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<document id="A4BD8A83EB4D9874FAF0459327D00DAE" ID-CLB-Dataset="113070" ID-DOI="10.1093/zoolinnean/zlab043" ID-GBIF-Dataset="e070d52f-d008-4f08-988a-a8b17c27418f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6959516" ID-ZooBank="1C89556A-12CB-43B7-9B49-E02EFF1543D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1653991014928" checkinUser="plazi" docAuthor="Medrano, Miguel, Kury, Adriano Brilhante &amp; Mendes, Amanda Cruz" docDate="2022" docId="03CA878CFFA5FFCAFEE9FDE2FAA727F6" docLanguage="en" docName="zlab043.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Cosmetinae Koch 1839" docType="treatment" docVersion="8" lastPageNumber="601" masterDocId="FFF3FFF4FFB5FFDAFFFEFFB7FFA72171" masterDocTitle="Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)" masterLastPageNumber="672" masterPageNumber="585" pageNumber="601" updateTime="1699338404186" updateUser="ExternalLinkService">
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<mods:title id="2B65C35507D5E8B354A2D833128694F7">Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)</mods:title>
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<mods:namePart id="0266A2252CA50A49085B6EEE9B348668">Medrano, Miguel</mods:namePart>
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<mods:namePart id="AECAFB0DA65BA593E52E4F6B252FBF68">Kury, Adriano Brilhante</mods:namePart>
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<mods:namePart id="D4E25AFB90755E036D6AD2FFF6772DF6">Mendes, Amanda Cruz</mods:namePart>
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<mods:title id="A82749ADCCB580CCA29F1E7D9C92C564">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="B17F972CBB8A0AB7FD656319DF1811B1">2022</mods:date>
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<mods:number id="B0F9C955B3B65F122582C177D4421123">2021-08-30</mods:number>
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<mods:number id="9271A891FF7F82CCD9DD99865B585282">195</mods:number>
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<treatment id="03CA878CFFA5FFCAFEE9FDE2FAA727F6" ID-DOI="http://doi.org/10.5281/zenodo.6963829" ID-GBIF-Taxon="198579188" ID-Zenodo-Dep="6963829" LSID="urn:lsid:plazi:treatment:03CA878CFFA5FFCAFEE9FDE2FAA727F6" httpUri="http://treatment.plazi.org/id/03CA878CFFA5FFCAFEE9FDE2FAA727F6" lastPageNumber="601" pageId="16" pageNumber="601">
<subSubSection id="C3796511FFA5FFCAFEE9FDE2FDD5231F" box="[279,626,597,622]" pageId="16" pageNumber="601" type="nomenclature">
<paragraph id="8BDC369AFFA5FFCAFEE9FDE2FDD5231F" blockId="16.[279,626,597,622]" box="[279,626,597,622]" pageId="16" pageNumber="601">
<taxonomicName id="4C634D19FFA5FFCAFEE9FDE2FDD5231F" ID-CoL="JDP" authorityName="Koch" authorityYear="1839" box="[279,626,597,622]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="596" phylum="Arthropoda" rank="subFamily" subFamily="Cosmetinae">
COSMETINAE C.L.
<bibRefCitation id="EFF24B6BFFA5FFCAFE14FDE2FDD5231F" author="Koch CL" box="[490,626,597,622]" pageId="16" pageNumber="601" refId="ref52151" refString="Koch CL. 1839. Ubersicht des Arachnidensystems. Zweites Heft. Nurnberg: C. H. Zeh'schen Buchhandlung." type="book" year="1839">KOCH, 1839</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="C3796511FFA5FFCAFF6FFDCAFF78223A" pageId="16" pageNumber="601" type="multiple">
<paragraph id="8BDC369AFFA5FFCAFF6FFDCAFF78223A" blockId="16.[145,762,637,904]" pageId="16" pageNumber="601">
Cosmetides C. L.
<bibRefCitation id="EFF24B6BFFA5FFCAFE89FDCAFDEE23E3" author="Koch CL" box="[375,585,637,659]" pageId="16" pageNumber="601" refId="ref52151" refString="Koch CL. 1839. Ubersicht des Arachnidensystems. Zweites Heft. Nurnberg: C. H. Zeh'schen Buchhandlung." type="book" year="1839">Koch, 1839: 19</bibRefCitation>
(vernacular ‘Breittasterige Kanker’; incl.
<taxonomicName id="4C634D19FFA5FFCAFDF9FD2BFDDB23C0" authorityName="Perty" authorityYear="1833" box="[519,636,668,689]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFDF9FD2BFDDB23C0" box="[519,636,668,689]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFD71FD2BFD5523C0" box="[655,754,668,689]" class="Arachnida" family="Cosmetidae" genus="Cynorta" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFD71FD2BFD5523C0" box="[655,754,668,689]" italics="true" pageId="16" pageNumber="601">Cynorta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFF6FFD0CFEB223A1" authorityName="Perty" authorityYear="1833" box="[145,277,699,720]" class="Anthozoa" family="Discosomidae" genus="Discosoma" kingdom="Animalia" order="Corallimorpharia" pageId="16" pageNumber="601" phylum="Cnidaria" rank="genus">
<emphasis id="B917EA88FFA5FFCAFF6FFD0CFEB223A1" box="[145,277,699,720]" italics="true" pageId="16" pageNumber="601">Discosoma</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFED7FD0DFED823BE" baseAuthorityName="Kury &amp; Garcia" baseAuthorityYear="2016" box="[297,383,698,719]" class="Arachnida" family="Cosmetidae" genus="Flirtea" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFED7FD0DFED823BE" box="[297,383,698,719]" italics="true" pageId="16" pageNumber="601">Flirtea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFE6CFD0DFE4E23BE" authorityName="Koch" authorityYear="1839" box="[402,489,698,719]" class="Magnoliopsida" family="Thymelaeaceae" genus="Gnidia" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="601" phylum="Tracheophyta" rank="genus">
<emphasis id="B917EA88FFA5FFCAFE6CFD0DFE4E23BE" box="[402,489,698,719]" italics="true" pageId="16" pageNumber="601">Gnidia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFE03FD0DFD2C23BE" baseAuthorityName="Kury &amp; Medrano" baseAuthorityYear="2018" box="[509,651,698,719]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFE03FD0DFD2C23BE" box="[509,651,698,719]" italics="true" pageId="16" pageNumber="601">Paecilaema</emphasis>
</taxonomicName>
). Junior subjective synonym of
<taxonomicName id="4C634D19FFA5FFCAFE65FD6EFD5E239F" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[411,761,729,751]" class="Arachnida" family="Gonyleptidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="family">Gonyleptidae Sundevall, 1833</taxonomicName>
by
<bibRefCitation id="EFF24B6BFFA5FFCAFF4BFD4FFED5227F" author="Sorensen WE" box="[181,370,760,782]" pageId="16" pageNumber="601" pagination="489 - 525" refId="ref55435" refString="Sorensen WE. 1873. Bidrag til Phalangidernes Morphologi og Systematik samt Beskrivelse af nogle nye, herhen horende Former. Naturhistorisk Tidsskrift, KjObenhavn, ser. 3 8: 489 - 525, pl. 15." type="journal article" year="1873">Sørensen (1873)</bibRefCitation>
; synonymy disclaimed by almost every subsequent author, including Sørensen himself (1884).
</paragraph>
</subSubSection>
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<paragraph id="8BDC369AFFA5FFCAFF57FCE3FE9A22F6" blockId="16.[145,762,637,904]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFF57FCE3FE9A22F6" italics="true" pageId="16" pageNumber="601">
<taxonomicName id="4C634D19FFA5FFCAFF57FCE3FE822218" ID-CoL="JDP" authorityName="Koch" authorityYear="1839" box="[169,293,852,873]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="596" phylum="Arthropoda" rank="subFamily" subFamily="Cosmetinae">Cosmetinae</taxonomicName>
<treatmentCitation id="0AC2108BFFA5FFCAFED3FCE3FD2C2218" author="Pickard-Cambridge FO" box="[301,651,852,873]" page="549" pageId="16" pageNumber="601" year="1904">
<bibRefCitation id="EFF24B6BFFA5FFCAFED3FCE3FD2C2218" author="Pickard-Cambridge FO" box="[301,651,852,873]" pageId="16" pageNumber="601" refId="ref53892" refString="Pickard-Cambridge FO. 1904. Order Opiliones [1 st part]. In: Godman FD, Salvin O, eds. Biologia Centrali-Americana. Vol. 2. Arachnida. Araneidea and Opiliones. London: Porter / Dulau &amp; Co., 545 - 560, pl. 52." type="journal volume" year="1904">Pickard-Cambridge, 1904: 549</bibRefCitation>
</treatmentCitation>
. First use as a subfamily:
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFF6FFC06FC27218B" pageId="16" pageNumber="601" type="discussion">
<paragraph id="8BDC369AFFA5FFCAFF6FFC06FC27218B" blockId="16.[145,762,945,1856]" lastBlockId="16.[809,1425,197,250]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFF6FFC06FEA322B7" box="[145,260,945,966]" italics="true" pageId="16" pageNumber="601">Comment:</emphasis>
The monophylum labelled ‘clade C’ is herein called
<taxonomicName id="4C634D19FFA5FFCAFF21FC67FECB2297" authorityName="Koch" authorityYear="1839" box="[223,364,976,998]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="596" phylum="Arthropoda" rank="subFamily" subFamily="Cosmetinae">Cosmetinae</taxonomicName>
, with a radically restricted sense as opposed to traditional sense in which it included 95% of the species in the family. In our analysis it comprises 14 terminals, representing three genera and roughly 20 species.
<taxonomicName id="4C634D19FFA5FFCAFE65FBFCFD2C2511" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[411,651,1099,1120]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFE65FBFCFDAF2511" box="[411,520,1099,1120]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFDF0FBFCFD2C2511" author="Perty JAM" box="[526,651,1099,1120]" pageId="16" pageNumber="601" pagination="201 - 209" refId="ref53820" refString="Perty JAM. 1833. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX [1817 - 1820] jussu et auspiciis Maximiliani Josephi I Bavariae Regis augustissimi peracto, collegerunt Dr. In: de Spix JB, de Martius CF-Ph, eds. Vol. 3. [Opiliones, pp 201 - 209]. [' 1830 - 1834 ']. Munich: Fleischer." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
is critical because it is the onomatophore of the family and the nominal subfamily. Medrano et al., (in press) deal with this genus (which has 22 species) and discuss its clear heterogeneity. Herein we included three better-known species from
<collectingCountry id="F374760AFFA5FFCAFEDBFB53FEC9258B" box="[293,366,1252,1274]" name="Brazil" pageId="16" pageNumber="601">Brazil</collectingCountry>
:
<taxonomicName id="4C634D19FFA5FFCAFE87FB53FF752469" authority="(Mello-Leitao, 1940)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1940" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="arietinus">
<emphasis id="B917EA88FFA5FFCAFE87FB53FDF42588" box="[377,595,1252,1273]" italics="true" pageId="16" pageNumber="601">Cosmetus arietinus</emphasis>
(
<bibRefCitation id="EFF24B6BFFA5FFCAFD9DFB53FF6F2469" author="de Mello-Leitao CF" pageId="16" pageNumber="601" pagination="39 - 42" refId="ref53404" refString="de Mello-Leitao CF. 1940. Quatro novos generos de Laniatores do Brasil. Papeis Avulsos do Departamento de Zoologia 1: 39 - 42." type="journal article" year="1940">Mello-Leitão, 1940</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFF22FAB4FD602469" authority="(Soares &amp; Soares, 1946)" baseAuthorityName="Soares &amp; Soares" baseAuthorityYear="1946" box="[220,711,1282,1304]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="B917EA88FFA5FFCAFF22FAB4FE122466" box="[220,437,1282,1304]" italics="true" pageId="16" pageNumber="601">Cosmetus delicatus</emphasis>
(
<bibRefCitation id="EFF24B6BFFA5FFCAFE3AFAB4FD672469" author="Soares, BAM &amp; Soares HEM" box="[452,704,1282,1304]" pageId="16" pageNumber="601" pagination="227 - 232" refId="ref55332" refString="Soares, BAM, Soares HEM. 1946. Um novo genero e duas novas especies de opilioes (Opiliones - Cosmetidae, Gonyleptidae). Papeis Avulsos do Departamento de Zoologia 7: 227 - 232." type="journal article" year="1946">Soares &amp; Soares, 1946</bibRefCitation>
)
</taxonomicName>
and *
<taxonomicName id="4C634D19FFA5FFCAFF60FA96FE562447" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[158,497,1313,1335]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="varius">
<emphasis id="B917EA88FFA5FFCAFF60FA96FEC42446" box="[158,355,1313,1335]" italics="true" pageId="16" pageNumber="601">Cosmetus varius</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFE92FA95FE562447" author="Perty JAM" box="[364,497,1313,1335]" pageId="16" pageNumber="601" pagination="201 - 209" refId="ref53820" refString="Perty JAM. 1833. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX [1817 - 1820] jussu et auspiciis Maximiliani Josephi I Bavariae Regis augustissimi peracto, collegerunt Dr. In: de Spix JB, de Martius CF-Ph, eds. Vol. 3. [Opiliones, pp 201 - 209]. [' 1830 - 1834 ']. Munich: Fleischer." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
, which are deemed to represent a natural nucleus of six or seven Brazilian species of the genus. Species from
<collectingCountry id="F374760AFFA5FFCAFDE4FAE8FDDD2405" box="[538,634,1375,1396]" name="Panama" pageId="16" pageNumber="601">Panama</collectingCountry>
,
<collectingCountry id="F374760AFFA5FFCAFD7AFAE9FD5E2405" box="[644,761,1374,1396]" name="Venezuela" pageId="16" pageNumber="601">Venezuela</collectingCountry>
and the Andes should probably not be included here (Medrano
<emphasis id="B917EA88FFA5FFCAFEFDFA2BFE9624C0" box="[259,305,1436,1457]" italics="true" pageId="16" pageNumber="601">et al</emphasis>
., in press). Following the restricted sense proposed in
<bibRefCitation id="EFF24B6BFFA5FFCAFEE0FA0CFD9424A1" author="Medrano M &amp; Kury AB" box="[286,563,1466,1488]" pageId="16" pageNumber="601" pagination="1206 - 1233" refId="ref53005" refString="Medrano M, Kury AB. 2018. Relationships and cladistic analysis of Roquettea with description of two new species and notes about evolution of stylus in Cosmetidae (Opiliones, Grassatores). Invertebrate Systematics 32: 1206 - 1233." type="journal article" year="2018">Kury &amp; Medrano (2018)</bibRefCitation>
,
<taxonomicName id="4C634D19FFA5FFCAFDBEFA0DFE8C249F" authority="Roewer, 1928" authorityName="Roewer" authorityYear="1928" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFDBEFA0DFD5F24BE" box="[576,760,1466,1487]" italics="true" pageId="16" pageNumber="601">Metavononoides</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFF6FFA6EFE8C249F" author="Roewer CF" box="[145,299,1497,1518]" pageId="16" pageNumber="601" pagination="527 - 632" refId="ref54532" refString="Roewer CF. 1928 [' 1927 ']. Weitere Weberknechte II. (2. Erganzung der Weberknechte der Erde, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen 26: 527 - 632." type="journal article" year="1928">Roewer, 1928</bibRefCitation>
</taxonomicName>
is represented in our analysis by three terminals: *
<taxonomicName id="4C634D19FFA5FFCAFEDAFA4FFD3D277C" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1923" box="[292,666,1528,1549]" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="barbacenensis">
<emphasis id="B917EA88FFA5FFCAFEDAFA4FFD3D277C" box="[292,666,1528,1549]" italics="true" pageId="16" pageNumber="601">Metavononoides barbacenensis</emphasis>
</taxonomicName>
(Mello- Leitão, 1923),
<taxonomicName id="4C634D19FFA5FFCAFED7F9A1FF75273B" authority="(Mello-Leitao, 1935)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1935" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="guttulosus">
<emphasis id="B917EA88FFA5FFCAFED7F9A1FDFF275A" box="[297,600,1558,1579]" italics="true" pageId="16" pageNumber="601">Metavononoides guttulosus</emphasis>
(
<bibRefCitation id="EFF24B6BFFA5FFCAFD9DF9A1FF6D273A" author="de Mello-Leitao CF" pageId="16" pageNumber="601" pagination="369 - 411" refId="ref53287" refString="de Mello-Leitao CF. 1935. A proposito de alguns opilioes novos. Memorias do Instituto Butantan 9: 369 - 411." type="journal article" year="1935">Mello-Leitão, 1935</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C634D19FFA5FFCAFEEFF982FF752718" authority="(Mello-Leitao, 1923)" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1923" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B917EA88FFA5FFCAFEEFF982FDEE273B" box="[273,585,1589,1610]" italics="true" pageId="16" pageNumber="601">Metavononoides orientalis</emphasis>
(
<bibRefCitation id="EFF24B6BFFA5FFCAFDA2F982FF6F2718" author="de Mello-Leitao C" pageId="16" pageNumber="601" pagination="107 - 197" refId="ref53172" refString="de Mello-Leitao C. 1923. Opiliones Laniatores do Brasil. Archivos do Museu Nacional 24: 107 - 197." type="journal article" year="1923">Mello-Leitão, 1923</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C634D19FFA5FFCAFF22F9E3FDA92718" authority="C. L. Koch, 1839" authorityName="C. L. Koch" authorityYear="1839" box="[220,526,1620,1642]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFF22F9E3FEFA2718" box="[220,349,1620,1641]" italics="true" pageId="16" pageNumber="601">Paecilaema</emphasis>
C.L.
<bibRefCitation id="EFF24B6BFFA5FFCAFE6FF9E3FDA92718" author="Koch CL" box="[401,526,1620,1642]" pageId="16" pageNumber="601" refId="ref52151" refString="Koch CL. 1839. Ubersicht des Arachnidensystems. Zweites Heft. Nurnberg: C. H. Zeh'schen Buchhandlung." type="book" year="1839">Koch, 1839</bibRefCitation>
</taxonomicName>
currently includes
<emphasis id="B917EA88FFA5FFCAFD16F9E3FD542718" box="[744,755,1620,1641]" italics="true" pageId="16" pageNumber="601">c</emphasis>
. 100 species (
<bibRefCitation id="EFF24B6BFFA5FFCAFEDFF9C4FE7927F6" author="Kury AB &amp; Mendes AC &amp; Cardoso L &amp; Kury MS &amp; de Granado AA &amp; Giribet G" box="[289,478,1650,1672]" pageId="16" pageNumber="601" refId="ref52813" refString="Kury AB, Mendes AC, Cardoso L, Kury MS, de Granado AA, Giribet G. 2021. World catalogue of Opiliones. WCO-Lite v. 1.2.1. Available at: https: // wcolite. com /. Accessed 3 February 2021." type="url" year="2021">
Kury
<emphasis id="B917EA88FFA5FFCAFE9CF9C4FE3D27F6" box="[354,410,1650,1672]" italics="true" pageId="16" pageNumber="601">et al.</emphasis>
, 2021
</bibRefCitation>
) of which only a fraction may represent a monophylum (as hypothesized by
<bibRefCitation id="EFF24B6BFFA5FFCAFF6FF907FE3927B4" author="Medrano M &amp; Kury AB" box="[145,414,1712,1734]" pageId="16" pageNumber="601" pagination="1206 - 1233" refId="ref53005" refString="Medrano M, Kury AB. 2018. Relationships and cladistic analysis of Roquettea with description of two new species and notes about evolution of stylus in Cosmetidae (Opiliones, Grassatores). Invertebrate Systematics 32: 1206 - 1233." type="journal article" year="2018">Kury &amp; Medrano, 2018</bibRefCitation>
). Eight species of
<taxonomicName id="4C634D19FFA5FFCAFD8BF907FD5E27B4" authorityName="C. L. Koch" authorityYear="1839" box="[629,761,1712,1733]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFD8BF907FD5E27B4" box="[629,761,1712,1733]" italics="true" pageId="16" pageNumber="601">Paecilaema</emphasis>
</taxonomicName>
are included in our analysis. Neither
<taxonomicName id="4C634D19FFA5FFCAFDBEF979FD5F2792" authorityName="Roewer" authorityYear="1928" box="[576,760,1742,1763]" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFDBEF979FD5F2792" box="[576,760,1742,1763]" italics="true" pageId="16" pageNumber="601">Metavononoides</emphasis>
</taxonomicName>
nor
<taxonomicName id="4C634D19FFA5FFCAFF40F95AFE982673" authorityName="C. L. Koch" authorityYear="1839" box="[190,319,1773,1794]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFF40F95AFE982673" box="[190,319,1773,1794]" italics="true" pageId="16" pageNumber="601">Paecilaema</emphasis>
</taxonomicName>
resolve here unambiguously as clades, but this subfamily clearly needs more focused study to determine their composition and which possible extra genera might still be added or created to allocate some species.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FEACFAB02248" pageId="16" pageNumber="601" type="diagnosis">
<paragraph id="8BDC369AFFA5FFCAFCD7FEACFAB02248" blockId="16.[809,1426,282,825]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FEACFC002041" box="[809,935,283,304]" italics="true" pageId="16" pageNumber="601">Diagnosis:</emphasis>
Small and delicate cosmetids with long and unarmed legs (
<figureCitation id="13582A1FFFA5FFCAFBE6FE8EFB35203F" box="[1048,1170,313,335]" captionStart="Figure 4" captionStartId="4.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1490]" captionTargetId="figure-133@4.[145,1425,195,1490]" captionTargetPageId="4" captionText="Figure 4. Habitus in vivo of some non-discosomaticinae. A, Eucynortella cryptograma from Manabí (Ecuador). B, Vononana adrik from the biological research station Panguana (Peru). C, Taito spaceinvaders from Humaitá (Brazil). D, Flirtea valida from Pará (Brazil). E, Eulibitia scalaris from Boyacá (Colombia). F, Metalibitia sp. from Bolivian Amazon. G, Acantholibitia pluriarcuata from Piauí (Brazil). H, Libitioides albolineata from North Carolina (USA). Photographs of Martin Mandák (A), Dieter Schulten (B), Leonardo Carvalho (C, D, G), Miguel Medrano (E), Arthur Anker (F) and Jason Crockwell (H)." figureDoi="http://doi.org/10.5281/zenodo.6959530" httpUri="https://zenodo.org/record/6959530/files/figure.png" pageId="16" pageNumber="601">Fig. 4B, C</figureCitation>
). Area I unarmed or with small, rounded tubercles and area III with two paramedian spines (
<figureCitation id="13582A1FFFA5FFCAFBEFFEC1FB2720FA" box="[1041,1152,374,396]" captionStart="Figure 4" captionStartId="4.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1490]" captionTargetId="figure-133@4.[145,1425,195,1490]" captionTargetPageId="4" captionText="Figure 4. Habitus in vivo of some non-discosomaticinae. A, Eucynortella cryptograma from Manabí (Ecuador). B, Vononana adrik from the biological research station Panguana (Peru). C, Taito spaceinvaders from Humaitá (Brazil). D, Flirtea valida from Pará (Brazil). E, Eulibitia scalaris from Boyacá (Colombia). F, Metalibitia sp. from Bolivian Amazon. G, Acantholibitia pluriarcuata from Piauí (Brazil). H, Libitioides albolineata from North Carolina (USA). Photographs of Martin Mandák (A), Dieter Schulten (B), Leonardo Carvalho (C, D, G), Miguel Medrano (E), Arthur Anker (F) and Jason Crockwell (H)." figureDoi="http://doi.org/10.5281/zenodo.6959530" httpUri="https://zenodo.org/record/6959530/files/figure.png" pageId="16" pageNumber="601">Fig. 4B, C</figureCitation>
) that may be fused into one thick spine (
<taxonomicName id="4C634D19FFA5FFCAFC08FE22FBCF20DB" authorityName="Perty" authorityYear="1833" box="[1014,1128,405,426]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC08FE22FBCF20DB" box="[1014,1128,405,426]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
</taxonomicName>
). Other areas unarmed. Dorsal scutum (DS) beta-type, constriction between carapace and abdomen (scutal groove) marked in lateral view (
<figureCitation id="13582A1FFFA5FFCAFC30FE46FBEF2377" box="[974,1096,497,519]" captionStart="Figure 4" captionStartId="4.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1490]" captionTargetId="figure-133@4.[145,1425,195,1490]" captionTargetPageId="4" captionText="Figure 4. Habitus in vivo of some non-discosomaticinae. A, Eucynortella cryptograma from Manabí (Ecuador). B, Vononana adrik from the biological research station Panguana (Peru). C, Taito spaceinvaders from Humaitá (Brazil). D, Flirtea valida from Pará (Brazil). E, Eulibitia scalaris from Boyacá (Colombia). F, Metalibitia sp. from Bolivian Amazon. G, Acantholibitia pluriarcuata from Piauí (Brazil). H, Libitioides albolineata from North Carolina (USA). Photographs of Martin Mandák (A), Dieter Schulten (B), Leonardo Carvalho (C, D, G), Miguel Medrano (E), Arthur Anker (F) and Jason Crockwell (H)." figureDoi="http://doi.org/10.5281/zenodo.6959530" httpUri="https://zenodo.org/record/6959530/files/figure.png" pageId="16" pageNumber="601">Fig. 4B, C</figureCitation>
). Chelicerae monomorphic without a basal spine in the movable finger. Yellowish spots in the form of a lyre (except in some members of
<taxonomicName id="4C634D19FFA5FFCAFCD7FDFAFC312313" authorityName="Perty" authorityYear="1833" box="[809,918,589,610]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFCD7FDFAFC312313" box="[809,918,589,610]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
</taxonomicName>
) or as large dots, sparse all over the scutum (
<figureCitation id="13582A1FFFA5FFCAFCCFFDDBFC0223F0" box="[817,933,620,642]" captionStart="Figure 4" captionStartId="4.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1490]" captionTargetId="figure-133@4.[145,1425,195,1490]" captionTargetPageId="4" captionText="Figure 4. Habitus in vivo of some non-discosomaticinae. A, Eucynortella cryptograma from Manabí (Ecuador). B, Vononana adrik from the biological research station Panguana (Peru). C, Taito spaceinvaders from Humaitá (Brazil). D, Flirtea valida from Pará (Brazil). E, Eulibitia scalaris from Boyacá (Colombia). F, Metalibitia sp. from Bolivian Amazon. G, Acantholibitia pluriarcuata from Piauí (Brazil). H, Libitioides albolineata from North Carolina (USA). Photographs of Martin Mandák (A), Dieter Schulten (B), Leonardo Carvalho (C, D, G), Miguel Medrano (E), Arthur Anker (F) and Jason Crockwell (H)." figureDoi="http://doi.org/10.5281/zenodo.6959530" httpUri="https://zenodo.org/record/6959530/files/figure.png" pageId="16" pageNumber="601">Fig. 4B, C</figureCitation>
). Smooth claws in legs III and IV (except for some
<taxonomicName id="4C634D19FFA5FFCAFC6FFD3CFC5923D1" authorityName="Perty" authorityYear="1833" box="[913,1022,651,672]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC6FFD3CFC5923D1" box="[913,1022,651,672]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
</taxonomicName>
). Lateral part of coxa IV uniformly brown, but yellow spots may be present in
<taxonomicName id="4C634D19FFA5FFCAFAE2FD1EFA2D23CF" authorityName="Perty" authorityYear="1833" box="[1308,1418,681,702]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFAE2FD1EFA2D23CF" box="[1308,1418,681,702]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
</taxonomicName>
. Basitarsomeres slightly enlarged in males, but not inflated. Ventral plate of the penis not elongated with enormous MS-C contrasting with other MS, flattened and curved. The MS-A and MS-D are short.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FCECFB0B2201" box="[809,1196,858,880]" pageId="16" pageNumber="601" type="type_taxon">
<paragraph id="8BDC369AFFA5FFCAFCD7FCECFB0B2201" blockId="16.[809,1196,858,880]" box="[809,1196,858,880]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FCECFC0B2201" box="[809,940,859,880]" italics="true" pageId="16" pageNumber="601">
<typeStatus id="54D88838FFA5FFCAFCD7FCECFCF92201" box="[809,862,859,880]" pageId="16" pageNumber="601">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="4C634D19FFA5FFCAFC4DFCECFB02221E" ID-CoL="7P4ZF" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[947,1189,858,880]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC4DFCECFB872201" box="[947,1056,859,880]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFBD8FCECFB02221E" author="Perty JAM" box="[1062,1189,858,880]" pageId="16" pageNumber="601" pagination="201 - 209" refId="ref53820" refString="Perty JAM. 1833. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX [1817 - 1820] jussu et auspiciis Maximiliani Josephi I Bavariae Regis augustissimi peracto, collegerunt Dr. In: de Spix JB, de Martius CF-Ph, eds. Vol. 3. [Opiliones, pp 201 - 209]. [' 1830 - 1834 ']. Munich: Fleischer." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FC26FA9422B4" pageId="16" pageNumber="601" type="synonymic_list">
<paragraph id="8BDC369AFFA5FFCAFCD7FC26FA9422B4" blockId="16.[809,1425,913,965]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FC26FC4522D6" box="[809,994,913,935]" italics="true" pageId="16" pageNumber="601">Included genera:</emphasis>
<taxonomicName id="4C634D19FFA5FFCAFC18FC26FB7622D7" authority="Perty, 1833" authorityName="Perty" authorityYear="1833" box="[998,1233,913,934]" class="Arachnida" family="Cosmetidae" genus="Cosmetus" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC18FC26FBF422D7" box="[998,1107,913,934]" italics="true" pageId="16" pageNumber="601">Cosmetus</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFBA8FC26FB7622D7" author="Perty JAM" box="[1110,1233,913,934]" pageId="16" pageNumber="601" pagination="201 - 209" refId="ref53820" refString="Perty JAM. 1833. Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCXVII-MDCCCXX [1817 - 1820] jussu et auspiciis Maximiliani Josephi I Bavariae Regis augustissimi peracto, collegerunt Dr. In: de Spix JB, de Martius CF-Ph, eds. Vol. 3. [Opiliones, pp 201 - 209]. [' 1830 - 1834 ']. Munich: Fleischer." type="book chapter" year="1833">Perty, 1833</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFB25FC26FC1822B4" authority="Roewer, 1928" authorityName="Roewer" authorityYear="1928" class="Arachnida" family="Cosmetidae" genus="Metavononoides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFB25FC26FA3622D7" box="[1243,1425,913,934]" italics="true" pageId="16" pageNumber="601">Metavononoides</emphasis>
<bibRefCitation id="EFF24B6BFFA5FFCAFCD7FC07FC1822B4" author="Roewer CF" box="[809,959,944,965]" pageId="16" pageNumber="601" pagination="527 - 632" refId="ref54532" refString="Roewer CF. 1928 [' 1927 ']. Weitere Weberknechte II. (2. Erganzung der Weberknechte der Erde, 1923). Abhandlungen der Naturwissenschaftlichen Verein zu Bremen 26: 527 - 632." type="journal article" year="1928">Roewer, 1928</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C634D19FFA5FFCAFC06FC18FA8B22B4" authority="C. L. Koch, 1839" authorityName="C. L. Koch" authorityYear="1839" box="[1016,1324,943,965]" class="Arachnida" family="Cosmetidae" genus="Paecilaema" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC06FC18FBDE22B5" box="[1016,1145,943,964]" italics="true" pageId="16" pageNumber="601">Paecilaema</emphasis>
C.L.
<bibRefCitation id="EFF24B6BFFA5FFCAFB50FC18FA8B22B4" author="Koch CL" box="[1198,1324,943,965]" pageId="16" pageNumber="601" refId="ref52151" refString="Koch CL. 1839. Ubersicht des Arachnidensystems. Zweites Heft. Nurnberg: C. H. Zeh'schen Buchhandlung." type="book" year="1839">Koch, 1839</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FC51FB96256B" pageId="16" pageNumber="601" type="distribution">
<paragraph id="8BDC369AFFA5FFCAFCD7FC51FB96256B" blockId="16.[809,1425,998,1051]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FC51FB9B228A" box="[809,1084,998,1019]" italics="true" pageId="16" pageNumber="601">Combined distribution:</emphasis>
Central portion of Brazilian Atlantic Forest (
<figureCitation id="13582A1FFFA5FFCAFC1DFBB2FB85256B" box="[995,1058,1029,1051]" captionStart="Figure 8" captionStartId="13.[163,242,1616,1638]" captionTargetBox="[163,1443,197,1575]" captionTargetId="figure-138@13.[163,1443,195,1576]" captionTargetPageId="13" captionText="Figure 8. Strict consensus of trees obtained with K = 6.2878 and 7.4809, indicating larger groups as defined herein, with asserted distributions juxtaposed (cont.). * indicates type species of the genera. Navajo rugs equal as Figure 7." figureDoi="http://doi.org/10.5281/zenodo.6959538" httpUri="https://zenodo.org/record/6959538/files/figure.png" pageId="16" pageNumber="601">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFBD7FBE2FB36251F" box="[1065,1169,1109,1134]" pageId="16" pageNumber="601" type="multiple">
<paragraph id="8BDC369AFFA5FFCAFBD7FBE2FB36251F" blockId="16.[1065,1169,1109,1134]" box="[1065,1169,1109,1134]" pageId="16" pageNumber="601">CLADE D</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FB27FA812592" pageId="16" pageNumber="601" type="discussion">
<paragraph id="8BDC369AFFA5FFCAFCD7FB27FA812592" blockId="16.[809,1424,1168,1251]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FB27FC1525D7" box="[809,946,1168,1190]" italics="true" pageId="16" pageNumber="601">Comment:</emphasis>
This monophylum includes the
<taxonomicName id="4C634D19FFA5FFCAFCD7FB19FC5E25B5" authorityName="Roewer" authorityYear="1923" box="[809,1017,1198,1220]" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="596" rank="subFamily" subFamily="Discosomaticinae">Discosomaticinae</taxonomicName>
(which is treated later in greater detail than the other clades) and the clade F.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFBD4FAA0FB282441" box="[1066,1167,1303,1328]" pageId="16" pageNumber="601" type="multiple">
<paragraph id="8BDC369AFFA5FFCAFBD4FAA0FB282441" blockId="16.[1066,1167,1303,1328]" box="[1066,1167,1303,1328]" pageId="16" pageNumber="601">CLADE F</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7FAE7FBFA24D2" pageId="16" pageNumber="601" type="discussion">
<paragraph id="8BDC369AFFA5FFCAFCD7FAE7FBFA24D2" blockId="16.[809,1425,1360,1443]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7FAE7FC382414" box="[809,927,1360,1381]" italics="true" pageId="16" pageNumber="601">Comment:</emphasis>
This monophylum includes the clades G and H, each of which includes two subfamilies and sundry unassigned genera.
</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFBD7FA7AFB362497" box="[1065,1169,1485,1510]" pageId="16" pageNumber="601" type="multiple">
<paragraph id="8BDC369AFFA5FFCAFBD7FA7AFB362497" blockId="16.[1065,1169,1485,1510]" box="[1065,1169,1485,1510]" pageId="16" pageNumber="601">CLADE G</paragraph>
</subSubSection>
<subSubSection id="C3796511FFA5FFCAFCD7F9A1FAA727F6" pageId="16" pageNumber="601" type="discussion">
<paragraph id="8BDC369AFFA5FFCAFCD7F9A1FAA727F6" blockId="16.[809,1426,1557,1671]" pageId="16" pageNumber="601">
<emphasis id="B917EA88FFA5FFCAFCD7F9A1FC3B275A" box="[809,924,1558,1579]" italics="true" pageId="16" pageNumber="601">Comment:</emphasis>
This clade includes two well-supported new subfamilies:
<taxonomicName id="4C634D19FFA5FFCAFC3EF983FBED273B" authority="Medrano &amp; Kury &amp; Mendes, 2022" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" box="[960,1098,1588,1610]" pageId="16" pageNumber="601" rank="subFamily" status="subfam. nov." subFamily="Ferkeriinae">Ferkeriinae</taxonomicName>
<taxonomicNameLabel id="A22457F3FFA5FFCAFBACF983FB412738" box="[1106,1254,1588,1610]" pageId="16" pageNumber="601" rank="subFamily">subfam. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C634D19FFA5FFCAFADFF983FA37273B" authority="Medrano &amp; Kury &amp; Mendes, 2022" authorityName="Medrano &amp; Kury &amp; Mendes" authorityYear="2022" box="[1313,1424,1588,1610]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Taitoinae">Taitoinae</taxonomicName>
<taxonomicNameLabel id="A22457F3FFA5FFCAFCD7F9E5FC1F2719" box="[809,952,1618,1640]" pageId="16" pageNumber="601" rank="subFamily">subfam. nov.</taxonomicNameLabel>
, plus four isolated genera:
<taxonomicName id="4C634D19FFA5FFCAFB19F9E5FA2C2716" authorityName="Mello-Leitao" authorityYear="1928" box="[1255,1419,1618,1639]" class="Arachnida" family="Cosmetidae" genus="Acantholibitia" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFB19F9E5FA2C2716" box="[1255,1419,1618,1639]" italics="true" pageId="16" pageNumber="601">Acantholibitia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFCD7F9C6FC3727F7" authorityName="Roewer" authorityYear="1912" box="[809,912,1649,1670]" class="Arachnida" family="Cosmetidae" genus="Eulibitia" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFCD7F9C6FC3727F7" box="[809,912,1649,1670]" italics="true" pageId="16" pageNumber="601">Eulibitia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C634D19FFA5FFCAFC63F9C6FBB027F7" authorityName="Roewer" authorityYear="1912" box="[925,1047,1649,1670]" class="Arachnida" family="Cosmetidae" genus="Libitioides" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFC63F9C6FBB027F7" box="[925,1047,1649,1670]" italics="true" pageId="16" pageNumber="601">Libitioides</emphasis>
</taxonomicName>
and
<taxonomicName id="4C634D19FFA5FFCAFBAEF9C6FB5D27F7" authorityName="Medrano, Kury &amp; Mendes" authorityYear="2022" box="[1104,1274,1649,1670]" class="Arachnida" family="Cosmetidae" genus="Qarikichkauru" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
<emphasis id="B917EA88FFA5FFCAFBAEF9C6FB5D27F7" box="[1104,1274,1649,1670]" italics="true" pageId="16" pageNumber="601">Qarikichkauru</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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