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<document id="2C5AF71E3304FB84A83CBD083A984A8C" ID-DOI="10.11646/zootaxa.5351.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8392232" ID-ZooBank="1BD2A231-4D66-47A7-8EE6-4A1C241C4190" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695992926648" checkinUser="plazi" docAuthor="Zhao, Fangyu, Yang, Lan, Li, Shuqiang, Zheng, Guo &amp; Yao, Zhiyuan" docDate="2023" docId="03CA8799E72DFFC112C1D1ECDD78F982" docLanguage="en" docName="zootaxa.5351.5.3.pdf" docOrigin="Zootaxa 5351 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana daxiangi Li, Zheng &amp; Yao 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="549" masterDocId="FFF3FFE1E72BFFC71256D03EDE72FF83" masterDocTitle="A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China" masterLastPageNumber="558" masterPageNumber="543" pageNumber="549" updateTime="1695993610286" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4D0E9D9776DF42014C8D98840E96A848">A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China</mods:title>
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<mods:namePart id="ED518A77F4B27A12E9BC7D6AF5498EB5">Zhao, Fangyu</mods:namePart>
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<mods:namePart id="8A9A58E552D8F786374EB291392ABB93">Yang, Lan</mods:namePart>
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<mods:namePart id="6A768FCD2BD6305F3547107235B4232D">Li, Shuqiang</mods:namePart>
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<mods:namePart id="9D8DE19D5AC259DC274D712E1F6B4BEC">Zheng, Guo</mods:namePart>
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<heading id="D09481E3E72DFFC112C1D1ECDCDCFE6E" bold="true" box="[151,686,466,493]" fontSize="11" level="1" pageId="6" pageNumber="549" reason="1">
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<taxonomicName id="4C634D0CE72DFFC112C1D1ECDC3BFE6E" authority="Li, Zheng &amp; Yao" authorityName="Li, Zheng &amp; Yao" authorityYear="2023" box="[151,585,466,493]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="daxiangi" status="sp. nov.">
<emphasis id="B917EA9DE72DFFC112C1D1ECDF00FE6E" bold="true" box="[151,370,466,493]" italics="true" pageId="6" pageNumber="549">Belisana daxiangi</emphasis>
Li, Zheng &amp; Yao
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<taxonomicNameLabel id="A22457E6E72DFFC11007D1EDDCDCFE6E" box="[593,686,467,493]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3796504E72DFFC112C1D1C9DF3BFD92" box="[151,329,502,529]" pageId="6" pageNumber="549" type="description">
<paragraph id="8BDC368FE72DFFC112C1D1C9DF3BFD92" blockId="6.[151,686,466,529]" box="[151,329,502,529]" pageId="6" pageNumber="549">
<figureCitation id="13582A0AE72DFFC112C1D1C9DE89FD92" box="[151,251,502,529]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="7.[151,250,1862,1886]" captionStartId-1="8.[151,250,1943,1967]" captionTargetBox-0="[249,1338,237,1839]" captionTargetBox-1="[203,1384,181,1919]" captionTargetId-0="figure-37@7.[249,1338,237,1839]" captionTargetId-1="figure-22@8.[203,1384,181,1919]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." captionText-1="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi-0="http://doi.org/10.5281/zenodo.8392244" figureDoi-1="http://doi.org/10.5281/zenodo.8392248" httpUri-0="https://zenodo.org/record/8392244/files/figure.png" httpUri-1="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Figs 45</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11350D1C9DF3BFD92" box="[262,329,503,529]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9CD</figureCitation>
</paragraph>
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<paragraph id="8BDC368FE72DFFC112C1D201DEB3FD22" blockId="6.[151,1437,574,1537]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC112C1D201DFB4FDDA" bold="true" box="[151,454,574,601]" pageId="6" pageNumber="549">
Type material.
<typeStatus id="54D8882DE72DFFC11319D201DFB2FDDA" box="[335,448,575,601]" pageId="6" pageNumber="549">Holotype</typeStatus>
:
</emphasis>
Male (SYNU-Ar00310), near
<taxonomicName id="4C634D0CE72DFFC11173D201DDA4FDDA" authority="Village" authorityName="Village" box="[805,982,575,601]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="bubeng">Bubeng Village</taxonomicName>
(
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,
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, elevation
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),
<taxonomicName id="4C634D0CE72DFFC112A0D25DDC11FDFE" authority="County, Xishuangbanna" authorityName="County, Xishuangbanna" box="[246,611,611,637]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="mengla">Mengla County, Xishuangbanna</taxonomicName>
,
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<collectingRegion id="49A7F86DE72DFFC11038D25DDCBEFDFE" box="[622,716,611,637]" country="China" name="Yunnan" pageId="6" pageNumber="549">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="B917EA9DE72DFFC1108CD25CDD51FDFF" bold="true" box="[730,803,610,636]" pageId="6" pageNumber="549">
<collectingCountry id="F374761FE72DFFC1108CD25CDD51FDFF" box="[730,803,610,636]" name="China" pageId="6" pageNumber="549">China</collectingCountry>
</emphasis>
,
<date id="FFDD104FE72DFFC11164D25DDD95FDFE" box="[818,999,611,637]" pageId="6" pageNumber="549" value="2012-08-12">12 August 2012</date>
, Guo Zheng, Xue Li, and Wenyue Zhu leg.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E72DFFC11291D295DFBEFD6A" pageId="6" pageNumber="549" type="etymology">
<paragraph id="8BDC368FE72DFFC11291D295DFBEFD6A" blockId="6.[151,1437,574,1537]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D295DF3CFD46" bold="true" box="[199,334,683,709]" pageId="6" pageNumber="549">Etymology.</emphasis>
The specific name is a patronym in honor of the late Chinese arachnologist Daxiang Song; noun (name) in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E72DFFC11291D2CCDF02FC42" pageId="6" pageNumber="549" type="diagnosis">
<paragraph id="8BDC368FE72DFFC11291D2CCDF02FC42" blockId="6.[151,1437,574,1537]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D2CCDF30FC8F" bold="true" box="[199,322,754,780]" pageId="6" pageNumber="549">Diagnosis.</emphasis>
The new species resembles
<taxonomicName id="4C634D0CE72DFFC110DFD2CDDD64FC8E" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" box="[649,790,755,781]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fengzheni" status="sp. nov.">
<emphasis id="B917EA9DE72DFFC110DFD2CDDD64FC8E" box="[649,790,755,781]" italics="true" pageId="6" pageNumber="549">B. fengzheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457E6E72DFFC11149D2CDDD0AFC8E" box="[799,888,755,781]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13582A0AE72DFFC111DED2CDDD9DFC8E" box="[904,1007,755,781]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="9.[151,250,1856,1880]" captionStartId-1="10.[151,250,1856,1880]" captionTargetBox-0="[233,1355,181,1832]" captionTargetBox-1="[233,1355,181,1832]" captionTargetId-0="figure-18@9.[233,1355,181,1832]" captionTargetId-1="figure-22@10.[233,1355,181,1832]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 6. Belisana fengzheni sp. nov., holotype male (SYNU-Ar00311). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." captionText-1="FIGURE 7. Belisana fengzheni sp. nov., holotype male (C, EF; SYNU-Ar00311), paratype male (D, SYNU-Ar00312) and paratype female (AB, GH; SYNU-Ar00315). A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.8392254" figureDoi-1="http://doi.org/10.5281/zenodo.8392258" httpUri-0="https://zenodo.org/record/8392254/files/figure.png" httpUri-1="https://zenodo.org/record/8392258/files/figure.png" pageId="6" pageNumber="549">
Figs 6
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</figureCitation>
,
<figureCitation id="13582A0AE72DFFC111ABD2CDDA48FC8E" box="[1021,1082,755,781]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9EF</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11611D2CDDAEBFC8E" box="[1095,1177,755,781]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="6" pageNumber="549">10CD</figureCitation>
) having similar bulbal apophyses (
<figureCitation id="13582A0AE72DFFC11348D329DF0BFCB2" box="[286,377,790,817]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-22@8.[203,1384,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.8392248" httpUri="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Fig. 5A</figureCitation>
) and male chelicerae (
<figureCitation id="13582A0AE72DFFC110D4D329DCAFFCB2" box="[642,733,790,817]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-22@8.[203,1384,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.8392248" httpUri="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Fig. 5B</figureCitation>
), but it can be distinguished by prolatero-distal membranous lamella of procursus with weakly sclerotized lamella (arrow
<quantity id="4C9B9B6AE72DFFC11112D305DD02FCD6" box="[836,880,827,853]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="549" unit="in" value="1.0">1 in</quantity>
<figureCitation id="13582A0AE72DFFC11121D305DDA0FCD6" box="[887,978,827,853]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11188D305DA70FCD6" box="[990,1026,827,853]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
; vs. absent in
<taxonomicName id="4C634D0CE72DFFC116F4D305DB5FFCD6" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" box="[1186,1325,827,853]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fengzheni" status="sp. nov.">
<emphasis id="B917EA9DE72DFFC116F4D305DB5FFCD6" box="[1186,1325,827,853]" italics="true" pageId="6" pageNumber="549">B. fengzheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457E6E72DFFC11763D305DBE2FCD6" box="[1333,1424,827,853]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
), by procursus with two spine-shaped dorso-distal apophyses (arrows 2,
<quantity id="4C9B9B6AE72DFFC111E3D361DD92FCFA" box="[949,992,863,889]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="549" unit="in" value="3.0">3 in</quantity>
<figureCitation id="13582A0AE72DFFC111B1D361DA30FCFA" box="[999,1090,863,889]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11618D361DA00FCFA" box="[1102,1138,863,889]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
; vs. only one apophysis in
<taxonomicName id="4C634D0CE72DFFC112C1D3BDDF53FC1E" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" box="[151,289,899,925]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fengzheni" status="sp. nov.">
<emphasis id="B917EA9DE72DFFC112C1D3BDDF53FC1E" box="[151,289,899,925]" italics="true" pageId="6" pageNumber="549">B. fengzheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457E6E72DFFC1137ED3BDDFF0FC1E" box="[296,386,899,925]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
), and by procursus with ventral membranous lamella (arrow
<quantity id="4C9B9B6AE72DFFC1167DD3BDDA24FC1E" box="[1067,1110,899,925]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="549" unit="in" value="4.0">4 in</quantity>
<figureCitation id="13582A0AE72DFFC1160BD3BDDAC5FC1E" box="[1117,1207,899,925]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11695D3BDDA95FC1E" box="[1219,1255,899,925]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
; vs. absent in
<taxonomicName id="4C634D0CE72DFFC117D2D3BDDF70FC42" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="549" phylum="Arthropoda" rank="species" species="fengzheni" status="sp. nov.">
<emphasis id="B917EA9DE72DFFC117D2D3BDDF70FC42" italics="true" pageId="6" pageNumber="549">B. fengzheni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22457E6E72DFFC1135FD399DF16FC42" box="[265,356,935,961]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C3796504E72DFFC11291D3F4DAE4FA5E" pageId="6" pageNumber="549" type="description">
<paragraph id="8BDC368FE72DFFC11291D3F4DF74FA16" blockId="6.[151,1437,574,1537]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D3F4DFD2FC66" bold="true" box="[199,416,970,997]" pageId="6" pageNumber="549">Description. Male</emphasis>
(
<emphasis id="B917EA9DE72DFFC113E5D3F5DC68FC66" bold="true" box="[435,538,971,997]" pageId="6" pageNumber="549">
<typeStatus id="54D8882DE72DFFC113E5D3F5DC68FC66" box="[435,538,971,997]" pageId="6" pageNumber="549">holotype</typeStatus>
</emphasis>
, SYNU-Ar00310): Total length 1.58 (1.66 with clypeus), prosoma 0.57 long, 0.55 wide, opisthosoma 1.01 long, 0.53 wide. Legs I and II missing, leg III: 5.05 (1.37 + 0.19 + 1.23 + 1.74 + 0.52), leg IV: (2.30 + 0.20 + 2.06 + + ). Eye interdistances and diameters: PME-PME 0.14, PME 0.06, PME-ALE 0.03, AME absent. Sternum width/length: 0.45/0.45. Habitus as in
<figureCitation id="13582A0AE72DFFC1119ED409DA36FBD2" box="[968,1092,1078,1105]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-22@8.[203,1384,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.8392248" httpUri="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Figs 5CE</figureCitation>
. Dorsal shield of prosoma and sternum whitish, without marks. Legs whitish, without darker rings. Opisthosoma whitish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="13582A0AE72DFFC110E0D441DD79FB1A" box="[694,779,1150,1177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-22@8.[203,1384,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.8392248" httpUri="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Fig. 5B</figureCitation>
) with pair of proximo-lateral apophyses and pair of curved distal apophyses (distance between tips: 0.29). Pedipalp as in
<figureCitation id="13582A0AE72DFFC1110ED49DDDA5FB3E" box="[856,983,1187,1213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4AB</figureCitation>
; trochanter as long as wide; femur with retrolatero-proximal apophysis (arrow in
<figureCitation id="13582A0AE72DFFC1103AD4F9DCB6FB62" box="[620,708,1223,1249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Fig. 4B</figureCitation>
); procursus (
<figureCitation id="13582A0AE72DFFC11101D4F9DDABFB62" box="[855,985,1223,1249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4AD</figureCitation>
,
<figureCitation id="13582A0AE72DFFC111B3D4F9DA58FB62" box="[997,1066,1223,1249]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9CD</figureCitation>
) simple proximally but complex distally, with prolatero-distal membranous lamella with weakly sclerotized lamella (arrow
<quantity id="4C9B9B6AE72DFFC116CAD4D5DABAFA86" box="[1180,1224,1259,1285]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="549" unit="in" value="1.0">1 in</quantity>
<figureCitation id="13582A0AE72DFFC11686D4D5DB5EFA86" box="[1232,1324,1259,1285]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC1176FD4D5DB2FFA86" box="[1337,1373,1259,1285]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
), two spine-shaped dorso-distal apophyses (arrows 2,
<quantity id="4C9B9B6AE72DFFC110EDD531DC9AFAAA" box="[699,744,1295,1321]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="549" unit="in" value="3.0">3 in</quantity>
<figureCitation id="13582A0AE72DFFC110A7D531DD3CFAAA" box="[753,846,1295,1321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC1110DD531DD0DFAAA" box="[859,895,1295,1321]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
), ventral membranous lamella (arrow
<quantity id="4C9B9B6AE72DFFC11760D531DB11FAAA" box="[1334,1379,1295,1321]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="6" pageNumber="549" unit="in" value="4.0">4 in</quantity>
<figureCitation id="13582A0AE72DFFC1173AD531DEC8FACE" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4C</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11290D50DDE98FACE" box="[198,234,1331,1357]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9C</figureCitation>
), retrolatero-distal membranous lamella (arrow in
<figureCitation id="13582A0AE72DFFC11149D50DDD0EFACE" box="[799,892,1331,1357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4D</figureCitation>
,
<figureCitation id="13582A0AE72DFFC111D1D50DDDDFFACE" box="[903,941,1331,1357]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9D</figureCitation>
), and retrolatero-subdistal membranous flap (f in
<figureCitation id="13582A0AE72DFFC11287D569DF5DFAF2" box="[209,303,1367,1393]" captionStart="FIGURE 4" captionStartId="7.[151,250,1862,1886]" captionTargetBox="[249,1338,237,1839]" captionTargetId="figure-37@7.[249,1338,237,1839]" captionTargetPageId="7" captionText="FIGURE 4. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392244" httpUri="https://zenodo.org/record/8392244/files/figure.png" pageId="6" pageNumber="549">Figs 4D</figureCitation>
,
<figureCitation id="13582A0AE72DFFC1136AD569DF10FAF2" box="[316,354,1367,1393]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="6" pageNumber="549">9D</figureCitation>
); bulb (
<figureCitation id="13582A0AE72DFFC113EDD569DC67FAF2" box="[443,533,1366,1393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-22@8.[203,1384,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Belisana daxiangi sp. nov., holotype male (SYNU-Ar00310).A. Bulb, prolateral view; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.03 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.8392248" httpUri="https://zenodo.org/record/8392248/files/figure.png" pageId="6" pageNumber="549">Fig. 5A</figureCitation>
) with hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi.
</paragraph>
<paragraph id="8BDC368FE72DFFC11291D5A1DFD7FA3A" blockId="6.[151,1437,574,1537]" box="[199,421,1439,1465]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D5A1DF5AFA3A" bold="true" box="[199,296,1439,1465]" pageId="6" pageNumber="549">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="8BDC368FE72DFFC11291D5FDDAE4FA5E" blockId="6.[151,1437,574,1537]" box="[199,1174,1474,1501]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D5FDDFF7FA5F" bold="true" box="[199,389,1474,1501]" pageId="6" pageNumber="549">Natural history.</emphasis>
The species was collected in a seasonal rainforest by canopy fogging.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E72DFFC11291D5D8DD78F982" box="[199,778,1510,1537]" pageId="6" pageNumber="549" type="distribution">
<paragraph id="8BDC368FE72DFFC11291D5D8DD78F982" blockId="6.[151,1437,574,1537]" box="[199,778,1510,1537]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D5D8DF13F983" bold="true" box="[199,353,1510,1536]" pageId="6" pageNumber="549">Distribution.</emphasis>
<collectingCountry id="F374761FE72DFFC11331D5D9DFD8F982" box="[359,426,1511,1537]" name="China" pageId="6" pageNumber="549">China</collectingCountry>
(
<collectingRegion id="49A7F86DE72DFFC113EFD5D9DC61F982" box="[441,531,1511,1537]" country="China" name="Yunnan" pageId="6" pageNumber="549">Yunnan</collectingRegion>
,
<typeStatus id="54D8882DE72DFFC11048D5D9DC3CF982" box="[542,590,1511,1537]" pageId="6" pageNumber="549">type</typeStatus>
locality;
<figureCitation id="13582A0AE72DFFC110EED5D9DC8BF982" box="[696,761,1511,1537]" captionStart="FIGURE 1" captionStartId="3.[151,250,957,981]" captionTargetBox="[151,1436,179,931]" captionTargetId="figure-18@3.[151,1436,179,931]" captionTargetPageId="3" captionText="FIGURE 1. Distribution records of the Belisana species discussed in this paper. 1, B. chuandiani sp. nov.; 2, B. daxiangi sp. nov.; 3, B. fengzheni sp. nov.; 45, B. zhengi." figureDoi="http://doi.org/10.5281/zenodo.8392236" httpUri="https://zenodo.org/record/8392236/files/figure.png" pageId="6" pageNumber="549">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>