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<document ID-DOI="http://doi.org/10.5281/zenodo.3789752" ID-GBIF-Dataset="7a68e13d-3c05-49d5-86a3-7bdb5c4af1ce" ID-GBIF-Taxon="163755352" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789752" ID-ZooBank="553C91CE-FEA1-4867-929B-2865647E4B2E" approvalRequired="14" approvalRequired_for_textStreams="14" checkinTime="1588002532591" checkinUser="plazi" docAuthor="Ballerio, Alberto &amp; Maruyama, Munetoshi" docDate="2010" docId="03CA87D50455FFF465FE48164A1B2B48" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="ZooKeys 34 (34)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@5da20b76" docStyleName="zookeys.2008.journal_article" docTitle="Madrasostes depressum Paulian 1992" docType="treatment" docVersion="4" lastPageNumber="85" masterDocId="FFF3FFAD045DFFFC65724A5A4E182815" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="8" pageNumber="85" updateTime="1643557208252" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ballerio, Alberto</mods:namePart>
<mods:affiliation>, Brescia, Italy</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maruyama, Munetoshi</mods:namePart>
<mods:affiliation>The Kyushu University Museum, Fukuoka, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-01-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>34</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>77</mods:start>
<mods:end>104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.34.268</mods:identifier>
<mods:identifier type="GBIF-Dataset">7a68e13d-3c05-49d5-86a3-7bdb5c4af1ce</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576606</mods:identifier>
<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789752" ID-GBIF-Taxon="163755352" ID-Zenodo-Dep="3789752" LSID="urn:lsid:plazi:treatment:03CA87D50455FFF465FE48164A1B2B48" httpUri="http://treatment.plazi.org/id/03CA87D50455FFF465FE48164A1B2B48" lastPageNumber="85" pageId="8" pageNumber="85">
<subSubSection pageId="8" pageNumber="85" type="nomenclature">
<paragraph blockId="8.[140,1108,588,862]" box="[140,698,588,615]" pageId="8" pageNumber="85">
<heading box="[140,698,588,615]" fontSize="11" level="2" pageId="8" pageNumber="85" reason="3">
<taxonomicName authority="Paulian, 1992" authorityName="Paulian" authorityYear="1992" box="[140,589,588,615]" class="Insecta" family="Hybosoridae" genus="Madrasostes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="85" phylum="Arthropoda" rank="species" species="depressum">
<emphasis bold="true" box="[140,589,588,615]" pageId="8" pageNumber="85">
<emphasis bold="true" box="[140,415,588,615]" italics="true" pageId="8" pageNumber="85">
<emphasis box="[140,286,588,615]" inLineHeading="true" pageId="8" pageNumber="85" reason="1">Madrasostes</emphasis>
depressum
</emphasis>
<bibRefCitation author="Paulian R" box="[422,589,589,615]" pageId="8" pageNumber="85" pagination="119 - 124" refId="ref11539" refString="Paulian R (1992) Notes sur les Coleopteres Scarabaeoidea du Museum de Geneve. V. Revue Suisse de Zoologie 99: 119 - 124." type="journal article" year="1992">Paulian, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[604,690,589,615]" captionStart="Figure 2" captionStartId="15.[140,212,1558,1580]" captionText="Figure 2. Ceratocanthinae of Ulu Gombak. A Madrasostes depressum habitus dorsal B Madrasostes boucomonti habitus dorsal C Madrasostes agostii sp. n. habitus dorsal D Madrasostes clypeale male habitus dorsal E Madrasostes malyanum habitus dorsal F Madrasostes mirificum sp. n. habitus dorsal G habitus lateral of male H fore body of female I male protibia dorsal view Į male protibia lateral view." pageId="8" pageNumber="85">Fig. 2A</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="8.[140,1108,588,862]" box="[185,846,624,651]" pageId="8" pageNumber="85">
Known distribution:
<collectingCountry box="[425,535,624,650]" name="Indonesia" pageId="8" pageNumber="85">Indonesia</collectingCountry>
(
<collectingRegion box="[550,648,624,650]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="85">Sumatra</collectingRegion>
) (
<bibRefCitation author="Paulian R" box="[671,834,624,650]" pageId="8" pageNumber="85" pagination="119 - 124" refId="ref11539" refString="Paulian R (1992) Notes sur les Coleopteres Scarabaeoidea du Museum de Geneve. V. Revue Suisse de Zoologie 99: 119 - 124." type="journal article" year="1992">Paulian, 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="85" type="discussion">
<paragraph blockId="8.[140,1108,588,862]" pageId="8" pageNumber="85">
Remarks. First record for West
<collectingCountry box="[530,629,659,685]" name="Malaysia" pageId="8" pageNumber="85">Malaysia</collectingCountry>
(
<collectingRegion box="[643,747,659,686]" country="Malaysia" name="Selangor" pageId="8" pageNumber="85">Selangor</collectingRegion>
). This species has two distinctive paired thick sclerites in the internal sac of aedeagus. The shape of these sclerites in the Ulu Gombak material is slightly different from the shape of the sclerites of the
<typeStatus pageId="8" pageNumber="85">holotype</typeStatus>
, however, due also to the circumstance that the Sumatran population is known by the type only, we were unable to find further differences that could allow a taxonomic separation of the Sumatran populations from Peninsular Malaysian specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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