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<mods:title id="B9890A579E3EA0A907E2A8229A6EEFB7">Two new species and two new records of the genus Tenuibaetis Kang &amp; Yang, 1994 (Ephemeroptera: Baetidae) from Indian Himalaya</mods:title>
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<emphasis id="B90CBD7FFFC4FFA21FBDFDC6FED3FF4B" bold="true" box="[151,383,151,178]" italics="true" pageId="11" pageNumber="522">Tenuibaetis arduus</emphasis>
(
<bibRefCitation id="EFE91C9CFFC4FFA21EA4FDC6FD28FF48" author="Kang, S. - C. &amp; Chang, H. - C. &amp; Yang, C. - T." box="[398,644,151,177]" pageId="11" pageNumber="522" pagination="9 - 44" refId="ref11047" refString="Kang, S. - C., Chang, H. - C. &amp; Yang, C. - T. (1994) A revision of the genus Baetis in Taiwan. Journal of Taiwan Museum, 47, 9 - 44." type="journal article" year="1994">Kang &amp; Yang, 1994</bibRefCitation>
)
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(
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,
<figureCitation id="13437DE8FFC4FFA21E03FDEAFEC5FF2C" box="[297,361,187,213]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">8AF</figureCitation>
,
<figureCitation id="13437DE8FFC4FFA21E5EFDEAFE17FF2C" box="[372,443,187,213]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="11" pageNumber="522">9AG</figureCitation>
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<emphasis id="B90CBD7FFFC4FFA21FBDFC51FF7AFEEE" box="[151,214,256,279]" italics="true" pageId="11" pageNumber="522">Baetis</emphasis>
(
<emphasis id="B90CBD7FFFC4FFA21FCFFC51FEF5FEE1" box="[229,345,256,280]" italics="true" pageId="11" pageNumber="522">Tenuibaetis</emphasis>
)
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in Kang
<emphasis id="B90CBD7FFFC4FFA21D21FC51FD94FEE1" box="[523,568,256,280]" italics="true" pageId="11" pageNumber="522">et al</emphasis>
n. sp. 1994: 29
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<emphasis id="B90CBD7FFFC4FFA21FBDFC4FFE99FECF" box="[151,309,286,310]" italics="true" pageId="11" pageNumber="522">Baetiella ardua</emphasis>
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Waltz &amp; McCafferty, 1997:
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<emphasis id="B90CBD7FFFC4FFA21FBDFC6CFEFAFEAC" box="[151,342,317,341]" italics="true" pageId="11" pageNumber="522">Tenuibaetis arduus</emphasis>
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Fujitani, Hirowatari &amp; Tanida, 2003a:
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<emphasis id="B90CBD7FFFC4FFA21FBDFCD2FE2CFE64" bold="true" box="[151,384,387,413]" pageId="11" pageNumber="522">Material examined.</emphasis>
<materialsCitation id="3B106B30FFC4FFA21EACFCD2FBEDFE38" ID-GBIF-Occurrence="3944667302" collectingDate="2018-03-11" collectorName="Coll. Bikramjit Sinha &amp; Reg. No. I" country="India" county="West Kameng district" latitude="27.422222" location="Dirang Valley" longLatPrecision="20" longitude="92.288055" municipality="Chakparang" pageId="11" pageNumber="522" specimenCount="1" specimenCount-larva="1" stateProvince="Arunachal Pradesh">
<specimenCount id="9D7EAAE4FFC4FFA21EACFCD2FE79FE64" box="[390,469,387,413]" pageId="11" pageNumber="522" type="larva">1 larva</specimenCount>
,
<collectingCountry id="F36F21FDFFC4FFA21EF5FCD2FD9DFE64" box="[479,561,387,413]" name="India" pageId="11" pageNumber="522">INDIA</collectingCountry>
,
<collectingRegion id="49BCAF8FFFC4FFA21D10FCD2FCBCFE64" box="[570,784,387,413]" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="522">Arunachal Pradesh</collectingRegion>
,
<collectingCounty id="62A619E1FFC4FFA21C30FCD5FBA1FE64" box="[794,1037,387,414]" pageId="11" pageNumber="522">West Kameng district</collectingCounty>
,
<collectingMunicipality id="6BA3FB17FFC4FFA21B3DFCD2FB32FE64" box="[1047,1182,387,413]" pageId="11" pageNumber="522">Chakparang</collectingMunicipality>
stream,
<location id="8EA737B6FFC4FFA21BD6FCD2FA34FE64" LSID="urn:lsid:plazi:treatment:03D1D07BFFC4FFA61FBDFDC6FA31FE70:8EA737B6FFC4FFA21BD6FCD2FA34FE64" box="[1276,1432,387,413]" country="India" county="West Kameng district" latitude="27.422222" longLatPrecision="20" longitude="92.288055" municipality="Chakparang" name="Dirang Valley" pageId="11" pageNumber="522" stateProvince="Arunachal Pradesh">Dirang Valley</location>
,
<geoCoordinate id="EE4C07AAFFC4FFA21FBDFCF8FEB3FE3D" box="[151,287,425,452]" degrees="27" direction="north" minutes="25" orientation="latitude" pageId="11" pageNumber="522" precision="15" seconds="20" value="27.422222">27°2520″N</geoCoordinate>
,
<geoCoordinate id="EE4C07AAFFC4FFA21E1AFCF8FE19FE3D" box="[304,437,425,452]" degrees="92" direction="east" minutes="17" orientation="longitude" pageId="11" pageNumber="522" precision="15" seconds="17" value="92.288055">92°1717″E</geoCoordinate>
,
<date id="FFC647ADFFC4FFA21EECFCF6FDEAFE38" box="[454,582,423,449]" pageId="11" pageNumber="522" value="2018-03-11">
<collectingDate id="EF82BE45FFC4FFA21EECFCF6FDEAFE38" box="[454,582,423,449]" pageId="11" pageNumber="522" value="2018-03-11">11.III.2018</collectingDate>
</date>
,
<collectorName id="268D04BBFFC4FFA21D7DFCF6FCF8FE38" box="[599,852,423,449]" pageId="11" pageNumber="522">Coll. Bikramjit Sinha</collectorName>
(
<collectorName id="268D04BBFFC4FFA21C42FCF6FC46FE38" box="[872,1002,423,450]" pageId="11" pageNumber="522">Reg. No. I</collectorName>
/E/331)
</materialsCitation>
;
<materialsCitation id="3B106B30FFC4FFA21B79FCF6FE84FDF3" ID-GBIF-Occurrence="3944667306" collectingDate="2018-03-07" collectorName="Coll. Bikramjit Sinha &amp; Reg. No. I" country="India" county="Tawang district" latitude="27.589724" location="Jang" longLatPrecision="20" longitude="91.98167" municipality="Nuranang falls" pageId="11" pageNumber="522" specimenCount="2" specimenCount-larva="2" stateProvince="Arunachal Pradesh">
<specimenCount id="9D7EAAE4FFC4FFA21B79FCF6FB19FE38" box="[1107,1205,423,449]" pageId="11" pageNumber="522" type="larva">2 larvae</specimenCount>
,
<collectingCountry id="F36F21FDFFC4FFA21BECFCF6FAB4FE38" box="[1222,1304,423,449]" name="India" pageId="11" pageNumber="522">INDIA</collectingCountry>
,
<collectingRegion id="49BCAF8FFFC4FFA21A02FCF6FF5FFE1C" country="India" name="Arunachal Pradesh" pageId="11" pageNumber="522">Arunachal Pradesh</collectingRegion>
,
<collectingCounty id="62A619E1FFC4FFA21FD4FC9DFE02FE1C" box="[254,430,459,486]" pageId="11" pageNumber="522">Tawang district</collectingCounty>
,
<collectingMunicipality id="6BA3FB17FFC4FFA21E93FC9DFDCEFE1C" box="[441,610,459,486]" pageId="11" pageNumber="522">Nuranang falls</collectingMunicipality>
,
<location id="8EA737B6FFC4FFA21D47FC9DFD0FFE1F" LSID="urn:lsid:plazi:treatment:03D1D07BFFC4FFA61FBDFDC6FA31FE70:8EA737B6FFC4FFA21D47FC9DFD0FFE1F" box="[621,675,460,486]" country="India" county="Tawang district" latitude="27.589724" longLatPrecision="20" longitude="91.98167" municipality="Nuranang falls" name="Jang" pageId="11" pageNumber="522" stateProvince="Arunachal Pradesh">Jang</location>
,
<geoCoordinate id="EE4C07AAFFC4FFA21D85FC9CFC92FE1F" box="[687,830,460,488]" degrees="27" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="522" precision="15" seconds="23" value="27.589724">27°3523″ N</geoCoordinate>
,
<geoCoordinate id="EE4C07AAFFC4FFA21C60FC9CFC63FE11" box="[842,975,461,488]" degrees="91" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="522" precision="15" seconds="54" value="91.98167">91°5854″E</geoCoordinate>
,
<date id="FFC647ADFFC4FFA21CF0FC9AFBF6FE1C" box="[986,1114,459,485]" pageId="11" pageNumber="522" value="2018-03-07">
<collectingDate id="EF82BE45FFC4FFA21CF0FC9AFBF6FE1C" box="[986,1114,459,485]" pageId="11" pageNumber="522" value="2018-03-07">07.III.2018</collectingDate>
</date>
,
<collectorName id="268D04BBFFC4FFA21B4DFC9AFAF4FE1C" box="[1127,1368,459,485]" pageId="11" pageNumber="522">Coll. Bikramjit Sinha</collectorName>
(
<collectorName id="268D04BBFFC4FFA21A4DFC9AFF7CFDF3" pageId="11" pageNumber="522">Reg. No. I</collectorName>
/E/539)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFC4FFA51FEDFF42FA31FDC4" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522" type="description">
<paragraph id="8BC7616DFFC4FFA21FEDFF42FCB2FDA8" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFF42FEF5FDD4" bold="true" box="[199,345,531,557]" pageId="11" pageNumber="522">Description.</emphasis>
Mature larva (
<figureCitation id="13437DE8FFC4FFA21ED4FF42FDD2FDD4" box="[510,638,531,557]" captionStart="FIGURES 7" captionStartId="12.[151,264,1974,1998]" captionTargetBox="[151,1436,597,1951]" captionTargetId="figure-216@12.[151,1436,597,1951]" captionTargetPageId="12" captionText="FIGURES 7. Larva: Tenuibaetis arduus, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view; C, color pattern, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="11" pageNumber="522">Figs 7AC</figureCitation>
). Body length
<quantity id="4C80CC88FFC4FFA21C08FF42FC00FDD7" box="[802,940,531,558]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="5.6" metricValueMin="5.3" pageId="11" pageNumber="522" unit="mm" value="5.449999999999999" valueMax="5.6" valueMin="5.3">5.35.6 mm</quantity>
; cerci length
<quantity id="4C80CC88FFC4FFA21B68FF42FB60FDD7" box="[1090,1228,531,558]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="11" pageNumber="522" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
; paracercus length
<quantity id="4C80CC88FFC4FFA21FBDFF66FE8DFDAB" box="[151,289,567,594]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="11" pageNumber="522" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
; antenna 1.1× as long as head capsule length.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDFF0AFEFAFCD0" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFF0AFEE4FD8C" bold="true" box="[199,328,603,629]" pageId="11" pageNumber="522">Coloration</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E7CFF0AFE79FD8C" box="[342,469,603,629]" captionStart="FIGURES 7" captionStartId="12.[151,264,1974,1998]" captionTargetBox="[151,1436,597,1951]" captionTargetId="figure-216@12.[151,1436,597,1951]" captionTargetPageId="12" captionText="FIGURES 7. Larva: Tenuibaetis arduus, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view; C, color pattern, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="11" pageNumber="522">Figs 7AC</figureCitation>
). Head dark brown medially pale with a pair of dark spots; base of clypeus pale yellow. Thorax brown with bright median, dorsal suture, mesothorax medially with bright striation and abdomen dorsally dark brown with two dark spots in medial area, abdominal segment IX with U-shaped pale white striation, segment X brownish white. Head, thorax and abdomen ventrally brown, somewhat brighter than dorsally. Legs light brown; femur dorsally pale and apical margin dark brown, distomedially dark brown; claws distally dark brown; caudal filaments brown.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDFE62FDE2FC8B" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFE62FECDFCB4" bold="true" box="[199,353,819,845]" pageId="11" pageNumber="522">Morphology.</emphasis>
<emphasis id="B90CBD7FFFC4FFA21E4CFE62FE6FFCB4" box="[358,451,819,845]" italics="true" pageId="11" pageNumber="522">Antenna</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21EF8FE62FD86FCB4" box="[466,554,819,845]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8A</figureCitation>
). Scape and pedicel sub-cylindrical, flagellum with rounded spines apically and fine, simple setae on apex of segments.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDFE2AFE1CFBDC" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFE2AFE8CFC6C" box="[199,288,891,917]" italics="true" pageId="11" pageNumber="522">Labrum</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E18FE2AFE21FC6C" box="[306,397,891,917]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8B</figureCitation>
). Subrectangular with length 0.7× maximum width. Medial emargination of distal margin with a small, apically pointed process. Dorsally with many medium, fine, simple setae; submarginal arc of setae composed of 1+3 simple setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid, pectinate setae; ventral surface with two short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDF97EFE4FFB6B" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDF97EFEDAFBB0" box="[199,374,1071,1097]" italics="true" pageId="11" pageNumber="522">Right mandible</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21EA0F97EFE4BFBB0" box="[394,487,1071,1097]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8D</figureCitation>
). Incisors fused. Outer and inner set of denticles with four denticles respectively. Prostheca stick-like, apically and distolaterally denticulate. Margin between prostheca and mola straight. Tuft of setae at apex of mola present.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDF9CAFBA5FABC" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDF9CAFECCFB4C" box="[199,352,1179,1205]" italics="true" pageId="11" pageNumber="522">Left mandible</emphasis>
. Incisors fused. Outer and inner set of denticles with three denticles respectively, plus one-minute, intermediate denticle. Prostheca robust, apically with small denticles and combshape structure. Margin straight between prostheca and mola. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically slender. Apex of mola tuft of setae present. Both mandibles with lateral margins almost straight; basal half with fine, simple setae scattered over dorsal surface.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDF81EFBFCFA48" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDF81EFEF3FA90" box="[199,351,1359,1385]" italics="true" pageId="11" pageNumber="522">Hypopharynx</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E45F81EFE6BFA90" box="[367,455,1359,1385]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8C</figureCitation>
). Lingua about as long as superlingua; lingua longer than wide; apically triangular with medial tuft of stout setae; distal half not expanded. Superlingua with distal margin straight; lateral margins rounded; fine, long, simple setae along distal margin; short, pointed setae along lateral margin.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDF8EAFA89F99F" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDF8EAFEB2FA2C" box="[199,286,1467,1493]" italics="true" pageId="11" pageNumber="522">Maxilla</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E04F8EAFE29FA2C" box="[302,389,1467,1493]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8E</figureCitation>
). Galea-lacinia with one simple, robust apical seta under the crown. Inner dorsal row of setae with three dentisetae, distal dentiseta, tooth-like; middle dentiseta biserrate and proximal dentiseta, furcation strongly developed and abducted. Medially with one feathered spine-like seta and 45 long, simple setae. Maxillary palp 1.5× as long as length of galea-lacinia; two segmented. Segment I equal in length to segment II. Apex of segment II rounded without nipple. Setae on maxillary palp fine and simple setae over surface of segments I and II.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDFB3EFA89F87C" blockId="11.[151,1437,387,2034]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFB3EFEB1F970" box="[199,285,1647,1673]" italics="true" pageId="11" pageNumber="522">Labium</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E07FB3EFE28F970" box="[301,388,1647,1673]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="11" pageNumber="522">Fig. 8F</figureCitation>
). Base of glossa broad, towards narrowing apex; paraglossa shorter; inner margin with eight spine-like setae increasing the length of distally; apex with three long and one medium, robust, pectinate setae; outer margin with five long, spine-like setae; ventral surface with fine, short setae. Paraglossa subrectangular, slightly curved inward; apex rounded, with three rows of long, simple, robust, pectinate setae apically; dorsally with three medium, simple setae; ventrally long setae near inner margin. Labial palp 3 segmented; segment I 0.7× of length of segments II and III combined. Segment II slightly produced inward to form a moderately expanded lobe at distal corner, dorsally with a row of 56 long, simple setae; Segment III conical, slightly pointed apically; covered with short, spine-like simple setae. Segments I, II and III with fine simple setae scattered over dorsal surface.
</paragraph>
<paragraph id="8BC7616DFFC4FFA21FEDFADEFD35F850" blockId="11.[151,1437,387,2034]" box="[199,665,1935,1962]" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFADEFE2CF850" box="[199,384,1935,1961]" italics="true" pageId="11" pageNumber="522">Hind wing pads.</emphasis>
Present, well developed.
</paragraph>
<paragraph id="8BC7616DFFC4FFA51FEDFAE2FD40FE9C" blockId="11.[151,1437,387,2034]" lastBlockId="12.[151,1437,151,573]" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522">
<emphasis id="B90CBD7FFFC4FFA21FEDFAE2FEB3F834" box="[199,287,1971,1997]" italics="true" pageId="11" pageNumber="522">Foreleg</emphasis>
(
<figureCitation id="13437DE8FFC4FFA21E05FAE2FE25F834" box="[303,393,1971,1997]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="11" pageNumber="522">Fig. 9A</figureCitation>
). Ratio of foreleg segments 1.4:1.0:0.8:0.2.
<emphasis id="B90CBD7FFFC4FFA21C50FAE5FC66F834" box="[890,970,1972,1997]" italics="true" pageId="11" pageNumber="522">Femur.</emphasis>
Length ca. 2.5× maximum width. Dorsal margin with a row of ca 2122 long, curved, apically rounded setae; length of setae 0.22× maximum width of femur. Apex rounded with a row of short, stout, pointed setae, many stout lanceolate setae on lateral surface. Villopore well developed. Dorsal margin of tibia with a row of fine simple setae; ventral margin with a row of 69 spine-like setae and a tuft of long, fine, simple setae; anterior surface scattered with stout, spatulate setae.
<emphasis id="B90CBD7FFFC3FFA51BB1FD8EFB70FF00" box="[1179,1244,223,249]" italics="true" pageId="12" pageNumber="523">Tibia.</emphasis>
Anterior surface with scattered stout setae, lanceolate. Tibio-patellar suture present on basal 2/3 area.
<emphasis id="B90CBD7FFFC3FFA51B7FFC52FB0AFEE5" box="[1109,1190,259,284]" italics="true" pageId="12" pageNumber="523">Tarsus.</emphasis>
Dorsal margin with a row of short, stout setae. Inner margin with a row of spine-like setae. Claw distally pointed and curved (
<figureCitation id="13437DE8FFC3FFA51A16FC76FA38FEB8" box="[1340,1428,295,321]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="12" pageNumber="523">Fig. 9C</figureCitation>
) with one row of 11 denticles and 56 stripes apically.
</paragraph>
<paragraph id="8BC7616DFFC3FFA51FEDFC3EFB2BFE54" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="523">
<emphasis id="B90CBD7FFFC3FFA51FEDFC3EFEAAFE71" box="[199,262,367,392]" italics="true" pageId="12" pageNumber="523">Terga</emphasis>
(
<figureCitation id="13437DE8FFC3FFA51E3CFC3EFEDCFE70" box="[278,368,367,393]" captionStart="FIGURES 7" captionStartId="12.[151,264,1974,1998]" captionTargetBox="[151,1436,597,1951]" captionTargetId="figure-216@12.[151,1436,597,1951]" captionTargetPageId="12" captionText="FIGURES 7. Larva: Tenuibaetis arduus, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view; C, color pattern, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="12" pageNumber="523">Fig. 7A</figureCitation>
). Surface with scattered V-shaped scale bases and scattered fine, simple setae. Posterior margin of each segment a pair of small spots; tergum IX pale white with row of V- shaped spines.
</paragraph>
<paragraph id="8BC7616DFFC3FFA51FEDFCE6FD9FFE0C" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="523">
<emphasis id="B90CBD7FFFC3FFA51FEDFCE6FF51FE28" box="[199,253,439,465]" italics="true" pageId="12" pageNumber="523">Gills</emphasis>
(
<figureCitation id="13437DE8FFC3FFA51E27FCE6FE27FE28" box="[269,395,439,465]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="12" pageNumber="523">Figs 9DF</figureCitation>
). Single lamellate gills present on segments IVII. Margin with small denticles intercalating fine, simple setae. Tracheae obscure.
</paragraph>
<paragraph id="8BC7616DFFC3FFA51FEDFCAEFA31FDC4" blockId="12.[151,1437,151,573]" pageId="12" pageNumber="523">
<emphasis id="B90CBD7FFFC3FFA51FEDFCAEFE95FDE1" box="[199,313,511,536]" italics="true" pageId="12" pageNumber="523">Paraproct</emphasis>
(
<figureCitation id="13437DE8FFC3FFA51E62FCAEFE0EFDE0" box="[328,418,511,537]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="12" pageNumber="523">Fig. 9G</figureCitation>
). Distally not expanded, with 1314 marginal stout spines. Surface scattered with V-shaped scale bases, micropores fine, simple setae, with a patch of notched scales. Cercotractor with medium, marginal spines.
</paragraph>
</subSubSection>
<caption id="DF0731E5FFC3FFA51FBDFAE7FDAAF80B" ID-DOI="http://doi.org/10.5281/zenodo.7235716" ID-Zenodo-Dep="7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="12" pageNumber="523" startId="12.[151,264,1974,1998]" targetBox="[151,1436,597,1951]" targetPageId="12">
<paragraph id="8BC7616DFFC3FFA51FBDFAE7FDAAF80B" blockId="12.[151,1436,1974,2034]" pageId="12" pageNumber="523">
<emphasis id="B90CBD7FFFC3FFA51FBDFAE7FE8FF837" bold="true" box="[151,291,1974,1998]" pageId="12" pageNumber="523">FIGURES 7.</emphasis>
Larva:
<taxonomicName id="4C781AEEFFC3FFA51E5FFAE7FD9AF837" authorityName="Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" baseAuthorityName="Fujitani, Hirowatari &amp; Tanida" baseAuthorityYear="1994" box="[373,566,1974,1998]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="523" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B90CBD7FFFC3FFA51E5FFAE7FD9AF837" box="[373,566,1974,1998]" italics="true" pageId="12" pageNumber="523">Tenuibaetis arduus</emphasis>
</taxonomicName>
, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view;
<emphasis id="B90CBD7FFFC3FFA51FFCFA8BFF43F80B" bold="true" box="[214,239,2010,2034]" pageId="12" pageNumber="523">C,</emphasis>
color pattern, ventral view.
</paragraph>
</caption>
<caption id="DF0731E5FFC2FFA41FBDFAC6FD3CF82D" ID-DOI="http://doi.org/10.5281/zenodo.7235718" ID-Zenodo-Dep="7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="13" pageNumber="524" startId="13.[151,264,1943,1967]" targetBox="[195,1393,181,1917]" targetPageId="13">
<paragraph id="8BC7616DFFC2FFA41FBDFAC6FD3CF82D" blockId="13.[151,1436,1943,2004]" pageId="13" pageNumber="524">
<emphasis id="B90CBD7FFFC2FFA41FBDFAC6FE88F856" bold="true" box="[151,292,1943,1967]" pageId="13" pageNumber="524">FIGURES 8.</emphasis>
Larva:
<taxonomicName id="4C781AEEFFC2FFA41E53FAC9FD97F849" authorityName="Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" baseAuthorityName="Fujitani, Hirowatari &amp; Tanida" baseAuthorityYear="1994" box="[377,571,1944,1968]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="524" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B90CBD7FFFC2FFA41E53FAC9FD97F849" box="[377,571,1944,1968]" italics="true" pageId="13" pageNumber="524">Tenuibaetis arduus</emphasis>
</taxonomicName>
, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium.
</paragraph>
</caption>
<caption id="DF0731E5FFC1FFA71FBDFAC7FF52F82B" ID-DOI="http://doi.org/10.5281/zenodo.7235720" ID-Zenodo-Dep="7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="14" pageNumber="525" startId="14.[151,264,1942,1966]" targetBox="[154,1437,181,1917]" targetPageId="14">
<paragraph id="8BC7616DFFC1FFA71FBDFAC7FF52F82B" blockId="14.[151,1436,1942,2002]" pageId="14" pageNumber="525">
<emphasis id="B90CBD7FFFC1FFA71FBDFAC7FE88F857" bold="true" box="[151,292,1942,1966]" pageId="14" pageNumber="525">FIGURES 9.</emphasis>
Larva:
<taxonomicName id="4C781AEEFFC1FFA71E52FAC7FD97F857" authorityName="Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" baseAuthorityName="Fujitani, Hirowatari &amp; Tanida" baseAuthorityYear="1994" box="[376,571,1942,1966]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="525" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B90CBD7FFFC1FFA71E52FAC7FD97F857" box="[376,571,1942,1966]" italics="true" pageId="14" pageNumber="525">Tenuibaetis arduus</emphasis>
</taxonomicName>
, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct.
</paragraph>
</caption>
<subSubSection id="C36232E6FFC0FFA61FEDFDC7FD02FEE4" pageId="15" pageNumber="526" type="diagnosis">
<paragraph id="8BC7616DFFC0FFA61FEDFDC7FD02FEE4" blockId="15.[151,1437,150,393]" pageId="15" pageNumber="526">
<emphasis id="B90CBD7FFFC0FFA61FEDFDC7FEE8FF49" bold="true" box="[199,324,150,176]" pageId="15" pageNumber="526">Diagnosis:</emphasis>
<taxonomicName id="4C781AEEFFC0FFA61E64FDC6FD89FF48" authorityName="Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" baseAuthorityName="Fujitani, Hirowatari &amp; Tanida" baseAuthorityYear="1994" box="[334,549,151,177]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="526" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B90CBD7FFFC0FFA61E64FDC6FD89FF48" box="[334,549,151,177]" italics="true" pageId="15" pageNumber="526">Tenuibaetis arduus</emphasis>
</taxonomicName>
can be differentiated by the following combination of characters: (i) labrum deeply cleft without “V” shaped marking medially (
<figureCitation id="13437DE8FFC0FFA61DCCFDEAFC93FF2C" box="[742,831,187,213]" captionStart="FIGURES 8" captionStartId="13.[151,264,1943,1967]" captionTargetBox="[195,1393,181,1917]" captionTargetId="figure-17@13.[195,1393,181,1920]" captionTargetPageId="13" captionText="FIGURES 8. Larva: Tenuibaetis arduus, A, head, frontal view; dorsal view: B, labrum; C, hypopharynx; D, left mandible (arrow indicated denticles); E, maxilla; F, labium." figureDoi="http://doi.org/10.5281/zenodo.7235718" httpUri="https://zenodo.org/record/7235718/files/figure.png" pageId="15" pageNumber="526">Fig. 8B</figureCitation>
); (ii) abdominal posterior marginal spines blunt (
<figureCitation id="13437DE8FFC0FFA61A45FDEAFF11FF00" captionStart="FIGURES 7" captionStartId="12.[151,264,1974,1998]" captionTargetBox="[151,1436,597,1951]" captionTargetId="figure-216@12.[151,1436,597,1951]" captionTargetPageId="12" captionText="FIGURES 7. Larva: Tenuibaetis arduus, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view; C, color pattern, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="15" pageNumber="526">Fig. 7A</figureCitation>
); (iii) color pattern of thorax and abdomen distinct (
<figureCitation id="13437DE8FFC0FFA61C20FD8EFC27FF00" box="[778,907,223,249]" captionStart="FIGURES 7" captionStartId="12.[151,264,1974,1998]" captionTargetBox="[151,1436,597,1951]" captionTargetId="figure-216@12.[151,1436,597,1951]" captionTargetPageId="12" captionText="FIGURES 7. Larva: Tenuibaetis arduus, A, color pattern, dorsal view (arrow indicated U shaped spines); B, habitus, lateral view; C, color pattern, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7235716" httpUri="https://zenodo.org/record/7235716/files/figure.png" pageId="15" pageNumber="526">Figs 7AC</figureCitation>
); (iv) tracheation of gills obscure (
<figureCitation id="13437DE8FFC0FFA61A38FD8EFA23FF00" box="[1298,1423,223,249]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="15" pageNumber="526">Figs 9DF</figureCitation>
); (v) claw with one row of 11 denticles (
<figureCitation id="13437DE8FFC0FFA61D63FC52FD0DFEE4" box="[585,673,259,285]" captionStart="FIGURES 9" captionStartId="14.[151,264,1942,1966]" captionTargetBox="[154,1437,181,1917]" captionTargetId="figure-21@14.[151,1437,181,1918]" captionTargetPageId="14" captionText="FIGURES 9. Larva: Tenuibaetis arduus, dorsal view: A, foreleg; B, hind leg; C, claw; D, gill I; E, gill IV; F, gill VII; G, paraproct." figureDoi="http://doi.org/10.5281/zenodo.7235720" httpUri="https://zenodo.org/record/7235720/files/figure.png" pageId="15" pageNumber="526">Fig. 9C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFC0FFA61FEDFC77FC8DFEB8" box="[199,801,294,321]" pageId="15" pageNumber="526" type="distribution">
<paragraph id="8BC7616DFFC0FFA61FEDFC77FC8DFEB8" blockId="15.[151,1437,150,393]" box="[199,801,294,321]" pageId="15" pageNumber="526">
<emphasis id="B90CBD7FFFC0FFA61FEDFC77FECFFEB9" bold="true" box="[199,355,294,320]" pageId="15" pageNumber="526">Distribution:</emphasis>
<collectingCountry id="F36F21FDFFC0FFA61E43FC76FE17FEB8" box="[361,443,295,321]" name="Taiwan" pageId="15" pageNumber="526">Taiwan</collectingCountry>
and
<collectingCountry id="F36F21FDFFC0FFA61ED8FC76FD80FEB8" box="[498,556,295,321]" name="India" pageId="15" pageNumber="526">India</collectingCountry>
(
<collectingRegion id="49BCAF8FFFC0FFA61D10FC76FCB9FEB8" box="[570,789,295,321]" country="India" name="Arunachal Pradesh" pageId="15" pageNumber="526">Arunachal Pradesh</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36232E6FFC0FFA61FEDFC1AFA31FE70" pageId="15" pageNumber="526" type="discussion">
<paragraph id="8BC7616DFFC0FFA61FEDFC1AFA31FE70" blockId="15.[151,1437,150,393]" pageId="15" pageNumber="526">
<emphasis id="B90CBD7FFFC0FFA61FEDFC1AFE91FE9C" bold="true" box="[199,317,331,357]" pageId="15" pageNumber="526">Remarks:</emphasis>
This species was originally described from
<collectingCountry id="F36F21FDFFC0FFA61C34FC1AFCDCFE9C" box="[798,880,331,357]" name="Taiwan" pageId="15" pageNumber="526">Taiwan</collectingCountry>
based on larvae (
<bibRefCitation id="EFE91C9CFFC0FFA61B06FC1AFAA8FE9C" author="Kang, S. - C. &amp; Chang, H. - C. &amp; Yang, C. - T." box="[1068,1284,331,357]" pageId="15" pageNumber="526" pagination="9 - 44" refId="ref11047" refString="Kang, S. - C., Chang, H. - C. &amp; Yang, C. - T. (1994) A revision of the genus Baetis in Taiwan. Journal of Taiwan Museum, 47, 9 - 44." type="journal article" year="1994">Kang &amp;Yang 1994</bibRefCitation>
). Present new record of the larvae
<taxonomicName id="4C781AEEFFC0FFA61E53FC3EFDE2FE70" authorityName="Fujitani, Hirowatari &amp; Tanida" authorityYear="2003" baseAuthorityName="Fujitani, Hirowatari &amp; Tanida" baseAuthorityYear="1994" box="[377,590,367,393]" class="Insecta" family="Baetidae" genus="Tenuibaetis" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="526" phylum="Arthropoda" rank="species" species="arduus">
<emphasis id="B90CBD7FFFC0FFA61E53FC3EFDE2FE70" box="[377,590,367,393]" italics="true" pageId="15" pageNumber="526">Tenuibaetis arduus</emphasis>
</taxonomicName>
from Arunachal Pradesh is an extension distributional range of the species.
</paragraph>
</subSubSection>
</treatment>
</document>