treatments-xml/data/03/CA/87/03CA87CD14383C521D427D62FBEAFC68.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.5643376" ID-GBIF-Dataset="3f24224e-3ff9-474c-a138-adb332383ac3" ID-GBIF-Taxon="189892284" ID-ISSN="1030-1887" ID-Zenodo-Dep="5643376" approvalRequired="2" approvalRequired_for_document="1" approvalRequired_for_treatments="1" checkinTime="1635948658041" checkinUser="diego" docAuthor="Callmander, Martin W., Gallaher, Timothy, Buerki, Sven, Zich, Frank A. &amp; Field, Ashley R." docDate="2021" docId="03CA87CD14383C521D427D62FBEAFC68" docLanguage="en" docName="AustSystBot.34.327-335.pdf" docOrigin="Australian Systematic Botany 34" docSource="https://doi.org/10.1071/SB20033" docTitle="Pandanus grayorum Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field 2021, sp. nov." docType="treatment" docVersion="4" lastPageId="6" lastPageNumber="333" masterDocId="FFF3FFB514393C541E687969FFEBFF94" masterDocTitle="Pandanus grayorum (Pandanaceae), a new species endemic to north-eastern Queensland (Australia)" masterLastPageNumber="335" masterPageNumber="327" pageId="1" pageNumber="328" updateTime="1635956109901" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Pandanus grayorum (Pandanaceae), a new species endemic to north-eastern Queensland (Australia)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Callmander, Martin W.</mods:namePart>
<mods:affiliation>Conservatoire et Jardin botaniques de Genève, Chemin de lImpératrice 1, CP 71, CH- 1292 Chambésy, Switzerland.</mods:affiliation>
<mods:nameIdentifier type="email">martin.callmander@ville-ge.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gallaher, Timothy</mods:namePart>
<mods:affiliation>Herbarium Pacificum, Bishop Museum, Department of Natural Sciences, 1525 Bernice Street, Honolulu, HI 96817 - 2704, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Buerki, Sven</mods:namePart>
<mods:affiliation>Department of Biological Sciences, Boise State University, Boise, ID 83725, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zich, Frank A.</mods:namePart>
<mods:affiliation>Australian Tropical Herbarium, James Cook University, Cairns Campus, McGregor Road, Smithfield, Qld 4878, Australia. &amp; National Research Collections Australia, Commonwealth Industrial and Scientific Research Organisation (CSIRO), GPO Box 1700, Canberra, ACT 2601, Australia.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Field, Ashley R.</mods:namePart>
<mods:affiliation>Australian Tropical Herbarium, James Cook University, Cairns Campus, McGregor Road, Smithfield, Qld 4878, Australia. &amp; Department of Biological Sciences, Boise State University, Boise, ID 83725, USA.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Australian Systematic Botany</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-06-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>327</mods:start>
<mods:end>335</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>https://doi.org/10.1071/SB20033</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1071/SB20033</mods:identifier>
<mods:identifier type="GBIF-Dataset">3f24224e-3ff9-474c-a138-adb332383ac3</mods:identifier>
<mods:identifier type="ISSN">1030-1887</mods:identifier>
<mods:identifier type="Zenodo-Dep">5643372</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5643376" ID-GBIF-Taxon="189892284" ID-Zenodo-Dep="5643376" LSID="urn:lsid:plazi:treatment:03CA87CD14383C521D427D62FBEAFC68" httpUri="http://treatment.plazi.org/id/03CA87CD14383C521D427D62FBEAFC68" lastPageId="6" lastPageNumber="333" pageId="1" pageNumber="328">
<subSubSection pageId="1" pageNumber="328" type="nomenclature">
<paragraph blockId="1.[810,1419,1035,1098]" pageId="1" pageNumber="328">
<heading centered="true" fontSize="9" level="3" pageId="1" pageNumber="328" reason="4">
<taxonomicName authority="Callm., Buerki &amp; Gallaher" authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[810,1322,1035,1058]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="328" phylum="Tracheophyta" rank="species" species="grayorum" status="sp. nov.">
<emphasis bold="true" box="[810,1029,1035,1058]" italics="true" pageId="1" pageNumber="328">Pandanusgrayorum</emphasis>
Callm., Buerki &amp; Gallaher
</taxonomicName>
,
<taxonomicNameLabel box="[1333,1419,1035,1058]" pageId="1" pageNumber="328" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation box="[1080,1141,1076,1098]" captionStart="Fig" captionStartId="2.[155,186,1734,1753]" captionTargetBox="[159,1480,212,1703]" captionTargetId="figure-1@2.[156,842,214,1260]" captionTargetPageId="2" captionText="Fig. 1. Pandanus grayorum. A. Habitat. B. Phalanges with pileus. C. Syncarp with axillary plantlets. D. Staminate flowers. E. Plantlet. F. Staminate inflorescence. Images of Field &amp; Zich 5017 by A. Field (A). Images of Gray 9976 (type) by B. Gray (B, C). Images of Field &amp; Zich 5019 by A. Field (D, F). Image of Field &amp; Zich 5015 by A. Field (E)." figureDoi="http://doi.org/10.5281/zenodo.5643378" httpUri="https://zenodo.org/record/5643378/files/figure.png" pageId="1" pageNumber="328">Fig. 1</figureCitation>
.)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="328" type="materials_examined">
<paragraph blockId="1.[795,1433,1132,1271]" pageId="1" pageNumber="328">
<materialsCitation ID-GBIF-Occurrence="3397196306" collectingDate="2018-10-04" collectionCode="BISH, BRI, CANB, CNS" collectorName="B. Gray" country="Australia" elevation="8" location="Jubilee Road, off Flying Fish Point Road, Innisfail" municipality="Cook District" pageId="1" pageNumber="328" specimenCode="G 00446800" specimenCount="2" stateProvince="Queensland" typeStatus="type">
<emphasis box="[795,853,1133,1154]" italics="true" pageId="1" pageNumber="328">
<typeStatus box="[795,847,1133,1154]" pageId="1" pageNumber="328">Type</typeStatus>
:
</emphasis>
<collectingCountry box="[856,1004,1132,1154]" name="Australia" pageId="1" pageNumber="328" value="Australia">AUSTRALIA</collectingCountry>
.
<collectingRegion box="[1012,1137,1132,1154]" country="Australia" name="Queensland" pageId="1" pageNumber="328">Queensland</collectingRegion>
:
<collectingMunicipality box="[1145,1286,1132,1154]" pageId="1" pageNumber="328">Cook District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03CA87CD14383C521D427D62FBEAFC68:8EBC600014383C551B677D05FB64FB0B" country="Australia" municipality="Cook District" name="Jubilee Road, off Flying Fish Point Road, Innisfail" pageId="1" pageNumber="328" stateProvince="Queensland">Jubilee Road, off Flying Fish Point Road, Innisfail</location>
,
<geoCoordinate box="[1176,1290,1157,1183]" pageId="1" pageNumber="328">
17
<superScript attach="right" box="[1202,1211,1157,1173]" fontSize="7" pageId="1" pageNumber="328">°</superScript>
<superScript attach="none" box="[1211,1273,1157,1183]" fontSize="7" pageId="1" pageNumber="328">290460</superScript>
S
</geoCoordinate>
,
<geoCoordinate box="[1300,1429,1157,1183]" pageId="1" pageNumber="328">
146
<superScript attach="right" box="[1339,1348,1157,1173]" fontSize="7" pageId="1" pageNumber="328">°</superScript>
<superScript attach="none" box="[1347,1410,1157,1183]" fontSize="7" pageId="1" pageNumber="328">020260</superScript>
E
</geoCoordinate>
,
<quantity box="[795,836,1190,1212]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="328" unit="m" value="8.0">
<elevation box="[795,836,1190,1212]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="328" unit="m" value="8.0">8 m</elevation>
</quantity>
,
<date box="[847,939,1190,1212]" pageId="1" pageNumber="328" value="2018-10-04">
<collectingDate box="[847,939,1190,1212]" pageId="1" pageNumber="328" value="2018-10-04">4.x.2018</collectingDate>
</date>
, fr.,
<emphasis box="[987,1125,1190,1213]" italics="true" pageId="1" pageNumber="328">
<collectorName box="[987,1067,1191,1213]" pageId="1" pageNumber="328">B. Gray</collectorName>
9976
</emphasis>
(holo:
<specimenCode box="[1200,1333,1190,1212]" pageId="1" pageNumber="328">G 00446800</specimenCode>
! [3 sheets + carpological
<specimenCount box="[964,1067,1220,1242]" pageId="1" pageNumber="328" type="generic">specimen</specimenCount>
]; iso:
<collectionCode box="[1142,1208,1220,1242]" pageId="1" pageNumber="328">BISH</collectionCode>
!,
<collectionCode box="[1229,1274,1220,1241]" pageId="1" pageNumber="328">BRI</collectionCode>
,
<collectionCode box="[1292,1365,1220,1242]" pageId="1" pageNumber="328">CANB</collectionCode>
,
<collectionCode box="[1383,1432,1220,1242]" pageId="1" pageNumber="328">CNS</collectionCode>
148706! [3 sheets + carpological
<specimenCount box="[1149,1253,1249,1271]" pageId="1" pageNumber="328" type="generic">specimen</specimenCount>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="328" type="reference_group">
<paragraph blockId="1.[795,1432,1302,1348]" pageId="1" pageNumber="328">
<taxonomicName box="[795,921,1302,1322]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="328" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[795,885,1302,1322]" italics="true" pageId="1" pageNumber="328">Pandanus</emphasis>
sp.
</taxonomicName>
(Russell River) in W.E.Cooper &amp; W.T.Cooper,
<emphasis italics="true" pageId="1" pageNumber="328">Fruits Austral. Trop. Rainforest</emphasis>
390 (2004).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="328" type="diagnosis">
<paragraph blockId="1.[795,900,1395,1414]" box="[795,900,1395,1414]" pageId="1" pageNumber="328">
<emphasis box="[795,900,1395,1414]" italics="true" pageId="1" pageNumber="328">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="1.[795,1434,1434,1719]" pageId="1" pageNumber="328">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[795,1012,1434,1456]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="328" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[795,1012,1434,1456]" italics="true" pageId="1" pageNumber="328">Pandanus grayorum</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName authorityName="H. St. John" box="[1268,1399,1434,1456]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="328" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[1268,1399,1434,1456]" italics="true" pageId="1" pageNumber="328">P. gemmifer</emphasis>
</taxonomicName>
in having larger crown leaves (210
<emphasis box="[1138,1151,1466,1484]" italics="true" pageId="1" pageNumber="328"></emphasis>
280 ×
<quantity box="[1227,1306,1463,1485]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="328" unit="cm" value="7.5" valueMax="8.0" valueMin="7.0">
7
<emphasis box="[1240,1253,1466,1484]" italics="true" pageId="1" pageNumber="328"></emphasis>
8 cm
</quantity>
<emphasis box="[1316,1327,1464,1485]" italics="true" pageId="1" pageNumber="328">v</emphasis>
. 170
<emphasis box="[1382,1395,1466,1484]" italics="true" pageId="1" pageNumber="328"></emphasis>
230 ×
<quantity box="[820,896,1492,1514]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="328" unit="cm" value="5.5" valueMax="6.0" valueMin="5.0">
5
<emphasis box="[833,846,1495,1513]" italics="true" pageId="1" pageNumber="328"></emphasis>
6 cm
</quantity>
in
<taxonomicName authorityName="H. St. John" box="[927,1051,1492,1514]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="328" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[927,1051,1492,1514]" italics="true" pageId="1" pageNumber="328">P. gemmifer</emphasis>
</taxonomicName>
) and syncarps (35
<emphasis box="[1239,1252,1495,1513]" italics="true" pageId="1" pageNumber="328"></emphasis>
40 ×
<quantity box="[1308,1409,1492,1514]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="1" pageNumber="328" unit="cm" value="20.0" valueMax="22.0" valueMin="18.0">
18
<emphasis box="[1334,1347,1495,1513]" italics="true" pageId="1" pageNumber="328"></emphasis>
22 cm
</quantity>
<date pageId="1" pageNumber="328" value="1925-05-23">
<emphasis box="[1415,1426,1493,1514]" italics="true" pageId="1" pageNumber="328">v</emphasis>
. 23
<emphasis box="[821,834,1524,1542]" italics="true" pageId="1" pageNumber="328"></emphasis>
25
</date>
×
<quantity box="[893,999,1521,1543]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="1" pageNumber="328" unit="cm" value="14.0" valueMax="18.0" valueMin="10.0">
10
<emphasis box="[918,931,1524,1542]" italics="true" pageId="1" pageNumber="328"></emphasis>
18 cm
</quantity>
); apex of phalanges flat or slightly domed at apex, divided by apical sinuses ~
<quantity box="[1179,1253,1550,1572]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="328" unit="cm" value="0.5">0.5 cm</quantity>
deep into small, often truncate pyramids as many as there are carpels (
<emphasis box="[803,814,1610,1631]" italics="true" pageId="1" pageNumber="328">v</emphasis>
. rounded to the stigmatic cluster,
<quantity box="[1197,1312,1609,1631]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="1" pageNumber="328" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">
12
<emphasis box="[1222,1235,1612,1630]" italics="true" pageId="1" pageNumber="328"></emphasis>
15 mm
</quantity>
wide with narrow apical central sinuses restricted to the central cluster); stigma on apex or on the edge of the slightly truncate apical face of each pyramid (
<emphasis box="[969,980,1698,1719]" italics="true" pageId="1" pageNumber="328">v</emphasis>
. on proximal face of the central apex cluster).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="330" pageId="1" pageNumber="328" type="description">
<paragraph blockId="1.[795,920,1754,1773]" box="[795,920,1754,1773]" pageId="1" pageNumber="328">
<emphasis box="[795,920,1754,1773]" italics="true" pageId="1" pageNumber="328">Description</emphasis>
</paragraph>
<paragraph blockId="1.[795,1433,1793,1932]" lastBlockId="3.[110,748,215,1260]" lastPageId="3" lastPageNumber="330" pageId="1" pageNumber="328">
<emphasis box="[795,842,1794,1815]" italics="true" pageId="1" pageNumber="328">Tree</emphasis>
up to
<quantity box="[913,954,1793,1815]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="328" unit="m" value="9.0">9 m</quantity>
tall, stem with conical nodules, prop roots at base up to
<quantity box="[909,973,1822,1844]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="328" unit="cm" value="50.0">50 cm</quantity>
high, upper branches (and sometimes stem) producing axillary sprouts (plantlets).
<emphasis box="[1215,1286,1852,1873]" italics="true" pageId="1" pageNumber="328">Leaves</emphasis>
<quantity box="[1299,1433,1851,1873]" metricMagnitude="0" metricUnit="m" metricValue="2.45" metricValueMax="2.8" metricValueMin="2.1" pageId="1" pageNumber="328" unit="cm" value="245.0" valueMax="280.0" valueMin="210.0">
210
<emphasis box="[1337,1350,1855,1873]" italics="true" pageId="1" pageNumber="328"></emphasis>
280 cm
</quantity>
long,
<quantity box="[861,945,1881,1903]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="1" pageNumber="328" unit="cm" value="7.5" valueMax="8.0" valueMin="7.0">
7
<emphasis box="[874,887,1884,1902]" italics="true" pageId="1" pageNumber="328"></emphasis>
8 cm
</quantity>
wide in the middle, ~
<quantity box="[1207,1265,1881,1903]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="328" unit="cm" value="9.0">9 cm</quantity>
wide near the sheath, gradually attenuate in the upper part, coriaceous; longitudinal veins visible on both surfaces; marginal prickles beginning at
<quantity box="[242,318,244,266]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="3" pageNumber="330" unit="cm" value="8.0" valueMax="9.0" valueMin="7.0">
7
<emphasis box="[255,268,247,265]" italics="true" pageId="3" pageNumber="330"></emphasis>
9 cm
</quantity>
above the base and extending to the apex, antrorse, 5.5(
<emphasis box="[248,261,276,294]" italics="true" pageId="3" pageNumber="330"></emphasis>
6) mm in the lower third, 5(
<emphasis box="[558,571,276,294]" italics="true" pageId="3" pageNumber="330"></emphasis>
10) mm apart, to
<quantity box="[110,192,302,324]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="3" pageNumber="330" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">
2
<emphasis box="[122,135,306,324]" italics="true" pageId="3" pageNumber="330"></emphasis>
5 mm
</quantity>
long in the mid-third;
<quantity box="[419,501,302,324]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="330" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">
3
<emphasis box="[432,445,306,324]" italics="true" pageId="3" pageNumber="330"></emphasis>
5 mm
</quantity>
apart, up to ~
<quantity box="[642,696,302,324]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="330" unit="mm" value="1.0">1 mm</quantity>
long in the distal third,
<quantity box="[307,393,332,354]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="330" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">
2
<emphasis box="[320,333,335,353]" italics="true" pageId="3" pageNumber="330"></emphasis>
3 mm
</quantity>
apart; midrib armed in the upper 1/4, prickles
<quantity box="[242,325,361,383]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="330" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">
4
<emphasis box="[255,268,364,382]" italics="true" pageId="3" pageNumber="330"></emphasis>
5 mm
</quantity>
long), irregularly disposed (
<quantity box="[616,714,361,383]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="3" pageNumber="330" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">
3
<emphasis box="[629,642,364,382]" italics="true" pageId="3" pageNumber="330"></emphasis>
10 mm
</quantity>
) in the middle, then regularly disposed, spaced (
<quantity box="[572,659,390,412]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="3" pageNumber="330" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">
2
<emphasis box="[585,598,393,411]" italics="true" pageId="3" pageNumber="330"></emphasis>
5 mm
</quantity>
) as long as on the marginal in the distal 1/3; sheath
<quantity box="[544,620,419,441]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="3" pageNumber="330" unit="cm" value="7.0" valueMax="9.0" valueMin="5.0">
5
<emphasis box="[557,570,423,441]" italics="true" pageId="3" pageNumber="330"></emphasis>
9 cm
</quantity>
long, ~
<quantity box="[697,747,419,441]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="330" unit="cm" value="9.0">9 cm</quantity>
wide at apex, ~
<quantity box="[265,366,449,471]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="3" pageNumber="330" unit="cm" value="10.5" valueMax="11.0" valueMin="10.0">
10
<emphasis box="[291,304,452,470]" italics="true" pageId="3" pageNumber="330"></emphasis>
11 cm
</quantity>
wide at base.
<emphasis box="[511,649,449,471]" italics="true" pageId="3" pageNumber="330">Infrutescence</emphasis>
terminal, a solitary syncarp on a curved peduncle; syncarp 35
<emphasis box="[678,691,481,499]" italics="true" pageId="3" pageNumber="330"></emphasis>
40 ×
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18
<emphasis box="[135,148,510,528]" italics="true" pageId="3" pageNumber="330"></emphasis>
22 cm
</quantity>
, oblong to obovate, bright red at maturity; peduncle ~
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60
<emphasis box="[149,162,540,558]" italics="true" pageId="3" pageNumber="330"></emphasis>
70 cm
</quantity>
long,
<quantity box="[294,345,536,558]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="330" unit="cm" value="2.0">2 cm</quantity>
wide at apex, trigonous, veins visible, first bract borne just above the base of syncarp, 8 bract scars along peduncle and the remaining bracts crowded at the base of the syncarp. Phalanges, 60
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80 per syncarp, 6
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8(
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9) cm high, 4
<emphasis box="[122,135,656,674]" italics="true" pageId="3" pageNumber="330"></emphasis>
5(
<emphasis box="[156,169,656,674]" italics="true" pageId="3" pageNumber="330"></emphasis>
6) cm wide,
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2.5
<emphasis box="[335,348,656,674]" italics="true" pageId="3" pageNumber="330"></emphasis>
3.5 cm
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thick, 4
<emphasis box="[506,519,656,674]" italics="true" pageId="3" pageNumber="330"></emphasis>
6 angled; carpels (8
<emphasis box="[726,739,656,674]" italics="true" pageId="3" pageNumber="330"></emphasis>
) 12
<emphasis box="[135,148,686,704]" italics="true" pageId="3" pageNumber="330"></emphasis>
18(
<emphasis box="[182,195,686,704]" italics="true" pageId="3" pageNumber="330"></emphasis>
20), incompletely united, free in the upper ~1/3
<emphasis box="[695,708,686,704]" italics="true" pageId="3" pageNumber="330"></emphasis>
1/2, tapering to the base; pileus flat or slightly domed at the apex, divided by apical sinuses ~
<quantity box="[375,443,741,763]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="330" unit="cm" value="0.5">0.5 cm</quantity>
deep into small, often truncate pyramidsasmanyastherearecarpels.
<emphasis box="[479,548,771,792]" italics="true" pageId="3" pageNumber="330">Stigma</emphasis>
1 percarpel,oblique to vertical, generally reniform, sometimes deltoid on apex or on the edge of the slightly truncate apical face of each pyramid; endocarpbony, inlower 2/3, marginsup to
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wide,
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1.2
<emphasis box="[670,683,861,879]" italics="true" pageId="3" pageNumber="330"></emphasis>
2.5 cm
</quantity>
long in the centre,
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1
<emphasis box="[312,325,890,908]" italics="true" pageId="3" pageNumber="330"></emphasis>
2 cm
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wide; seed locule oblong, 1.5
<emphasis box="[677,690,890,908]" italics="true" pageId="3" pageNumber="330"></emphasis>
2.5 ×
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0.5
<emphasis box="[141,153,919,937]" italics="true" pageId="3" pageNumber="330"></emphasis>
1 cm
</quantity>
, superior and inferior mesocarp thick and fibrous.
<emphasis box="[110,353,946,967]" italics="true" pageId="3" pageNumber="330">Staminate inflorescence</emphasis>
terminal, pendent, creamy white, up to
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long, a three-sided raceme of spikes; peduncle fleshy, flaccid and soon disintegrating; lowest floral bract 67 ×
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, margins finely toothed, apex long-subulate, median bract 31 ×
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, with fine marginal prickles, basal floral bracts extending longer than distal floral bracts; spikes dense, cylindric, 6
<emphasis box="[289,301,1124,1142]" italics="true" pageId="3" pageNumber="330"></emphasis>
10 ×
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2.5
<emphasis box="[386,398,1124,1142]" italics="true" pageId="3" pageNumber="330"></emphasis>
4 cm
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, with manystaminate flowers; each spike of a fleshy central axis with stemonophore
<quantity box="[640,747,1150,1172]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="3" pageNumber="330" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">
10
<emphasis box="[665,677,1153,1171]" italics="true" pageId="3" pageNumber="330"></emphasis>
13 mm
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longeach bearing 20
<emphasis box="[313,325,1183,1201]" italics="true" pageId="3" pageNumber="330"></emphasis>
26 stamens; stamens with filament
<quantity box="[666,747,1179,1201]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="330" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">
1
<emphasis box="[679,691,1183,1201]" italics="true" pageId="3" pageNumber="330"></emphasis>
2 mm
</quantity>
long, anthers 2.5
<emphasis box="[288,300,1212,1230]" italics="true" pageId="3" pageNumber="330"></emphasis>
4.5 ×
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0.8
<emphasis box="[407,420,1212,1230]" italics="true" pageId="3" pageNumber="330"></emphasis>
1.2 mm
</quantity>
, including an apiculus
<quantity box="[110,234,1238,1260]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="330" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">
0.3
<emphasis box="[141,153,1241,1259]" italics="true" pageId="3" pageNumber="330"></emphasis>
0.5 mm
</quantity>
. (
<figureCitation box="[251,310,1238,1260]" captionStart="Fig" captionStartId="2.[155,186,1734,1753]" captionTargetBox="[159,1480,212,1703]" captionTargetId="figure-1@2.[156,842,214,1260]" captionTargetPageId="2" captionText="Fig. 1. Pandanus grayorum. A. Habitat. B. Phalanges with pileus. C. Syncarp with axillary plantlets. D. Staminate flowers. E. Plantlet. F. Staminate inflorescence. Images of Field &amp; Zich 5017 by A. Field (A). Images of Gray 9976 (type) by B. Gray (B, C). Images of Field &amp; Zich 5019 by A. Field (D, F). Image of Field &amp; Zich 5015 by A. Field (E)." figureDoi="http://doi.org/10.5281/zenodo.5643378" httpUri="https://zenodo.org/record/5643378/files/figure.png" pageId="3" pageNumber="330">Fig. 1</figureCitation>
.)
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5643378" ID-Zenodo-Dep="5643378" httpUri="https://zenodo.org/record/5643378/files/figure.png" pageId="2" pageNumber="329" startId="2.[155,186,1734,1753]" targetBox="[159,1480,212,1703]" targetPageId="2">
<paragraph blockId="2.[155,1478,1733,1806]" pageId="2" pageNumber="329">
<emphasis bold="true" box="[155,211,1734,1753]" pageId="2" pageNumber="329">Fig. 1.</emphasis>
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[232,411,1734,1753]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="329" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[232,411,1734,1753]" italics="true" pageId="2" pageNumber="329">Pandanus grayorum</emphasis>
</taxonomicName>
. A. Habitat. B. Phalanges with pileus. C. Syncarp with axillary plantlets. D. Staminate flowers. E. Plantlet. F. Staminate inflorescence. Images of
<emphasis box="[383,555,1760,1779]" italics="true" pageId="2" pageNumber="329">Field &amp; Zich 5017</emphasis>
by A. Field (A). Images of
<emphasis box="[823,920,1760,1779]" italics="true" pageId="2" pageNumber="329">Gray 9976</emphasis>
(type) by B. Gray (B, C). Images of
<emphasis box="[1275,1447,1760,1779]" italics="true" pageId="2" pageNumber="329">Field &amp; Zich 5019</emphasis>
by A. Field (D, F). Image of
<emphasis box="[390,556,1787,1806]" italics="true" pageId="2" pageNumber="329">Field &amp; Zich 5015</emphasis>
by A. Field (E).
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="330" type="distribution">
<paragraph blockId="3.[110,238,1302,1321]" box="[110,238,1302,1321]" pageId="3" pageNumber="330">
<emphasis box="[110,238,1302,1321]" italics="true" pageId="3" pageNumber="330">Distribution</emphasis>
</paragraph>
<paragraph blockId="3.[110,748,1340,1742]" pageId="3" pageNumber="330">
This species is known from the Wet Tropics of north-eastern Queensland on and near the banks of the lower reaches of the Mulgrave, Russell, Alice, Johnstone and Moresby rivers and associated subcoastal flood plains (
<figureCitation box="[470,531,1428,1450]" captionStart="Fig" captionStartId="4.[403,434,1839,1858]" captionTargetBox="[257,1375,210,1810]" captionTargetId="figure-1@4.[257,1376,210,1810]" captionTargetPageId="4" captionText="Fig. 2. Distribution map of Pandanus grayorum and P. gemmifer in north-eastern Queensland." figureDoi="http://doi.org/10.5281/zenodo.5643380" httpUri="https://zenodo.org/record/5643380/files/figure.png" pageId="3" pageNumber="330">Fig. 2</figureCitation>
). Plants identified as
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[110,243,1458,1479]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[110,243,1458,1479]" italics="true" pageId="3" pageNumber="330">P. grayorum</emphasis>
</taxonomicName>
have been observed in the Ella Bay, Moresby Range and Russell River National Parks and Etty Bay Conservation Reserve but most of the distribution occurs in fragmented remnants among agricultural land, suggesting its previous distribution may have been wider. It occurs in sympatry with
<taxonomicName box="[281,481,1603,1625]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="solms-laubachii">
<emphasis box="[281,481,1603,1625]" italics="true" pageId="3" pageNumber="330">P. solms-laubachii</emphasis>
</taxonomicName>
, but is parapatric with
<taxonomicName authorityName="H. St. John" box="[110,244,1632,1654]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[110,244,1632,1654]" italics="true" pageId="3" pageNumber="330">P. gemmifer</emphasis>
</taxonomicName>
, which occurs in the mid-upper reaches of the adjacent Barron, Tully and Herbert River systems to the north and south, on the Atherton Tableland, and on the McIlwraith Range on Cape York Peninsula.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="330" type="biology_ecology">
<paragraph blockId="3.[110,331,1784,1803]" box="[110,331,1784,1803]" pageId="3" pageNumber="330">
<emphasis box="[110,331,1784,1803]" italics="true" pageId="3" pageNumber="330">Habitat and ecology</emphasis>
</paragraph>
<paragraph blockId="3.[110,748,1822,1932]" lastBlockId="3.[795,1434,215,412]" pageId="3" pageNumber="330">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[110,320,1822,1844]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[110,320,1822,1844]" italics="true" pageId="3" pageNumber="330">Pandanus grayorum</emphasis>
</taxonomicName>
occurs in lowland rainforest, specifically simple to complex mesophyll vine forest, riparian gallery forest and rainforest margins with mangrove estuaries or
<emphasis box="[110,226,1910,1932]" italics="true" pageId="3" pageNumber="330">Melaleuca-</emphasis>
dominated woodlands. It occurs near large river banks and associated small creeks, moist gullies and tidal estuaries near sea level. Reproduction is both via seed and from vegetative propagules, which form a localised colony of trees. The vegetative propagules float and can be dispersed in fresh and tidal waters. They are covered in sharp curved hooks and are potentially zoochorous.
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[1120,1329,361,383]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[1120,1329,361,383]" italics="true" pageId="3" pageNumber="330">Pandanus grayorum</emphasis>
</taxonomicName>
can occur as a subcanopy, canopy or in open areas as an emergent tree.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="330" type="description">
<paragraph blockId="3.[795,909,454,473]" box="[795,909,454,473]" pageId="3" pageNumber="330">
<emphasis box="[795,909,454,473]" italics="true" pageId="3" pageNumber="330">Phenology</emphasis>
</paragraph>
<paragraph blockId="3.[795,1434,492,661]" pageId="3" pageNumber="330">Flowering occurs in October to January, with fruit developing over 1 year to ripen the following October through December. Field observations suggest that there are locally male- or female-biased stands, with an overall female bias across the whole population. Vegetative propagules are produced all-year round.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="330" type="conservation">
<paragraph blockId="3.[795,1006,702,722]" box="[795,1006,702,722]" pageId="3" pageNumber="330">
<emphasis box="[795,1006,702,722]" italics="true" pageId="3" pageNumber="330">Conservation status</emphasis>
</paragraph>
<paragraph blockId="3.[795,1433,741,1143]" pageId="3" pageNumber="330">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[795,1012,741,763]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[795,1012,741,763]" italics="true" pageId="3" pageNumber="330">Pandanus grayorum</emphasis>
</taxonomicName>
is known from few collections from several populations along rivers banks and small creeks. On the basis of collection and observation records, it has an extent of occurrence (EOO) of
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<superScript attach="left" box="[1140,1149,823,838]" fontSize="6" pageId="3" pageNumber="330">2</superScript>
and an area of occupancy (AOO) of
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<superScript attach="left" box="[978,987,852,867]" fontSize="6" pageId="3" pageNumber="330">2</superScript>
. Most records are from localities outside national parks. No threats were observed at the known localities; however, adjacent areas are road reserves and agricultural landscapes where disturbance and clearing has occurred and may continue into the future, so it is conceivable that populations may be damaged by future activities. A preliminary assessment suggests the IUCN rating of Vulnerable [VU B1ab(iii, iv)+B2ab(iii, iv)], using the IUCN Red List Categories and Criteria (International
<bibRefCitation author="International Union for Conservation of Nature" journalOrPublisher="IUCN Species Survival Commission: Gland, Switzerland; and Cambridge, UK" pageId="3" pageNumber="330" refId="ref5535" refString="International Union for Conservation of Nature (2012) ' IUCN Red List Categories and Criteria: version 3.1 ', 2 nd edn. (IUCN Species Survival Commission: Gland, Switzerland; and Cambridge, UK)" title="IUCN Red List Categories and Criteria: version 3.1 ', 2 nd edn" type="url" year="2012">Union for Conservation of Nature 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="330" type="etymology">
<paragraph blockId="3.[795,908,1185,1204]" box="[795,908,1185,1204]" pageId="3" pageNumber="330">
<emphasis box="[795,908,1185,1204]" italics="true" pageId="3" pageNumber="330">Etymology</emphasis>
</paragraph>
<paragraph blockId="3.[795,1434,1223,1625]" pageId="3" pageNumber="330">
This new species is named in honour of Joy and Bruce Gray. Bruce was formerly a technical officer with CSIRO in Atherton and has collected extensively in remote areas of northern Queensland. Over his career, Bruce has made a significant contribution to the knowledge of the northern Queensland flora through his herbarium collections, discovered many new species, and made the
<typeStatus box="[1389,1433,1399,1420]" pageId="3" pageNumber="330">type</typeStatus>
collections of 114 species, in addition to the one described here. Joy and Bruce welcomed the first author with a wonderful hospitality in Atherton in 2017 and Joy s support of Bruce has in effect enabled so many of Bruce s endeavours, including an epic trip to the tip of Cape York Peninsula in 2017. The first author finally dedicates this new
<taxonomicName box="[1331,1433,1574,1596]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1331,1433,1574,1596]" italics="true" pageId="3" pageNumber="330">Pandanus</emphasis>
</taxonomicName>
species for Bruce s 80th birthday.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="330" type="discussion">
<paragraph blockId="3.[795,859,1668,1686]" box="[795,859,1668,1686]" pageId="3" pageNumber="330">
<emphasis box="[795,859,1668,1686]" italics="true" pageId="3" pageNumber="330">Notes</emphasis>
</paragraph>
<paragraph blockId="3.[795,1433,1705,1932]" pageId="3" pageNumber="330">
Vegetative propagation in the genus
<taxonomicName authorityName="Parkinson" box="[1217,1319,1705,1727]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="genus">
<emphasis box="[1217,1319,1705,1727]" italics="true" pageId="3" pageNumber="330">Pandanus</emphasis>
</taxonomicName>
has been recorded in Australia and Madagascar: small branchlets develop near (the) apex of branches, then drop off; these act as vegetative propagules (
<bibRefCitation author="Stone BC" box="[1138,1262,1793,1815]" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="3" pageNumber="330" pagination="97 - 131" part="63" refId="ref5620" refString="Stone BC (1970) Observations on the genus Pandanus in Madagascar. Botanical Journal of the Linnean Society 63, 97 - 131." title="Observations on the genus Pandanus in Madagascar" type="journal article" year="1970">Stone 1970</bibRefCitation>
, p. 127). This vivipary is known in three species in Madagascar (
<bibRefCitation author="Guillaumet J-L" journalOrPublisher="Adansonia" pageId="3" pageNumber="330" pagination="429 - 431" part="12" refId="ref5513" refString="Guillaumet J-L (1972) Un procede de multiplication vegetative chez les Pandanus malgaches. Adansonia 12, 429 - 431." title="Un procede de multiplication vegetative chez les Pandanus malgaches" type="journal article" year="1972">Guillaumet 1972</bibRefCitation>
), but it remains rare. The active vegetative propagation in
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[795,923,1882,1903]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[795,923,1882,1903]" italics="true" pageId="3" pageNumber="330">P. grayorum</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="H. St. John" box="[972,1095,1881,1903]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="330" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[972,1095,1881,1903]" italics="true" pageId="3" pageNumber="330">P. gemmifer</emphasis>
</taxonomicName>
is so vigorous on aerial branches that they can easily be identified in the field.
</paragraph>
</subSubSection>
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<paragraph blockId="4.[403,1231,1839,1858]" box="[403,1231,1839,1858]" pageId="4" pageNumber="331">
<subSubSection box="[403,459,1839,1858]" pageId="4" pageNumber="331" type="description">
<heading bold="true" box="[403,459,1839,1858]" fontSize="8" level="2" pageId="4" pageNumber="331" reason="2">
<emphasis bold="true" box="[403,459,1839,1858]" pageId="4" pageNumber="331">Fig. 2.</emphasis>
</heading>
</subSubSection>
Distribution map of
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[656,831,1839,1858]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="4" pageNumber="331" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[656,831,1839,1858]" italics="true" pageId="4" pageNumber="331">Pandanus grayorum</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="H. St. John" box="[873,978,1839,1858]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="4" pageNumber="331" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[873,978,1839,1858]" italics="true" pageId="4" pageNumber="331">P. gemmifer</emphasis>
</taxonomicName>
in north-eastern
<collectingRegion box="[1124,1227,1839,1858]" country="Australia" name="Queensland" pageId="4" pageNumber="331">Queensland</collectingRegion>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5643384" ID-Zenodo-Dep="5643384" httpUri="https://zenodo.org/record/5643384/files/figure.png" pageId="5" pageNumber="332" startId="5.[128,159,1885,1904]" targetBox="[128,1413,210,1855]" targetPageId="5">
<paragraph blockId="5.[128,1414,1885,1931]" pageId="5" pageNumber="332">
<emphasis bold="true" box="[128,183,1885,1904]" pageId="5" pageNumber="332">Fig. 3.</emphasis>
<taxonomicName authorityName="H. St. John" box="[205,378,1885,1904]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="5" pageNumber="332" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[205,378,1885,1904]" italics="true" pageId="5" pageNumber="332">Pandanus gemmifer</emphasis>
</taxonomicName>
. A. Habitat. B. Axillary plantlets. C. Syncarp. D. Phalanges with pileus. All images of
<emphasis box="[1149,1242,1885,1904]" italics="true" pageId="5" pageNumber="332">Gray 9928</emphasis>
: by M. Callmander (A, B) and by B. Gray (C, D).
</paragraph>
</caption>
<caption ID-Table-UUID="DF1C6653143F3C521FE479ABFB3EFF41" box="[396,1237,194,213]" httpUri="http://table.plazi.org/id/DF1C6653143F3C521FE479ABFB3EFF41" pageId="6" pageNumber="333" startId="6.[396,448,194,213]" targetBox="[155,1478,239,593]" targetIsTable="true" targetPageId="6">
<paragraph blockId="6.[396,1237,194,213]" box="[396,1237,194,213]" pageId="6" pageNumber="333">
Table 1. Diagnostic characters for
<taxonomicName authorityName="H. St. John" box="[727,906,194,213]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis bold="true" box="[727,906,194,213]" italics="true" pageId="6" pageNumber="333">Pandanus gemmifer</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[917,1027,194,213]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis bold="true" box="[917,1027,194,213]" italics="true" pageId="6" pageNumber="333">P. grayorum</emphasis>
</taxonomicName>
and
<taxonomicName box="[1074,1237,194,213]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="solms-laubachii">P. solms-laubachii</taxonomicName>
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="333">
<table box="[155,1478,239,593]" gridcols="4" gridrows="13" pageId="6" pageNumber="333">
<tr box="[155,1478,239,258]" gridrow="0" pageId="6" pageNumber="333">
<th box="[155,373,239,258]" gridcol="0" gridrow="0" pageId="6" pageNumber="333">Character</th>
<th box="[423,742,239,258]" gridcol="1" gridrow="0" pageId="6" pageNumber="333">
<taxonomicName authorityName="H. St. John" box="[423,527,239,258]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[423,527,239,258]" italics="true" pageId="6" pageNumber="333">P. gemmifer</emphasis>
</taxonomicName>
</th>
<th box="[791,1078,239,258]" gridcol="2" gridrow="0" pageId="6" pageNumber="333">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[791,899,239,258]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[791,899,239,258]" italics="true" pageId="6" pageNumber="333">P. grayorum</emphasis>
</taxonomicName>
</th>
<th box="[1159,1478,239,258]" gridcol="3" gridrow="0" pageId="6" pageNumber="333">
<taxonomicName box="[1159,1319,239,258]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="solms-laubachii">
<emphasis box="[1159,1319,239,258]" italics="true" pageId="6" pageNumber="333">P. solms-laubachii</emphasis>
</taxonomicName>
</th>
</tr>
<tr box="[155,1478,281,301]" gridrow="1" pageId="6" pageNumber="333">
<th box="[155,373,281,301]" gridcol="0" gridrow="1" pageId="6" pageNumber="333">Leaf dimensions (cm)</th>
<td box="[423,742,281,301]" gridcol="1" gridrow="1" pageId="6" pageNumber="333">
170
<emphasis box="[455,466,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
230 × 5
<emphasis box="[536,547,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
6
</td>
<td box="[791,1078,281,301]" gridcol="2" gridrow="1" pageId="6" pageNumber="333">
210
<emphasis box="[823,834,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
280 × 7
<emphasis box="[904,915,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
8
</td>
<td box="[1159,1478,281,301]" gridcol="3" gridrow="1" pageId="6" pageNumber="333">
200
<emphasis box="[1191,1202,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
350 × 7
<emphasis box="[1272,1283,284,300]" italics="true" pageId="6" pageNumber="333"></emphasis>
12
</td>
</tr>
<tr box="[155,1478,308,327]" gridrow="2" pageId="6" pageNumber="333">
<th box="[155,373,308,327]" gridcol="0" gridrow="2" pageId="6" pageNumber="333">Syncarp (shape)</th>
<td box="[423,742,308,327]" gridcol="1" gridrow="2" pageId="6" pageNumber="333">Cylindric</td>
<td box="[791,1078,308,327]" gridcol="2" gridrow="2" pageId="6" pageNumber="333">Oblong to obovate</td>
<td box="[1159,1478,308,327]" gridcol="3" gridrow="2" pageId="6" pageNumber="333">Globose</td>
</tr>
<tr box="[155,1478,334,354]" gridrow="3" pageId="6" pageNumber="333">
<th box="[155,373,334,354]" gridcol="0" gridrow="3" pageId="6" pageNumber="333">Syncarp dimension (cm)</th>
<td box="[423,742,334,354]" gridcol="1" gridrow="3" pageId="6" pageNumber="333">
23
<emphasis box="[444,455,337,353]" italics="true" pageId="6" pageNumber="333"></emphasis>
25 × 10
<emphasis box="[526,537,337,353]" italics="true" pageId="6" pageNumber="333"></emphasis>
18
</td>
<td box="[791,1078,334,354]" gridcol="2" gridrow="3" pageId="6" pageNumber="333">
35
<emphasis box="[812,823,337,353]" italics="true" pageId="6" pageNumber="333"></emphasis>
40 × 18
<emphasis box="[894,905,337,353]" italics="true" pageId="6" pageNumber="333"></emphasis>
22
</td>
<td box="[1159,1478,334,354]" gridcol="3" gridrow="3" pageId="6" pageNumber="333">
25
<emphasis box="[1180,1191,337,353]" italics="true" pageId="6" pageNumber="333"></emphasis>
30 in diam.
</td>
</tr>
<tr box="[155,1478,361,380]" gridrow="4" pageId="6" pageNumber="333">
<th box="[155,373,361,380]" gridcol="0" gridrow="4" pageId="6" pageNumber="333">Phalange dimension (cm)</th>
<td box="[423,742,361,380]" gridcol="1" gridrow="4" pageId="6" pageNumber="333">
4.2
<emphasis box="[449,460,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
5.6 × 3.2
<emphasis box="[541,552,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
4.4 × 2.9
<emphasis box="[633,644,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
3.7
</td>
<td box="[791,1078,361,380]" gridcol="2" gridrow="4" pageId="6" pageNumber="333">
6
<emphasis box="[802,813,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
8(
<emphasis box="[830,841,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
9) × 4
<emphasis box="[897,908,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
5(
<emphasis box="[926,937,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
6) × 2.5
<emphasis box="[1009,1020,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
3.5
</td>
<td box="[1159,1478,361,380]" gridcol="3" gridrow="4" pageId="6" pageNumber="333">
5
<emphasis box="[1170,1181,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
7(
<emphasis box="[1198,1209,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
8) × 2
<emphasis box="[1265,1276,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
4(
<emphasis box="[1294,1305,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
5) × 2.5
<emphasis box="[1377,1388,364,380]" italics="true" pageId="6" pageNumber="333"></emphasis>
4.5
</td>
</tr>
<tr box="[155,1478,388,407]" gridrow="5" pageId="6" pageNumber="333">
<th box="[155,373,388,407]" gridcol="0" gridrow="5" pageId="6" pageNumber="333">Pileus (sides)</th>
<td box="[423,742,388,407]" gridcol="1" gridrow="5" pageId="6" pageNumber="333">Smooth to sulcate</td>
<td box="[791,1078,388,407]" gridcol="2" gridrow="5" pageId="6" pageNumber="333">Smooth to sulcate</td>
<td box="[1159,1478,388,407]" gridcol="3" gridrow="5" pageId="6" pageNumber="333">With a rib-and-groove pattern</td>
</tr>
<tr box="[155,1478,415,434]" gridrow="6" pageId="6" pageNumber="333">
<th box="[155,373,415,434]" gridcol="0" gridrow="6" pageId="6" pageNumber="333">Pileus (shape)</th>
<td box="[423,742,415,434]" gridcol="1" gridrow="6" pageId="6" pageNumber="333">Apex rounded to the stigmatic cluster</td>
<td box="[791,1078,415,434]" gridcol="2" gridrow="6" pageId="6" pageNumber="333">Flat or slightly domed at apex</td>
<td box="[1159,1478,415,434]" gridcol="3" gridrow="6" pageId="6" pageNumber="333">Convex-truncate</td>
</tr>
<tr box="[155,1478,441,460]" gridrow="7" pageId="6" pageNumber="333">
<th box="[155,373,441,460]" gridcol="0" gridrow="7" pageId="6" pageNumber="333">Pileus (sinuses) (cm)</th>
<td box="[423,742,441,460]" gridcol="1" gridrow="7" pageId="6" pageNumber="333">Apical central sinuses restricted to the</td>
<td box="[791,1078,441,460]" gridcol="2" gridrow="7" pageId="6" pageNumber="333">Narrow (~0.5) apical sinuses</td>
<td box="[1159,1478,441,460]" gridcol="3" gridrow="7" pageId="6" pageNumber="333">
Deep (~0.6
<emphasis box="[1255,1266,443,459]" italics="true" pageId="6" pageNumber="333"></emphasis>
1.2) apical sinuses
</td>
</tr>
<tr box="[155,1478,468,487]" gridrow="8" pageId="6" pageNumber="333" rowspan-0="1">
<td box="[423,742,468,487]" gridcol="1" gridrow="8" pageId="6" pageNumber="333">central stigmatic cluster</td>
<td box="[791,1078,468,487]" gridcol="2" gridrow="8" pageId="6" pageNumber="333">forming small often truncate</td>
<td box="[1159,1478,468,487]" gridcol="3" gridrow="8" pageId="6" pageNumber="333">forming cones on the whole surface</td>
</tr>
<tr box="[155,1478,494,513]" gridrow="9" pageId="6" pageNumber="333" rowspan-0="1" rowspan-1="1">
<td box="[791,1078,494,513]" gridcol="2" gridrow="9" pageId="6" pageNumber="333">pyramid on the whole surface</td>
<td box="[1159,1478,494,513]" gridcol="3" gridrow="9" pageId="6" pageNumber="333">of the apex</td>
</tr>
<tr box="[155,1478,521,540]" gridrow="10" pageId="6" pageNumber="333" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td box="[791,1078,521,540]" gridcol="2" gridrow="10" pageId="6" pageNumber="333">of the apex</td>
</tr>
<tr box="[155,1478,547,566]" gridrow="11" pageId="6" pageNumber="333">
<th box="[155,373,547,566]" gridcol="0" gridrow="11" pageId="6" pageNumber="333">Stigma (position)</th>
<td box="[423,742,547,566]" gridcol="1" gridrow="11" pageId="6" pageNumber="333">On proximal face of the central cluster</td>
<td box="[791,1078,547,566]" gridcol="2" gridrow="11" pageId="6" pageNumber="333">On the edge of a slightly truncate</td>
<td box="[1159,1478,547,566]" gridcol="3" gridrow="11" pageId="6" pageNumber="333">On summital part of cones</td>
</tr>
<tr box="[155,1478,574,593]" gridrow="12" pageId="6" pageNumber="333" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td box="[791,1078,574,593]" gridcol="2" gridrow="12" pageId="6" pageNumber="333">pyramid</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="6" pageNumber="333" type="distribution">
<paragraph blockId="6.[155,793,709,1725]" pageId="6" pageNumber="333">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[187,409,709,731]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[187,409,709,731]" italics="true" pageId="6" pageNumber="333">Pandanus grayorum</emphasis>
</taxonomicName>
occurs in close parapatry with
<taxonomicName authorityName="H. St. John" box="[155,282,738,760]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[155,282,738,760]" italics="true" pageId="6" pageNumber="333">P. gemmifer</emphasis>
</taxonomicName>
(
<figureCitation box="[301,366,738,760]" captionStart="Fig" captionStartId="4.[403,434,1839,1858]" captionTargetBox="[257,1375,210,1810]" captionTargetId="figure-1@4.[257,1376,210,1810]" captionTargetPageId="4" captionText="Fig. 2. Distribution map of Pandanus grayorum and P. gemmifer in north-eastern Queensland." figureDoi="http://doi.org/10.5281/zenodo.5643380" httpUri="https://zenodo.org/record/5643380/files/figure.png" pageId="6" pageNumber="333">Fig. 2</figureCitation>
, 3), with
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[476,608,739,760]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[476,608,739,760]" italics="true" pageId="6" pageNumber="333">P. grayorum</emphasis>
</taxonomicName>
occurring on the lower reaches of the Mulgrave, Russell, Alice, Johnstone and Moresby river systems, whereas
<taxonomicName authorityName="H. St. John" box="[508,637,797,819]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[508,637,797,819]" italics="true" pageId="6" pageNumber="333">P. gemmifer</emphasis>
</taxonomicName>
occurs on the Barron River system to the north and the Tully and Herbert river systems and nearby drainages to the south (
<figureCitation box="[710,778,855,877]" captionStart="Fig" captionStartId="4.[403,434,1839,1858]" captionTargetBox="[257,1375,210,1810]" captionTargetId="figure-1@4.[257,1376,210,1810]" captionTargetPageId="4" captionText="Fig. 2. Distribution map of Pandanus grayorum and P. gemmifer in north-eastern Queensland." figureDoi="http://doi.org/10.5281/zenodo.5643380" httpUri="https://zenodo.org/record/5643380/files/figure.png" pageId="6" pageNumber="333">Fig. 2</figureCitation>
). The two species are distinguished by the dimensions of their leaves and syncarps and the morphology of their phalanges (
<tableCitation box="[273,355,943,965]" captionStart="Table 1" captionStartId="6.[396,448,194,213]" captionTargetPageId="6" captionText="Table 1. Diagnostic characters for Pandanus gemmifer, P. grayorum and P. solms-laubachii" httpUri="http://table.plazi.org/id/DF1C6653143F3C521FE479ABFB3EFF41" pageId="6" pageNumber="333" tableUuid="DF1C6653143F3C521FE479ABFB3EFF41">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="6.[155,793,709,1725]" pageId="6" pageNumber="333">
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[187,393,972,994]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[187,393,972,994]" italics="true" pageId="6" pageNumber="333">Pandanus grayorum</emphasis>
</taxonomicName>
occurs in close sympatry with
<taxonomicName class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="solmslaubachii">
<emphasis italics="true" pageId="6" pageNumber="333">P. solmslaubachii</emphasis>
</taxonomicName>
(
<figureCitation box="[277,350,1001,1023]" captionStart="Fig" captionStartId="7.[110,141,1823,1842]" captionTargetBox="[110,1434,210,1795]" captionTargetId="figure-3@7.[107,821,210,1255]" captionTargetPageId="7" captionText="Fig. 4. Pandanus solms-laubachii. A. Habitat. B. Axillary plantlets. C. Syncarp. D. Phalanges with pileus. All images of Gray 9926: by M. Callmander (A, B) and by B. Gray (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5643386" httpUri="https://zenodo.org/record/5643386/files/figure.png" pageId="6" pageNumber="333">Fig. 4</figureCitation>
) and shares a somewhat similar tree architecture.
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[291,502,1031,1053]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[291,502,1031,1053]" italics="true" pageId="6" pageNumber="333">Pandanusgrayorum</emphasis>
</taxonomicName>
is readily identified by the presence of axillary buds developing into plantlets, which are absent in
<emphasis box="[249,655,1089,1111]" italics="true" pageId="6" pageNumber="333">
<taxonomicName box="[249,441,1089,1111]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="solms-laubachii">P. solms-laubachii</taxonomicName>
.
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[449,655,1089,1111]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">Pandanus grayorum</taxonomicName>
</emphasis>
also lacks the dense ascending rootlets present in
<taxonomicName box="[509,698,1118,1140]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="solms-laubachii">
<emphasis box="[509,698,1118,1140]" italics="true" pageId="6" pageNumber="333">P. solms-laubachii</emphasis>
</taxonomicName>
. The two species aredistinguished by thedimensions of their leaves, shape of syncarps and the morphology of their phalanges (
<tableCitation box="[162,245,1206,1228]" captionStart="Table 1" captionStartId="6.[396,448,194,213]" captionTargetPageId="6" captionText="Table 1. Diagnostic characters for Pandanus gemmifer, P. grayorum and P. solms-laubachii" httpUri="http://table.plazi.org/id/DF1C6653143F3C521FE479ABFB3EFF41" pageId="6" pageNumber="333" tableUuid="DF1C6653143F3C521FE479ABFB3EFF41">Table 1</tableCitation>
,
<figureCitation box="[261,325,1206,1228]" captionStart="Fig" captionStartId="2.[155,186,1734,1753]" captionTargetBox="[159,1480,212,1703]" captionTargetId="figure-1@2.[156,842,214,1260]" captionTargetPageId="2" captionText="Fig. 1. Pandanus grayorum. A. Habitat. B. Phalanges with pileus. C. Syncarp with axillary plantlets. D. Staminate flowers. E. Plantlet. F. Staminate inflorescence. Images of Field &amp; Zich 5017 by A. Field (A). Images of Gray 9976 (type) by B. Gray (B, C). Images of Field &amp; Zich 5019 by A. Field (D, F). Image of Field &amp; Zich 5015 by A. Field (E)." figureDoi="http://doi.org/10.5281/zenodo.5643378" httpUri="https://zenodo.org/record/5643378/files/figure.png" pageId="6" pageNumber="333">Fig. 1</figureCitation>
,
<figureCitation box="[342,357,1206,1228]" captionStart="Fig" captionStartId="7.[110,141,1823,1842]" captionTargetBox="[110,1434,210,1795]" captionTargetId="figure-3@7.[107,821,210,1255]" captionTargetPageId="7" captionText="Fig. 4. Pandanus solms-laubachii. A. Habitat. B. Axillary plantlets. C. Syncarp. D. Phalanges with pileus. All images of Gray 9926: by M. Callmander (A, B) and by B. Gray (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5643386" httpUri="https://zenodo.org/record/5643386/files/figure.png" pageId="6" pageNumber="333">4</figureCitation>
). The leaf form of
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[570,703,1207,1228]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[570,703,1207,1228]" italics="true" pageId="6" pageNumber="333">P. grayorum</emphasis>
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is more arcuate and the leaves are glossy, whereas in
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<emphasis box="[604,792,1235,1257]" italics="true" pageId="6" pageNumber="333">P. solms-laubachii</emphasis>
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the leaves are more straight and folded and are moderately glaucous.
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The genetic and morphological similarities among
<taxonomicName authorityName="H. St. John" box="[155,286,1352,1374]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="gemmifer">
<emphasis box="[155,286,1352,1374]" italics="true" pageId="6" pageNumber="333">P. gemmifer</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Callmander &amp; Gallaher &amp; Buerki &amp; Zich &amp; Field" authorityYear="2021" box="[306,442,1353,1374]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="333" phylum="Tracheophyta" rank="species" species="grayorum">
<emphasis box="[306,442,1353,1374]" italics="true" pageId="6" pageNumber="333">P. grayorum</emphasis>
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and
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<emphasis box="[504,702,1352,1374]" italics="true" pageId="6" pageNumber="333">P. solms-laubachii</emphasis>
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suggest that hybrid speciation may have played a role in the diversification of these species (
<bibRefCitation author="Gallaher T &amp; Callmander MW &amp; Buerki S &amp; Keeley SC" box="[534,778,1410,1432]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="6" pageNumber="333" pagination="20 - 32" part="83" refId="ref5408" refString="Gallaher T, Callmander MW, Buerki S, Keeley SC (2015) A long distance dispersal hypothesis for the Pandanaceae and the origins of the Pandanus tectorius complex. Molecular Phylogenetics and Evolution 83, 20 - 32. doi: 10.1016 / j. ympev. 2014.11.002" title="A long distance dispersal hypothesis for the Pandanaceae and the origins of the Pandanus tectorius complex" type="journal article" year="2015">
Gallaher
<emphasis box="[643,701,1410,1432]" italics="true" pageId="6" pageNumber="333">et al</emphasis>
. 2015
</bibRefCitation>
).
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<emphasis box="[155,373,1440,1462]" italics="true" pageId="6" pageNumber="333">Pandanus grayorum</emphasis>
</taxonomicName>
is the second taxon to be described that shares characteristics of both
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<emphasis box="[593,695,1469,1491]" italics="true" pageId="6" pageNumber="333">Pandanus</emphasis>
section
<emphasis box="[155,281,1498,1520]" italics="true" pageId="6" pageNumber="333">Austrokeura</emphasis>
</taxonomicName>
and
<taxonomicName box="[350,508,1498,1520]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="6" pageNumber="328" phylum="Tracheophyta" rank="section" section="Australibrassia">
<emphasis box="[350,508,1498,1520]" italics="true" pageId="6" pageNumber="333">Australibrassia</emphasis>
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(however, these sections were not retrieved as monophyletic groups in the molecular phylogeny, which might suggest that these characters evolved independently in this group;
<bibRefCitation author="Buerki S &amp; Callmander MW &amp; Devey DS &amp; Chappell L &amp; Gallaher T &amp; Munzinger J &amp; Haevermans T &amp; Forest F" box="[449,634,1586,1608]" journalOrPublisher="Taxon" pageId="6" pageNumber="333" pagination="1010 - 1020" part="61" refId="ref5124" refString="Buerki S, Callmander MW, Devey DS, Chappell L, Gallaher T, Munzinger J, Haevermans T, Forest F (2012) Straightening out the screw-pines: a first step in understanding phylogenetic relationships within Pandanaceae. Taxon 61, 1010 - 1020. doi: 10.1002 / tax. 615008" title="Straightening out the screw-pines: a first step in understanding phylogenetic relationships within Pandanaceae" type="journal article" year="2012">
Buerki
<emphasis box="[525,570,1586,1608]" italics="true" pageId="6" pageNumber="333">et al</emphasis>
. 2012
</bibRefCitation>
;
<bibRefCitation author="Gallaher T &amp; Callmander MW &amp; Buerki S &amp; Keeley SC" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="6" pageNumber="333" pagination="20 - 32" part="83" refId="ref5408" refString="Gallaher T, Callmander MW, Buerki S, Keeley SC (2015) A long distance dispersal hypothesis for the Pandanaceae and the origins of the Pandanus tectorius complex. Molecular Phylogenetics and Evolution 83, 20 - 32. doi: 10.1016 / j. ympev. 2014.11.002" title="A long distance dispersal hypothesis for the Pandanaceae and the origins of the Pandanus tectorius complex" type="journal article" year="2015">
Gallaher
<emphasis box="[741,786,1586,1608]" italics="true" pageId="6" pageNumber="333">et al</emphasis>
. 2015
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). Both phylogenomic and population genetic studies, coupled with field observations, are recommended to clarify species delimitations within this challenging group of
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.
</paragraph>
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<emphasis box="[155,490,1794,1813]" italics="true" pageId="6" pageNumber="333">Additional specimen examined</emphasis>
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.
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:
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<collectorName box="[414,503,1859,1878]" pageId="6" pageNumber="333">S.T. Blake</collectorName>
23631
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(
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</materialsCitation>
;
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<collectingMunicipality pageId="6" pageNumber="333">Bellenden Kerlanding</collectingMunicipality>
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NP,
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E
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,
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<elevation box="[639,671,1886,1905]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="333" unit="m" value="5.0">5 m</elevation>
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,fr.,
<emphasis italics="true" pageId="6" pageNumber="333">
<collectorName box="[706,773,1886,1905]" pageId="6" pageNumber="333">A.Field</collectorName>
&amp;
<collectorName box="[155,217,1912,1931]" pageId="6" pageNumber="333">F. Zich</collectorName>
5015
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(
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,
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,
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</materialsCitation>
;
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<collectingMunicipality box="[409,512,1912,1931]" pageId="6" pageNumber="333">Polly Creek</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03CA87CD14383C521D427D62FBEAFC68:8EBC6000143F3C521C607E11FB30FD4C" box="[520,1243,709,1931]" country="Australia" municipality="Polly Creek" name="Garadunga, in road reserve near junction of Polly Creek road and Jubilee Road" pageId="6" pageNumber="333" stateProvince="Queensland">Garadunga, in road reserve near junction of Polly Creek road and Jubilee Road</location>
,
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,
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<elevation box="[840,886,735,754]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="333" unit="m" value="12.0">12 m</elevation>
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, fr.,
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<collectorName box="[924,993,735,754]" pageId="6" pageNumber="333">A. Field</collectorName>
&amp;
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5017
</emphasis>
(
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,
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)
</materialsCitation>
;
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<collectingMunicipality box="[1275,1392,735,754]" pageId="6" pageNumber="333">Nind s Creek</collectingMunicipality>
,
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,
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S
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,
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1
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2 m
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, fr.,
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<collectorName box="[1190,1264,789,808]" pageId="6" pageNumber="333">A. Field</collectorName>
&amp;
<collectorName box="[1300,1366,789,808]" pageId="6" pageNumber="333">F. Zich</collectorName>
5018
</emphasis>
(
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,
<collectionCode box="[840,884,815,834]" collectionName="CNS" pageId="6" pageNumber="333">CNS</collectionCode>
,
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;
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,
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<elevation box="[1439,1474,842,861]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="333" unit="m" value="9.0">9 m</elevation>
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, staminate fl.,
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&amp;
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5019
</emphasis>
(
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,
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,
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</materialsCitation>
;
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,
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146
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,
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, fr.,
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<collectorName box="[1070,1172,895,914]" pageId="6" pageNumber="333">T. Gallaher</collectorName>
337
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(
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,
<collectionCode box="[1284,1339,895,914]" pageId="6" pageNumber="333">HAW</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397196305" collectingDate="2017-12-13" collectionCode="BRI, CNS, G" collectorName="B. Gray" country="Australia" elevation="7" location="Russell River, below Russell River boat camp" pageId="6" pageNumber="333" specimenCount="1" stateProvince="Queensland">
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,
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17
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,
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145
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<superScript attach="none" box="[1307,1364,918,940]" fontSize="6" pageId="6" pageNumber="333">5605400</superScript>
E
</geoCoordinate>
,
<quantity box="[1395,1435,921,940]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="333" unit="m" value="7.0">
<elevation box="[1395,1435,921,940]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="333" unit="m" value="7.0">7 m</elevation>
</quantity>
,
<date pageId="6" pageNumber="333" value="2017-12-13">
<collectingDate pageId="6" pageNumber="333" value="2017-12-13">13. XII.2017</collectingDate>
</date>
, fr.,
<emphasis box="[966,1088,948,967]" italics="true" pageId="6" pageNumber="333">
<collectorName box="[966,1037,948,967]" pageId="6" pageNumber="333">B. Gray</collectorName>
9955
</emphasis>
(
<collectionCode box="[1103,1142,948,967]" collectionName="BRI" pageId="6" pageNumber="333">BRI</collectionCode>
,
<collectionCode box="[1153,1197,948,967]" collectionName="CNS" pageId="6" pageNumber="333">CNS</collectionCode>
,
<collectionCode box="[1209,1227,948,967]" pageId="6" pageNumber="333">G</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3397196303" collectingDate="2003-10-29" collectionCode="BRI, CANB, CNS" collectorName="B. Gray" country="Australia" elevation="3" location="Russell River road" municipality="Russell River" pageId="6" pageNumber="333" specimenCount="1" stateProvince="Queensland">
<collectingMunicipality box="[1245,1365,948,967]" pageId="6" pageNumber="333">Russell River</collectingMunicipality>
NP,
<location LSID="urn:lsid:plazi:treatment:03CA87CD14383C521D427D62FBEAFC68:8EBC6000143F3C521BEE7ADDFC4CFC76" country="Australia" municipality="Russell River" name="Russell River road" pageId="6" pageNumber="333" stateProvince="Queensland">Russell River road</location>
,
<geoCoordinate box="[945,1045,971,994]" pageId="6" pageNumber="333">
17
<superScript attach="right" box="[967,974,971,984]" fontSize="6" pageId="6" pageNumber="333">°</superScript>
<superScript attach="none" box="[974,1031,971,994]" fontSize="6" pageId="6" pageNumber="333">1602800</superScript>
S
</geoCoordinate>
,
<geoCoordinate box="[1056,1169,971,994]" pageId="6" pageNumber="333">
145
<superScript attach="none" box="[1088,1152,971,994]" fontSize="6" pageId="6" pageNumber="333">°5605000</superScript>
E
</geoCoordinate>
,
<quantity box="[1179,1215,975,994]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="333" unit="m" value="3.0">
<elevation box="[1179,1215,975,994]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="333" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<date box="[1225,1317,975,994]" pageId="6" pageNumber="333" value="2003-10-29">
<collectingDate box="[1225,1317,975,994]" pageId="6" pageNumber="333" value="2003-10-29">29.X.2003</collectingDate>
</date>
, fr.,
<emphasis box="[1359,1478,975,994]" italics="true" pageId="6" pageNumber="333">
<collectorName box="[1359,1428,975,994]" pageId="6" pageNumber="333">B. Gray</collectorName>
8831
</emphasis>
(
<collectionCode box="[847,886,1001,1020]" collectionName="BRI" pageId="6" pageNumber="333">BRI</collectionCode>
,
<collectionCode box="[897,959,1001,1020]" collectionName="CANB" pageId="6" pageNumber="333">CANB</collectionCode>
,
<collectionCode box="[969,1014,1001,1020]" collectionName="CNS" pageId="6" pageNumber="333">CNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>