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<mods:title id="336C32DD93D5756F268A2DA5680B8976">Ghost shrimp Calliax de Saint Laurent, 1973 (Decapoda: Axiidea: Callianassidae) in the fossil record: systematics, palaeoecology and palaeobiogeography</mods:title>
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<mods:namePart id="26C84FE3D227A9765B4B3984D0ABEF29">Hyžný, Matúš</mods:namePart>
<mods:affiliation id="8707259D7F1125E113688EEE1F5A7C94">Geological-Paleontological Department, Natural History Museum, Vienna, Burgring 7, A- 1010 Vienna, Austria. &amp; Comenius University, Faculty of Natural Sciences, Department of Geology and Palaeontology, Mlynská dolina G 1, SVK- 842 15 Bratislava, Slovakia</mods:affiliation>
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<mods:affiliation id="035230293768CAD2D3978DF195C6E45B">Ljubljanska cesta 4 j, 1241 Kamnik, Slovenia. E-mail: rok. gasparic @ gmail. com</mods:affiliation>
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<heading id="D09481B0FFC6C81200A2FEE5FCC3B7D2" bold="true" box="[151,845,367,394]" fontSize="11" level="1" pageId="8" pageNumber="45" reason="1">
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<taxonomicName id="4C634D5FFFC6C81200A2FEE5FD31B7D2" authority="(A. Milne Edwards, 1860)" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[151,703,367,394]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC6C81200A2FEE5FEF8B7D1" bold="true" box="[151,374,367,393]" italics="true" pageId="8" pageNumber="45">Calliax michelottii</emphasis>
(A.
<bibRefCitation id="EFF24B2DFFC6C8120199FEFAFD39B7D2" author="Milne Edwards, A." box="[428,695,368,394]" pageId="8" pageNumber="45" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC6C81202F2FEFAFCC3B7D2" box="[711,845,368,394]" pageId="8" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="C3796557FFC6C81200A2FE18FD4FB7F3" box="[151,705,402,427]" pageId="8" pageNumber="45" type="description">
<paragraph id="8BDC36DCFFC6C81200A2FE18FD4FB7F3" blockId="8.[151,845,367,427]" box="[151,705,402,427]" pageId="8" pageNumber="45">
(
<figureCitation id="13582A59FFC6C81200AAFE18FE91B7F3" box="[159,287,402,427]" captionStart="FIGURE 5" captionStartId="7.[151,249,1290,1312]" captionTargetBox="[246,1336,525,1260]" captionTargetId="figure-517@7.[238,1348,519,1265]" captionTargetPageId="7" captionText="FIGURE 5. Callianassa michelottii A. Milne Edwards, 1860: holotype (MNHN-F-B32690): AB, refigured from A. Milne Edwards (1860: pl. 14, fig. 3); CD, photo of the holotype. A, C=inner surface; B, D=outer surface. Arrows indicate ridges. Photo by Lilian Cazes (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4919934" httpUri="https://zenodo.org/record/4919934/files/figure.png" pageId="8" pageNumber="45">Figs 5AD</figureCitation>
,
<figureCitation id="13582A59FFC6C812011FFE18FEE0B7F3" box="[298,366,402,427]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="8" pageNumber="45">6AL</figureCitation>
,
<figureCitation id="13582A59FFC6C812014CFE18FE4BB7F3" box="[377,453,402,427]" captionStart="FIGURE 7" captionStartId="10.[151,249,1261,1283]" captionTargetBox="[152,1435,416,1236]" captionTargetId="figure-467@10.[152,1435,416,1236]" captionTargetPageId="10" captionText="FIGURE 7. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Oligocene of the European plate: isolated propodi (AK) and carpus (LM). AB, NHMW 1874/0029/1154 a; C, D, G, NHMW 1874/0029/1154 a, G shows the specimen under a low-angle light to show the ridges on the fixed finger; EF, NHMW 1874/0029/1154 c; H, NHMW 1874/ 0029/1154 d; I, NHMW 1874/0029/1155; JK, NHMW 1874/0029/1154 e; LM, NHMW 1874/0029/1154 f. Specimens in AH, JM are from Sternberg (Germany), specimen in I is from Pinnow bei Schwerin (Germany). All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919942" httpUri="https://zenodo.org/record/4919942/files/figure.png" pageId="8" pageNumber="45">7AM</figureCitation>
,
<figureCitation id="13582A59FFC6C81201E5FE18FD9AB7F3" box="[464,532,402,427]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="8" pageNumber="45">8AL</figureCitation>
,
<figureCitation id="13582A59FFC6C812022AFE18FDD5B7F3" box="[543,603,402,427]" captionStart="FIGURE 9" captionStartId="13.[151,249,1148,1170]" captionTargetBox="[151,1436,339,1124]" captionTargetId="figure-285@13.[150,1436,339,1124]" captionTargetPageId="13" captionText="FIGURE 9. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Middle Miocene of Kamnik, Slovenia. A, RGA/SMNH 0754, right propodus; B, RGA/SMNH 0779, left propodus; C, RGA/SMNH 0864, right propodus; D, RGA/ SMNH 0773, left propodus; E, RGA/SMNH 1191, left major chela consisting of propodus, carpus and merus; F, RGA/SMNH 1075, left propodus articulated with dactylus; G, RGA/SMNH 1187, left propodus articulated with dactylus; H, RGA/SMNH 1069, left major chela consisting of dactylus, propodus and carpus; I, GBA 2009/014/0027, right propodus. All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919950" httpUri="https://zenodo.org/record/4919950/files/figure.png" pageId="8" pageNumber="45">9AI</figureCitation>
,
<figureCitation id="13582A59FFC6C8120252FE18FD37B7F3" box="[615,697,402,427]" captionStart="FIGURE 10" captionStartId="13.[151,249,1728,1750]" captionTargetBox="[157,1427,1404,1703]" captionTargetId="figure-294@13.[150,1437,1395,1705]" captionTargetPageId="13" captionText="FIGURE 10. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., reconstruction: A, major chela; B, minor chela (based on Fritsch 1871: pl. 17, fig. 14). Both chelae are depicted in outer lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4919954" httpUri="https://zenodo.org/record/4919954/files/figure.png" pageId="8" pageNumber="45">10AB</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB7311F2FFC6C81200A2FE50FA8CB7A8" box="[151,1282,474,496]" pageId="8" pageNumber="45">
<treatmentCitation id="0AC210CDFFC6C81200A2FE50FD3DB7A8" author="Milne Edwards, A." box="[151,691,474,496]" page="341" pageId="8" pageNumber="45" year="1860">
<taxonomicName id="4C634D5FFFC6C81200A2FE50FD3DB7A8" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authority="Michelotti A. Milne Edwards, 1860: 341" authorityName="Michelotti A. Milne Edwards" authorityYear="1860" box="[151,691,474,496]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC6C81200A2FE50FEF0B7A8" box="[151,382,474,496]" italics="true" pageId="8" pageNumber="45">Callianassa Michelotti</emphasis>
A.
<bibRefCitation id="EFF24B2DFFC6C8120196FE50FD3DB7A8" author="Milne Edwards, A." box="[419,691,474,496]" pageId="8" pageNumber="45" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860: 341</bibRefCitation>
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, pl. 14, fig. 3; A.
<treatmentCitation id="0AC210CDFFC6C8120353FE50FBF8B7A8" author="Milne Edwards, A." box="[870,1142,474,496]" page="210" pageId="8" pageNumber="45" year="1860">
<bibRefCitation id="EFF24B2DFFC6C8120353FE50FBF8B7A8" author="Milne Edwards, A." box="[870,1142,474,496]" pageId="8" pageNumber="45" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860: 210</bibRefCitation>
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, pl. 14, fig. 3.
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</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FE73FD26B457" blockId="8.[151,1436,474,926]" box="[151,680,505,527]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FE73FD26B457" box="[151,680,505,527]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200A2FE73FE1FB457" box="[151,401,505,527]" italics="true" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C81200A2FE73FEF9B457" authorityName="Milne-Edwards" authorityYear="1860" box="[151,375,505,527]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelotti">Callianassa michelotti</taxonomicName>
</emphasis>
<treatmentCitation id="0AC210CDFFC6C81201A7FE73FD2AB457" author="Schweitzer, C. E. &amp; Feldmann, R. M. &amp; Garassino, A. &amp; Karasawa, H. &amp; Schweigert, G." box="[402,676,505,527]" page="35" pageId="8" pageNumber="45" year="2010">
<bibRefCitation id="EFF24B2DFFC6C81201A7FE73FD2AB457" author="Schweitzer, C. E. &amp; Feldmann, R. M. &amp; Garassino, A. &amp; Karasawa, H. &amp; Schweigert, G." box="[402,676,505,527]" pageId="8" pageNumber="45" pagination="1 - 222" refId="ref15270" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222. http: // dx. doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
Schweitzer
<emphasis id="B917EACEFFC6C812023CFE70FDBBB457" box="[521,565,505,527]" italics="true" pageId="8" pageNumber="45">et al</emphasis>
., 2010: 35
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.
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</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FD92FD22B414" blockId="8.[151,1436,474,926]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FD92FD22B414" pageId="8" pageNumber="45">
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<taxonomicName id="4C634D5FFFC6C81200A2FD92FE0AB476" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authority="Michelottii" authorityName="Michelotti A. Milne Edwards" authorityYear="1860" box="[151,388,536,558]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">Callianassa Michelottii</taxonomicName>
</emphasis>
<treatmentCitation id="0AC210CDFFC6C81201AAFD92FDEEB476" author="Fritsch, K. von" box="[415,608,536,558]" page="691" pageId="8" pageNumber="45" year="1871">
<bibRefCitation id="EFF24B2DFFC6C81201AAFD92FDEEB476" author="Fritsch, K. von" box="[415,608,536,558]" pageId="8" pageNumber="45" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch, 1871: 691</bibRefCitation>
</treatmentCitation>
, pl. 17, figs. 513;
<treatmentCitation id="0AC210CDFFC6C8120306FD92FC74B476" author="Noetling, F." box="[819,1018,536,558]" page="84" pageId="8" pageNumber="45" year="1886">
<bibRefCitation id="EFF24B2DFFC6C8120306FD92FC74B476" author="Noetling, F." box="[819,1018,536,558]" pageId="8" pageNumber="45" pagination="32 - 34" refId="ref14502" refString="Noetling, F. (1886) Uber Crustaceenreste aus dem Oberoligocanen Sternberger Gestein. Sitzingsberichte der Gesselschaft naturforschender Freunde Berlin, 1886, 32 - 34." type="journal article" year="1886">Noetling, 1886: 84</bibRefCitation>
</treatmentCitation>
, pl. 5, fig. 4;
<treatmentCitation id="0AC210CDFFC6C81204A5FD92FAC1B476" author="Crema, C." box="[1168,1359,536,558]" page="667" pageId="8" pageNumber="45" year="1895">
<bibRefCitation id="EFF24B2DFFC6C81204A5FD92FAC1B476" author="Crema, C." box="[1168,1359,536,558]" pageId="8" pageNumber="45" pagination="664 - 681" refId="ref11114" refString="Crema, C. (1895) Sopra alcuni decapodi terziarii del Piemonte. Atti Reale Accademia delle Scienze di Torino, 30, 664 - 681." type="journal article" year="1895">Crema, 1895: 667</bibRefCitation>
</treatmentCitation>
, fig. 3;
<treatmentCitation id="0AC210CDFFC6C81200F2FDBCFE5CB414" author="Glaessner, M. F." box="[199,466,566,588]" page="167" pageId="8" pageNumber="45" year="1928">
<bibRefCitation id="EFF24B2DFFC6C81200F2FDBCFE5CB414" author="Glaessner, M. F." box="[199,466,566,588]" pageId="8" pageNumber="45" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner, 1928: 167168</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AC210CDFFC6C81201E9FDBCFD29B414" author="Glaessner, M. F." box="[476,679,566,588]" page="84" pageId="8" pageNumber="45" year="1929">
<bibRefCitation id="EFF24B2DFFC6C81201E9FDBCFD29B414" author="Glaessner, M. F." box="[476,679,566,588]" pageId="8" pageNumber="45" pagination="1 - 464" refId="ref12515" refString="Glaessner, M. F. (1929) Crustacea Decapoda. In. Pompeckj, F. J. (Ed.) Fossilium catalogus, 1. Animalium, 41, pp. 1 - 464. [W. Junk, Berlin]" type="journal article" year="1929">Glaessner, 1929: 84</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FDDFFD02B433" blockId="8.[151,1436,474,926]" box="[151,652,597,619]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FDDFFD02B433" box="[151,652,597,619]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200A2FDDFFE1FB433" box="[151,401,597,619]" italics="true" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C81200A2FDDFFEF9B433" box="[151,375,597,619]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelotii">Callianassa michelotii</taxonomicName>
</emphasis>
<treatmentCitation id="0AC210CDFFC6C81201A7FDDFFD06B433" author="Beurlen, K." box="[402,648,597,619]" page="111" pageId="8" pageNumber="45" year="1931">
<bibRefCitation id="EFF24B2DFFC6C81201A7FDDFFD06B433" author="Beurlen, K." box="[402,648,597,619]" pageId="8" pageNumber="45" pagination="111 - 112" refId="ref10830" refString="Beurlen, K. (1931) Die Decapodenfauna aus dem Rupelton des Mainzer Beckens. Senckenbergiana, 13, 111 - 112." type="journal article" year="1931">Beurlen, 1931: 111112</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FDFEFC19B4D2" blockId="8.[151,1436,474,926]" box="[151,919,628,650]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FDFEFC19B4D2" box="[151,919,628,650]" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C8120097FDFEFE95B4D2" ID-CoL="f45854b2-a302-4361-bd40-0423151d5e6a" authorityName="Michelotti A. Milne Edwards" authorityYear="1860" box="[162,283,628,650]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC6C8120097FDFEFE95B4D2" box="[162,283,628,650]" italics="true" pageId="8" pageNumber="45">Callianassa</emphasis>
</taxonomicName>
<taxonomicName id="4C634D5FFFC6C8120119FDFEFE1FB4D2" authorityName="Milne-Edwards" authorityYear="1860" box="[300,401,628,650]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelotti">
<emphasis id="B917EACEFFC6C8120119FDFEFE1FB4D2" box="[300,401,628,650]" italics="true" pageId="8" pageNumber="45">michelotti</emphasis>
</taxonomicName>
<treatmentCitation id="0AC210CDFFC6C8120199FDFEFD0FB4D2" author="Polkowsky, S." box="[428,641,628,650]" page="17" pageId="8" pageNumber="45" year="2005">
<bibRefCitation id="EFF24B2DFFC6C8120199FDFEFD0FB4D2" author="Polkowsky, S." box="[428,641,628,650]" pageId="8" pageNumber="45" pagination="1 - 126" refId="ref14603" refString="Polkowsky, S. (2005) Decapode Krebse aus dem oberoligozanen Sternberger Gestein von Kobrow (Mecklenburg). Tassados, 1, 1 - 126." type="journal article" year="2005">Polkowsky, 2005: 17</bibRefCitation>
</treatmentCitation>
, figs. 813, pl. 1, figs. 18.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FD18FCC9B4F0" blockId="8.[151,1436,474,926]" box="[151,839,658,680]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FD18FCC9B4F0" box="[151,839,658,680]" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C81200A2FD18FE2CB4F0" authorityName="A. Milne Edwards" authorityYear="1860" box="[151,418,658,680]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC6C81200A2FD18FE9EB4F0" box="[151,272,658,680]" italics="true" pageId="8" pageNumber="45">Callianassa</emphasis>
cf.
<emphasis id="B917EACEFFC6C8120103FD18FE2CB4F0" box="[310,418,658,680]" italics="true" pageId="8" pageNumber="45">michelottii</emphasis>
</taxonomicName>
<treatmentCitation id="0AC210CDFFC6C8120188FD18FD49B4F0" author="Moths, H. &amp; Montag, A." box="[445,711,658,680]" page="7" pageId="8" pageNumber="45" year="2002">
<bibRefCitation id="EFF24B2DFFC6C8120188FD18FD49B4F0" author="Moths, H. &amp; Montag, A." box="[445,711,658,680]" pageId="8" pageNumber="45" pagination="3 - 30" refId="ref14327" refString="Moths, H. &amp; Montag, A. (2002) Tertiare dekapode Krebse aus Geschieben und dem Anstehenden Noddeutschlands und Danemarks. Der Geschiebesammler, 35, 3 - 30." type="journal article" year="2002">Moths &amp; Montag, 2002: 7</bibRefCitation>
</treatmentCitation>
, pl. 4, fig. 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FD3BFCBEB49F" blockId="8.[151,1436,474,926]" box="[151,816,689,711]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FD3BFCBEB49F" box="[151,816,689,711]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200A2FD3BFEAFB49F" box="[151,289,689,711]" italics="true" pageId="8" pageNumber="45">ʻCallianassaʼ</emphasis>
<treatmentCitation id="0AC210CDFFC6C8120112FD3BFDB8B49F" author="Muller, P." box="[295,566,689,711]" page="53" pageId="8" pageNumber="45" year="1984">
<taxonomicName id="4C634D5FFFC6C8120112FD3BFDB8B49F" authority="Muller, 1984: 53" authorityName="Muller" authorityPageNumber="53" authorityYear="1984" box="[295,566,689,711]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC6C8120112FD3BFE09B49F" box="[295,391,689,711]" italics="true" pageId="8" pageNumber="45">szobensis</emphasis>
<bibRefCitation id="EFF24B2DFFC6C81201B8FD3BFDB8B49F" author="Muller, P." box="[397,566,689,711]" pageId="8" pageNumber="45" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984: 53</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(partim), pl. 7, figs.34.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FD5AFD2BB4BE" blockId="8.[151,1436,474,926]" box="[151,677,720,742]" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C81200A2FD5AFEF8B4BE" authorityName="Muller" authorityYear="1984" box="[151,374,720,742]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC6C81200A2FD5AFEF8B4BE" box="[151,374,720,742]" italics="true" pageId="8" pageNumber="45">Callianassa szobensis</emphasis>
</taxonomicName>
<bibRefCitation id="EFF24B2DFFC6C81201A5FD5AFDF4B4BE" author="Schweitzer, C. E. &amp; Feldmann, R. M. &amp; Garassino, A. &amp; Karasawa, H. &amp; Schweigert, G." box="[400,634,720,742]" pageId="8" pageNumber="45" pagination="1 - 222" refId="ref15270" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10, 1 - 222. http: // dx. doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
Schweitzer
<emphasis id="B917EACEFFC6C8120233FD5BFDBCB4BE" box="[518,562,720,742]" italics="true" pageId="8" pageNumber="45">et al</emphasis>
., 2010
</bibRefCitation>
, 37.
</paragraph>
</subSubSection>
<subSubSection id="C3796557FFC6C81200A2FD65FD0CB55C" box="[151,642,750,772]" pageId="8" pageNumber="45" type="description">
<paragraph id="8BDC36DCFFC6C81200A2FD65FD0CB55C" blockId="8.[151,1436,474,926]" box="[151,642,750,772]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C8120097FD64FE01B55C" box="[162,399,750,772]" italics="true" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C8120097FD64FEACB55C" authorityName="Michelotti A. Milne Edwards" authorityYear="1860" box="[162,290,750,772]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">Callianassa</taxonomicName>
<taxonomicName id="4C634D5FFFC6C812011AFD64FE01B55C" authorityName="Muller" authorityYear="1984" box="[303,399,750,772]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="szobensis">szobensis</taxonomicName>
</emphasis>
<bibRefCitation id="EFF24B2DFFC6C812019FFD64FDA0B55C" author="Hyzny, M." box="[426,558,750,772]" pageId="8" pageNumber="45" pagination="37 - 46" refId="ref13434" refString="Hyzny, M. (2011) In situ mud shrimps (Decapoda: Axiidea: Callianassidae) preserved within their burrows from the middle Miocene of the Central Paratethys. Bulletin of the Mizunami Fossil Museum, 37, 37 - 46." type="journal article" year="2011">Hyžný, 2011</bibRefCitation>
: table 2.
</paragraph>
</subSubSection>
<subSubSection id="C3796557FFC6C81200A2FC87FD40B583" pageId="8" pageNumber="45" type="reference_group">
<paragraph id="8BDC36DCFFC6C81200A2FC87FCDBB57B" blockId="8.[151,1436,474,926]" box="[151,853,781,803]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FC87FCDBB57B" box="[151,853,781,803]" pageId="8" pageNumber="45">
<taxonomicName id="4C634D5FFFC6C81200A2FC87FEBEB57B" box="[151,304,781,803]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B917EACEFFC6C81200A2FC87FE9EB57B" box="[151,272,781,803]" italics="true" pageId="8" pageNumber="45">Callianassa</emphasis>
sp.
</taxonomicName>
—Houša
<emphasis id="B917EACEFFC6C81201A6FC84FE29B57B" box="[403,423,782,803]" italics="true" pageId="8" pageNumber="45">in</emphasis>
<treatmentCitation id="0AC210CDFFC6C8120198FC87FD10B57B" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." box="[429,670,781,803]" page="734" pageId="8" pageNumber="45" year="1965">
<bibRefCitation id="EFF24B2DFFC6C8120198FC87FD10B57B" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." box="[429,670,781,803]" pageId="8" pageNumber="45" pagination="704 - 746" refId="ref15452" refString="Spinar, Z., Cepek, P., Boucek, B., Eliasova-Frajova, H., Havlicek, V., Horny, R., Housa, V., Kalabis, V., Kukalova, J., Marek, L., Nekvasilova, O., Pokorny, V., Prokop, R., Pribyl, A., Ruzicka, B., Snajdr, M., Stemprokova, D. &amp; Zikmundova, J. (1965) Systematicka paleontologie bezobratlych. Academia, Praha, 1049 pp. [pp. 704 - 746.]" type="book chapter" year="1965">
Špinar
<emphasis id="B917EACEFFC6C81201C0FC84FDACB57B" box="[501,546,781,803]" italics="true" pageId="8" pageNumber="45">et al</emphasis>
., 1965: 734
</bibRefCitation>
</treatmentCitation>
, figs. X-184185.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FCA6FC91B51A" blockId="8.[151,1436,474,926]" box="[151,799,812,834]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FCA6FC91B51A" box="[151,799,812,834]" pageId="8" pageNumber="45">
?
<taxonomicName id="4C634D5FFFC6C812009DFCA6FE07B51A" authority="Michelotti" authorityName="Michelotti" box="[168,393,812,834]" class="Malacostraca" family="Callianassidae" genus="Calianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC6C812009DFCA6FE07B51A" box="[168,393,812,834]" italics="true" pageId="8" pageNumber="45">Calianassa Michelotti</emphasis>
</taxonomicName>
<treatmentCitation id="0AC210CDFFC6C8120197FCA6FD11B51A" author="Wagner-Klett, W." box="[418,671,812,834]" page="107" pageId="8" pageNumber="45" year="1919">
<bibRefCitation id="EFF24B2DFFC6C8120197FCA6FD11B51A" author="Wagner-Klett, W." box="[418,671,812,834]" pageId="8" pageNumber="45" pagination="73 - 118" refId="ref15729" refString="Wagner-Klett, W. (1919) Das Tertiar von Wiesloch in Baden. Ein Beitrag zu seiner tektonischen, stratigraphischen und palaontologischen Kenntnis. Jahresberichte und Mitteilungen des Oberrheinischen Geologischen Vereins, 8, 73 - 118. http: // dx. doi. org / 10.1127 / jmogv / 8 / 1919 / 73" type="journal article" year="1919">Wagner-Klett, 1919: 107</bibRefCitation>
</treatmentCitation>
, pl. 2, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FCC0FCC0B538" blockId="8.[151,1436,474,926]" box="[151,846,842,864]" pageId="8" pageNumber="45">
?
<emphasis id="B917EACEFFC6C812009DFCC0FEBCB538" box="[168,306,842,864]" italics="true" pageId="8" pageNumber="45">ʻCallianassaʼ</emphasis>
<taxonomicName id="4C634D5FFFC6C812010AFCC0FDC0B538" authority="Muller, 1984: 53" authorityName="Muller" authorityPageNumber="53" authorityYear="1984" box="[319,590,842,864]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC6C812010AFCC0FE11B538" box="[319,415,842,864]" italics="true" pageId="8" pageNumber="45">szobensis</emphasis>
<bibRefCitation id="EFF24B2DFFC6C8120190FCC0FDC0B538" author="Muller, P." box="[421,590,842,864]" pageId="8" pageNumber="45" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984: 53</bibRefCitation>
</taxonomicName>
(partim); pl. 7, figs. 56.
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FCE3FCEEB527" blockId="8.[151,1436,474,926]" box="[151,864,873,895]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FCE3FCEEB527" box="[151,864,873,895]" pageId="8" pageNumber="45">
?
<taxonomicName id="4C634D5FFFC6C812009DFCE3FECFB527" box="[168,321,873,895]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B917EACEFFC6C812009DFCE3FEAFB527" box="[168,289,873,895]" italics="true" pageId="8" pageNumber="45">Callianassa</emphasis>
sp.
</taxonomicName>
<treatmentCitation id="0AC210CDFFC6C812016EFCE3FD10B527" author="Philippe, M. &amp; Secretan, S." box="[347,670,873,895]" page="128" pageId="8" pageNumber="45" year="1971">
<bibRefCitation id="EFF24B2DFFC6C812016EFCE3FD10B527" author="Philippe, M. &amp; Secretan, S." box="[347,670,873,895]" pageId="8" pageNumber="45" pagination="117 - 134" refId="ref14530" refString="Philippe, M. &amp; Secretan, S. (1971) Crustaces decapods du Burdigalien des Courennes (Vaucluse). Annales de Paleontologie (Invertebres), 57, 117 - 134." type="journal article" year="1971">Philippe &amp; Secretan, 1971: 128</bibRefCitation>
</treatmentCitation>
, pl. C, figs. 1314.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FC02FCF3B5C6" blockId="8.[151,1436,474,926]" box="[151,893,904,926]" pageId="8" pageNumber="45">
? Eine nicht bestimmbare Hand—
<bibRefCitation id="EFF24B2DFFC6C81201DAFC02FD44B5C6" author="Lorenthey, I." box="[495,714,904,926]" pageId="8" pageNumber="45" pagination="202 - 259" refId="ref14078" refString="Lorenthey, I. (1907) Beitrage zur Tertiaren Dekapodenfauna Sardiniens. Mathematischen und Naturwissenchaftlichen Berichte aus Ungarn, 25, 202 - 259, 2 plates." type="book chapter" year="1907">Lőrenthey, 1907: 212</bibRefCitation>
, 222, pl. 1, fig. 5.
</paragraph>
<paragraph id="8BDC36DCFFC6C81200A2FC4CFD40B583" blockId="8.[151,718,965,987]" box="[151,718,965,987]" pageId="8" pageNumber="45">
<treatmentCitationGroup id="AB7311F2FFC6C81200A2FC4CFD40B583" box="[151,718,965,987]" pageId="8" pageNumber="45">
non
<treatmentCitation id="0AC210CDFFC6C81200F6FC4FFDFAB583" author="Muller, P." box="[195,628,965,987]" page="7" pageId="8" pageNumber="45" year="1993">
<taxonomicName id="4C634D5FFFC6C81200F6FC4FFDFAB583" authority="Muller, 1993: 7" authorityName="Muller" authorityPageNumber="7" authorityYear="1993" box="[195,628,965,987]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC6C81200F6FC4FFEB2B583" box="[195,316,965,987]" italics="true" pageId="8" pageNumber="45">Callianassa</emphasis>
cf.
<emphasis id="B917EACEFFC6C8120156FC4FFE41B583" box="[355,463,965,987]" italics="true" pageId="8" pageNumber="45">michelottii</emphasis>
<bibRefCitation id="EFF24B2DFFC6C81201E0FC4FFDFAB583" author="Muller, P." box="[469,628,965,987]" pageId="8" pageNumber="45" pagination="1 - 39" refId="ref14453" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller, 1993: 7</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 3A.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3796557FFC6C81200A2FB8DFB9DB2A5" pageId="8" pageNumber="45" type="diagnosis">
<paragraph id="8BDC36DCFFC6C81200A2FB8DFB9DB2A5" blockId="8.[151,1437,1031,2013]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200A2FB8DFE9FB278" bold="true" box="[151,273,1031,1056]" inLineHeading="true" pageId="8" pageNumber="45" reason="1">Diagnosis.</emphasis>
Major cheliped massive; merus longer than high (L/H=2.2) with convex upper margin, carpus higher than long (L/H=0.45), about half the length of manus or shorter, upper margin straight, proximolower margin regularly rounded and smooth in outline; propodus rectangular, longer than high, exhibiting two morphotypes (L/ H=1.11.2; 1.31.5); outer lateral surface of manus smooth, adorned with several tubercles; fixed finger shorter than manus, triangular, with two parallel ridges extending onto manus, lower (less developed) ridge positioned close to the lower propodal margin; occlusal margin of fixed finger serrated and adorned with a blunt tooth pointing up and forward; dactylus slender, with serrated occlusal margin and pointed tip.
</paragraph>
</subSubSection>
<subSubSection id="C3796557FFC6C81300F2FA83FD1AB41D" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45" type="description">
<paragraph id="8BDC36DCFFC6C81200F2FA83FB7EB0CA" blockId="8.[151,1437,1031,2013]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200F2FA83FED9B37A" bold="true" box="[199,343,1289,1314]" inLineHeading="true" pageId="8" pageNumber="45" reason="1">Description.</emphasis>
Major cheliped massive. Merus longer than high (L/H=2.2), with prominent keel along the midline; upper margin unarmed, convex; lower margin concave, serrated. Carpus higher than long (L/H=0.45), upper margin straight or concave, proximolower margin regularly rounded and smooth in outline. Propodus rectangular, longer than high (L/H=1.11.5), distinctly longer than carpus; upper and lower margins straight, parallel to each other, forming distinct keels on the inner sides, rounded proximally; distal margin straight, parallel to proximal margin, the base of the fixed finger may form a notch. Outer lateral surface of manus smooth, adorned with several tubercles (at the bases of setal pores) lying in a row positioned obliquely. Inner lateral surface of manus concave, with distinct depression at the base of fixed finger. Fixed finger shorter than manus, triangular, with two ridges extending onto manus, lower (less developed) ridge positioned close to the lower propodal margin, upper one (strongly developed) accompanied with several tubercles proximally; occlusal margin serrated and adorned with a blunt tooth pointing up and forward. Dactylus slender, with serrated occlusal margin and pointed tip.
</paragraph>
<paragraph id="8BDC36DCFFC6C81200F2F914FAB4B1CC" blockId="8.[151,1437,1031,2013]" pageId="8" pageNumber="45">
<emphasis id="B917EACEFFC6C81200F2F914FEC4B0EF" bold="true" box="[199,330,1694,1719]" inLineHeading="true" pageId="8" pageNumber="45" reason="1">Variations.</emphasis>
Major propodi of
<taxonomicName id="4C634D5FFFC6C8120218F914FC8AB0EF" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[557,772,1694,1719]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC6C8120218F914FC8AB0EF" box="[557,772,1694,1719]" italics="true" pageId="8" pageNumber="45">Calliax michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC6C8120325F914FC1FB0EE" box="[784,913,1694,1719]" pageId="8" pageNumber="45" rank="species">comb. nov.</taxonomicNameLabel>
exhibit a certain degree of variation. When examining a large number of specimens, one can observe differences in the manus length/height ratio attaining values in two intervals, i.e. 1.11.2 (shorter morph) and 1.31.5 (longer morph). Unfortunately it is difficult to state whether it is intraspecific variation or if it mirrors sexual dimorphism.
<bibRefCitation id="EFF24B2DFFC6C81203FCF886FB14B17D" author="Polkowsky, S." box="[969,1178,1804,1829]" pageId="8" pageNumber="45" pagination="1 - 126" refId="ref14603" refString="Polkowsky, S. (2005) Decapode Krebse aus dem oberoligozanen Sternberger Gestein von Kobrow (Mecklenburg). Tassados, 1, 1 - 126." type="journal article" year="2005">Polkowsky (2005)</bibRefCitation>
tentatively interpreted two morphotypes of
<taxonomicName id="4C634D5FFFC6C81201BFF8B8FDABB112" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[394,549,1841,1866]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC6C81201BFF8B8FDABB112" box="[394,549,1841,1866]" italics="true" pageId="8" pageNumber="45">C. michelottii</emphasis>
</taxonomicName>
as sexual dimorphs. Some propodi may possess a relatively well developed notch at the base of the fixed finger. Its development seems to be at least partly correlated with the size, but as already pointed out by
<bibRefCitation id="EFF24B2DFFC6C81201ACF8F1FDE4B1CC" author="Polkowsky, S." box="[409,618,1915,1940]" pageId="8" pageNumber="45" pagination="1 - 126" refId="ref14603" refString="Polkowsky, S. (2005) Decapode Krebse aus dem oberoligozanen Sternberger Gestein von Kobrow (Mecklenburg). Tassados, 1, 1 - 126." type="journal article" year="2005">Polkowsky (2005)</bibRefCitation>
, the notch in the longer morphotype is usually better developed.
</paragraph>
<paragraph id="8BDC36DCFFC6C81300A2F815FDF0B68D" blockId="8.[151,1437,1031,2013]" lastBlockId="9.[151,1437,151,581]" lastPageId="9" lastPageNumber="46" pageId="8" pageNumber="45">Development of the ridges is a stable character, i.e. there are always two ridges present, and the lower one is rather faint. What differs in various propodi is the shape of the upper ridge in its proximalmost part. In some specimens it is straight, but in others it is curved downwards. The development of tubercles accompanying the upper ridge also varies, depending on the size of the animal.</paragraph>
<paragraph id="8BDC36DCFFC7C81300F2FF6BFE53B7EA" blockId="9.[151,1437,151,581]" pageId="9" pageNumber="46">
<bibRefCitation id="EFF24B2DFFC7C81300F2FF6BFE09B6A2" author="Glaessner, M. F." box="[199,391,225,250]" pageId="9" pageNumber="46" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner (1928)</bibRefCitation>
reported strong ornamentation on the isolated propodi from Jarenina,
<collectingCountry id="F374764CFFC7C81304ABFF6BFA8EB6A2" box="[1182,1280,225,250]" name="Slovenia" pageId="9" pageNumber="46">Slovenia</collectingCountry>
(
<figureCitation id="13582A59FFC7C8130525FF6BFA1EB6A2" box="[1296,1424,225,250]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="9" pageNumber="46">Figs 8AB</figureCitation>
). This can, however, be ascribed mainly to preservational aspects. Specimens from Želiezovce,
<collectingCountry id="F374764CFFC7C81304F6FE8CFAABB747" box="[1219,1317,262,287]" name="Slovakia" pageId="9" pageNumber="46">Slovakia</collectingCountry>
(
<bibRefCitation id="EFF24B2DFFC7C8130500FE8CFF79B71C" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." pageId="9" pageNumber="46" pagination="704 - 746" refId="ref15452" refString="Spinar, Z., Cepek, P., Boucek, B., Eliasova-Frajova, H., Havlicek, V., Horny, R., Housa, V., Kalabis, V., Kukalova, J., Marek, L., Nekvasilova, O., Pokorny, V., Prokop, R., Pribyl, A., Ruzicka, B., Snajdr, M., Stemprokova, D. &amp; Zikmundova, J. (1965) Systematicka paleontologie bezobratlych. Academia, Praha, 1049 pp. [pp. 704 - 746.]" type="book chapter" year="1965">
Špinar
<emphasis id="B917EACEFFC7C81305BDFE8DFF23B71C" italics="true" pageId="9" pageNumber="46">et al</emphasis>
. 1965
</bibRefCitation>
: fig. X-184; refigured here as
<figureCitation id="13582A59FFC7C813027CFEA1FD12B71C" box="[585,668,299,324]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="9" pageNumber="46">Fig. 8L</figureCitation>
) and Szob,
<collectingCountry id="F374764CFFC7C8130317FEA1FC0AB71B" box="[802,900,299,323]" name="Hungary" pageId="9" pageNumber="46">Hungary</collectingCountry>
(
<bibRefCitation id="EFF24B2DFFC7C81303A7FEA1FBADB71C" author="Muller, P." box="[914,1059,299,324]" pageId="9" pageNumber="46" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller 1984</bibRefCitation>
: pl. 7, figs. 3, 4; refigured here as
<figureCitation id="13582A59FFC7C81300A2FEC5FE84B730" box="[151,266,335,360]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="9" pageNumber="46">Figs 8JK</figureCitation>
) exhibit a similar pattern of ornamentation, although not so pronounced as in the material from Jarenina. Possibly it is related to calcification of the cuticle. Experimental data, which are lacking at this point of time, would help in elucidating this issue.
</paragraph>
<paragraph id="8BDC36DCFFC7C81300F2FE34FD1AB41D" blockId="9.[151,1437,151,581]" pageId="9" pageNumber="46">
Major P1carpus may have a concave upper margin (
<bibRefCitation id="EFF24B2DFFC7C813032DFE34FC22B78F" author="Fritsch, K. von" box="[792,940,446,471]" pageId="9" pageNumber="46" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch 1871</bibRefCitation>
: pl. 17, fig. 9; refigured here as
<figureCitation id="13582A59FFC7C813052EFE34FAFDB78F" box="[1307,1395,446,471]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="9" pageNumber="46">Fig. 6E</figureCitation>
). A similarly concave upper margin was also observed in extant
<taxonomicName id="4C634D5FFFC7C8130378FE69FB84B7A4" authorityName="Sakai" authorityYear="1999" box="[845,1034,483,508]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="doerjesti">
<emphasis id="B917EACEFFC7C8130378FE69FB84B7A4" box="[845,1034,483,508]" italics="true" pageId="9" pageNumber="46">Calliax doerjesti</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF24B2DFFC7C813042EFE69FB2EB7A4" author="Sakai, K." box="[1051,1184,483,508]" pageId="9" pageNumber="46" pagination="1 - 152" refId="ref15001" refString="Sakai, K. (1999) Synopsis of the family Callianassidae, with keys to subfamilies, genera and species, and the description of new taxa (Crustacea: Decapoda: Thalassinidea). Zoologische Verhandelingen, Leiden, 326, 1 - 152." type="journal article" year="1999">Sakai 1999</bibRefCitation>
: fig. 28, 29;
<figureCitation id="13582A59FFC7C8130501FE69FA01B7A4" box="[1332,1423,483,508]" captionStart="FIGURE 3" captionStartId="4.[151,249,1131,1153]" captionTargetBox="[165,1429,185,1107]" captionTargetId="figure-274@4.[150,1437,183,1107]" captionTargetPageId="4" captionText="FIGURE 3. Extant Calliax species: AB, Calliax doerjesti Sakai, 1999, note the discrepancy in figuring lower margin of major P1 merus in „A“ and „a“ (redrawn after Sakai 1999: figs 2829); CD, Calliax lobata (de Gaillande &amp; Lagardère, 1966) (redrawn after Ngoc-Ho 2003: figs 17DE); EF, Calliax cf. C. lobata, line drawings of NHMW 25511 (Fig. 2). Left column=major chelae (outer surface), right column=minor chelae (outer surface). Setae are omitted. From two ridges at the base of the fixed finger only the more developed one is depicted in the sketches (see text for details). Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919928" httpUri="https://zenodo.org/record/4919928/files/figure.png" pageId="9" pageNumber="46">Fig. 3A</figureCitation>
). This feature may be related to age and/or size of the individual. For resolving this issue in
<taxonomicName id="4C634D5FFFC7C81304B5FD8DFADFB478" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[1152,1361,519,544]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC7C81304B5FD8DFADFB478" box="[1152,1361,519,544]" italics="true" pageId="9" pageNumber="46">Calliax michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC7C8130562FD8DFF49B41C" pageId="9" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
more preserved carpi must be examined.
</paragraph>
</subSubSection>
<caption id="DF1C6654FFC7C81300A2FA55FD6AB0BD" ID-DOI="http://doi.org/10.5281/zenodo.4919938" ID-Zenodo-Dep="4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="9" pageNumber="46" startId="9.[151,249,1503,1525]" targetBox="[160,1420,654,1470]" targetPageId="9">
<paragraph id="8BDC36DCFFC7C81300A2FA55FD6AB0BD" blockId="9.[151,1436,1503,1766]" pageId="9" pageNumber="46">
<emphasis id="B917EACEFFC7C81300A2FA55FE9DB3AC" bold="true" box="[151,275,1503,1525]" pageId="9" pageNumber="46">FIGURE 6.</emphasis>
<taxonomicName id="4C634D5FFFC7C813012EFA55FD61B3AD" authority="(A. Milne Edwards, 1860)" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[283,751,1503,1525]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC7C813012EFA55FE57B3AD" box="[283,473,1503,1525]" italics="true" pageId="9" pageNumber="46">Calliax michelottii</emphasis>
(A.
<bibRefCitation id="EFF24B2DFFC7C813023FFA55FD66B3AD" author="Milne Edwards, A." box="[522,744,1503,1525]" pageId="9" pageNumber="46" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC7C81302C2FA55FCE5B3AD" box="[759,875,1503,1525]" pageId="9" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
, occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to
<taxonomicName id="4C634D5FFFC7C813044BF9EDFA87B025" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[1150,1289,1639,1661]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC7C813044BF9EDFA87B025" box="[1150,1289,1639,1661]" italics="true" pageId="9" pageNumber="46">C. michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC7C8130527F9EDFA0AB025" box="[1298,1412,1639,1661]" pageId="9" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from
<bibRefCitation id="EFF24B2DFFC7C81302B9F926FC91B09A" author="Fritsch, K. von" box="[652,799,1708,1730]" pageId="9" pageNumber="46" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871)</bibRefCitation>
, H is refigured from
<bibRefCitation id="EFF24B2DFFC7C81303C3F926FB25B09A" author="Lorenthey, I." box="[1014,1195,1708,1730]" pageId="9" pageNumber="46" pagination="202 - 259" refId="ref14078" refString="Lorenthey, I. (1907) Beitrage zur Tertiaren Dekapodenfauna Sardiniens. Mathematischen und Naturwissenchaftlichen Berichte aus Ungarn, 25, 202 - 259, 2 plates." type="book chapter" year="1907">Lőrenthey (1907)</bibRefCitation>
, IL are refigured from
<bibRefCitation id="EFF24B2DFFC7C81300A2F95AFEB2B0BE" author="Noetling, F." box="[151,316,1744,1766]" pageId="9" pageNumber="46" pagination="32 - 34" refId="ref14502" refString="Noetling, F. (1886) Uber Crustaceenreste aus dem Oberoligocanen Sternberger Gestein. Sitzingsberichte der Gesselschaft naturforschender Freunde Berlin, 1886, 32 - 34." type="journal article" year="1886">Noetling (1886)</bibRefCitation>
. AH are to scale, scale bar equals 5 mm.
</paragraph>
</caption>
<subSubSection id="C3796557FFC7C81000F2F89DFD92B03E" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="8BDC36DCFFC7C81000F2F89DFA80B730" blockId="9.[151,1436,1815,2024]" lastBlockId="10.[151,1436,151,360]" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<emphasis id="B917EACEFFC7C81300F2F89DFE38B168" bold="true" box="[199,438,1815,1840]" inLineHeading="true" pageId="9" pageNumber="46" reason="1">Material examined.</emphasis>
Only remains of major chelipeds have been examined: the
<typeStatus id="54D8887EFFC7C81304B1F89DFB68B168" box="[1156,1254,1815,1840]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C634D5FFFC7C8130523F89DFE81B10D" authorityName="Muller" authorityYear="1993" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC7C8130523F89DFE81B10D" italics="true" pageId="9" pageNumber="46">Callianassa michelottii</emphasis>
</taxonomicName>
from Superga at Turin,
<collectingCountry id="F374764CFFC7C8130203F8B6FDE4B10D" box="[566,618,1852,1877]" name="Italy" pageId="9" pageNumber="46">Italy</collectingCountry>
(MNHN-F-B32690;
<figureCitation id="13582A59FFC7C8130351F8B6FC23B10D" box="[868,941,1852,1877]" captionStart="FIGURE 5" captionStartId="7.[151,249,1290,1312]" captionTargetBox="[246,1336,525,1260]" captionTargetId="figure-517@7.[238,1348,519,1265]" captionTargetPageId="7" captionText="FIGURE 5. Callianassa michelottii A. Milne Edwards, 1860: holotype (MNHN-F-B32690): AB, refigured from A. Milne Edwards (1860: pl. 14, fig. 3); CD, photo of the holotype. A, C=inner surface; B, D=outer surface. Arrows indicate ridges. Photo by Lilian Cazes (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4919934" httpUri="https://zenodo.org/record/4919934/files/figure.png" pageId="9" pageNumber="46">Fig. 5</figureCitation>
); 21 fragmentary propodi and one carpus (
<collectionCode id="ED72AE19FFC7C81300ABF8EBFF72B122" box="[158,252,1889,1914]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46" type="Museum">NHMW</collectionCode>
1874/0029/1154 av) from Sternberg,
<collectingCountry id="F374764CFFC7C81302E3F8EBFCB0B122" box="[726,830,1889,1914]" name="Germany" pageId="9" pageNumber="46">Germany</collectingCountry>
(
<figureCitation id="13582A59FFC7C8130360F8EBFC2EB122" box="[853,928,1889,1914]" captionStart="FIGURE 7" captionStartId="10.[151,249,1261,1283]" captionTargetBox="[152,1435,416,1236]" captionTargetId="figure-467@10.[152,1435,416,1236]" captionTargetPageId="10" captionText="FIGURE 7. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Oligocene of the European plate: isolated propodi (AK) and carpus (LM). AB, NHMW 1874/0029/1154 a; C, D, G, NHMW 1874/0029/1154 a, G shows the specimen under a low-angle light to show the ridges on the fixed finger; EF, NHMW 1874/0029/1154 c; H, NHMW 1874/ 0029/1154 d; I, NHMW 1874/0029/1155; JK, NHMW 1874/0029/1154 e; LM, NHMW 1874/0029/1154 f. Specimens in AH, JM are from Sternberg (Germany), specimen in I is from Pinnow bei Schwerin (Germany). All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919942" httpUri="https://zenodo.org/record/4919942/files/figure.png" pageId="9" pageNumber="46">Fig. 7</figureCitation>
); one isolated propodus from Pinnow bei Schwerin,
<collectingCountry id="F374764CFFC7C8130123F80CFEF0B1C7" box="[278,382,1926,1951]" name="Germany" pageId="9" pageNumber="46">Germany</collectingCountry>
(
<collectionCode id="ED72AE19FFC7C81301A0F80CFE7DB1C7" box="[405,499,1926,1951]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46" type="Museum">NHMW</collectionCode>
1874/0029/1155;
<figureCitation id="13582A59FFC7C81302FAF80CFCAAB1C7" box="[719,804,1926,1951]" captionStart="FIGURE 7" captionStartId="10.[151,249,1261,1283]" captionTargetBox="[152,1435,416,1236]" captionTargetId="figure-467@10.[152,1435,416,1236]" captionTargetPageId="10" captionText="FIGURE 7. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Oligocene of the European plate: isolated propodi (AK) and carpus (LM). AB, NHMW 1874/0029/1154 a; C, D, G, NHMW 1874/0029/1154 a, G shows the specimen under a low-angle light to show the ridges on the fixed finger; EF, NHMW 1874/0029/1154 c; H, NHMW 1874/ 0029/1154 d; I, NHMW 1874/0029/1155; JK, NHMW 1874/0029/1154 e; LM, NHMW 1874/0029/1154 f. Specimens in AH, JM are from Sternberg (Germany), specimen in I is from Pinnow bei Schwerin (Germany). All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919942" httpUri="https://zenodo.org/record/4919942/files/figure.png" pageId="9" pageNumber="46">Fig. 7I</figureCitation>
); one right propodus articulated with dactylus from Pucking,
<collectingCountry id="F374764CFFC7C813013CF821FED2B19C" box="[265,348,1963,1988]" name="Austria" pageId="9" pageNumber="46">Austria</collectingCountry>
(
<collectionCode id="ED72AE19FFC7C8130146F821FE5FB19C" box="[371,465,1963,1988]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46" type="Museum">NHMW</collectionCode>
2003/0026/0913;
<figureCitation id="13582A59FFC7C813029AF821FC81B19C" box="[687,783,1963,1988]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="9" pageNumber="46">Fig. 8C</figureCitation>
); entire articulated right cheliped from Neuhofen bei Tettenweis (
<collectionCode id="ED72AE19FFC7C8130115F845FEF0B1B0" box="[288,382,1999,2024]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="9" pageNumber="46" type="Museum">NHMW</collectionCode>
2010/0089/0001;
<figureCitation id="13582A59FFC7C813027BF845FD13B1B0" box="[590,669,1999,2024]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="9" pageNumber="46">Fig. 8I</figureCitation>
); two isolated left propodi (
<collectionCode id="ED72AE19FFC7C81303EEF845FB99B1B0" box="[987,1047,1999,2024]" pageId="9" pageNumber="46">RGA</collectionCode>
/
<collectionCode id="ED72AE19FFC7C8130415F845FBFFB1B0" box="[1056,1137,1999,2024]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="9" pageNumber="46">SMNH</collectionCode>
0773, 0779), two isolated right propodi (
<collectionCode id="ED72AE19FFC4C8100102FF1DFEFDB6E8" box="[311,371,151,176]" pageId="10" pageNumber="47">RGA</collectionCode>
/
<collectionCode id="ED72AE19FFC4C8100149FF1DFE43B6E8" box="[380,461,151,176]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="10" pageNumber="47">SMNH</collectionCode>
0754, 0864), four articulated chelae consisting of propodus and dactylus (
<collectionCode id="ED72AE19FFC4C8100533FF1DFACCB6E8" box="[1286,1346,151,176]" pageId="10" pageNumber="47">RGA</collectionCode>
/
<collectionCode id="ED72AE19FFC4C810057EFF1DFA12B6E8" box="[1355,1436,151,176]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="10" pageNumber="47">SMNH</collectionCode>
1075, 1187) and even carpus and merus (
<collectionCode id="ED72AE19FFC4C8100258FF36FD27B68D" box="[621,681,188,213]" pageId="10" pageNumber="47">RGA</collectionCode>
/
<collectionCode id="ED72AE19FFC4C8100284FF36FC8CB68D" box="[689,770,188,213]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="10" pageNumber="47">SMNH</collectionCode>
1069, 1191) from Kamnik-Košiše,
<collectingCountry id="F374764CFFC4C81004AFFF36FB72B68D" box="[1178,1276,188,213]" name="Slovenia" pageId="10" pageNumber="47">Slovenia</collectingCountry>
(
<figureCitation id="13582A59FFC4C8100539FF36FA1EB68D" box="[1292,1424,188,213]" captionStart="FIGURE 9" captionStartId="13.[151,249,1148,1170]" captionTargetBox="[151,1436,339,1124]" captionTargetId="figure-285@13.[150,1436,339,1124]" captionTargetPageId="13" captionText="FIGURE 9. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Middle Miocene of Kamnik, Slovenia. A, RGA/SMNH 0754, right propodus; B, RGA/SMNH 0779, left propodus; C, RGA/SMNH 0864, right propodus; D, RGA/ SMNH 0773, left propodus; E, RGA/SMNH 1191, left major chela consisting of propodus, carpus and merus; F, RGA/SMNH 1075, left propodus articulated with dactylus; G, RGA/SMNH 1187, left propodus articulated with dactylus; H, RGA/SMNH 1069, left major chela consisting of dactylus, propodus and carpus; I, GBA 2009/014/0027, right propodus. All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919950" httpUri="https://zenodo.org/record/4919950/files/figure.png" pageId="10" pageNumber="47">Figs 9AH</figureCitation>
); right isolated propodus from the unknown site at
<collectingRegion id="49A7F83EFFC4C810028FFF6BFC9AB6A2" box="[698,788,225,250]" country="Slovenia" name="Kamnik" pageId="10" pageNumber="47">Kamnik</collectingRegion>
(Stein in Krain),
<collectingCountry id="F374764CFFC4C81003E3FF6BFBB6B6A2" box="[982,1080,225,250]" name="Slovenia" pageId="10" pageNumber="47">Slovenia</collectingCountry>
(
<collectionCode id="ED72AE19FFC4C8100473FF6BFB0DB6A2" box="[1094,1155,225,250]" pageId="10" pageNumber="47">GBA</collectionCode>
2009/014/0027;
<figureCitation id="13582A59FFC4C8100574FF6BFA00B6A2" box="[1345,1422,225,250]" captionStart="FIGURE 9" captionStartId="13.[151,249,1148,1170]" captionTargetBox="[151,1436,339,1124]" captionTargetId="figure-285@13.[150,1436,339,1124]" captionTargetPageId="13" captionText="FIGURE 9. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Middle Miocene of Kamnik, Slovenia. A, RGA/SMNH 0754, right propodus; B, RGA/SMNH 0779, left propodus; C, RGA/SMNH 0864, right propodus; D, RGA/ SMNH 0773, left propodus; E, RGA/SMNH 1191, left major chela consisting of propodus, carpus and merus; F, RGA/SMNH 1075, left propodus articulated with dactylus; G, RGA/SMNH 1187, left propodus articulated with dactylus; H, RGA/SMNH 1069, left major chela consisting of dactylus, propodus and carpus; I, GBA 2009/014/0027, right propodus. All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919950" httpUri="https://zenodo.org/record/4919950/files/figure.png" pageId="10" pageNumber="47">Fig. 9I</figureCitation>
); two isolated propodi from Jarenina (Jahring),
<collectingCountry id="F374764CFFC4C8100284FE8CFC9DB747" box="[689,787,262,287]" name="Slovenia" pageId="10" pageNumber="47">Slovenia</collectingCountry>
already reported (but not figured) by
<bibRefCitation id="EFF24B2DFFC4C81004EDFE8CFA12B747" author="Glaessner, M. F." box="[1240,1436,262,287]" pageId="10" pageNumber="47" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner (1928)</bibRefCitation>
(
<collectionCode id="ED72AE19FFC4C81000ABFEA1FF72B71C" box="[158,252,299,324]" pageId="10" pageNumber="47">UMJGP</collectionCode>
7787377874;
<figureCitation id="13582A59FFC4C8100187FEA1FDBDB71C" box="[434,563,299,324]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="10" pageNumber="47">Figs 8AB</figureCitation>
); one right fragmentary propodus from Rohožník,
<collectingCountry id="F374764CFFC4C810044BFEA1FB6EB71C" box="[1150,1248,299,324]" name="Slovakia" pageId="10" pageNumber="47">Slovakia</collectingCountry>
(
<collectionCode id="ED72AE19FFC4C81004C7FEA1FA12B71C" box="[1266,1436,299,324]" pageId="10" pageNumber="47">KGP-MH RO-</collectionCode>
001;
<figureCitation id="13582A59FFC4C81000E5FEC5FEC1B730" box="[208,335,335,360]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="10" pageNumber="47">Figs 8DH</figureCitation>
);
<typeStatus id="54D8887EFFC4C8100156FEC5FE4BB730" box="[355,453,335,360]" pageId="10" pageNumber="47" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C634D5FFFC4C81001DEFEC5FD6FB730" authorityName="Muller" authorityYear="1984" box="[491,737,335,360]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC4C81001DEFEC5FD6FB730" box="[491,737,335,360]" italics="true" pageId="10" pageNumber="47">Callianassa szobensis</emphasis>
</taxonomicName>
from Szob,
<collectingCountry id="F374764CFFC4C8100359FEDAFC40B730" box="[876,974,336,360]" name="Hungary" pageId="10" pageNumber="47">Hungary</collectingCountry>
(M.2004.158.1;
<figureCitation id="13582A59FFC4C81004BEFEC5FA8FB730" box="[1163,1281,335,360]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="10" pageNumber="47">Figs 8JK</figureCitation>
).
</paragraph>
<caption id="DF1C6654FFC4C81000A2FB67FB90B3F7" ID-DOI="http://doi.org/10.5281/zenodo.4919942" ID-Zenodo-Dep="4919942" httpUri="https://zenodo.org/record/4919942/files/figure.png" pageId="10" pageNumber="47" startId="10.[151,249,1261,1283]" targetBox="[152,1435,416,1236]" targetPageId="10">
<paragraph id="8BDC36DCFFC4C81000A2FB67FB90B3F7" blockId="10.[151,1436,1261,1455]" pageId="10" pageNumber="47">
<emphasis id="B917EACEFFC4C81000A2FB67FE9BB35A" bold="true" box="[151,277,1261,1283]" pageId="10" pageNumber="47">FIGURE 7.</emphasis>
<taxonomicName id="4C634D5FFFC4C810012AFB67FD70B35B" authority="(A. Milne Edwards, 1860)" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[287,766,1261,1283]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC4C810012AFB67FE51B35B" box="[287,479,1261,1283]" italics="true" pageId="10" pageNumber="47">Calliax michelottii</emphasis>
(A.
<bibRefCitation id="EFF24B2DFFC4C8100221FB67FD79B35B" author="Milne Edwards, A." box="[532,759,1261,1283]" pageId="10" pageNumber="47" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC4C8100332FB67FCF5B35B" box="[775,891,1261,1283]" pageId="10" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
from the Oligocene of the European plate: isolated propodi (AK) and carpus (LM). AB, NHMW 1874/0029/1154 a; C, D, G, NHMW 1874/0029/1154 a, G shows the specimen under a low-angle light to show the ridges on the fixed finger; EF, NHMW 1874/0029/1154 c; H, NHMW 1874/ 0029/1154 d; I, NHMW 1874/0029/1155; JK, NHMW 1874/0029/1154 e; LM, NHMW 1874/0029/1154 f. Specimens in AH, JM are from Sternberg (Germany), specimen in I is from Pinnow bei Schwerin (Germany). All specimens are to scale and were covered with ammonium chloride prior to photography. Scale bar equals 5 mm.
</paragraph>
</caption>
<paragraph id="8BDC36DCFFC4C81000F2FA6BFD92B03E" blockId="10.[151,1437,1505,2034]" pageId="10" pageNumber="47">
<emphasis id="B917EACEFFC4C81000F2FA6BFED7B3A2" bold="true" box="[199,345,1505,1530]" inLineHeading="true" pageId="10" pageNumber="47" reason="1">Occurrence.</emphasis>
Oligocene (Rupelian)Middle Miocene (Serravallian) of Europe. The oldest occurrence of the species is known from the Rupelian of the Mainz Basin,
<collectingCountry id="F374764CFFC4C8100326F98FFCF5B046" box="[787,891,1541,1566]" name="Germany" pageId="10" pageNumber="47">Germany</collectingCountry>
(
<bibRefCitation id="EFF24B2DFFC4C81003BFF98FFB92B046" author="Fritsch, K. von" box="[906,1052,1541,1566]" pageId="10" pageNumber="47" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch 1871</bibRefCitation>
;
<figureCitation id="13582A59FFC4C810041DF98FFB27B046" box="[1064,1193,1541,1566]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="10" pageNumber="47">Figs 6AG</figureCitation>
). The youngest one is from the MiddleLate Badenian (Serravallian) of
<collectingCountry id="F374764CFFC4C81002FDF9A3FCA4B01A" box="[712,810,1577,1602]" name="Slovakia" pageId="10" pageNumber="47">Slovakia</collectingCountry>
(herein). All confirmed occurrences are summarized in
<tableCitation id="C6E10367FFC4C81000A2F9C7FF63B03E" box="[151,237,1613,1638]" captionStart="TABLE 2" captionStartId="11.[151,239,512,535]" captionTargetPageId="11" captionText="TABLE 2. Synopsis of confirmed occurrences of Calliax michelottii (A. Milne Edwards, 1860) comb. nov. The occurrences are arranged according to their stratigraphic age." httpUri="http://table.plazi.org/id/DF1C6654FFC5C81100A2FD8AFCACB46F" pageId="10" pageNumber="47" tableUuid="DF1C6654FFC5C81100A2FD8AFCACB46F">Table 2</tableCitation>
. For details see text below.
</paragraph>
</subSubSection>
<subSubSection id="C3796557FFC4C81700F2F9FBFBF2B745" lastPageId="13" lastPageNumber="50" pageId="10" pageNumber="47" type="discussion">
<paragraph id="8BDC36DCFFC4C81000F2F9FBFB3DB166" blockId="10.[151,1437,1505,2034]" pageId="10" pageNumber="47">
<emphasis id="B917EACEFFC4C81000F2F9FBFEB5B0D2" bold="true" box="[199,315,1649,1674]" inLineHeading="true" pageId="10" pageNumber="47" reason="1">Remarks.</emphasis>
<taxonomicName id="4C634D5FFFC4C810017DF9FBFDDDB0D2" authorityName="Muller" authorityYear="1993" box="[328,595,1649,1674]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC4C810017DF9FBFDDDB0D2" box="[328,595,1649,1674]" italics="true" pageId="10" pageNumber="47">Callianassa michelottii</emphasis>
</taxonomicName>
was originally described based on isolated propodi (
<figureCitation id="13582A59FFC4C81004E3F9FBFA91B0D2" box="[1238,1311,1649,1674]" captionStart="FIGURE 5" captionStartId="7.[151,249,1290,1312]" captionTargetBox="[246,1336,525,1260]" captionTargetId="figure-517@7.[238,1348,519,1265]" captionTargetPageId="7" captionText="FIGURE 5. Callianassa michelottii A. Milne Edwards, 1860: holotype (MNHN-F-B32690): AB, refigured from A. Milne Edwards (1860: pl. 14, fig. 3); CD, photo of the holotype. A, C=inner surface; B, D=outer surface. Arrows indicate ridges. Photo by Lilian Cazes (MNHN)." figureDoi="http://doi.org/10.5281/zenodo.4919934" httpUri="https://zenodo.org/record/4919934/files/figure.png" pageId="10" pageNumber="47">Fig. 5</figureCitation>
) from the Miocene of Superga hill near Turin,
<collectingCountry id="F374764CFFC4C810020DF91FFDE2B0F6" box="[568,620,1685,1710]" name="Italy" pageId="10" pageNumber="47">Italy</collectingCountry>
(A.
<bibRefCitation id="EFF24B2DFFC4C8100294F91FFC1DB0F6" author="Milne Edwards, A." box="[673,915,1685,1710]" pageId="10" pageNumber="47" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards 1860</bibRefCitation>
). A.
<bibRefCitation id="EFF24B2DFFC4C81003F9F91FFB49B0F6" author="Milne Edwards, A." box="[972,1223,1685,1710]" pageId="10" pageNumber="47" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards (1860</bibRefCitation>
: p. 211) noted that the presence of the keel on the fixed finger accompanied by the tubercles above it is unique not only for fossil species, but this character distinguishes the species from all extant callianassids known at that time as well. Indeed, the
<typeStatus id="54D8887EFFC4C81000F7F888FF7CB142" box="[194,242,1794,1818]" pageId="10" pageNumber="47">type</typeStatus>
species of
<taxonomicName id="4C634D5FFFC4C8100141F88BFE49B142" authorityName="de Saint Laurent" authorityYear="1973" box="[372,455,1793,1818]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC4C8100141F88BFE49B142" box="[372,455,1793,1818]" italics="true" pageId="10" pageNumber="47">Calliax</emphasis>
</taxonomicName>
was described more than a century later by
<bibRefCitation id="EFF24B2DFFC4C81003F0F88BFACAB142" author="de Gaillande, D. &amp; Lagardere, J. - P." box="[965,1348,1793,1818]" pageId="10" pageNumber="47" pagination="259 - 265" refId="ref12440" refString="de Gaillande, D. &amp; Lagardere, J. - P. (1966) Description de Callianassa (Callichirus) lobata nov. sp. (Crustacea Decapoda Callianassidae). Recueil des Travaux de la Station Marine d'Endoume, 40, 259 - 265." type="journal article" year="1966">de Gaillande &amp; Lagardère (1966)</bibRefCitation>
.
<bibRefCitation id="EFF24B2DFFC4C8100564F88BFF6FB166" author="Crema, C." pageId="10" pageNumber="47" pagination="664 - 681" refId="ref11114" refString="Crema, C. (1895) Sopra alcuni decapodi terziarii del Piemonte. Atti Reale Accademia delle Scienze di Torino, 30, 664 - 681." type="journal article" year="1895">Crema (1895)</bibRefCitation>
reported
<taxonomicName id="4C634D5FFFC4C8100178F8ACFE6BB166" authorityName="Muller" authorityYear="1993" box="[333,485,1829,1854]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC4C8100178F8ACFE6BB166" box="[333,485,1829,1854]" italics="true" pageId="10" pageNumber="47">C. michelottii</emphasis>
</taxonomicName>
from the Middle Miocene of nearby localities in the Turin hills.
</paragraph>
<paragraph id="8BDC36DCFFC4C81100F2F8C3FB5CB788" blockId="10.[151,1437,1505,2034]" lastBlockId="11.[151,1437,151,464]" lastPageId="11" lastPageNumber="48" pageId="10" pageNumber="47">
<bibRefCitation id="EFF24B2DFFC4C81000F2F8C3FEE9B13A" author="Fritsch, K. von" box="[199,359,1865,1890]" pageId="10" pageNumber="47" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871)</bibRefCitation>
reported
<taxonomicName id="4C634D5FFFC4C81001E0F8C0FDE0B13A" authorityName="Muller" authorityYear="1993" box="[469,622,1865,1890]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC4C81001E0F8C0FDE0B13A" box="[469,622,1865,1890]" italics="true" pageId="10" pageNumber="47">C. michelottii</emphasis>
</taxonomicName>
from the “Middle” Oligocene (Rupelian) “Septarienthon” of the Mainz Basin (W
<collectingCountry id="F374764CFFC4C8100133F8E7FEFCB1DE" box="[262,370,1901,1926]" name="Germany" pageId="10" pageNumber="47">Germany</collectingCountry>
). Interestingly,
<bibRefCitation id="EFF24B2DFFC4C810022EF8E7FD3DB1DE" author="Fritsch, K. von" box="[539,691,1901,1926]" pageId="10" pageNumber="47" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871</bibRefCitation>
: pl. 17, figs 513) reported and also figured a small chela preserved together with the major P1 propodus (
<figureCitation id="13582A59FFC4C810020CF81BFD1CB1F2" box="[569,658,1937,1962]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="10" pageNumber="47">Fig. 6G</figureCitation>
). He tentatively interpreted it as remains of P2, but it in fact represents a minor P1 chela. The description of
<bibRefCitation id="EFF24B2DFFC4C810021AF83FFC8BB196" author="Fritsch, K. von" box="[559,773,1973,1998]" pageId="10" pageNumber="47" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871: 696)</bibRefCitation>
is very clear in this sense (“Der bewegliche Finger steht auf einem weit nach vorn vorspringenden Theil des Ballens. Vom unbeweglichen Finger ist nu rein ganz geringer Theil sichtbar, warscheinlich war derselbe indess kurz, die hervortretenden Leisten desselben Gliedes der Vorderhand fehlen.” English translation: Dactylus is located on the manus portion projecting forward. Only very small portion of the index is visible. It was probably short, although the tip is missing.)
<bibRefCitation id="EFF24B2DFFC5C81103EEFF55FB37B6A0" author="Fritsch, K. von" box="[987,1209,223,248]" pageId="11" pageNumber="48" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871: 692)</bibRefCitation>
mentioned also the presence of the cephalothorax (“Wohl liegt ein Exemplar vor, an welchem man Theile des kleinen dünnschaligen Cephalothorax und der seitlich zusammengedrückten hinteren Fusspaare erkennt, doch is dasselbe für eine Charakteristik des Thieres zu ungenügend erhalten.” English translation: There is a sample in which the individual shows the preserved small cephalothorax with thin cuticle and the laterally compressed “rear” pereiopod. However, the preservation of both characters is insufficient.), but he did not figure it. The material of
<bibRefCitation id="EFF24B2DFFC5C8110494FE1EFACEB7F5" author="Fritsch, K. von" box="[1185,1344,404,429]" pageId="11" pageNumber="48" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871)</bibRefCitation>
was reexamined by
<bibRefCitation id="EFF24B2DFFC5C811011BFE3DFE55B788" author="Beurlen, K." box="[302,475,439,464]" pageId="11" pageNumber="48" pagination="111 - 112" refId="ref10830" refString="Beurlen, K. (1931) Die Decapodenfauna aus dem Rupelton des Mainzer Beckens. Senckenbergiana, 13, 111 - 112." type="journal article" year="1931">Beurlen (1931)</bibRefCitation>
; he confirmed much of what was already done in the original work.
</paragraph>
<caption id="DF1C6654FFC5C81100A2FD8AFCACB46F" ID-Table-UUID="DF1C6654FFC5C81100A2FD8AFCACB46F" httpUri="http://table.plazi.org/id/DF1C6654FFC5C81100A2FD8AFCACB46F" pageId="11" pageNumber="48" startId="11.[151,239,512,535]" targetBox="[159,1325,594,1184]" targetIsTable="true" targetPageId="11">
<paragraph id="8BDC36DCFFC5C81100A2FD8AFCACB46F" blockId="11.[151,1435,511,567]" pageId="11" pageNumber="48">
<emphasis id="B917EACEFFC5C81100A2FD8AFE81B44F" bold="true" box="[151,271,512,535]" pageId="11" pageNumber="48">TABLE 2.</emphasis>
Synopsis of confirmed occurrences of
<taxonomicName id="4C634D5FFFC5C81102E2FE75FB52B44F" authority="(A. Milne Edwards, 1860)" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[727,1244,511,535]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC5C81102E2FE75FC2AB44F" box="[727,932,511,535]" italics="true" pageId="11" pageNumber="48">Calliax michelottii</emphasis>
(A.
<bibRefCitation id="EFF24B2DFFC5C81103D4FD8AFB58B44F" author="Milne Edwards, A." box="[993,1238,512,535]" pageId="11" pageNumber="48" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC5C81104DCFD8AFAE8B44F" box="[1257,1382,512,535]" pageId="11" pageNumber="48" rank="species">comb. nov.</taxonomicNameLabel>
The occurrences are arranged according to their stratigraphic age.
</paragraph>
</caption>
<paragraph id="8BDC36DCFFC5C81100AAFDD8FAA3B2F8" pageId="11" pageNumber="48">
<table id="F963C47CFFC537E500AAFDD8FAA3B2F8" box="[159,1325,594,1184]" gridcols="3" gridrows="14" pageId="11" pageNumber="48">
<tr id="3553349EFFC537E500AAFDD8FAA3B430" box="[159,1325,594,616]" gridrow="0" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFDD8FE2FB430" box="[159,417,594,616]" gridcol="0" gridrow="0" pageId="11" pageNumber="48">
<emphasis id="B917EACEFFC5C81100AAFDD8FF76B430" bold="true" box="[159,248,594,616]" pageId="11" pageNumber="48">Locality</emphasis>
</th>
<th id="76825DE2FFC537E501F4FDD8FC93B430" box="[449,797,594,616]" gridcol="1" gridrow="0" pageId="11" pageNumber="48">
<emphasis id="B917EACEFFC5C81101F4FDD8FE65B430" bold="true" box="[449,491,594,616]" pageId="11" pageNumber="48">Age</emphasis>
</th>
<th id="76825DE2FFC537E503AAFDD8FAA3B430" box="[927,1325,594,616]" gridcol="2" gridrow="0" pageId="11" pageNumber="48">
<emphasis id="B917EACEFFC5C81103AAFDD8FB85B430" bold="true" box="[927,1035,594,616]" pageId="11" pageNumber="48">Reference</emphasis>
</th>
</tr>
<tr id="3553349EFFC537E500AAFDF6FAA3B4CA" box="[159,1325,636,658]" gridrow="1" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFDF6FE2FB4CA" box="[159,417,636,658]" gridcol="0" gridrow="1" pageId="11" pageNumber="48">Flörsheim (D)</th>
<td id="76825DE2FFC537E501F4FDF6FC93B4CA" box="[449,797,636,658]" gridcol="1" gridrow="1" pageId="11" pageNumber="48">Early Oligocene</td>
<td id="76825DE2FFC537E503AAFDF6FAA3B4CA" box="[927,1325,636,658]" gridcol="2" gridrow="1" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFDF6FBBCB4CA" author="Fritsch, K. von" box="[927,1074,636,658]" pageId="11" pageNumber="48" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871)</bibRefCitation>
,
<bibRefCitation id="EFF24B2DFFC5C8110409FDF6FB58B4CA" author="Beurlen, K." box="[1084,1238,636,658]" pageId="11" pageNumber="48" pagination="111 - 112" refId="ref10830" refString="Beurlen, K. (1931) Die Decapodenfauna aus dem Rupelton des Mainzer Beckens. Senckenbergiana, 13, 111 - 112." type="journal article" year="1931">Beurlen (1931)</bibRefCitation>
</td>
</tr>
<tr id="3553349EFFC537E500AAFD2FFAA3B482" box="[159,1325,677,730]" gridrow="2" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFD2FFE2FB482" box="[159,417,677,730]" gridcol="0" gridrow="2" pageId="11" pageNumber="48">Offenbach bei Frankfurt (D)</th>
<td id="76825DE2FFC537E501F4FD2FFC93B482" box="[449,797,677,730]" gridcol="1" gridrow="2" pageId="11" pageNumber="48">Early Oligocene</td>
<td id="76825DE2FFC537E503AAFD2FFAA3B482" box="[927,1325,677,730]" gridcol="2" gridrow="2" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFD2FFBA1B4E3" author="Fritsch, K. von" box="[927,1071,677,699]" pageId="11" pageNumber="48" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch (1871)</bibRefCitation>
</td>
</tr>
<tr id="3553349EFFC537E500AAFD67FAA3B55B" box="[159,1325,749,771]" gridrow="3" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFD67FE2FB55B" box="[159,417,749,771]" gridcol="0" gridrow="3" pageId="11" pageNumber="48">Sternberg (D)</th>
<td id="76825DE2FFC537E501F4FD67FC93B55B" box="[449,797,749,771]" gridcol="1" gridrow="3" pageId="11" pageNumber="48">Late Oligocene</td>
<td id="76825DE2FFC537E503AAFD67FAA3B55B" box="[927,1325,749,771]" gridcol="2" gridrow="3" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFC9CFAA3B574" box="[159,1325,790,812]" gridrow="4" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFC9CFE2FB574" box="[159,417,790,812]" gridcol="0" gridrow="4" pageId="11" pageNumber="48">Pinnow bei Schwerin (D)</th>
<td id="76825DE2FFC537E501F4FC9CFC93B574" box="[449,797,790,812]" gridcol="1" gridrow="4" pageId="11" pageNumber="48">Late Oligocene</td>
<td id="76825DE2FFC537E503AAFC9CFAA3B574" box="[927,1325,790,812]" gridcol="2" gridrow="4" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFCCAFAA3B50E" box="[159,1325,832,854]" gridrow="5" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFCCAFE2FB50E" box="[159,417,832,854]" gridcol="0" gridrow="5" pageId="11" pageNumber="48">Kobrow (D)</th>
<td id="76825DE2FFC537E501F4FCCAFC93B50E" box="[449,797,832,854]" gridcol="1" gridrow="5" pageId="11" pageNumber="48">Late Oligocene</td>
<td id="76825DE2FFC537E503AAFCCAFAA3B50E" box="[927,1325,832,854]" gridcol="2" gridrow="5" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFCCAFBCDB50E" author="Noetling, F." box="[927,1091,832,854]" pageId="11" pageNumber="48" pagination="32 - 34" refId="ref14502" refString="Noetling, F. (1886) Uber Crustaceenreste aus dem Oberoligocanen Sternberger Gestein. Sitzingsberichte der Gesselschaft naturforschender Freunde Berlin, 1886, 32 - 34." type="journal article" year="1886">Noetling (1886)</bibRefCitation>
,
<bibRefCitation id="EFF24B2DFFC5C811047BFCCAFA86B50E" author="Polkowsky, S." box="[1102,1288,832,854]" pageId="11" pageNumber="48" pagination="1 - 126" refId="ref14603" refString="Polkowsky, S. (2005) Decapode Krebse aus dem oberoligozanen Sternberger Gestein von Kobrow (Mecklenburg). Tassados, 1, 1 - 126." type="journal article" year="2005">Polkowsky (2005)</bibRefCitation>
</td>
</tr>
<tr id="3553349EFFC537E500AAFCE3FAA3B527" box="[159,1325,873,895]" gridrow="6" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFCE3FE2FB527" box="[159,417,873,895]" gridcol="0" gridrow="6" pageId="11" pageNumber="48">Pucking (A)</th>
<td id="76825DE2FFC537E501F4FCE3FC93B527" box="[449,797,873,895]" gridcol="1" gridrow="6" pageId="11" pageNumber="48">Early Miocene (Late Egerian)</td>
<td id="76825DE2FFC537E503AAFCE3FAA3B527" box="[927,1325,873,895]" gridcol="2" gridrow="6" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFC18FAA3B5F0" box="[159,1325,914,936]" gridrow="7" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFC18FE2FB5F0" box="[159,417,914,936]" gridcol="0" gridrow="7" pageId="11" pageNumber="48">Neuhofen (D)</th>
<td id="76825DE2FFC537E501F4FC18FC93B5F0" box="[449,797,914,936]" gridcol="1" gridrow="7" pageId="11" pageNumber="48">Early Miocene (Late Ottnangian)</td>
<td id="76825DE2FFC537E503AAFC18FAA3B5F0" box="[927,1325,914,936]" gridcol="2" gridrow="7" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFC36FAA3B58A" box="[159,1325,956,978]" gridrow="8" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFC36FE2FB58A" box="[159,417,956,978]" gridcol="0" gridrow="8" pageId="11" pageNumber="48">Jarenina (SLO)</th>
<td id="76825DE2FFC537E501F4FC36FC93B58A" box="[449,797,956,978]" gridcol="1" gridrow="8" pageId="11" pageNumber="48">Early Miocene (Karpatian)</td>
<td id="76825DE2FFC537E503AAFC36FAA3B58A" box="[927,1325,956,978]" gridcol="2" gridrow="8" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFC36FBC2B58A" author="Glaessner, M. F." box="[927,1100,956,978]" pageId="11" pageNumber="48" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner (1928)</bibRefCitation>
</td>
</tr>
<tr id="3553349EFFC537E500AAFC6FFAA3B5A3" box="[159,1325,997,1019]" gridrow="9" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFC6FFE2FB5A3" box="[159,417,997,1019]" gridcol="0" gridrow="9" pageId="11" pageNumber="48">Szob (H)</th>
<td id="76825DE2FFC537E501F4FC6FFC93B5A3" box="[449,797,997,1019]" gridcol="1" gridrow="9" pageId="11" pageNumber="48">Middle Miocene (Early Badenian)</td>
<td id="76825DE2FFC537E503AAFC6FFAA3B5A3" box="[927,1325,997,1019]" gridcol="2" gridrow="9" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFC6FFBA0B5A3" author="Muller, P." box="[927,1070,997,1019]" pageId="11" pageNumber="48" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984)</bibRefCitation>
</td>
</tr>
<tr id="3553349EFFC537E500AAFB84FAA3B27C" box="[159,1325,1038,1060]" gridrow="10" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFB84FE2FB27C" box="[159,417,1038,1060]" gridcol="0" gridrow="10" pageId="11" pageNumber="48">Rohožník (SVK)</th>
<td id="76825DE2FFC537E501F4FB84FC93B27C" box="[449,797,1038,1060]" gridcol="1" gridrow="10" pageId="11" pageNumber="48">Middle Miocene (Late Badenian)</td>
<td id="76825DE2FFC537E503AAFB84FAA3B27C" box="[927,1325,1038,1060]" gridcol="2" gridrow="10" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFBB2FAA3B216" box="[159,1325,1080,1102]" gridrow="11" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFBB2FE2FB216" box="[159,417,1080,1102]" gridcol="0" gridrow="11" pageId="11" pageNumber="48">Kamnik, Košiše (SLO)</th>
<td id="76825DE2FFC537E501F4FBB2FC93B216" box="[449,797,1080,1102]" gridcol="1" gridrow="11" pageId="11" pageNumber="48">Middle Miocene (Badenian)</td>
<td id="76825DE2FFC537E503AAFBB2FAA3B216" box="[927,1325,1080,1102]" gridcol="2" gridrow="11" pageId="11" pageNumber="48">herein</td>
</tr>
<tr id="3553349EFFC537E500AAFBEBFAA3B22F" box="[159,1325,1121,1143]" gridrow="12" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFBEBFE2FB22F" box="[159,417,1121,1143]" gridcol="0" gridrow="12" pageId="11" pageNumber="48">Želiezovce (SVK)</th>
<td id="76825DE2FFC537E501F4FBEBFC93B22F" box="[449,797,1121,1143]" gridcol="1" gridrow="12" pageId="11" pageNumber="48">Middle Miocene (Badenian)</td>
<td id="76825DE2FFC537E503AAFBEBFAA3B22F" box="[927,1325,1121,1143]" gridcol="2" gridrow="12" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81103AAFBEBFBEBB22F" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." box="[927,1125,1121,1143]" pageId="11" pageNumber="48" pagination="704 - 746" refId="ref15452" refString="Spinar, Z., Cepek, P., Boucek, B., Eliasova-Frajova, H., Havlicek, V., Horny, R., Housa, V., Kalabis, V., Kukalova, J., Marek, L., Nekvasilova, O., Pokorny, V., Prokop, R., Pribyl, A., Ruzicka, B., Snajdr, M., Stemprokova, D. &amp; Zikmundova, J. (1965) Systematicka paleontologie bezobratlych. Academia, Praha, 1049 pp. [pp. 704 - 746.]" type="book chapter" year="1965">Špinar et al. (1965)</bibRefCitation>
, herein
</td>
</tr>
<tr id="3553349EFFC537E500AAFB00FAA3B2F8" box="[159,1325,1162,1184]" gridrow="13" pageId="11" pageNumber="48">
<th id="76825DE2FFC537E500AAFB00FE2FB2F8" box="[159,417,1162,1184]" gridcol="0" gridrow="13" pageId="11" pageNumber="48">Superga, Turin hills (I)</th>
<td id="76825DE2FFC537E501F4FB00FC93B2F8" box="[449,797,1162,1184]" gridcol="1" gridrow="13" pageId="11" pageNumber="48">Middle Miocene</td>
<td id="76825DE2FFC537E503AAFB00FAA3B2F8" box="[927,1325,1162,1184]" gridcol="2" gridrow="13" pageId="11" pageNumber="48">
A.
<bibRefCitation id="EFF24B2DFFC5C811038BFB00FB28B2F8" author="Milne Edwards, A." box="[958,1190,1162,1184]" pageId="11" pageNumber="48" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards (1860)</bibRefCitation>
,
<bibRefCitation id="EFF24B2DFFC5C8110485FB00FAA3B2F8" author="Crema, C." box="[1200,1325,1162,1184]" pageId="11" pageNumber="48" pagination="664 - 681" refId="ref11114" refString="Crema, C. (1895) Sopra alcuni decapodi terziarii del Piemonte. Atti Reale Accademia delle Scienze di Torino, 30, 664 - 681." type="journal article" year="1895">Crema 1895</bibRefCitation>
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDC36DCFFC5C81100F2FB68FA13B366" blockId="11.[151,1437,1250,2023]" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81100F2FB68FEF4B2A3" author="Noetling, F." box="[199,378,1250,1275]" pageId="11" pageNumber="48" pagination="32 - 34" refId="ref14502" refString="Noetling, F. (1886) Uber Crustaceenreste aus dem Oberoligocanen Sternberger Gestein. Sitzingsberichte der Gesselschaft naturforschender Freunde Berlin, 1886, 32 - 34." type="journal article" year="1886">Noetling (1886</bibRefCitation>
: pl. 5, fig. 4; refigured here as
<figureCitation id="13582A59FFC5C81102C0FB68FCF8B2A3" box="[757,886,1250,1275]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="11" pageNumber="48">Figs 6HK</figureCitation>
) reported the species from the Late Oligocene “Sternberger Gestein”of Kobrow,
<collectingCountry id="F374764CFFC5C811022FFA8EFD0AB345" box="[538,644,1284,1309]" name="Germany" pageId="11" pageNumber="48">Germany</collectingCountry>
. The latest contribution on
<taxonomicName id="4C634D5FFFC5C811038AFA8EFB48B345" authorityName="Muller" authorityYear="1993" box="[959,1222,1284,1309]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC5C811038AFA8EFB48B345" box="[959,1222,1284,1309]" italics="true" pageId="11" pageNumber="48">Callianassa michelottii</emphasis>
</taxonomicName>
from this facies is that by
<bibRefCitation id="EFF24B2DFFC5C81100DFFAAFFE39B366" author="Polkowsky, S." box="[234,439,1317,1342]" pageId="11" pageNumber="48" pagination="1 - 126" refId="ref14603" refString="Polkowsky, S. (2005) Decapode Krebse aus dem oberoligozanen Sternberger Gestein von Kobrow (Mecklenburg). Tassados, 1, 1 - 126." type="journal article" year="2005">Polkowsky (2005)</bibRefCitation>
who discussed at length the variation of the species as well as its geographic distribution.
</paragraph>
<paragraph id="8BDC36DCFFC5C81100F2FACCFD7DB04B" blockId="11.[151,1437,1250,2023]" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81100F2FACCFE06B307" author="Lorenthey, I." box="[199,392,1350,1375]" pageId="11" pageNumber="48" pagination="202 - 259" refId="ref14078" refString="Lorenthey, I. (1907) Beitrage zur Tertiaren Dekapodenfauna Sardiniens. Mathematischen und Naturwissenchaftlichen Berichte aus Ungarn, 25, 202 - 259, 2 plates." type="book chapter" year="1907">Lőrenthey (1907</bibRefCitation>
: pl. 1, fig. 5; refigured in
<figureCitation id="13582A59FFC5C811029AFACCFC8AB307" box="[687,772,1350,1375]" captionStart="FIGURE 6" captionStartId="9.[151,249,1503,1525]" captionTargetBox="[160,1420,654,1470]" captionTargetId="figure-436@9.[151,1435,637,1479]" captionTargetPageId="9" captionText="FIGURE 6. Calliax michelottii (A. Milne Edwards, 1860) comb. nov., occurrences published previously. AG: Oligocene of the Mainz Basin, Flörsheim and Offenbach at Frankfurt (Germany). AC, right major P1 propodus articulated with dactylus from inner, outer and upper view, respectively; D, cross section of major P1 propodus; E, right major P1 merus articulated with carpus; F, right major P1 propodus articulated with carpus in inner view; G, left major P1 propodus associated with right minor P1 chela (propodus articulated with dactylus). H: Miocene of Sardinia (Italy), the assignment to C. michelottii comb. nov. is questionable. IL: Oligocene of „Sternberger Gestein“, Kobrow (Germany), left major P1 propodus in inner (I), outer (J), lower (K) and upper view (L). AG are refigured from Fritsch (1871), H is refigured from Lőrenthey (1907), IL are refigured from Noetling (1886). AH are to scale, scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919938" httpUri="https://zenodo.org/record/4919938/files/figure.png" pageId="11" pageNumber="48">Fig. 6L</figureCitation>
) reported and figured undetermined isolated propodi from the Miocene of Cagliari,
<collectingRegion id="49A7F83EFFC5C811018CFAE0FD94B3DB" box="[441,538,1386,1411]" country="Italy" name="Sardegna" pageId="11" pageNumber="48">Sardinia</collectingRegion>
. With two parallel ridges on the fixed finger, the specimen shows affinities to
<taxonomicName id="4C634D5FFFC5C81100A2FA04FEE4B3FF" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[151,362,1422,1447]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC5C81100A2FA04FEE4B3FF" box="[151,362,1422,1447]" italics="true" pageId="11" pageNumber="48">Calliax michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC5C8110147FA04FE7FB3FE" box="[370,497,1422,1447]" pageId="11" pageNumber="48" rank="species">comb. nov.</taxonomicNameLabel>
, but possesses rather convex upper propodal margin, which is unusual for
<taxonomicName id="4C634D5FFFC5C8110577FA04FA1AB3FF" authorityName="de Saint Laurent" authorityYear="1973" box="[1346,1428,1422,1447]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC5C8110577FA04FA1AB3FF" box="[1346,1428,1422,1447]" italics="true" pageId="11" pageNumber="48">Calliax</emphasis>
</taxonomicName>
, although the character itself may reflect mere intraspecific variation. The material (collection of Lovisato) is considered lost (
<bibRefCitation id="EFF24B2DFFC5C8110162FA5CFD38B3B7" author="De Angeli, A. &amp; Garassino, A." box="[343,694,1494,1519]" pageId="11" pageNumber="48" pagination="3 - 95" refId="ref11194" refString="De Angeli, A. &amp; Garassino, A. (2006) Catalog and bibliography of the fossil Stomatopoda and Decapoda from Italy. Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35 (1), 3 - 95." type="journal article" year="2006">De Angeli &amp; Garassino 2006</bibRefCitation>
); thus, the re-examination of the material is not possible. As a consequence, we treat the occurrence as questionable.
</paragraph>
<paragraph id="8BDC36DCFFC5C81100F2F994FBD3B0FB" blockId="11.[151,1437,1250,2023]" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81100F2F994FE21B06F" author="Wagner-Klett, W." box="[199,431,1566,1591]" pageId="11" pageNumber="48" pagination="73 - 118" refId="ref15729" refString="Wagner-Klett, W. (1919) Das Tertiar von Wiesloch in Baden. Ein Beitrag zu seiner tektonischen, stratigraphischen und palaontologischen Kenntnis. Jahresberichte und Mitteilungen des Oberrheinischen Geologischen Vereins, 8, 73 - 118. http: // dx. doi. org / 10.1127 / jmogv / 8 / 1919 / 73" type="journal article" year="1919">Wagner-Klett (1919)</bibRefCitation>
reported
<taxonomicName id="4C634D5FFFC5C811022FF994FC90B06F" authorityName="Muller" authorityYear="1993" box="[538,798,1566,1591]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC5C811022FF994FC90B06F" box="[538,798,1566,1591]" italics="true" pageId="11" pageNumber="48">Callianassa michelottii</emphasis>
</taxonomicName>
from the Oligocene Septarienton of Wiesloch,
<collectingCountry id="F374764CFFC5C811051AF994FA17B06F" box="[1327,1433,1566,1591]" name="Germany" pageId="11" pageNumber="48">Germany</collectingCountry>
. The figure he provided (
<bibRefCitation id="EFF24B2DFFC5C811018DF9C8FD16B003" author="Wagner-Klett, W." box="[440,664,1602,1627]" pageId="11" pageNumber="48" pagination="73 - 118" refId="ref15729" refString="Wagner-Klett, W. (1919) Das Tertiar von Wiesloch in Baden. Ein Beitrag zu seiner tektonischen, stratigraphischen und palaontologischen Kenntnis. Jahresberichte und Mitteilungen des Oberrheinischen Geologischen Vereins, 8, 73 - 118. http: // dx. doi. org / 10.1127 / jmogv / 8 / 1919 / 73" type="journal article" year="1919">Wagner-Klett 1919</bibRefCitation>
: pl. 2, fig. 1), however, does not conform to the diagnosis of the species. The specimen consisting of an articulated carpus, propodus, and dactylus shows no ridges on the fixed finger. The material was either not properly figured or it represents a different species.
</paragraph>
<paragraph id="8BDC36DCFFC5C81100F2F924FC11B157" blockId="11.[151,1437,1250,2023]" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81100F2F924FE03B09F" author="Glaessner, M. F." box="[199,397,1710,1735]" pageId="11" pageNumber="48" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner (1928)</bibRefCitation>
reported the species from the Helvetian (Early Miocene) of Jahring (today's Jarenina,
<collectingCountry id="F374764CFFC5C81100A2F958FF75B0B3" box="[151,251,1746,1771]" name="Slovenia" pageId="11" pageNumber="48">Slovenia</collectingCountry>
) in the Styrian Basin. He, however, did not provide figures. The re-examination of the material by the senior author confirms the identification of the material (
<figureCitation id="13582A59FFC5C8110326F97CFC1DB157" box="[787,915,1782,1807]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="11" pageNumber="48">Figs 8AB</figureCitation>
).
</paragraph>
<paragraph id="8BDC36DCFFC5C81100F2F890FCF2B1BF" blockId="11.[151,1437,1250,2023]" pageId="11" pageNumber="48">
<bibRefCitation id="EFF24B2DFFC5C81100F2F890FE30B16B" author="Beurlen, K." box="[199,446,1818,1843]" pageId="11" pageNumber="48" pagination="135 - 160" refId="ref10854" refString="Beurlen, K. (1939) Neue Decapoden-Krebse aus dem ungarischen Tertiar. Palaontologische Zeitschrift, 21, 135 - 160, 1 plate. http: // dx. doi. org / 10.1007 / bf 03043252" type="book chapter" year="1939">Beurlen (1939: 143)</bibRefCitation>
when discussing affinities of
<taxonomicName id="4C634D5FFFC5C8110377F890FA96B16B" authority="Beurlen, 1939" authorityName="Beurlen" authorityYear="1939" box="[834,1304,1818,1843]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="brevimanus">
<emphasis id="B917EACEFFC5C8110377F890FBD5B16B" box="[834,1115,1818,1843]" italics="true" pageId="11" pageNumber="48">Callianassa brevimanus</emphasis>
<bibRefCitation id="EFF24B2DFFC5C8110459F890FA96B16B" author="Beurlen, K." box="[1132,1304,1818,1843]" pageId="11" pageNumber="48" pagination="135 - 160" refId="ref10854" refString="Beurlen, K. (1939) Neue Decapoden-Krebse aus dem ungarischen Tertiar. Palaontologische Zeitschrift, 21, 135 - 160, 1 plate. http: // dx. doi. org / 10.1007 / bf 03043252" type="book chapter" year="1939">Beurlen, 1939</bibRefCitation>
</taxonomicName>
, (currently recognized as a member of
<taxonomicName id="4C634D5FFFC5C81101EFF8B4FCC4B10F" authority="Holmes, 1904" authorityName="Holmes" authorityYear="1904" box="[474,842,1854,1879]" class="Malacostraca" family="Callianassidae" genus="Lepidophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC5C81101EFF8B4FD13B10F" box="[474,669,1854,1879]" italics="true" pageId="11" pageNumber="48">Lepidophthalmus</emphasis>
<bibRefCitation id="EFF24B2DFFC5C8110293F8B4FCC4B10F" author="Holmes, S. J." box="[678,842,1854,1879]" pageId="11" pageNumber="48" pagination="307 - 328" refId="ref13391" refString="Holmes, S. J. (1904) On some new or imperfectly known species of West American Crustacea. Proceedings of the California Academy of Sciences, Series 3, 3, 307 - 328, Plates 35 - 37." type="journal article" year="1904">Holmes, 1904</bibRefCitation>
</taxonomicName>
; see Hyžný &amp; Dulai in press) briefly mentioned
<taxonomicName id="4C634D5FFFC5C81105B7F8B5FE81B123" authorityName="Muller" authorityYear="1993" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC5C81105B7F8B5FE81B123" italics="true" pageId="11" pageNumber="48">C. michelottii</emphasis>
</taxonomicName>
as coming from the Oligocene of the Mainz Basin and Miocene of
<collectingCountry id="F374764CFFC5C8110435F8E8FBBDB123" box="[1024,1075,1890,1915]" name="Italy" pageId="11" pageNumber="48">Italy</collectingCountry>
and the Vienna Basin, however, without any further reference. It is supposed here that
<bibRefCitation id="EFF24B2DFFC5C8110337F80CFC20B1C7" author="Beurlen, K." box="[770,942,1926,1951]" pageId="11" pageNumber="48" pagination="135 - 160" refId="ref10854" refString="Beurlen, K. (1939) Neue Decapoden-Krebse aus dem ungarischen Tertiar. Palaontologische Zeitschrift, 21, 135 - 160, 1 plate. http: // dx. doi. org / 10.1007 / bf 03043252" type="book chapter" year="1939">Beurlen (1939)</bibRefCitation>
mentioned the occurrence from the Vienna Basin erroneously, as there is no such published report known to the authors. He might have been referring to the published occurrence from the Styrian Basin by
<bibRefCitation id="EFF24B2DFFC5C8110281F844FCF6B1BF" author="Glaessner, M. F." box="[692,888,1998,2023]" pageId="11" pageNumber="48" pagination="161 - 219" refId="ref12488" refString="Glaessner, M. F. (1928) Die Dekapodenfauna des osterreichischen Jungtertiars. Jahrbuch der Geologischen Bundesanstalt Wien, 78, 161 - 219." type="journal article" year="1928">Glaessner (1928)</bibRefCitation>
.
</paragraph>
<caption id="DF1C6654FFC2C81600A2FBBAFBDFB357" ID-DOI="http://doi.org/10.5281/zenodo.4919946" ID-Zenodo-Dep="4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="12" pageNumber="49" startId="12.[151,249,1072,1094]" targetBox="[155,1436,195,1047]" targetPageId="12">
<paragraph id="8BDC36DCFFC2C81600A2FBBAFBDFB357" blockId="12.[151,1436,1072,1295]" pageId="12" pageNumber="49">
<emphasis id="B917EACEFFC2C81600A2FBBAFE9BB21D" bold="true" box="[151,277,1072,1094]" pageId="12" pageNumber="49">FIGURE 8.</emphasis>
<taxonomicName id="4C634D5FFFC2C816012AFBBAFD73B21E" authority="(A. Milne Edwards, 1860)" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[287,765,1072,1094]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC2C816012AFBBAFE51B21E" box="[287,479,1072,1094]" italics="true" pageId="12" pageNumber="49">Calliax michelottii</emphasis>
(A.
<bibRefCitation id="EFF24B2DFFC2C8160226FBBAFD78B21E" author="Milne Edwards, A." box="[531,758,1072,1094]" pageId="12" pageNumber="49" pagination="129 - 357" refId="ref14289" refString="Milne Edwards, A. (1860) Histoire des Crustaces podophthalmaires fossiles et Monographie des Decapodes Macroures de la famille des Thalassinens. Annales des Sciences Naturelles, 4 e Serie 14, 129 - 357." type="journal article" year="1860">Milne Edwards, 1860</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC2C8160332FBBAFCF5B21D" box="[775,891,1072,1094]" pageId="12" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of
<taxonomicName id="4C634D5FFFC2C8160516FB3FFE06B2B4" authority="Muller, 1984" authorityName="Muller" authorityYear="1984" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC2C8160516FB3FFF79B2B4" italics="true" pageId="12" pageNumber="49">Callianassa szobensis</emphasis>
<bibRefCitation id="EFF24B2DFFC2C8160134FB5CFE06B2B4" author="Muller, P." box="[257,392,1238,1260]" pageId="12" pageNumber="49" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
</taxonomicName>
from Szob (Hungary), refigured from
<bibRefCitation id="EFF24B2DFFC2C8160310FB5CFC3DB2B4" author="Muller, P." box="[805,947,1238,1260]" pageId="12" pageNumber="49" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller (1984</bibRefCitation>
: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from
<bibRefCitation id="EFF24B2DFFC2C8160228FB73FD53B357" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." box="[541,733,1273,1295]" pageId="12" pageNumber="49" pagination="704 - 746" refId="ref15452" refString="Spinar, Z., Cepek, P., Boucek, B., Eliasova-Frajova, H., Havlicek, V., Horny, R., Housa, V., Kalabis, V., Kukalova, J., Marek, L., Nekvasilova, O., Pokorny, V., Prokop, R., Pribyl, A., Ruzicka, B., Snajdr, M., Stemprokova, D. &amp; Zikmundova, J. (1965) Systematicka paleontologie bezobratlych. Academia, Praha, 1049 pp. [pp. 704 - 746.]" type="book chapter" year="1965">
Špinar
<emphasis id="B917EACEFFC2C8160250FB70FD1FB357" box="[613,657,1273,1295]" italics="true" pageId="12" pageNumber="49">et al</emphasis>
. (1965
</bibRefCitation>
: fig. X-184). Scale bar equals 5 mm.
</paragraph>
</caption>
<paragraph id="8BDC36DCFFC2C81600F2FAB5FE1AB39D" blockId="12.[151,1436,1343,2016]" pageId="12" pageNumber="49">
Several chelae from the Middle Miocene (“Badenian”) of Želiezovce (Slovak part of the Danube Basin) figured by Houša
<emphasis id="B917EACEFFC2C8160154FAEFFEF9B324" box="[353,375,1381,1404]" italics="true" pageId="12" pageNumber="49">in</emphasis>
<bibRefCitation id="EFF24B2DFFC2C816014BFAEEFDDDB325" author="Spinar, Z. &amp; Cepek, P. &amp; Boucek, B. &amp; Eliasova-Frajova, H. &amp; Havlicek, V. &amp; Horny, R. &amp; Housa, V. &amp; Kalabis, V. &amp; Kukalova, J. &amp; Marek, L. &amp; Nekvasilova, O. &amp; Pokorny, V. &amp; Prokop, R. &amp; Pribyl, A. &amp; Ruzicka, B. &amp; Snajdr, M. &amp; Stemprokova, D. &amp; Zikmundova, J." box="[382,595,1380,1405]" pageId="12" pageNumber="49" pagination="704 - 746" refId="ref15452" refString="Spinar, Z., Cepek, P., Boucek, B., Eliasova-Frajova, H., Havlicek, V., Horny, R., Housa, V., Kalabis, V., Kukalova, J., Marek, L., Nekvasilova, O., Pokorny, V., Prokop, R., Pribyl, A., Ruzicka, B., Snajdr, M., Stemprokova, D. &amp; Zikmundova, J. (1965) Systematicka paleontologie bezobratlych. Academia, Praha, 1049 pp. [pp. 704 - 746.]" type="book chapter" year="1965">
Špinar
<emphasis id="B917EACEFFC2C81601FBFAEFFD8EB325" box="[462,512,1380,1405]" italics="true" pageId="12" pageNumber="49">et al</emphasis>
. (1965
</bibRefCitation>
: figs. X-184185, refigured herein as 8L) as
<taxonomicName id="4C634D5FFFC2C8160479FAEEFB77B323" box="[1100,1273,1380,1405]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B917EACEFFC2C8160479FAEEFB5CB325" box="[1100,1234,1380,1405]" italics="true" pageId="12" pageNumber="49">Callianassa</emphasis>
sp.
</taxonomicName>
can be clearly assigned to
<taxonomicName id="4C634D5FFFC2C816012EFA0DFD91B3F8" authorityName="Muller" authorityYear="1993" box="[283,543,1415,1440]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C816012EFA0DFD91B3F8" box="[283,543,1415,1440]" italics="true" pageId="12" pageNumber="49">Callianassa michelottii</emphasis>
</taxonomicName>
. Unfortunately, the repeated search for the material was unsuccessful, thus, it is herein considered lost.
</paragraph>
<paragraph id="8BDC36DCFFC2C81600F2FA45FEE1B02F" blockId="12.[151,1436,1343,2016]" pageId="12" pageNumber="49">
<bibRefCitation id="EFF24B2DFFC2C81600F2FA45FE73B3B0" author="Philippe, M. &amp; Secretan, S." box="[199,509,1487,1512]" pageId="12" pageNumber="49" pagination="117 - 134" refId="ref14530" refString="Philippe, M. &amp; Secretan, S. (1971) Crustaces decapods du Burdigalien des Courennes (Vaucluse). Annales de Paleontologie (Invertebres), 57, 117 - 134." type="journal article" year="1971">Philippe &amp; Secretan (1971)</bibRefCitation>
reported 10 fragmentary propodi from the Burdigalian of SE
<collectingCountry id="F374764CFFC2C8160489FA5AFA82B3BF" box="[1212,1292,1488,1511]" name="France" pageId="12" pageNumber="49">France</collectingCountry>
. The figured specimens (Pl. C, figs. 13, 14) show the keeled fixed finger accompanied with furrows, thus, pointing to attribution of the material to
<taxonomicName id="4C634D5FFFC2C816016AF99DFE3CB068" authorityName="de Saint Laurent" authorityYear="1973" box="[351,434,1559,1584]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC2C816016AF99DFE3CB068" box="[351,434,1559,1584]" italics="true" pageId="12" pageNumber="49">Calliax</emphasis>
</taxonomicName>
. The age and location of the specimens would speak for indentification as
<taxonomicName id="4C634D5FFFC2C81604C9F992FA1BB068" authorityName="Muller" authorityYear="1993" box="[1276,1429,1559,1584]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C81604C9F992FA1BB068" box="[1276,1429,1559,1584]" italics="true" pageId="12" pageNumber="49">C. michelottii</emphasis>
</taxonomicName>
. Without personal re-examination of the material, however, we are reluctant to treat the specimens as conspecific with
<taxonomicName id="4C634D5FFFC2C81600FAF9EAFEE6B020" authorityName="Muller" authorityYear="1993" box="[207,360,1631,1656]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C81600FAF9EAFEE6B020" box="[207,360,1631,1656]" italics="true" pageId="12" pageNumber="49">C. michelottii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDC36DCFFC2C81600F2F90EFA8BB15F" blockId="12.[151,1436,1343,2016]" pageId="12" pageNumber="49">
The description and figures of “
<taxonomicName id="4C634D5FFFC2C8160205F90EFD38B0C5" authorityName="Michelotti A. Milne Edwards" authorityYear="1860" box="[560,694,1668,1693]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B917EACEFFC2C8160205F90EFD38B0C5" box="[560,694,1668,1693]" italics="true" pageId="12" pageNumber="49">Callianassa</emphasis>
</taxonomicName>
<taxonomicName id="4C634D5FFFC2C81602FFF90EFC43B0C5" authority="Muller, 1984" authorityName="Muller" authorityYear="1984" box="[714,973,1668,1693]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC2C81602FFF90EFCBAB0C5" box="[714,820,1668,1693]" italics="true" pageId="12" pageNumber="49">szobensis</emphasis>
<bibRefCitation id="EFF24B2DFFC2C8160309F90EFC43B0C5" author="Muller, P." box="[828,973,1668,1693]" pageId="12" pageNumber="49" pagination="3 - 317" refId="ref14428" refString="Muller, P. (1984) Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica, 42, 3 - 317." type="journal article" year="1984">Müller, 1984</bibRefCitation>
</taxonomicName>
from the Middle Miocene (Badenian) of
<collectingCountry id="F374764CFFC2C81600A2F922FF77B098" box="[151,249,1704,1728]" name="Hungary" pageId="12" pageNumber="49">Hungary</collectingCountry>
fit the diagnosis and variations of
<taxonomicName id="4C634D5FFFC2C81602A9F922FCB6B098" authorityName="Muller" authorityYear="1993" box="[668,824,1703,1728]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C81602A9F922FCB6B098" box="[668,824,1703,1728]" italics="true" pageId="12" pageNumber="49">C. michelottii</emphasis>
</taxonomicName>
. The propodus is longer than high, the fixed finger possesses two ridges parallel to one another, and there are tubercles on the lateral surface of the propodus (
<figureCitation id="13582A59FFC2C816055FF946FF59B150" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="12" pageNumber="49">Figs 8JK</figureCitation>
). As a consequence, “
<emphasis id="B917EACEFFC2C81601F9F97AFE68B15F" box="[460,486,1776,1799]" italics="true" pageId="12" pageNumber="49">C.</emphasis>
<taxonomicName id="4C634D5FFFC2C81601CCF965FDEDB150" authorityName="Muller" authorityYear="1984" box="[505,611,1775,1800]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="szobensis">
<emphasis id="B917EACEFFC2C81601CCF965FDEDB150" box="[505,611,1775,1800]" italics="true" pageId="12" pageNumber="49">szobensis</emphasis>
</taxonomicName>
is considered a junior subjective synonym of
<taxonomicName id="4C634D5FFFC2C8160453F97AFB70B150" authorityName="Muller" authorityYear="1993" box="[1126,1278,1775,1800]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C8160453F97AFB70B150" box="[1126,1278,1775,1800]" italics="true" pageId="12" pageNumber="49">C. michelottii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDC36DCFFC2C81600F2F89EFDF0B1C0" blockId="12.[151,1436,1343,2016]" pageId="12" pageNumber="49">
From the Miocene of Spain (
<collectingRegion id="49A7F83EFFC2C816023EF89EFDF2B175" box="[523,636,1812,1837]" country="Spain" name="Catalunya" pageId="12" pageNumber="49">Catalonia</collectingRegion>
),
<bibRefCitation id="EFF24B2DFFC2C81602A5F89EFCA7B175" author="Muller, P." box="[656,809,1812,1837]" pageId="12" pageNumber="49" pagination="1 - 39" refId="ref14453" refString="Muller, P. (1993) Neogene decapod crustaceans from Catalonia. Scripta Musei Geologici Seminarii Barcinonensis, 225, 1 - 39." type="journal article" year="1993">Müller (1993</bibRefCitation>
: fig. 3A) reported and figured an isolated propodus of a callianassid shrimp. He classified it as
<taxonomicName id="4C634D5FFFC2C8160273F8BDFCE3B108" authorityName="Muller" authorityYear="1993" box="[582,877,1847,1872]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C8160273F8BDFD42B108" box="[582,716,1847,1872]" italics="true" pageId="12" pageNumber="49">Callianassa</emphasis>
cf.
<emphasis id="B917EACEFFC2C81602C0F8BDFCE3B108" box="[757,877,1847,1872]" italics="true" pageId="12" pageNumber="49">michelottii</emphasis>
</taxonomicName>
. The specimen, however, is rather dissimilar to
<taxonomicName id="4C634D5FFFC2C81605B7F8B2FE81B12D" authorityName="Muller" authorityYear="1993" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii">
<emphasis id="B917EACEFFC2C81605B7F8B2FA1BB117" box="[1410,1429,1848,1871]" italics="true" pageId="12" pageNumber="49">C</emphasis>
.
<emphasis id="B917EACEFFC2C81600A2F8D6FE81B12D" box="[151,271,1884,1909]" italics="true" pageId="12" pageNumber="49">michelottii</emphasis>
</taxonomicName>
; it does not possess a double ridge on the fixed finger and has the upper margin converging proximally, which is very atypical for the latter species.
</paragraph>
<paragraph id="8BDC36DCFFC2C81700F2F82EFBF2B745" blockId="12.[151,1436,1343,2016]" lastBlockId="13.[151,1437,151,285]" lastPageId="13" lastPageNumber="50" pageId="12" pageNumber="49">
<taxonomicName id="4C634D5FFFC2C81600F2F82EFE15B1E5" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[199,411,1956,1981]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC2C81600F2F82EFE15B1E5" box="[199,411,1956,1981]" italics="true" pageId="12" pageNumber="49">Calliax michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC2C8160191F82EFD87B1E5" box="[420,521,1956,1981]" pageId="12" pageNumber="49" rank="species">n. comb.</taxonomicNameLabel>
is morphologically very close to its extant congeners. The upper margin of the major P1 merus is, however, more convex in
<taxonomicName id="4C634D5FFFC2C81602A6F842FCA5B1B8" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[659,811,1991,2016]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC2C81602A6F842FCA5B1B8" box="[659,811,1991,2016]" italics="true" pageId="12" pageNumber="49">C. michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC2C8160307F842FC1AB1B8" box="[818,916,1991,2016]" pageId="12" pageNumber="49" rank="species">n. comb.</taxonomicNameLabel>
(
<bibRefCitation id="EFF24B2DFFC2C8160397F84DFBBAB1B8" author="Fritsch, K. von" box="[930,1076,1991,2016]" pageId="12" pageNumber="49" pagination="679 - 701" refId="ref12298" refString="Fritsch, K. von (1871) Ueber einige fossile Crustaceen aus dem Septarienthon des Mainzer Beckens. Zeitschrift der Deutschen geologischen Gesselschaft, 23, 679 - 701, plates 16 - 17." type="journal article" year="1871">Fritsch 1871</bibRefCitation>
: pl. 17, fig. 9; see also
<figureCitation id="13582A59FFC2C8160503F84DFA00B1B8" box="[1334,1422,1991,2016]" captionStart="FIGURE 8" captionStartId="12.[151,249,1072,1094]" captionTargetBox="[155,1436,195,1047]" captionTargetId="figure-449@12.[150,1436,195,1047]" captionTargetPageId="12" captionText="FIGURE 8. Calliax michelottii (A. Milne Edwards, 1860) comb. nov. from the Alpine-Carpathian orogenic system. AB, UMJGP 7787377874, isolated propodi from Jarenina (Slovenia); C, NHMW 2003/0026/0913, right propodus articulated with dactylus from Pucking (Austria); DH, KGP-MH RO-001, fragmentary right propodus from Rohožník (Slovakia), note large setal pores on the outer lateral surface (E), arrows (in F) indicate ridges; I, NHMW 2010/0089/0001, counterpart of the major left cheliped from Neuhofen bei Tettenweis (Germany), arrows indicate ridges; JK, M.2004.158.1, holotype of Callianassa szobensis Müller, 1984 from Szob (Hungary), refigured from Müller (1984: pl. 7, figs 34); L, left major propodus from Želiezovce (Slovakia), refigured from Špinar et al. (1965: fig. X-184). Scale bar equals 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4919946" httpUri="https://zenodo.org/record/4919946/files/figure.png" pageId="12" pageNumber="49">Figs. 8I</figureCitation>
), whereas it is rather straight in both extant species,
<taxonomicName id="4C634D5FFFC3C81702EDFF12FCD2B6E8" authorityName="Sakai" authorityYear="1999" box="[728,860,151,176]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="doerjesti">
<emphasis id="B917EACEFFC3C81702EDFF12FCD2B6E8" box="[728,860,151,176]" italics="true" pageId="13" pageNumber="50">C. doerjesti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D5FFFC3C81703A0FF12FC70B6E8" baseAuthorityName="de Gaillande &amp; Lagardere" baseAuthorityYear="1966" box="[917,1022,151,176]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="B917EACEFFC3C81703A0FF12FC70B6E8" box="[917,1022,151,176]" italics="true" pageId="13" pageNumber="50">C. lobata</emphasis>
</taxonomicName>
(
<figureCitation id="13582A59FFC3C817043AFF1DFBE9B6E8" box="[1039,1127,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,249,1131,1153]" captionTargetBox="[165,1429,185,1107]" captionTargetId="figure-274@4.[150,1437,183,1107]" captionTargetPageId="4" captionText="FIGURE 3. Extant Calliax species: AB, Calliax doerjesti Sakai, 1999, note the discrepancy in figuring lower margin of major P1 merus in „A“ and „a“ (redrawn after Sakai 1999: figs 2829); CD, Calliax lobata (de Gaillande &amp; Lagardère, 1966) (redrawn after Ngoc-Ho 2003: figs 17DE); EF, Calliax cf. C. lobata, line drawings of NHMW 25511 (Fig. 2). Left column=major chelae (outer surface), right column=minor chelae (outer surface). Setae are omitted. From two ridges at the base of the fixed finger only the more developed one is depicted in the sketches (see text for details). Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919928" httpUri="https://zenodo.org/record/4919928/files/figure.png" pageId="13" pageNumber="50">Fig. 3A</figureCitation>
and
<figureCitation id="13582A59FFC3C8170495FF1DFB79B6E8" box="[1184,1271,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,249,1131,1153]" captionTargetBox="[165,1429,185,1107]" captionTargetId="figure-274@4.[150,1437,183,1107]" captionTargetPageId="4" captionText="FIGURE 3. Extant Calliax species: AB, Calliax doerjesti Sakai, 1999, note the discrepancy in figuring lower margin of major P1 merus in „A“ and „a“ (redrawn after Sakai 1999: figs 2829); CD, Calliax lobata (de Gaillande &amp; Lagardère, 1966) (redrawn after Ngoc-Ho 2003: figs 17DE); EF, Calliax cf. C. lobata, line drawings of NHMW 25511 (Fig. 2). Left column=major chelae (outer surface), right column=minor chelae (outer surface). Setae are omitted. From two ridges at the base of the fixed finger only the more developed one is depicted in the sketches (see text for details). Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4919928" httpUri="https://zenodo.org/record/4919928/files/figure.png" pageId="13" pageNumber="50">Fig. 3C</figureCitation>
, respectively). The development of spines on the lower margin of the major P1 merus in
<taxonomicName id="4C634D5FFFC3C81703FEFF36FB12B68D" authorityName="Hyžný &amp; Gašparič" authorityYear="2014" baseAuthorityName="A. Milne Edwards" baseAuthorityYear="1860" box="[971,1180,188,213]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="michelottii" status="comb. nov.">
<emphasis id="B917EACEFFC3C81703FEFF36FB12B68D" box="[971,1180,188,213]" italics="true" pageId="13" pageNumber="50">Calliax michelottii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457B5FFC3C8170497FF36FA90B68B" box="[1186,1310,188,213]" pageId="13" pageNumber="50" rank="species">comb. nov.</taxonomicNameLabel>
is closer to that of
<taxonomicName id="4C634D5FFFC3C81700DCFF6AFEDDB6A0" baseAuthorityName="de Gaillande &amp; Lagardere" baseAuthorityYear="1966" box="[233,339,223,248]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="50" phylum="Arthropoda" rank="species" species="lobata">
<emphasis id="B917EACEFFC3C81700DCFF6AFEDDB6A0" box="[233,339,223,248]" italics="true" pageId="13" pageNumber="50">C. lobata</emphasis>
</taxonomicName>
. The tooth formula of the occlusal margin of the fixed finger looks different in all three species, but this may be a matter of variation and is not considered taxonomically important here.
</paragraph>
</subSubSection>
</treatment>
</document>