFCCFFAA1CEDFA5EFB81F8D0" box="[967,1069,1807,1833]" captionStart="FIGURE 14" captionStartId="20.[151,250,1477,1501]" captionTargetBox="[151,1436,561,1454]" captionTargetId="figure-225@20.[151,1436,561,1454]" captionTargetPageId="20" captionText="FIGURE 14. Distribution map of the genus Tenuibaetis in India." figureDoi="http://doi.org/10.5281/zenodo.7235734" httpUri="https://zenodo.org/record/7235734/files/figure.png" pageId="3" pageNumber="514">Fig. 14A</figureCitation>
) (from 
<quantity id="4C80CC88FFCCFFAA1BAEFA5FFB4DF8D0" box="[1156,1249,1806,1833]" metricMagnitude="0" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="3" pageNumber="514" unit="m" value="4.25" valueMax="5.0" valueMin="3.5">3.5–5 m</quantity>
wide and 
<quantity id="4C80CC88FFCCFFAA1A7CFA5EFF15F8B4" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="3" pageNumber="514" unit="cm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 cm</quantity>
depth), Sangla Valley. This river is characterized by medium temperature 15–18 
<superScript id="7C0DCC25FFCCFFAA1B04FA63FB99F8B9" attach="right" box="[1070,1077,1842,1856]" fontSize="6" pageId="3" pageNumber="514">°</superScript>
C and relative humidity 30–33% at the time of sampling, average current velocity (
<quantity id="4C80CC88FFCCFFAA1DEFFA07FCABF888" box="[709,775,1878,1905]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="514" unit="m" value="0.5">0.5 m</quantity>
/sec) and mainly gravel and pebbles at the bottom of river.
</paragraph>
</subSubSection>
</treatment>
</document>