259 lines
39 KiB
XML
259 lines
39 KiB
XML
<document id="960486D02E3C883091D4B21CF3CB1088" ID-CLB-Dataset="7531" ID-DOI="10.5852/ejt.2021.742.1291" ID-GBIF-Dataset="62f9dc57-bade-4fbc-8af7-f92ff107593c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4655404" ID-ZooBank="23424EC3-8440-4C72-AD9E-C1EC3087F7CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617262825393" checkinUser="plazi" docAuthor="De Oliveira, Luiz Fernando M. & Brescovit, Antonio Domingos" docDate="2021" docId="03CF87ADFF86661EFD17947DFE32FB96" docLanguage="en" docName="ejt-742_1-77.pdf" docOrigin="European Journal of Taxonomy 742" docSource="http://dx.doi.org/10.5852/ejt.2021.742.1291" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tafana arawak De Oliveira & Brescovit 2021, sp. nov." docType="treatment" docUuid="7BF3C0A5-9472-46CF-80F3-AEEDFB9EA022" docUuidSource="ZooBank" docVersion="8" lastPageNumber="49" masterDocId="FFF6FFD5FFA8662EFF99965EFF99FFCF" masterDocTitle="Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="47" updateTime="1698935806814" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DEE78A6F870AF84A4BD457671AF98633">Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species</mods:title>
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<mods:roleTerm id="42582FDC91FDD14162B13770DC9DB579">Author</mods:roleTerm>
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<mods:namePart id="A424435641BBD55E8BD401D19FAE387A">De Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:affiliation id="3F1E8F66B856ADB737E2426116C92D74">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. & Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="1D5EF3D89A5D01D788F402D379BB2AEC" type="email">luiz.moura.lfm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9B33F2C77AFC57BA5D7A2CDCE0418AD6">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="9E490D9AF227CDA7C5A4583DBB0CEE4D">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="3E2F78554BAC8590B462760ED2C8B3B3" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:title id="027B6388E5CCD14750F382DF32C22F90">European Journal of Taxonomy</mods:title>
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<mods:part id="AEA55BE63E52DE514A9073384EA2F846">
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<mods:date id="96D6D6C2ABBC07649B640F24F7D21906">2021</mods:date>
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<mods:number id="F4299FE54A6011241F075CB86AFDFCEF">2021-03-31</mods:number>
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<treatment id="03CF87ADFF86661EFD17947DFE32FB96" ID-DOI="http://doi.org/10.5281/zenodo.4658767" ID-GBIF-Taxon="180784550" ID-Zenodo-Dep="4658767" LSID="urn:lsid:zoobank.org:act:7BF3C0A5-9472-46CF-80F3-AEEDFB9EA022" httpUri="http://treatment.plazi.org/id/03CF87ADFF86661EFD17947DFE32FB96" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
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<subSubSection id="C37C6530FF866600FD17947DFC3FFDF1" box="[654,934,547,574]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8BD936BBFF866600FD17947DFC3FFDF1" blockId="46.[362,1225,547,645]" box="[654,934,547,574]" pageId="46" pageNumber="47">
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<heading id="D09181D7FF866600FD17947DFC3FFDF1" box="[654,934,547,574]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
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<taxonomicName id="4C664D38FF866600FD17947DFCDDFDF1" ID-CoL="98Z2X" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[654,836,547,574]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="arawak" status="sp. nov.">
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<emphasis id="B912EAA9FF866600FD17947DFCDDFDF1" bold="true" box="[654,836,547,574]" italics="true" pageId="46" pageNumber="47">Tafana arawak</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FF866600FCD2947AFC3FFDF1" box="[843,934,548,574]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6530FF866600FEF39419FCE3FD4A" pageId="46" pageNumber="47" type="description">
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<paragraph id="8BD936BBFF866600FEF39419FB50FDAD" blockId="46.[362,1225,547,645]" box="[362,1225,583,610]" pageId="46" pageNumber="47">
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<uri id="FFF73AB9FF866600FEF39419FB50FDAD" box="[362,1225,583,610]" pageId="46" pageNumber="47">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC00C6EFF866600FD119419FB50FDAD" box="[648,1225,583,610]" pageId="46" pageNumber="47">7BF3C0A5-9472-46CF-80F3-AEEDFB9EA022</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BD936BBFF866600FD209434FCE3FD4A" blockId="46.[362,1225,547,645]" box="[697,890,618,645]" pageId="46" pageNumber="47">
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<figureCitation id="135D2A3EFF866600FD209434FC8AFD4B" box="[697,787,618,644]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="46" pageNumber="47">
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Figs
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<geoCoordinate id="EE52507CFF866600FD6A9434FC8AFD4B" box="[755,787,618,644]" degrees="27, 33" direction="east" orientation="longitude" pageId="46" pageNumber="47" precision="555" value="27.33">27</geoCoordinate>
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</figureCitation>
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,
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<figureCitation id="135D2A3EFF866600FC869434FCC8FD4B" box="[799,849,618,644]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: A–H =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="46" pageNumber="47">33E</figureCitation>
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,
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<figureCitation id="135D2A3EFF866600FCC49434FCE3FD4A" box="[861,890,618,645]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="46" pageNumber="47">35</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6530FF866600FF2494F2FC7AFC90" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="8BD936BBFF866600FF2494F2FEAFFD09" blockId="46.[189,310,684,710]" box="[189,310,684,710]" pageId="46" pageNumber="47">
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||
<heading id="D09181D7FF866600FF2494F2FEAFFD09" bold="true" box="[189,310,684,710]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="B912EAA9FF866600FF2494F2FEAFFD09" bold="true" box="[189,310,684,710]" pageId="46" pageNumber="47">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD936BBFF866600FF249485FC7AFC90" blockId="46.[188,1399,731,863]" pageId="46" pageNumber="47">
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Males of
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<taxonomicName id="4C664D38FF866600FEB19485FE4FFD3A" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[296,470,731,757]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="arawak" status="sp. nov.">
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<emphasis id="B912EAA9FF866600FEB19485FE4FFD3A" box="[296,470,731,757]" italics="true" pageId="46" pageNumber="47">Tafana arawak</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FF866600FE449485FDAEFD3A" box="[477,567,731,757]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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resemble others of the
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<taxonomicName id="4C664D38FF866600FCD99485FC02FD3A" authorityName="MNHN" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[832,923,731,757]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="silhavyi">
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<emphasis id="B912EAA9FF866600FCD99485FC02FD3A" box="[832,923,731,757]" italics="true" pageId="46" pageNumber="47">silhavyi</emphasis>
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</taxonomicName>
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group by a laminar and coiled apophysis at the base of the embolus, but differ by the retrolateral tibial apophysis bifid (
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<figureCitation id="135D2A3EFF866600FBF794A0FB66FCD7" box="[1134,1279,766,792]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="46" pageNumber="47">Fig. 27C–E</figureCitation>
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). Females resemble those of
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<taxonomicName id="4C664D38FF866600FE0D957CFD95FCF4" authorityName="MNHN" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[404,524,801,827]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="silhavyi">
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<emphasis id="B912EAA9FF866600FE0D957CFD95FCF4" box="[404,524,801,827]" italics="true" pageId="46" pageNumber="47">T. silhavyi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C664D38FF866600FDDF957CFD53FCF4" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[582,714,802,827]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="maracay" status="sp. nov.">
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<emphasis id="B912EAA9FF866600FDDF957CFD53FCF4" box="[582,714,802,827]" italics="true" pageId="46" pageNumber="47">T. maracay</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22157D2FF866600FD48957CFCB5FCF3" box="[721,812,802,828]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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by the epigynal plate with wide atrium, but differ by the atrium extremely wide and the large hood (
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<figureCitation id="135D2A3EFF866600FC9F951BFC0EFC90" box="[774,919,837,863]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="46" pageNumber="47">Figs 27F–G</figureCitation>
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,
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<figureCitation id="135D2A3EFF866600FC3B951BFC4CFC90" box="[930,981,837,863]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: A–H =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="46" pageNumber="47">33E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37C6530FF866600FF2495D9FD73FBD9" pageId="46" pageNumber="47" type="etymology">
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<paragraph id="8BD936BBFF866600FF2495D9FEDDFC6E" blockId="46.[189,324,903,929]" box="[189,324,903,929]" pageId="46" pageNumber="47">
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<heading id="D09181D7FF866600FF2495D9FEDDFC6E" bold="true" box="[189,324,903,929]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="B912EAA9FF866600FF2495D9FEDDFC6E" bold="true" box="[189,324,903,929]" pageId="46" pageNumber="47">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD936BBFF866600FF2495EBFD73FBD9" blockId="46.[189,1399,949,1047]" pageId="46" pageNumber="47">The specific name is derived from the Brazilian Portuguese word ‘Aruaques’ (in Spanish ‘Arahuaco’) for the name of an indigenous group from the Americas, named ‘Arawak’, meaning ‘flour eater’. The name should be treated as a noun in apposition.</paragraph>
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</subSubSection>
|
||
<subSubSection id="C37C6530FF866600FF249260FBBCFAFC" pageId="46" pageNumber="47" type="materials_examined">
|
||
<paragraph id="8BD936BBFF866600FF249260FE33FB97" blockId="46.[189,426,1086,1112]" box="[189,426,1086,1112]" pageId="46" pageNumber="47">
|
||
<heading id="D09181D7FF866600FF249260FE33FB97" bold="true" box="[189,426,1086,1112]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="B912EAA9FF866600FF249260FE33FB97" bold="true" box="[189,426,1086,1112]" pageId="46" pageNumber="47">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF739233FEF7FB01" blockId="46.[189,1398,1133,1230]" pageId="46" pageNumber="47">
|
||
<materialsCitation id="3B0E3CE6FF866600FF739233FEF7FB01" ID-GBIF-Occurrence="3066856381" collectingDate="no date" collectionCode="MNHN" collectorName="E. Simon" country="Venezuela" latitude="10.405555" location="Colonia Tovar" longLatPrecision="21" longitude="-67.289444" municipality="Colonia Tovar" pageId="46" pageNumber="47" specimenCode="MNHN 11320" specimenCount="1" specimenCount-male="1" stateProvince="Aragua" typeStatus="holotype">
|
||
<emphasis id="B912EAA9FF866600FF739233FEC5FB48" bold="true" box="[234,348,1133,1159]" pageId="46" pageNumber="47">
|
||
<typeStatus id="54DD8819FF866600FF739233FEC5FB48" box="[234,348,1133,1159]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F371762BFF866600FF2492CEFEF2FB65" box="[189,363,1168,1194]" name="Venezuela" pageId="46" pageNumber="47">VENEZUELA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D60FD32FF866600FE1092CEFE25FB64" box="[393,444,1168,1195]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49A2F859FF866600FE5392CEFDBAFB65" box="[458,547,1168,1194]" country="Venezuela" name="Aragua" pageId="46" pageNumber="47">Aragua</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBDACC1FF866600FDAA92CEFD7BFB64" box="[563,738,1168,1195]" pageId="46" pageNumber="47">Colonia Tovar</collectingMunicipality>
|
||
; [
|
||
<geoCoordinate id="EE52507CFF866600FD6192D0FC0DFB64" box="[760,916,1166,1195]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="20" value="10.405555">10°24′20″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE52507CFF866600FC3A92D0FBDDFB65" box="[931,1092,1166,1194]" degrees="67" direction="west" minutes="17" orientation="longitude" pageId="46" pageNumber="47" precision="15" seconds="22" value="-67.289444">67°17′22″ W</geoCoordinate>
|
||
];
|
||
<collectingDate id="EF9CE993FF866600FBC592CFFB21FB65" box="[1116,1208,1168,1195]" pageId="46" pageNumber="47">no date</collectingDate>
|
||
;
|
||
<collectorName id="2693536DFF866600FB5192CFFAA0FB65" box="[1224,1337,1168,1195]" pageId="46" pageNumber="47">E. Simon</collectorName>
|
||
leg.;
|
||
<specimenCode id="DBC09EC0FF866600FF2492EAFEF3FB01" box="[189,362,1204,1230]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="46" pageNumber="47">MNHN 11320</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF7392ABFBBCFAFC" blockId="46.[189,1061,1269,1331]" pageId="46" pageNumber="47">
|
||
<materialsCitation id="3B0E3CE6FF866600FF7392ABFBB8FAFC" ID-GBIF-Occurrence="3066856331" collectionCode="MNHN" collectorName="E. Simon" country="Venezuela" latitude="10.405555" location="VENEZUELA" longLatPrecision="21" longitude="-67.289444" pageId="46" pageNumber="47" specimenCode="MNHN 11320" specimenCount="1" specimenCount-female="1" stateProvince="Aragua" typeStatus="paratype">
|
||
<emphasis id="B912EAA9FF866600FF7392ABFEC5FAC0" bold="true" box="[234,348,1269,1295]" pageId="46" pageNumber="47">
|
||
<typeStatus id="54DD8819FF866600FF7392ABFEC5FAC0" box="[234,348,1269,1295]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F371762BFF866600FF249347FEF2FAFC" box="[189,363,1305,1331]" name="Venezuela" pageId="46" pageNumber="47">VENEZUELA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D60FD32FF866600FE1A9347FE29FAFC" box="[387,432,1305,1331]" pageId="46" pageNumber="47" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<specimenCode id="DBC09EC0FF866600FCED9347FBB8FAFC" box="[884,1057,1305,1331]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="46" pageNumber="47">MNHN 11320</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6530FF86661EFF249304FAEEFC27" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
|
||
<paragraph id="8BD936BBFF866600FF249304FED7FABB" blockId="46.[189,334,1370,1396]" box="[189,334,1370,1396]" pageId="46" pageNumber="47">
|
||
<heading id="D09181D7FF866600FF249304FED7FABB" bold="true" box="[189,334,1370,1396]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="B912EAA9FF866600FF249304FED7FABB" bold="true" box="[189,334,1370,1396]" pageId="46" pageNumber="47">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF7393D7FDFAFA6C" blockId="46.[189,1399,1417,1550]" box="[234,611,1417,1443]" pageId="46" pageNumber="47">
|
||
<emphasis id="B912EAA9FF866600FF7393D7FEB3FA6C" bold="true" box="[234,298,1417,1443]" pageId="46" pageNumber="47">Male</emphasis>
|
||
(
|
||
<typeStatus id="54DD8819FF866600FEA093D7FE3DFA6C" box="[313,420,1417,1443]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
, MNHN 11320)
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF2493F2FDF4F9C2" blockId="46.[189,1399,1417,1550]" pageId="46" pageNumber="47">
|
||
COLOR. Exemplar probably discolored. Carapace orange with cephalic region dark (
|
||
<figureCitation id="135D2A3EFF866600FB0B93F2FA99FA09" box="[1170,1280,1452,1478]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="46" pageNumber="47">Fig. 27A</figureCitation>
|
||
). Labium, sternum and endites yellow. Chelicerae reddish. Legs I–IV yellow, darkest at tarsi. Abdomen yellow with spots in dorsal view (
|
||
<figureCitation id="135D2A3EFF866600FE6B93ADFDF8F9C2" box="[498,609,1523,1549]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="46" pageNumber="47">Fig. 27A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF24906BFE4DF976" blockId="46.[188,1399,1589,1721]" pageId="46" pageNumber="47">MEASUREMENTS. Total length 6.1, carapace length 2.7, width 2.2. Clypeus height 0.08. Eye diameters and interdistances: AME 0.08, ALE 0.12, PME 0.1, PLE 0.12; AME–AME 0.04, AME–ALE 0.02, PME– PME 0.12, PME–PLE 0.06, ALE–PLE 0.04. Chelicerae 1.0 long, with four promarginal teeth and four retromarginal denticles.</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF2490BFFA88F8D0" blockId="46.[189,1398,1761,1823]" pageId="46" pageNumber="47">LEG MEASUREMENTS. I—femur 3.1/ patella 1.1/ tibia 3.8/ metatarsus 3.27 / tarsus 1.3/ total 12.57; II— 2.8/ 1.1/ 3.1 /2.8 / 1.1/ 10.9; III—2.0 /0.8 / 1.2/ 2.0/ 0.6/ 6.6; IV—2.8 / 0.75 /2.6 / 3.2/ 0.8/ 10.15.</paragraph>
|
||
<paragraph id="8BD936BBFF866600FF249118FB3DF84C" blockId="46.[188,1399,1862,1924]" pageId="46" pageNumber="47">LEG SPINATION. I—tibia v2-2-2, p1-1-1, r1-1-1, metatarsus v2-0-0, p0-1-0, r0-1-0; II—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus p0-1-0, r0-1-0; III—tibia v2-2-2, p1-1-0, r1-1-0; IV—tibia p1-1-0.</paragraph>
|
||
<paragraph id="8BD936BBFF866601FF2491F5FD6DFEA6" blockId="46.[189,1399,1963,2025]" lastBlockId="47.[189,1399,265,361]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
|
||
PALP. Retrolateral tibial apophysis bifid with anterior branch truncated and posterior branch thin; cymbium with retrolateral projection; apophysis at the base of the embolus laminar and coiled; embolus long with large base and narrow apex; median apophysis long and curved apically; ventral tegular process long and hyaline; basal protuberance of the tegulum short; sperm duct with secondary looping in oblique position in the tegulum (
|
||
<figureCitation id="135D2A3EFF876601FDC29711FD71FEA6" box="[603,744,335,361]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="47" pageNumber="48">Fig. 27C–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF196633FF876601FF2490A5FAFCF826" ID-DOI="http://doi.org/10.5281/zenodo.4655489" ID-Zenodo-Dep="4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="47" pageNumber="48" startId="47.[189,232,1787,1813]" targetBox="[189,1398,461,1746]" targetPageId="47">
|
||
<paragraph id="8BD936BBFF876601FF2490A5FAFCF826" blockId="47.[185,1399,1787,2026]" pageId="47" pageNumber="48">
|
||
<emphasis id="B912EAA9FF876601FF2490A5FE83F8DA" bold="true" box="[189,282,1787,1813]" pageId="47" pageNumber="48">Fig. 27.</emphasis>
|
||
<taxonomicName id="4C664D38FF876601FEBD90A2FE4FF8D9" authority="De Oliveira & Brescovit, 2021" authorityName="De Oliveira & Brescovit" authorityYear="2021" box="[292,470,1788,1814]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="arawak" status="sp. nov.">
|
||
<emphasis id="B912EAA9FF876601FEBD90A2FE4FF8D9" box="[292,470,1788,1814]" italics="true" pageId="47" pageNumber="48">Tafana arawak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22157D2FF876601FE4690A2FDA2F8D9" box="[479,571,1788,1814]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).
|
||
<emphasis id="B912EAA9FF876601FE839141FEB6F8F6" bold="true" box="[282,303,1823,1849]" pageId="47" pageNumber="48">A</emphasis>
|
||
.Habitus,dorsal view.
|
||
<emphasis id="B912EAA9FF876601FDB19141FDA5F8F6" bold="true" box="[552,572,1823,1849]" pageId="47" pageNumber="48">B</emphasis>
|
||
.Habitus,dorsal view.
|
||
<emphasis id="B912EAA9FF876601FCA09141FCD7F8F6" bold="true" box="[825,846,1823,1849]" pageId="47" pageNumber="48">C</emphasis>
|
||
. Left palp, ventral view.
|
||
<emphasis id="B912EAA9FF876601FBF89141FBEFF8F6" bold="true" box="[1121,1142,1823,1849]" pageId="47" pageNumber="48">D</emphasis>
|
||
.Left palp, retrolateral view.
|
||
<emphasis id="B912EAA9FF876601FE9E911CFE82F893" bold="true" box="[263,283,1858,1884]" pageId="47" pageNumber="48">E</emphasis>
|
||
. Left palp, dorsal view.
|
||
<emphasis id="B912EAA9FF876601FDD2911CFDC4F893" bold="true" box="[587,605,1858,1884]" pageId="47" pageNumber="48">F</emphasis>
|
||
. Epigynum, ventral view (arrow indicating character).
|
||
<emphasis id="B912EAA9FF876601FA99911CFA8EF893" bold="true" box="[1280,1303,1858,1884]" pageId="47" pageNumber="48">G</emphasis>
|
||
. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD936BBFF98661EFF249757FAEEFEEC" blockId="48.[189,1399,265,291]" box="[189,1399,265,291]" pageId="48" pageNumber="49">ABDOMEN. Length 3.2, epigastric furrow 0.63 from tracheal spiracle, spiracle 1.6 from base of spinnerets.</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF739715FDE4FEAA" blockId="48.[189,1223,331,392]" box="[234,637,331,357]" pageId="48" pageNumber="49">
|
||
<emphasis id="B912EAA9FF98661EFF739715FEDCFEAA" bold="true" box="[234,325,331,357]" pageId="48" pageNumber="49">Female</emphasis>
|
||
(
|
||
<typeStatus id="54DD8819FF98661EFECC9715FE26FEAA" box="[341,447,331,357]" pageId="48" pageNumber="49" type="paratype">paratype</typeStatus>
|
||
, MNHN 11320)
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF249730FB5EFE47" blockId="48.[189,1223,331,392]" box="[189,1223,366,392]" pageId="48" pageNumber="49">
|
||
COLOR. Exemplar probably discolored. Coloration as in males of this species (
|
||
<figureCitation id="135D2A3EFF98661EFBD79730FB20FE47" box="[1102,1209,366,392]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="48" pageNumber="49">Fig. 27B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF2497EEFE4CFDFC" blockId="48.[188,1398,431,563]" pageId="48" pageNumber="49">MEASUREMENTS. Total length 7.1, carapace length 2.9, width 2.2. Clypeus height 0.06. Eye diameters and interdistances: AME 0.08, ALE 0.1, PME 0.1, PLE 0.12; AME–AME 0.02, AME–ALE 0.02, PME– PME 0.14, PME–PLE 0.08, ALE–PLE 0.04. Chelicerae 1.0 long, with four promarginal teeth and six retromarginal denticles.</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF249405FAB9FD58" blockId="48.[189,1398,602,664]" pageId="48" pageNumber="49">LEG MEASUREMENTS. I—femur 2.9/ patella 1.1/ tibia 3.2/ metatarsus 2.59 / tarsus 1.1/ total 10.89; II— 2.6/ 1.0/ 2.75 / 2.3/ 1.0 /9.65; III—2.0 / 0.8/ 1.5 /1.7 /0.6 / 6.6; IV—2.8/ 0.82 / 2.5/ 3.1 /0.75 / 9.97.</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF2494E1FBDAFD33" blockId="48.[188,1399,703,764]" pageId="48" pageNumber="49">LEG SPINATION. I—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus v2-0-0, p0-1-0, r0-1-0; II—tibia v2-2-2, p1-1-0, r1-1-0, metatarsus p0-1-0, r0-1-0; III—tibia p1-1-0, r1-1-0; IV—tibia r1-1-0.</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF24957AFBF5FC68" blockId="48.[188,1399,803,935]" pageId="48" pageNumber="49">
|
||
EPIGYNUM. Hood triangular; wide atrium; epigynal plate covered with hyaline cuticle in the posterior region; lateral lobes sinuous. Internally, copulatory ducts extremely large in the anterior region; primary spermathecae oval and separated; secondary spermathecae small, lateral of the primary spermathecae, and fertilization ducts longer than the primary spermathecae (
|
||
<figureCitation id="135D2A3EFF98661EFC0995D3FBB9FC68" box="[912,1056,909,935]" captionStart="Fig" captionStartId="47.[189,232,1787,1813]" captionTargetBox="[189,1398,461,1746]" captionTargetId="figure-177@47.[189,1398,461,1746]" captionTargetPageId="47" captionText="Fig. 27. Tafana arawak sp. nov. (A, C–E= holotype, ♂, MNHN 11320; B, F–G =paratype,♀, MNHN 11320).A.Habitus,dorsalview.B.Habitus,dorsalview.C. Leftpalp, ventral view.D.Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view (arrow indicating character). G. Vulva, dorsal view. Abbreviations: A=atrium; aRTA=anterior branch of the retrolateral tibial apophysis; CD= copulatory ducts; CO=copulatory opening; Ch =character; E =embolus; FD =fertilization ducts; H =hood; LL=lateral lobes; pRTA= posterior branch of the retrolateral tibial apophysis; S1= primary spermathecae; S2 =secondary spermathecae; T=tegulum. Scale bars: A–B= 3.5 mm; C–G =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655489" httpUri="https://zenodo.org/record/4655489/files/figure.png" pageId="48" pageNumber="49">Figs 27F–G</figureCitation>
|
||
,
|
||
<figureCitation id="135D2A3EFF98661EFBB595D3FBC6FC68" box="[1068,1119,909,935]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: A–H =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="48" pageNumber="49">33E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF249591FAEEFC27" blockId="48.[189,1399,974,1001]" box="[189,1399,974,1001]" pageId="48" pageNumber="49">ABDOMEN. Length 4.0, epigastric furrow 0.8 from tracheal spiracle, spiracle 2.2 from base of spinnerets.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C6530FF98661EFF24924EFE32FB96" box="[189,427,1040,1113]" pageId="48" pageNumber="49" type="distribution">
|
||
<paragraph id="8BD936BBFF98661EFF24924EFECFFBE5" blockId="48.[189,342,1040,1066]" box="[189,342,1040,1066]" pageId="48" pageNumber="49">
|
||
<heading id="D09181D7FF98661EFF24924EFECFFBE5" bold="true" box="[189,342,1040,1066]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
|
||
<emphasis id="B912EAA9FF98661EFF24924EFECFFBE5" bold="true" box="[189,342,1040,1066]" pageId="48" pageNumber="49">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD936BBFF98661EFF249260FE32FB96" blockId="48.[189,427,1086,1113]" box="[189,427,1086,1113]" pageId="48" pageNumber="49">
|
||
<collectingCountry id="F371762BFF98661EFF249260FEAFFB97" box="[189,310,1086,1112]" name="Venezuela" pageId="48" pageNumber="49">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="135D2A3EFF98661EFEDF9260FE04FB96" box="[326,413,1086,1113]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="48" pageNumber="49">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |