131 lines
19 KiB
XML
131 lines
19 KiB
XML
<document id="BECA1DF696D41A1D1420A0D61E0DFE07" ID-CLB-Dataset="27341" ID-DOI="10.11646/zootaxa.4585.1.2" ID-GBIF-Dataset="8547d24d-5ff0-4946-9593-946f24086548" ID-ISSN="1175-5326" ID-Zenodo-Dep="2636187" ID-ZooBank="1D7B13B8-C6D0-44F4-AFD2-47733E01E08E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554964596229" checkinUser="plazi" docAuthor="Ulitzka, Manfred R." docDate="2019" docId="03C187ED4047FFA707A45927FD965D2A" docLanguage="en" docName="zootaxa.4585.1.2.pdf" docOrigin="Zootaxa 4585 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rohrthrips patrickmuelleri Ulitzka 2019, sp. n." docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="FFF8FF954042FFA107335F58FF98587F" masterDocTitle="Five new species of Rohrthrips (Thysanoptera: Rohrthripidae) from Burmese amber, and the evolution of Tubulifera wings" masterLastPageNumber="40" masterPageNumber="27" pageNumber="32" updateTime="1698719504840" updateUser="ExternalLinkService">
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<mods:title id="ECEB11A859897F638AEA65D048EA449B">Five new species of Rohrthrips (Thysanoptera: Rohrthripidae) from Burmese amber, and the evolution of Tubulifera wings</mods:title>
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<mods:namePart id="CACB5B93207AA226878FA8AB60050B3C">Ulitzka, Manfred R.</mods:namePart>
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<mods:title id="4A3D0604DDD4F752ABA85B672B13BF5C">Zootaxa</mods:title>
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<treatment id="03C187ED4047FFA707A45927FD965D2A" ID-DOI="http://doi.org/10.5281/zenodo.5933854" ID-GBIF-Taxon="156147058" ID-Zenodo-Dep="5933854" LSID="urn:lsid:plazi:treatment:03C187ED4047FFA707A45927FD965D2A" httpUri="http://treatment.plazi.org/id/03C187ED4047FFA707A45927FD965D2A" lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32">
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<subSubSection id="C37265704047FFA407A45927FDBD5EE5" box="[151,549,1663,1690]" pageId="5" pageNumber="32" type="nomenclature">
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<paragraph id="8BD736FB4047FFA407A45927FDBD5EE5" blockId="5.[151,549,1663,1723]" box="[151,549,1663,1690]" pageId="5" pageNumber="32">
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<heading id="D09F81974047FFA407A45927FDBD5EE5" bold="true" box="[151,549,1663,1690]" fontSize="11" level="1" pageId="5" pageNumber="32" reason="1">
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<emphasis id="B91CEAE94047FFA407A45927FDBD5EE5" bold="true" box="[151,549,1663,1690]" pageId="5" pageNumber="32">
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<taxonomicName id="4C684D784047FFA407A45927FE445EE6" authority="Ulitzka, 2019" authorityName="Ulitzka" authorityYear="2019" box="[151,476,1663,1689]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="patrickmuelleri" status="sp. nov.">
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<emphasis id="B91CEAE94047FFA407A45927FE445EE6" bold="true" box="[151,476,1663,1689]" italics="true" pageId="5" pageNumber="32">Rohrthrips patrickmuelleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F57924047FFA406D059D8FDBD5EE5" box="[483,549,1664,1690]" pageId="5" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265704047FFA707A459FAFC2D5D72" lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32" type="description">
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<paragraph id="8BD736FB4047FFA407A459FAFE925EC4" blockId="5.[151,549,1663,1723]" box="[151,266,1698,1723]" pageId="5" pageNumber="32">
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(
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<figureCitation id="13532A7E4047FFA407AC59FAFE9B5EC4" box="[159,259,1698,1723]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="5" pageNumber="32">Figs 6–8</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD736FB4047FFA407A459B3FB445F58" blockId="5.[151,1436,1770,2011]" pageId="5" pageNumber="32">
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<emphasis id="B91CEAE94047FFA407A459B3FF735F7C" box="[151,235,1771,1795]" italics="true" pageId="5" pageNumber="32">Female</emphasis>
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. Body fully extended; pronotum basally slightly dented; basal abdominal segments difficult to assess due to shiny refraction; legs folded under body except right mid leg; fore wings overlapping hind wings.
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</paragraph>
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<paragraph id="8BD736FB4047FFA407F4586AFC7E5F10" blockId="5.[151,1436,1770,2011]" pageId="5" pageNumber="32">
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Colour (
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<figureCitation id="13532A7E4047FFA40617586AFEFE5F34" box="[292,358,1842,1867]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="5" pageNumber="32">Fig. 6</figureCitation>
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) uniformly light brown including antennae and legs; all major setae and wing fringes dark; wing veins coloured pale greyish-brown, only a shade darker as wing membrane.
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</paragraph>
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<paragraph id="8BD736FB4047FFA707F45822FCD45BB8" blockId="5.[151,1436,1770,2011]" lastBlockId="6.[151,1437,151,1365]" lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32">
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Head (
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<figureCitation id="13532A7E4047FFA406215822FECE5FEC" box="[274,342,1914,1939]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="5" pageNumber="32">Fig. 6</figureCitation>
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) with reticulate sculpture particularly in the middle; about as long as wide and produced in front of anterior margin of eyes, with cheeks slightly rounded behind eyes; cheeks lacking setae; one pair of long but very fine ocellar setae close to base of antennae. Eyes not prolonged ventrally. Hind ocelli close to compound eyes; far apart from each other. Vertex with postocular setae far behind eyes (left seta missing). Antennae 9-segmented; segment I barrel-shaped; segment II basally asymmetric without elongated pedicle; III–VIII symmetrical, inverse conically shaped and tapering distally from level of sense cones; sensorium on segment II not visible; number of sense cones of the following segments difficult to assess but at least one inner and one outer on III–VI and one outer only on VII; segment IX conically shaped, at base narrower than segment VIII distally. Mouth cone short and rounded. Maxillary palps 2-segmented; labial palps minute, not assessable. Pronotum wider than long, trapezoidal; posteroangular setae long and pointed, other pronotal setae not developed or missing; epimeral setae long and pointed. Metanotum with a pair of fine setae far apart from the front margin of the plate. Prosternum with fernal plates wider than long; basantra not visible (very likely not developed); mesopresternum complete, very flat boatshaped; mesosternellum and metasternum fused. Fore wings (
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<figureCitation id="13532A7E4044FFA704615E84FC0F598A" box="[850,919,476,501]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="6" pageNumber="33">Fig. 6</figureCitation>
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) becoming progressively larger in distal half; anterior vein complete, reaching costa near apex; bearing two distal setae and one basal seta located anteriorly to the vein; straight except for the short bending close to its tip; second vein not developed; membrane without microtrichia. Fringe cilia straight; arising from sockets; duplicated cilia present around apical margin of wing; clavus with paired setiform processes at tip. Hind wings at base clearly with microtrichia (
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<figureCitation id="13532A7E4044FFA703875D34FB625AFB" box="[1204,1274,620,645]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="6" pageNumber="33">Fig. 8</figureCitation>
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); without any veins; base with recurved setae; sockets of wing fringes smaller as on fore wing (
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<figureCitation id="13532A7E4044FFA703725DD7FB105AD7" box="[1089,1160,655,680]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="6" pageNumber="33">Fig. 7</figureCitation>
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). Fore legs with femora stout; fore tarsi with two teeth. Mid and hind legs slender. All femora with one long ventral and all tibiae with one long dorsal seta; mid and hind tibiae terminally each with three spines, much smaller on mid tibiae; mid and hind tarsi two-segmented, with hamus. Abdominal tergites faintly sculptured reticulate (partly visible only), without wing-retaining setae; lateral setae long, pointed; setae s3 on IV–VII longer than other lateral setae and conspicuously bent inwards (
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<figureCitation id="13532A7E4044FFA706D75C1CFDB25B22" box="[484,554,836,861]" captionStart="FIGURES 5–8" captionStartId="8.[151,264,1494,1517]" captionTargetBox="[151,1436,195,1473]" captionTargetId="figure@8.[151,1436,195,1473]" captionTargetPageId="8" captionText="FIGURES 5–8. Rohrthrips spp. (5) R. maryae sp. n. holotype male, dorsal view. R. patrickmuelleri sp. n. holotype female 6– 8: (6) dorsal view; (7) surface pattern of wings (phase contrast), indicated red: longitudinally orientated surface pattern, indicated black: submarginal ridge; (8) basal part of wings (phase contrast), indicated white: wing vein, indicated red: longitudinally orientated surface pattern, indicated black: microtrichia." httpUri="https://zenodo.org/record/2636191/files/figure.png" pageId="6" pageNumber="33">Fig. 6</figureCitation>
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). Abdominal segment I (pelta) not assessable; IX conical, with setae s3 long and somewhat protruding tube; sternite IX with structures interpretable as gynosternal plates and fustis (cf.
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<bibRefCitation id="EFF94B0A4044FFA702645C3FFF795BDA" author="Bhatti, J. S." pageId="6" pageNumber="33" pagination="285 - 352" refId="ref10160" refString="Bhatti, J. S. (1998 b) New structural features in the Order Tubulifera (Insecta). 4. The ovispan and other abdominal structures. Zoology (Journal of Pure and Applied Zoology), 5, 285 - 352." type="journal article" year="1998">Bhatti 1998b</bibRefCitation>
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). Abdominal segment X (tube) conically extended at base; its sculpture not assessable; apparently without campaniform sensilla; with terminal crown of long anal setae.
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</paragraph>
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<paragraph id="8BD736FB4044FFA707F45C8CFE6C5C97" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="33">
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<emphasis id="B91CEAE94044FFA707F45C8CFEEE5B92" bold="true" box="[199,374,980,1005]" pageId="6" pageNumber="33">Measurements</emphasis>
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. Female (in microns): Body, length 1600. Head, length 202; width 200. Eyes, length 85; width 38. Hind ocelli, diameter 19; distance between the hind ocelli 50. Ocellar setae 32. Postocular setae 38. Prothorax, length 233; width 378; posteroangular setae 100; epimeral setae 107. Pterothorax, largest width 365. Metanotal median setae about 57. Abdomen, length 1000; largest width 461 (segments III–IV); segment X (tube), length 151; basal width 63. Setae s3 on tergite IX 220. Antennae, length 399; length (largest width) of segment
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<date id="FFD6103B4044FFA703C55B3DFABA5C03" box="[1270,1314,1124,1148]" pageId="6" pageNumber="33" value="1940-01">I 40</date>
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(25),
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<date id="FFD6103B4044FFA702555B3DFA045C03" box="[1382,1436,1124,1148]" pageId="6" pageNumber="33" value="1947-02">II 47</date>
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(34),
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<date id="FFD6103B4044FFA707E65BD1FE8E5CDF" box="[213,278,1159,1184]" pageId="6" pageNumber="33" value="1962-03">III 62</date>
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(34),
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<date id="FFD6103B4044FFA7066E5BD1FE385CDF" box="[349,416,1160,1184]" pageId="6" pageNumber="33" value="1947-04">IV 47</date>
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(31),
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<date id="FFD6103B4044FFA706D45BD1FDB85CDF" box="[487,544,1159,1184]" pageId="6" pageNumber="33" value="1953-05">V 53</date>
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(31),
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<date id="FFD6103B4044FFA705545BD1FD325CDF" box="[615,682,1159,1184]" pageId="6" pageNumber="33" value="1950-06">VI 50</date>
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(28),
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<date id="FFD6103B4044FFA705C25BD1FCA55CDF" box="[753,829,1160,1184]" pageId="6" pageNumber="33" value="1947-07">VII 47</date>
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(22),
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<date id="FFD6103B4044FFA704B75BD1FC415CDF" box="[900,985,1160,1184]" pageId="6" pageNumber="33" value="1934-08">VIII 34</date>
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(16),
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<date id="FFD6103B4044FFA703135BD1FBFB5CDF" box="[1056,1123,1160,1184]" pageId="6" pageNumber="33" value="1919-09">IX 19</date>
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(8). Sense cones, length of inner (outer) on segment
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<date id="FFD6103B4044FFA706845BF5FE605CBB" box="[439,504,1196,1220]" pageId="6" pageNumber="33" value="1920-03">III 20</date>
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(18),
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<date id="FFD6103B4044FFA7050D5BF5FD185CBB" box="[574,640,1196,1220]" pageId="6" pageNumber="33" value="1922-04">IV 22</date>
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(19),
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<date id="FFD6103B4044FFA705F45BF5FD675CBB" box="[711,767,1196,1220]" pageId="6" pageNumber="33" value="1922-05">V 22</date>
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(25),
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<date id="FFD6103B4044FFA704755BF5FC105CBA" box="[838,904,1196,1221]" pageId="6" pageNumber="33" value="1925-06">VI 25</date>
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(19), VII – (16); basal width 4–5. Fore wings, length 1030; largest width 206.
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</paragraph>
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<paragraph id="8BD736FB4044FFA707F45BACFC2D5D72" blockId="6.[151,1437,151,1365]" box="[199,949,1268,1293]" pageId="6" pageNumber="33">
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<emphasis id="B91CEAE94044FFA707F45BACFEF35D72" bold="true" box="[199,363,1268,1293]" pageId="6" pageNumber="33">Syninclusions</emphasis>
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. Remnants of another insect and a stellate trichome.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37265704044FFA707F45A4FFD965D2A" pageId="6" pageNumber="33" type="etymology">
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<paragraph id="8BD736FB4044FFA707F45A4FFD965D2A" blockId="6.[151,1437,151,1365]" pageId="6" pageNumber="33">
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<emphasis id="B91CEAE94044FFA707F45A4FFEDE5D4F" bold="true" box="[199,326,1303,1328]" pageId="6" pageNumber="33">Etymology</emphasis>
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. The species
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<taxonomicName id="4C684D784044FFA706C35A41FD5A5D4F" authorityName="Ulitzka" authorityYear="2019" box="[496,706,1304,1328]" class="Insecta" family="Phlaeothripidae" genus="Rohrthrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="patrickmuelleri">
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<emphasis id="B91CEAE94044FFA706C35A41FD5A5D4F" box="[496,706,1304,1328]" italics="true" pageId="6" pageNumber="33">R. patrickmuelleri</emphasis>
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</taxonomicName>
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is dedicated to Patrick Müller for his generous donations of Burmese amber thrips inclusions.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |