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<mods:title id="A280D3B395FEED67258354BEEF97A65A">Phloeocharis subtilissima Mannerheim (Staphylinidae: Phloeo­ charinae) and Cephennium gallicum Ganglbauer (Scydmaenidae) new to North America: a case study in the introduction of exotic Coleoptera to the port of Halifax, with new records of other species</mods:title>
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<treatment id="03C187EFFFD4D64FFE8CF945FD20FC49" ID-DOI="http://doi.org/10.5281/zenodo.6273424" ID-GBIF-Taxon="124556370" ID-Zenodo-Dep="6273424" LSID="urn:lsid:plazi:treatment:03C187EFFFD4D64FFE8CF945FD20FC49" httpUri="http://treatment.plazi.org/id/03C187EFFFD4D64FFE8CF945FD20FC49" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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<paragraph id="8BD736F9FFD4D64CFE8CF945FD4AF947" blockId="1.[264,690,1718,1744]" box="[264,690,1718,1744]" pageId="1" pageNumber="2">
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<taxonomicName id="4C684D7AFFD4D64CFE8CF945FD4AF947" ID-CoL="SF5Y" authority="Ganglbauer" authorityName="Ganglbauer" box="[264,690,1718,1744]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gallicum">
<emphasis id="B91CEAEBFFD4D64CFE8CF945FD4AF947" bold="true" box="[264,690,1718,1744]" pageId="1" pageNumber="2">
<emphasis id="B91CEAEBFFD4D64CFE8CF945FDEEF947" bold="true" box="[264,534,1718,1744]" italics="true" pageId="1" pageNumber="2">Cephennium gallicum</emphasis>
Ganglbauer
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8BD736F9FFD4D64FFE8CF8F2FC38FE39" blockId="1.[264,1323,1793,1899]" lastBlockId="2.[264,1324,284,990]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<taxonomicName id="4C684D7AFFD4D64CFE8CF8F2FD08F88C" ID-CoL="SF5Y" authority="Ganglbauer, 1899" authorityName="Ganglbauer" authorityYear="1899" box="[264,752,1793,1819]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="gallicum">
<emphasis id="B91CEAEBFFD4D64CFE8CF8F2FDF6F88D" box="[264,526,1793,1818]" italics="true" pageId="1" pageNumber="2">Cephennium gallicum</emphasis>
Ganglbauer, 1899
</taxonomicName>
(
<taxonomicName id="4C684D7AFFD4D64CFC80F8F2FC73F88C" ID-CoL="C2L" box="[772,907,1793,1819]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C684D7AFFD4D64CFC1EF8F2FBB1F88C" box="[922,1097,1793,1819]" class="Insecta" family="Scydmaenidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Scydmaenidae</taxonomicName>
:
<taxonomicName id="4C684D7AFFD4D64CFBDDF8F2FB12F88C" ID-CoL="8VR7K" box="[1113,1258,1793,1819]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Cephenniini">Cephenniini</taxonomicName>
) is a member of a diverse and widely distributed genus that is primarily Palearctic.
<bibRefCitation id="EFF94B08FFD4D64CFB13F8DAFEAEF8FC" pageId="1" pageNumber="13" refString="Freude, H., Harde, K. W. &amp; Lohse. G. A. (1971) Die Kafer Mitteleuropas: Band 3. Goecke &amp; Evers, Krefeld, Germany, 365 pp." type="book">Freude et al. (1971)</bibRefCitation>
discussed 14 species of
<taxonomicName id="4C684D7AFFD4D64CFDF3F8A2FCF3F8FD" ID-CoL="8VRKF" box="[631,779,1873,1898]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAEBFFD4D64CFDF3F8A2FCF3F8FD" box="[631,779,1873,1898]" italics="true" pageId="1" pageNumber="2">Cephennium</emphasis>
</taxonomicName>
from Central Europe, however, P. Jaloszynski (pers. comm.) indicates that there are many additional undescribed species and the genus is in need of extensive revision.
<bibRefCitation id="EFF94B08FFD7D64FFD12FEB7FCB7FEC9" box="[662,847,324,350]" pageId="2" pageNumber="14" refString="O'Keefe, S. T. (2001) Scydmaenidae Leach, 1815. In: Arnett, R. H. Jr. &amp; Thomas, M. C. [Eds]: American Beetles. Volume 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, USA. pp. 259 - 267." type="book chapter">O'Keefe (2001)</bibRefCitation>
reports that there are over 100 species described worldwide. Only one of these, the blind
<taxonomicName id="4C684D7AFFD7D64FFCD1FE9FFB6DFE11" authority="Brendel" authorityName="Brendel" box="[853,1173,364,390]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anopthalmicum">
<emphasis id="B91CEAEBFFD7D64FFCD1FE9FFBD6FE12" box="[853,1070,364,389]" italics="true" pageId="2" pageNumber="3">C. anopthalmicum</emphasis>
Brendel
</taxonomicName>
, known only from central coastal California, is Nearctic (
<bibRefCitation id="EFF94B08FFD7D64FFC8FFE67FC4BFE39" author="O'Keefe" box="[779,947,404,430]" pageId="2" pageNumber="14" refString="O'Keefe, S. T. (2001) Scydmaenidae Leach, 1815. In: Arnett, R. H. Jr. &amp; Thomas, M. C. [Eds]: American Beetles. Volume 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, USA. pp. 259 - 267." type="book chapter" year="2001">O'Keefe 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8BD736F9FFD7D64FFEBCFE4FFAD3FDE1" blockId="2.[264,1324,284,990]" pageId="2" pageNumber="3">
<taxonomicName id="4C684D7AFFD7D64FFEBCFE4FFDC6FE42" ID-CoL="SF5Y" box="[312,574,444,469]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gallicum">
<emphasis id="B91CEAEBFFD7D64FFEBCFE4FFDC6FE42" box="[312,574,444,469]" italics="true" pageId="2" pageNumber="3">Cephennium gallicum</emphasis>
</taxonomicName>
is a widely distributed species found in central and northern Europe.
<bibRefCitation id="EFF94B08FFD7D64FFEEAFE17FDA4FE69" box="[366,604,484,510]" pageId="2" pageNumber="13" refString="Freude, H., Harde, K. W. &amp; Lohse. G. A. (1971) Die Kafer Mitteleuropas: Band 3. Goecke &amp; Evers, Krefeld, Germany, 365 pp." type="book">Freude et al. (1971)</bibRefCitation>
report it from northern
<collectingCountry id="F37F7669FFD7D64FFCFAFE17FC3CFE69" box="[894,964,484,510]" name="Spain" pageId="2" pageNumber="3">Spain</collectingCountry>
,
<collectingCountry id="F37F7669FFD7D64FFC56FE17FBDDFE69" box="[978,1061,484,510]" name="France" pageId="2" pageNumber="3">France</collectingCountry>
, western
<collectingCountry id="F37F7669FFD7D64FFB1CFE17FADFFE69" box="[1176,1319,484,510]" name="Switzerland" pageId="2" pageNumber="3">Switzerland</collectingCountry>
, Rhineland,
<collectingCountry id="F37F7669FFD7D64FFE15FDFFFE01FDB1" box="[401,505,524,550]" name="Belgium" pageId="2" pageNumber="3">Belgium</collectingCountry>
,
<collectingCountry id="F37F7669FFD7D64FFD82FDFFFD90FDB1" box="[518,616,524,550]" name="Netherlands" pageId="2" pageNumber="3">Holland</collectingCountry>
,
<collectingCountry id="F37F7669FFD7D64FFDF0FDFFFD1DFDB1" box="[628,741,524,550]" name="Denmark" pageId="2" pageNumber="3">Denmark</collectingCountry>
, and southern
<collectingCountry id="F37F7669FFD7D64FFC17FDFFFC01FDB1" box="[915,1017,524,550]" name="United Kingdom" pageId="2" pageNumber="3">England</collectingCountry>
. It has also been found in Skane &amp; Uppland in
<collectingCountry id="F37F7669FFD7D64FFD9EFDC7FD8FFDD9" box="[538,631,564,590]" name="Sweden" pageId="2" pageNumber="3">Sweden</collectingCountry>
(MLU), Thüringen in
<collectingCountry id="F37F7669FFD7D64FFC18FDC7FBF1FDD9" box="[924,1033,564,590]" name="Germany" pageId="2" pageNumber="3">Germany</collectingCountry>
(
<bibRefCitation id="EFF94B08FFD7D64FFB9BFDC7FEBEFDE1" author="Kohler" pageId="2" pageNumber="13" refString="Kohler, F. &amp; Klausnitzer, B. (1998) Verzeichnis der Kafer Deutschlands. Entomlogische Nachrichten und Berichte - Beiheft 4. Dresden, Germany, 185 pp." type="book" year="1998">Köhler &amp; Klausnitzer 1998</bibRefCitation>
),
<collectingCountry id="F37F7669FFD7D64FFEDFFDAFFE56FDE1" box="[347,430,604,630]" name="Ireland" pageId="2" pageNumber="3">Ireland</collectingCountry>
(
<bibRefCitation id="EFF94B08FFD7D64FFE3AFDAFFDD6FDE1" author="Joy" box="[446,558,604,630]" pageId="2" pageNumber="13" refString="Joy, N. H. (1932) A Practical Handbook of British Beetles: Volume 1. Witherby, London, 622 pp." type="book" year="1932">Joy 1932</bibRefCitation>
),
<collectingCountry id="F37F7669FFD7D64FFDC7FDAFFD52FDE1" box="[579,682,604,630]" name="United Kingdom" pageId="2" pageNumber="3">Scotland</collectingCountry>
(
<bibRefCitation id="EFF94B08FFD7D64FFD3DFDAFFCCAFDE1" author="Ball" box="[697,818,604,630]" pageId="2" pageNumber="13" refString="Ball, S. G. (2002) Recorder 2002 database. Joint Nature Conservation Committee. Peterborough, United Kingdom." type="book chapter" year="2002">Ball 2002</bibRefCitation>
), and
<collectingCountry id="F37F7669FFD7D64FFCFCFDAFFC1AFDE1" box="[888,994,604,630]" name="Romania" pageId="2" pageNumber="3">Romania</collectingCountry>
(Jaloszynski, pers. comm.).
</paragraph>
<paragraph id="8BD736F9FFD7D64FFEBCFD77FD20FC49" blockId="2.[264,1324,284,990]" pageId="2" pageNumber="3">
Little information has been published on the bionomics of this species.
<bibRefCitation id="EFF94B08FFD7D64FFB2DFD77FAD3FD09" author="Joy" box="[1193,1323,644,670]" pageId="2" pageNumber="13" refString="Joy, N. H. (1932) A Practical Handbook of British Beetles: Volume 1. Witherby, London, 622 pp." type="book" year="1932">Joy (1932)</bibRefCitation>
reports it from &quot;old tree trunks,&quot; while Fowler (1890) says it is found &quot;in moss, etc.&quot; Jaloszynski (pers. comm.) noted that all species of
<taxonomicName id="4C684D7AFFD7D64FFCDAFD27FC0AFD7A" ID-CoL="8VRKF" box="[862,1010,724,749]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAEBFFD7D64FFCDAFD27FC0AFD7A" box="[862,1010,724,749]" italics="true" pageId="2" pageNumber="3">Cephennium</emphasis>
</taxonomicName>
are usually collected from leaf litter and rotten wood, most commonly in mountainous areas and in deciduous forests.
<bibRefCitation id="EFF94B08FFD7D64FFE8CFCD7FE09FCA9" box="[264,497,804,830]" pageId="2" pageNumber="13" refString="Freude, H., Harde, K. W. &amp; Lohse. G. A. (1971) Die Kafer Mitteleuropas: Band 3. Goecke &amp; Evers, Krefeld, Germany, 365 pp." type="book">Freude et al. (1971)</bibRefCitation>
state that species of
<taxonomicName id="4C684D7AFFD7D64FFD6CFCD7FC84FCAA" ID-CoL="8VRKF" box="[744,892,804,829]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAEBFFD7D64FFD6CFCD7FC84FCAA" box="[744,892,804,829]" italics="true" pageId="2" pageNumber="3">Cephennium</emphasis>
</taxonomicName>
are found particularly in decomposing plant materials and in humus at the base of the trees; sometimes also in decomposing wood. All species of
<taxonomicName id="4C684D7AFFD7D64FFD85FC87FD6DFC1A" ID-CoL="8VRKF" box="[513,661,884,909]" class="Insecta" family="Staphylinidae" genus="Cephennium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAEBFFD7D64FFD85FC87FD6DFC1A" box="[513,661,884,909]" italics="true" pageId="2" pageNumber="3">Cephennium</emphasis>
</taxonomicName>
are specialized predators of oribatid mites (Acari: Oribatei).
<bibRefCitation id="EFF94B08FFD7D64FFECFFC6FFE03FC21" author="Schmid" box="[331,507,924,950]" pageId="2" pageNumber="15" refString="Schmid, R. (1988) Morphologische Anpassungen in einem Rauber-Beute-System: Ameisenkafer (Scydmaenidae: Staphylinoidea) und gepanzerte Milben (Acari). Zoologische Jarbucher, Abteilung fur Systematik, Okologie und Geographie der Teire, 115, 207 - 228." type="journal article" year="1988">Schmid (1988)</bibRefCitation>
discusses the specialized morphology and behaviour of these beetles for feeding on these hard­shelled mites.
</paragraph>
</subSubSection>
</treatment>
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