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<mods:title id="3A08411EC9AB1558309437F29E0A672A">Additions to the taxonomy of the armadillo ants (Hymenoptera, Formicidae, Tatu i dri s)</mods:title>
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<paragraph id="8BD736D9AD30FF9FFF01FF40FD2AFF3B" blockId="8.[151,653,151,177]" box="[151,653,151,177]" pageId="8" pageNumber="69">
<heading id="D09F81B5AD30FF9FFF01FF40FD2AFF3B" bold="true" box="[151,653,151,177]" fontSize="11" level="1" pageId="8" pageNumber="69" reason="1">
<taxonomicName id="4C684D5AAD30FF9FFF01FF40FD2AFF3B" ID-CoL="54Z4K" authority="Brown and Kempf 1968" authorityName="Brown and Kempf" authorityYear="1968" box="[151,653,151,177]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD30FF9FFF01FF40FD2AFF3B" bold="true" box="[151,653,151,177]" pageId="8" pageNumber="69">
<emphasis id="B91CEACBAD30FF9FFF01FF40FEFCFF3B" bold="true" box="[151,347,151,177]" italics="true" pageId="8" pageNumber="69">Tatuidris tatusia</emphasis>
<bibRefCitation id="EFF94B28AD30FF9FFEF5FF40FD2AFF3B" author="Brown" box="[355,653,151,177]" pageId="8" pageNumber="69" refString="Brown, W. L. &amp; Kempf, W. W. (1968) Tatuidris, a remarkable new genus of Formicidae (Hymenoptera). Psyche, 74, 183 - 190." type="journal article" year="1968">Brown and Kempf 1968</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8BD736D9AD30FF9FFF01FF08FE25FEE5" blockId="8.[151,1436,222,367]" pageId="8" pageNumber="69">
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<taxonomicName id="4C684D5AAD30FF9FFF01FF08FDC2FF7E" ID-CoL="54Z4K" authority="Brown and Kempf 1968:187" authorityName="Brown and Kempf" authorityPageNumber="187" authorityYear="1968" box="[151,613,222,245]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD30FF9FFF01FF08FE9BFF7F" box="[151,316,223,245]" italics="true" pageId="8" pageNumber="69">Tatuidris tatusia</emphasis>
<treatmentCitation id="0AC910C8AD30FF9FFED4FF08FDC2FF7E" author="Brown" box="[322,613,222,244]" page="187" pageId="8" pageNumber="69" year="1968">
<bibRefCitation id="EFF94B28AD30FF9FFED4FF08FD9DFF7E" author="Brown" box="[322,570,222,244]" pageId="8" pageNumber="69" refString="Brown, W. L. &amp; Kempf, W. W. (1968) Tatuidris, a remarkable new genus of Formicidae (Hymenoptera). Psyche, 74, 183 - 190." type="journal article" year="1968">Brown and Kempf 1968</bibRefCitation>
:187
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, their figures 1–4. 
<typeStatus id="54D3887BAD30FF9FFCB7FF09FCD8FF7E" box="[801,895,222,244]" pageId="8" pageNumber="69" type="holotype">Holotype</typeStatus>
and 
<typeStatus id="54D3887BAD30FF9FFC39FF08FBAFFF7E" box="[943,1032,223,244]" pageId="8" pageNumber="69" type="paratype">Paratype</typeStatus>
workers: 
<collectingCountry id="F37F7649AD30FF9FFBFDFF09FB44FF7E" box="[1131,1251,222,244]" name="El Salvador" pageId="8" pageNumber="69">El Salvador</collectingCountry>
, La Libertad, 
<quantity id="4C909B3CAD30FF9FFAF9FF08FA3BFF7E" box="[1391,1436,222,244]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="8" pageNumber="69" unit="mi" value="2.0">2 mi</quantity>
S. Quetzaltepec, 
<date id="FFD61019AD30FF9FFEE6FF2AFE55FE99" box="[368,498,253,275]" pageId="8" pageNumber="69" value="1961-07-17">VII-17-1961</date>
(M.E. Irwin leg.) [
<typeStatus id="54D3887BAD30FF9FFD27FF2AFCA8FE99" box="[689,783,253,275]" pageId="8" pageNumber="69" type="holotype">holotype</typeStatus>
, UCDC] (image examined) [
<typeStatus id="54D3887BAD30FF9FFBA7FF29FB2AFE99" box="[1073,1165,254,275]" pageId="8" pageNumber="69" type="paratype">paratype</typeStatus>
, MCZC] (examined). 
<taxonomicName id="4C684D5AAD30FF9FFF01FECBFE40FEBB" authority="Lacau and Groc" authorityName="Lacau and Groc" box="[151,487,283,306]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD30FF9FFF01FECBFE9DFEB8" box="[151,314,284,306]" italics="true" pageId="8" pageNumber="69">Tatuidris kapasi</emphasis>
Lacau and Groc
</taxonomicName>
, 
<emphasis id="B91CEACBAD30FF9FFE64FECAFDA1FEB8" box="[498,518,285,306]" italics="true" pageId="8" pageNumber="69">in</emphasis>
<treatmentCitation id="0AC910C8AD30FF9FFD9BFECAFD72FEBB" author="Lacau" box="[525,725,284,306]" page="2" pageId="8" pageNumber="69" year="2012">
<bibRefCitation id="EFF94B28AD30FF9FFD9BFECAFD62FEBB" author="Lacau" box="[525,709,284,306]" pageId="8" pageNumber="69" refString="Lacau, S, Groc, S., Dejean, A. de Oliveira, M. L. &amp; Delabie J. C. H. (2012) Tatuidris kapasi sp. nov.: a new armadillo ant from French Guiana (Formicidae: Agroecomyrmecinae). Psyche, 2012, Article ID 926089, 6 pages, doi: 10.1155 / 2012 / 926089" type="book" year="2012">
Lacau 
<emphasis id="B91CEACBAD30FF9FFDC4FECAFDD8FEB8" box="[594,639,284,306]" italics="true" pageId="8" pageNumber="69">et al</emphasis>
. 2012
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:2
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, their figures 1–6. 
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worker: 
<collectingCountry id="F37F7649AD30FF9FFBC4FECBFB06FEBB" box="[1106,1185,284,305]" name="French Guiana" pageId="8" pageNumber="69">Guyane</collectingCountry>
Francaise, Montagne de Kaw, 0 4ο 38.21ˏ N; 0 52ο 17.36ˏ W, Alt. 
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.
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, 
<date id="FFD61019AD30FF9FFD4EFEEDFC82FEDA" box="[728,805,314,336]" pageId="8" pageNumber="69" value="2008-09">ix.2008</date>
(S. Groc, A. Dejean, and B. Corbara leg) [CPDC] (image examined). 
<emphasis id="B91CEACBAD30FF9FFEA8FE8DFE25FEE5" bold="true" box="[318,386,346,367]" italics="true" pageId="8" pageNumber="69">n. syn.</emphasis>
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Worker, male and gyne diagnosis: With same characters as in the genus description (
<figureCitation id="13532A5CAD30FF9FFBDEFE4BFB74FE3E" box="[1096,1235,412,436]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1499,1522]" captionStartId-1="3.[151,250,1951,1974]" captionStartId-2="7.[151,250,1941,1964]" captionStartId-3="8.[151,250,1455,1478]" captionTargetBox-0="[151,1436,555,1474]" captionTargetBox-1="[212,1380,189,1920]" captionTargetBox-2="[151,1436,1045,1927]" captionTargetBox-3="[151,1435,489,1429]" captionTargetId-0="figure@1.[151,1436,555,1475]" captionTargetId-1="figure@3.[212,1380,189,1934]" captionTargetId-2="figure@7.[151,1436,1045,1928]" captionTargetId-3="figure@8.[151,1436,489,1434]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 1. SEM images of Tatuidris tatusia external morphology. A) Head in (partial) full-face view; B) lateral view of the body; C) mandibular setae; D) close-up of mandible setae." captionText-1="FIGURE 2. Tatuidris tatusia type series, showing pilosity pattern A. A – D) Full face view, lateral view, dorsal view, and label of T. tatusia Holotype; E – G) Full face view, lateral view, dorsal view, and label of T. tatusia Paratype (MCZ collection Type locality: El Salvador [Brown and Kempf 1968]). Images A – D courtesy of antweb. org." captionText-2="FIGURE 3. Images of Tatuidris tatusia gyne from Otongachi, Ecuador. Specimen CASENT 0 178881. A) Dorsal view; B) Head in full face view; C) Detail of the wings; D) Lateral view of the body. All images courtesy of antweb. org." captionText-3="FIGURE 4. Images of Tatuidris tatusia male from Chiriqui, Panama. Specimen CASENT 0 102681. A) Dorsal view; B) Head in full face view; C) Detail of the wings; D) Lateral view of the body. All images courtesy of antweb. org." httpUri-0="https://zenodo.org/record/282480/files/figure.png" httpUri-1="https://zenodo.org/record/282481/files/figure.png" httpUri-2="https://zenodo.org/record/282482/files/figure.png" httpUri-3="https://zenodo.org/record/282483/files/figure.png" pageId="8" pageNumber="69">Figures 1–4</figureCitation>
).
</paragraph>
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<paragraph id="8BD736D9AD30FF9FFF01FA78FB66FA6E" blockId="8.[151,1435,1455,1508]" pageId="8" pageNumber="69">
<emphasis id="B91CEACBAD30FF9FFF01FA78FEB4FA4E" bold="true" box="[151,275,1455,1478]" pageId="8" pageNumber="69">FIGURE 4.</emphasis>
Images of 
<taxonomicName id="4C684D5AAD30FF9FFE10FA78FD8BFA4F" box="[390,556,1455,1477]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD30FF9FFE10FA78FD8BFA4F" box="[390,556,1455,1477]" italics="true" pageId="8" pageNumber="69">Tatuidris tatusia</emphasis>
</taxonomicName>
male from Chiriqui, Panama. Specimen CASENT 0 102681. A) Dorsal view; B) Head in full face view; C) Detail of the wings; D) Lateral view of the body. All images courtesy of antweb.org.
</paragraph>
</caption>
<paragraph id="8BD736D9AD30FF9FFF51F9C7FF72F8AE" blockId="8.[151,1437,1552,2009]" pageId="8" pageNumber="69">
<emphasis id="B91CEACBAD30FF9FFF51F9C7FE3EF9A2" box="[199,409,1552,1576]" italics="true" pageId="8" pageNumber="69">Pilosity variability</emphasis>
. Currently, four striking pilosity patterns are known to occur within 
<taxonomicName id="4C684D5AAD30FF9FFB23F9C7FAB0F9A2" box="[1205,1303,1552,1576]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD30FF9FFB23F9C7FAB0F9A2" box="[1205,1303,1552,1576]" italics="true" pageId="8" pageNumber="69">Tatuidris</emphasis>
</taxonomicName>
collections (
<figureCitation id="13532A5CAD30FF9FFF08F9E3FEA6F9C6" box="[158,257,1588,1612]" captionStart="FIGURE 5" captionStartId="9.[151,250,1529,1552]" captionTargetBox="[301,1279,559,1490]" captionTargetId="figure@9.[286,1300,519,1508]" captionTargetPageId="9" captionText="FIGURE 5. Visual representation of Tatuidris pilosity patterns. Images courtesy of Natalie Clay." httpUri="https://zenodo.org/record/282484/files/figure.png" pageId="8" pageNumber="69">Figure 5</figureCitation>
). Pilosity pattern A (
<figureCitation id="13532A5CAD30FF9FFE78F9E3FDF7F9C6" box="[494,592,1588,1612]" captionStart="FIGURE 6" captionStartId="11.[151,250,1487,1510]" captionTargetBox="[151,1435,183,1475]" captionTargetId="figure@11.[151,1436,183,1476]" captionTargetPageId="11" captionText="FIGURE 6. Tatuidris tatusia and specimens showing pilosity pattern A. A – C) Full face, lateral and dorsal view of T. tatusia from Matagalpa, Nicaragua; D – F) Full face, lateral and dorsal view of T. tatusia from Otongachi, Ecuador; G – I) Full face, lateral and dorsal view of T. tatusia from Maquipucuna, Ecuador; J – L) Full-face, lateral and dorsal view of T. tatusia from Cuzco, Peru. Images A – C and J – L courtesy of antweb. org." httpUri="https://zenodo.org/record/282485/files/figure.png" pageId="8" pageNumber="69">Figure 6</figureCitation>
) consists of a mix of both long flexuous and short appressed setae. This is the most common pilosity pattern and the one that most resembles the 
<typeStatus id="54D3887BAD30FF9FFC52F98EFC53F9FB" box="[964,1012,1625,1649]" pageId="8" pageNumber="69">type</typeStatus>
specimens from 
<collectingCountry id="F37F7649AD30FF9FFB2CF98FFA98F9FA" box="[1210,1343,1624,1648]" name="El Salvador" pageId="8" pageNumber="69">El Salvador</collectingCountry>
and the gyne from Otongachi, 
<collectingCountry id="F37F7649AD30FF9FFE0DF9ABFE5DF91E" box="[411,506,1660,1684]" name="Ecuador" pageId="8" pageNumber="69">Ecuador</collectingCountry>
. Pilosity pattern B (
<figureCitation id="13532A5CAD30FF9FFD76F9ABFCE4F91E" box="[736,835,1660,1684]" captionStart="FIGURE 7" captionStartId="12.[151,250,1425,1448]" captionTargetBox="[151,1435,437,1404]" captionTargetId="figure@12.[151,1436,437,1405]" captionTargetPageId="12" captionText="FIGURE 7. Tatuidris tatusia. Specimens showing pilosity pattern B. A – C) Full face, lateral and dorsal view of T. tatusia from Caves Branch, Belize; D – F) Full face, lateral and dorsal view of T. tatusia from Belmopan, Belize; G – I) Full-face, lateral and dorsal view of T. tatusia from Maquipucuna, Ecuador. Images G – I courtesy of antweb. org." httpUri="https://zenodo.org/record/282486/files/figure.png" pageId="8" pageNumber="69">Figure 7</figureCitation>
) is characterized by very short, fully appressed, and regular spaced setae arrayed homogeneously and equidistantly on the head, mesosoma, petiole, postpetiole and gaster. Pilosity pattern C (
<figureCitation id="13532A5CAD30FF9FFE54F913FD84F956" box="[450,547,1732,1756]" captionStart="FIGURE 8" captionStartId="13.[151,250,849,872]" captionTargetBox="[151,1435,337,828]" captionTargetId="figure@13.[151,1436,337,829]" captionTargetPageId="13" captionText="FIGURE 8. Tatuidris tatusia. Specimens showing pilosity pattern C. A) Full face and B) lateral view of T. tatusia from Puntarenas, Costa Rica." httpUri="https://zenodo.org/record/282487/files/figure.png" pageId="8" pageNumber="69">Figure 8</figureCitation>
) is characterized by dense lanose-looking setae. Pilosity pattern D (
<figureCitation id="13532A5CAD30FF9FFAA7F913FA33F956" box="[1329,1428,1732,1756]" captionStart="FIGURE 9" captionStartId="13.[151,250,1937,1960]" captionTargetBox="[151,1435,952,1916]" captionTargetId="figure@13.[151,1436,952,1916]" captionTargetPageId="13" captionText="FIGURE 9. Tatuidris tatusia. Specimens showing pilosity pattern D. A – C) Full face, lateral and dorsal view of T. tatusia from Los Tuxtlas, Mexico; D – F) Full face, lateral and dorsal view of T. tatusia from Chiapas, Mexico; G – I) Full face, lateral and dorsal view of T. tatusia from Magdalena, Colombia." httpUri="https://zenodo.org/record/282488/files/figure.png" pageId="8" pageNumber="69">Figure 9</figureCitation>
) consists of short and uniform decumbent (strongly inclined but not fully appressed) setae scattered throughout the body.
</paragraph>
<paragraph id="8BD736D9AD30FF9EFF51F8E7FEC8FE58" blockId="8.[151,1437,1552,2009]" lastBlockId="9.[151,1436,151,466]" lastPageId="9" lastPageNumber="70" pageId="8" pageNumber="69">
<emphasis id="B91CEACBAD30FF9FFF51F8E7FED1F8C2" box="[199,374,1840,1864]" italics="true" pageId="8" pageNumber="69">DNA Barcodes.</emphasis>
In total, 28 sequences (20 of them with full, 658bp, length) of the barcode gene were recovered from BOLD and included in the analyses (
<tableCitation id="C6EA0362AD30FF9FFDF8F883FD66F8E6" box="[622,705,1876,1900]" captionStart="TABLE 1" captionStartId="5.[151,239,439,463]" captionTargetBox="[165,1369,528,1959]" captionTargetPageId="5" captionText="TABLE 1. Genbank accession numbers. Species, BOLD accession numbers, simple ID and country of origin are provided." httpUri="http://table.plazi.org/id/DF176651AD3DFF92FF01FE60FF5AFE65" pageId="8" pageNumber="69" tableUuid="DF176651AD3DFF92FF01FE60FF5AFE65">Table 1</tableCitation>
). Among these sequences, 120 out of the 658 basepairs (18.23%) were variable. In general, specimens (excluding colony duplicates) presented very high levels of intraspecific pairwise differences (average = 5.36%, min = 0%, max=15.08%). The upper values of intraspecific pairwise differences encountered here are well above the 2% usually recovered for species differences in DNA barcode literature in general (
<bibRefCitation id="EFF94B28AD31FF9EFE0FFF40FCBDFF25" author="Ratnasingham" box="[409,794,151,175]" pageId="9" pageNumber="70" refString="Ratnasingham, S. &amp; Hebert, P. D. N. (2007) BOLD: The Barcode of Life Data System (www. barcodinglife. org). Molecular Ecology Notes, 7, 355 - 364." type="journal article" year="2007">Ratnasingham and Hebert 2007</bibRefCitation>
), and above the percentages of average intraspecific variability reported previously for ants (e.g. between the 1.67% and 6.37% encountered for 
<taxonomicName id="4C684D5AAD31FF9EFB67FF6BFAC1FF5E" box="[1265,1382,188,212]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD31FF9EFB67FF6BFAC1FF5E" box="[1265,1382,188,212]" italics="true" pageId="9" pageNumber="70">Anochetus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C684D5AAD31FF9EFF01FF37FEE6FF72" box="[151,321,224,248]" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD31FF9EFF01FF37FEE6FF72" box="[151,321,224,248]" italics="true" pageId="9" pageNumber="70">Odontomachus</emphasis>
</taxonomicName>
in 
<collectingCountry id="F37F7649AD31FF9EFEFCFF08FE56FF7D" box="[362,497,223,247]" name="Madagascar" pageId="9" pageNumber="70">Madagascar</collectingCountry>
by 
<bibRefCitation id="EFF94B28AD31FF9EFDB4FF08FC8AFF7D" author="Fisher" box="[546,813,223,247]" pageId="9" pageNumber="70" refString="Fisher, B. L. and Smith, M. A. (2008) A revision of malagasy species of Anochetus Mayr and Odontomachus Latreille (Hymenoptera: Formicidae). PLoS ONE, 3, e 1787." type="journal article" year="2008">Fisher and Smith 2008</bibRefCitation>
; see also 
<bibRefCitation id="EFF94B28AD31FF9EFC37FF08FB0CFF7D" author="Smith" box="[929,1195,223,247]" pageId="9" pageNumber="70" refString="Smith, M. A. &amp; Fisher, B. L. (2009) Invasions, DNA barcodes, and rapid biodiversity assessment using ants of Mauritius. Frontiers in Zoology, 6, 31." type="journal article" year="2009">Smith and Fisher 2009</bibRefCitation>
, 
<bibRefCitation id="EFF94B28AD31FF9EFB2CFF37FA36FF7D" author="Jansen" box="[1210,1425,223,248]" pageId="9" pageNumber="70" refString="Jansen, G., Savolainen, R. &amp; Vepsalainen, K. (2009) DNA barcoding as a heuristic tool for classifying undescribed Nearctic Myrmica ants (Hymenoptera: Formicidae). Zoologica Scripta, 38, 527 - 536." type="journal article" year="2009">
Jansen 
<emphasis id="B91CEACBAD31FF9EFA98FF36FAEDFF72" box="[1294,1354,224,248]" italics="true" pageId="9" pageNumber="70">et al.</emphasis>
2009
</bibRefCitation>
). Pilosity patterns present among individuals (excluding pilosity pattern “C” for which no sequence was available) were not in similar clusters as depicted on the NJ tree (
<figureCitation id="13532A5CAD31FF9EFC80FEF0FC2EFEB5" box="[790,905,295,319]" captionStart="FIGURE 10" captionStartId="14.[151,250,1139,1162]" captionTargetBox="[242,1345,195,1073]" captionTargetId="figure@14.[235,1352,195,1118]" captionTargetPageId="14" captionText="FIGURE 10. Neighbor joining tree based on K 2 P distances for 28 Tatuidris specimens and four other ant taxa (Paraponera clavata, Probolomyrmex sp., Proceratium avium and Strumigenys coveri) as outgroups. Labels consist of countries, first division, localities, and specimen IDs. Specimens with no color present a pilosity pattern “ A ”, similar to the type series. Specimens with pilosity pattern “ B ” are highlighted in light blue. Specimens with pilosity pattern “ D ” are highlighted in yellow. No specimen with pilosity pattern “ C ” was available for sequencing. Asterisks above nodes represent nodes with&gt; 70 % bootstrap support (999 repetitions). The tree is drawn to scale, with branch lengths in the same units as those of the genetic divergence inferred using the K 2 P model." httpUri="https://zenodo.org/record/282489/files/figure.png" pageId="9" pageNumber="70">Figure 10</figureCitation>
). Instead, four different clusters (groups) with strong geographic structure were observed: G1, 
<collectingCountry id="F37F7649AD31FF9EFD54FE9BFCB0FEEE" box="[706,791,332,356]" name="Mexico" pageId="9" pageNumber="70">Mexico</collectingCountry>
+ 
<collectingCountry id="F37F7649AD31FF9EFCB0FE9BFC32FEEE" box="[806,917,332,356]" name="Honduras" pageId="9" pageNumber="70">Honduras</collectingCountry>
+ 
<collectingCountry id="F37F7649AD31FF9EFC32FE9BFB86FEEE" box="[932,1057,332,356]" name="Guatemala" pageId="9" pageNumber="70">Guatemala</collectingCountry>
; G2, 
<collectingCountry id="F37F7649AD31FF9EFBF4FE9BFB47FEEE" box="[1122,1248,332,356]" name="Costa Rica" pageId="9" pageNumber="70">Costa Rica</collectingCountry>
+ 
<collectingCountry id="F37F7649AD31FF9EFB79FE9BFAC2FEEE" box="[1263,1381,332,356]" name="Nicaragua" pageId="9" pageNumber="70">Nicaragua</collectingCountry>
; G3, 
<collectingCountry id="F37F7649AD31FF9EFF01FEB8FF53FE0D" box="[151,244,367,391]" name="Ecuador" pageId="9" pageNumber="70">Ecuador</collectingCountry>
(West of Andes); and G4, 
<collectingCountry id="F37F7649AD31FF9EFDBCFEB8FD20FE0D" box="[554,647,367,391]" name="Ecuador" pageId="9" pageNumber="70">Ecuador</collectingCountry>
(East of Andes/Amazon Basin) (
<figureCitation id="13532A5CAD31FF9EFB97FEB8FBD4FE0D" box="[1025,1139,367,391]" captionStart="FIGURE 10" captionStartId="14.[151,250,1139,1162]" captionTargetBox="[242,1345,195,1073]" captionTargetId="figure@14.[235,1352,195,1118]" captionTargetPageId="14" captionText="FIGURE 10. Neighbor joining tree based on K 2 P distances for 28 Tatuidris specimens and four other ant taxa (Paraponera clavata, Probolomyrmex sp., Proceratium avium and Strumigenys coveri) as outgroups. Labels consist of countries, first division, localities, and specimen IDs. Specimens with no color present a pilosity pattern “ A ”, similar to the type series. Specimens with pilosity pattern “ B ” are highlighted in light blue. Specimens with pilosity pattern “ D ” are highlighted in yellow. No specimen with pilosity pattern “ C ” was available for sequencing. Asterisks above nodes represent nodes with&gt; 70 % bootstrap support (999 repetitions). The tree is drawn to scale, with branch lengths in the same units as those of the genetic divergence inferred using the K 2 P model." httpUri="https://zenodo.org/record/282489/files/figure.png" pageId="9" pageNumber="70">Figure 10</figureCitation>
). Furthermore, across this geographic range, pairwise barcode divergence among specimens was significantly related to distance (R2=0.37, p&lt;0.01, 
<figureCitation id="13532A5CAD31FF9EFF62FE6DFEC5FE58" box="[244,354,442,466]" captionStart="FIGURE 11" captionStartId="15.[151,250,1072,1095]" captionTargetBox="[265,1318,213,1046]" captionTargetId="figure@15.[255,1332,193,1051]" captionTargetPageId="15" captionText="FIGURE 11. Linear regression between genetic distance (pairwise divergence) and geographic distance." httpUri="https://zenodo.org/record/282490/files/figure.png" pageId="9" pageNumber="70">Figure 11</figureCitation>
).
</paragraph>
<caption id="DF176651AD31FF9EFF01FA2EFBD4F99A" box="[151,1139,1529,1552]" httpUri="https://zenodo.org/record/282484/files/figure.png" pageId="9" pageNumber="70" targetBox="[301,1279,559,1490]" targetPageId="9">
<paragraph id="8BD736D9AD31FF9EFF01FA2EFBD4F99A" blockId="9.[151,1139,1529,1552]" box="[151,1139,1529,1552]" pageId="9" pageNumber="70">
<emphasis id="B91CEACBAD31FF9EFF01FA2EFEB5F985" bold="true" box="[151,274,1529,1552]" pageId="9" pageNumber="70">FIGURE 5.</emphasis>
Visual representation of 
<taxonomicName id="4C684D5AAD31FF9EFD87FA2DFDCEF99A" box="[529,617,1530,1552]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD31FF9EFD87FA2DFDCEF99A" box="[529,617,1530,1552]" italics="true" pageId="9" pageNumber="70">Tatuidris</emphasis>
</taxonomicName>
pilosity patterns. Images courtesy of Natalie Clay.
</paragraph>
</caption>
<paragraph id="8BD736D9AD31FF9EFF51F9E9FF64F86A" blockId="9.[151,1436,1597,2016]" pageId="9" pageNumber="70">
<emphasis id="B91CEACBAD31FF9EFF51F9E9FF53F9DF" box="[199,244,1598,1621]" italics="true" pageId="9" pageNumber="70">Size</emphasis>
. Specimens of 
<taxonomicName id="4C684D5AAD31FF9EFE37F9E9FDAEF9DF" box="[417,521,1598,1621]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD31FF9EFE37F9E9FDAEF9DF" box="[417,521,1598,1621]" italics="true" pageId="9" pageNumber="70">T. tatusia</emphasis>
</taxonomicName>
are small (average WbL = 
<quantity id="4C909B3CAD31FF9EFCABF9EAFC39F9DF" box="[829,926,1597,1621]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="70" unit="mm" value="0.62">0.62mm</quantity>
), but specimens can vary greatly in size, with larger specimens being twice as large as the smaller ones (min WbL = 
<quantity id="4C909B3CAD31FF9EFC4CF9B7FBE4F9F2" box="[986,1091,1632,1656]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="70" unit="mm" value="0.45">0.45 mm</quantity>
, max WbL = 
<quantity id="4C909B3CAD31FF9EFB7CF9B7FAF1F9F2" box="[1258,1366,1632,1656]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="70" unit="mm" value="0.9">0.90 mm</quantity>
). Size variability within trap catches (possibly same colonies) may be considerable. For example, workers from one collection catch in 
<collectingCountry id="F37F7649AD31FF9EFEFAF97FFE78F94A" box="[364,479,1704,1728]" name="Nicaragua" pageId="9" pageNumber="70">Nicaragua</collectingCountry>
(collection series MGB#1179) varied 30% in size (WbL from 
<quantity id="4C909B3CAD31FF9EFB36F97FFAFAF94A" box="[1184,1373,1704,1728]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="9" pageNumber="70" unit="mm" value="0.75" valueMax="0.85" valueMin="0.65">0.65 to 0.85 mm</quantity>
). It is still unclear whether intra-colony size variation is due to the presence of morphological worker castes (e.g. minor and 
<taxonomicName id="4C684D5AAD31FF9EFF50F927FEA0F882" box="[198,263,1776,1800]" class="Insecta" family="Formicidae" genus="Cyphomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="70" phylum="Arthropoda" rank="species" species="major">major</taxonomicName>
castes) or continuous size variability. The PCA analysis revealed that most variability among specimens is related to size (proportion of variance explained by PC-1 = 0.915), with PC-2, PC-3 and PC-4 (e.g. shape) explaining little (0.033, 0.021 and 0.011, respectively) of total variation (
<tableCitation id="C6EA0362AD31FF9EFC55F8EFFBBDF8DA" box="[963,1050,1848,1872]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[223,1350,240,967]" captionTargetPageId="10" captionText="TABLE 2. First four PCA coefficients for morphological variables retained in this study. PC eigenvalues (% of variance explained) are also given." httpUri="http://table.plazi.org/id/DF176651AD32FF9DFF01FF40FE0EFF45" pageId="9" pageNumber="70" tableUuid="DF176651AD32FF9DFF01FF40FE0EFF45">Table 2</tableCitation>
). Eye length loaded positively and contrasted against all other variables, in PC2. Tibia length and tibia width loaded negatively and contrasted against most other variables, in PC3. Size variability summarized in this analysis was not related to pilosity patterns (
<figureCitation id="13532A5CAD31FF9EFF08F872FEB6F837" box="[158,273,1957,1981]" captionStart="FIGURE 12" captionStartId="16.[151,250,1462,1485]" captionTargetBox="[323,1269,540,1432]" captionTargetId="figure@16.[308,1278,519,1441]" captionTargetPageId="16" captionText="FIGURE 12. PC scores for specimens included in this analysis. Symbols represent different pilosity patterns. Circles = pilosity pattern A (similar to type series). Crosses = pilosity pattern B. Triangles = pilosity pattern C. X's = pilosity pattern D." httpUri="https://zenodo.org/record/282491/files/figure.png" pageId="9" pageNumber="70">Figure 12</figureCitation>
). In general no PC correlated with pilosity patterns. Hand drawings courtesy of Natalie Clay (
<figureCitation id="13532A5CAD31FF9EFAC7F872FF11F86A" captionStart="FIGURE 12" captionStartId="16.[151,250,1462,1485]" captionTargetBox="[323,1269,540,1432]" captionTargetId="figure@16.[308,1278,519,1441]" captionTargetPageId="16" captionText="FIGURE 12. PC scores for specimens included in this analysis. Symbols represent different pilosity patterns. Circles = pilosity pattern A (similar to type series). Crosses = pilosity pattern B. Triangles = pilosity pattern C. X's = pilosity pattern D." httpUri="https://zenodo.org/record/282491/files/figure.png" pageId="9" pageNumber="70">Figure 12</figureCitation>
).
</paragraph>
<caption id="DF176651AD32FF9DFF01FF40FE0EFF45" ID-Table-UUID="DF176651AD32FF9DFF01FF40FE0EFF45" httpUri="http://table.plazi.org/id/DF176651AD32FF9DFF01FF40FE0EFF45" pageId="10" pageNumber="71" targetBox="[223,1350,240,967]" targetIsTable="true" targetPageId="10">
<paragraph id="8BD736D9AD32FF9DFF01FF40FE0EFF45" blockId="10.[151,1435,151,207]" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF01FF40FEAEFF25" bold="true" box="[151,265,151,175]" pageId="10" pageNumber="71">TABLE 2.</emphasis>
First four PCA coefficients for morphological variables retained in this study. PC eigenvalues (% of variance explained) are also given.
</paragraph>
</caption>
<paragraph id="8BD736D9AD32FF9DFF69FF27FAE7FC4C" pageId="10" pageNumber="71">
<table id="F968C479AD320068FF49FF27FAE1FC4D" box="[223,1350,240,967]" gridcols="5" gridrows="16" pageId="10" pageNumber="71">
<tr id="3558349BAD320068FF49FF27FAE1FE8D" box="[223,1350,240,263]" gridrow="0" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FF27FEF0FE8D" box="[223,343,240,263]" gridcol="0" gridrow="0" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF69FF27FE90FE8D" bold="true" box="[255,311,240,263]" pageId="10" pageNumber="71">Trait</emphasis>
</th>
<th id="76895DE7AD320068FE6AFF27FDE5FE8D" box="[508,578,240,263]" gridcol="1" gridrow="0" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFD9EFF27FD91FE8D" bold="true" box="[520,566,240,263]" pageId="10" pageNumber="71">PC1</emphasis>
</th>
<th id="76895DE7AD320068FD63FF27FCE0FE8D" box="[757,839,240,263]" gridcol="2" gridrow="0" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFC91FF27FC92FE8D" bold="true" box="[775,821,240,263]" pageId="10" pageNumber="71">PC2</emphasis>
</th>
<th id="76895DE7AD320068FC62FF27FBE0FE8D" box="[1012,1095,240,263]" gridcol="3" gridrow="0" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFB90FF27FB93FE8D" bold="true" box="[1030,1076,240,263]" pageId="10" pageNumber="71">PC3</emphasis>
</th>
<th id="76895DE7AD320068FB65FF27FAE1FE8D" box="[1267,1350,240,263]" gridcol="4" gridrow="0" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFA93FF27FA94FE8D" bold="true" box="[1285,1331,240,263]" pageId="10" pageNumber="71">PC4</emphasis>
</th>
</tr>
<tr id="3558349BAD320068FF49FECCFAE1FEB8" box="[223,1350,283,306]" gridrow="1" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FECCFEF0FEB8" box="[223,343,283,306]" gridcol="0" gridrow="1" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF6BFECCFE9EFEB8" bold="true" box="[253,313,283,306]" pageId="10" pageNumber="71">AScL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFECCFDE5FEB8" box="[508,578,283,306]" gridcol="1" gridrow="1" pageId="10" pageNumber="71">0.2737</td>
<td id="76895DE7AD320068FD63FECCFCE0FEB8" box="[757,839,283,306]" gridcol="2" gridrow="1" pageId="10" pageNumber="71">–0.0619</td>
<td id="76895DE7AD320068FC62FECCFBE0FEB8" box="[1012,1095,283,306]" gridcol="3" gridrow="1" pageId="10" pageNumber="71">0.0329</td>
<td id="76895DE7AD320068FB65FECCFAE1FEB8" box="[1267,1350,283,306]" gridcol="4" gridrow="1" pageId="10" pageNumber="71">0.2525</td>
</tr>
<tr id="3558349BAD320068FF49FE9FFAE1FED5" box="[223,1350,328,351]" gridrow="2" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FE9FFEF0FED5" box="[223,343,328,351]" gridcol="0" gridrow="2" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF6FFE9FFE99FED5" bold="true" box="[249,318,328,351]" pageId="10" pageNumber="71">AScW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFE9FFDE5FED5" box="[508,578,328,351]" gridcol="1" gridrow="2" pageId="10" pageNumber="71">0.2730</td>
<td id="76895DE7AD320068FD63FE9FFCE0FED5" box="[757,839,328,351]" gridcol="2" gridrow="2" pageId="10" pageNumber="71">–0.1618</td>
<td id="76895DE7AD320068FC62FE9FFBE0FED5" box="[1012,1095,328,351]" gridcol="3" gridrow="2" pageId="10" pageNumber="71">–0.0559</td>
<td id="76895DE7AD320068FB65FE9FFAE1FED5" box="[1267,1350,328,351]" gridcol="4" gridrow="2" pageId="10" pageNumber="71">0.1040</td>
</tr>
<tr id="3558349BAD320068FF49FEA3FAE1FE00" box="[223,1350,372,394]" gridrow="3" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FEA3FEF0FE00" box="[223,343,372,394]" gridcol="0" gridrow="3" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE9CFEA3FE8BFE03" bold="true" box="[266,300,372,393]" pageId="10" pageNumber="71">EL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFEA3FDE5FE00" box="[508,578,372,394]" gridcol="1" gridrow="3" pageId="10" pageNumber="71">0.2135</td>
<td id="76895DE7AD320068FD63FEA3FCE0FE00" box="[757,839,372,394]" gridcol="2" gridrow="3" pageId="10" pageNumber="71">0.9362</td>
<td id="76895DE7AD320068FC62FEA3FBE0FE00" box="[1012,1095,372,394]" gridcol="3" gridrow="3" pageId="10" pageNumber="71">–0.1003</td>
<td id="76895DE7AD320068FB65FEA3FAE1FE00" box="[1267,1350,372,394]" gridcol="4" gridrow="3" pageId="10" pageNumber="71">–0.0722</td>
</tr>
<tr id="3558349BAD320068FF49FE77FAE1FE3C" box="[223,1350,416,438]" gridrow="4" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FE77FEF0FE3C" box="[223,343,416,438]" gridcol="0" gridrow="4" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE9FFE76FE8AFE3C" bold="true" box="[265,301,417,438]" pageId="10" pageNumber="71">HL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFE77FDE5FE3C" box="[508,578,416,438]" gridcol="1" gridrow="4" pageId="10" pageNumber="71">0.2761</td>
<td id="76895DE7AD320068FD63FE77FCE0FE3C" box="[757,839,416,438]" gridcol="2" gridrow="4" pageId="10" pageNumber="71">–0.0490</td>
<td id="76895DE7AD320068FC62FE77FBE0FE3C" box="[1012,1095,416,438]" gridcol="3" gridrow="4" pageId="10" pageNumber="71">0.0880</td>
<td id="76895DE7AD320068FB65FE77FAE1FE3C" box="[1267,1350,416,438]" gridcol="4" gridrow="4" pageId="10" pageNumber="71">0.2140</td>
</tr>
<tr id="3558349BAD320068FF49FE1BFAE1FE68" box="[223,1350,460,482]" gridrow="5" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FE1BFEF0FE68" box="[223,343,460,482]" gridcol="0" gridrow="5" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE93FE1AFE95FE68" bold="true" box="[261,306,461,482]" pageId="10" pageNumber="71">HW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFE1BFDE5FE68" box="[508,578,460,482]" gridcol="1" gridrow="5" pageId="10" pageNumber="71">0.2780</td>
<td id="76895DE7AD320068FD63FE1BFCE0FE68" box="[757,839,460,482]" gridcol="2" gridrow="5" pageId="10" pageNumber="71">–0.0043</td>
<td id="76895DE7AD320068FC62FE1BFBE0FE68" box="[1012,1095,460,482]" gridcol="3" gridrow="5" pageId="10" pageNumber="71">0.0557</td>
<td id="76895DE7AD320068FB65FE1BFAE1FE68" box="[1267,1350,460,482]" gridcol="4" gridrow="5" pageId="10" pageNumber="71">–0.0271</td>
</tr>
<tr id="3558349BAD320068FF49FE2FFAE1FD84" box="[223,1350,504,526]" gridrow="6" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FE2FFEF0FD84" box="[223,343,504,526]" gridcol="0" gridrow="6" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE92FE2FFE95FD87" bold="true" box="[260,306,504,525]" pageId="10" pageNumber="71">IAD</emphasis>
</th>
<td id="76895DE7AD320068FE6AFE2FFDE5FD84" box="[508,578,504,526]" gridcol="1" gridrow="6" pageId="10" pageNumber="71">0.2749</td>
<td id="76895DE7AD320068FD63FE2FFCE0FD84" box="[757,839,504,526]" gridcol="2" gridrow="6" pageId="10" pageNumber="71">–0.1594</td>
<td id="76895DE7AD320068FC62FE2FFBE0FD84" box="[1012,1095,504,526]" gridcol="3" gridrow="6" pageId="10" pageNumber="71">0.0246</td>
<td id="76895DE7AD320068FB65FE2FFAE1FD84" box="[1267,1350,504,526]" gridcol="4" gridrow="6" pageId="10" pageNumber="71">0.1754</td>
</tr>
<tr id="3558349BAD320068FF49FDF3FAE1FDB0" box="[223,1350,548,570]" gridrow="7" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FDF3FEF0FDB0" box="[223,343,548,570]" gridcol="0" gridrow="7" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE9DFDF2FE8CFDB0" bold="true" box="[267,299,549,570]" pageId="10" pageNumber="71">PL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFDF3FDE5FDB0" box="[508,578,548,570]" gridcol="1" gridrow="7" pageId="10" pageNumber="71">0.2679</td>
<td id="76895DE7AD320068FD63FDF3FCE0FDB0" box="[757,839,548,570]" gridcol="2" gridrow="7" pageId="10" pageNumber="71">–0.1677</td>
<td id="76895DE7AD320068FC62FDF3FBE0FDB0" box="[1012,1095,548,570]" gridcol="3" gridrow="7" pageId="10" pageNumber="71">0.1582</td>
<td id="76895DE7AD320068FB65FDF3FAE1FDB0" box="[1267,1350,548,570]" gridcol="4" gridrow="7" pageId="10" pageNumber="71">–0.4504</td>
</tr>
<tr id="3558349BAD320068FF49FD87FAE1FDEC" box="[223,1350,592,614]" gridrow="8" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FD87FEF0FDEC" box="[223,343,592,614]" gridcol="0" gridrow="8" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE92FD86FE95FDEC" bold="true" box="[260,306,593,614]" pageId="10" pageNumber="71">PpL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFD87FDE5FDEC" box="[508,578,592,614]" gridcol="1" gridrow="8" pageId="10" pageNumber="71">0.2643</td>
<td id="76895DE7AD320068FD63FD87FCE0FDEC" box="[757,839,592,614]" gridcol="2" gridrow="8" pageId="10" pageNumber="71">0.0462</td>
<td id="76895DE7AD320068FC62FD87FBE0FDEC" box="[1012,1095,592,614]" gridcol="3" gridrow="8" pageId="10" pageNumber="71">0.4339</td>
<td id="76895DE7AD320068FB65FD87FAE1FDEC" box="[1267,1350,592,614]" gridcol="4" gridrow="8" pageId="10" pageNumber="71">–0.3384</td>
</tr>
<tr id="3558349BAD320068FF49FDABFAE1FD18" box="[223,1350,636,658]" gridrow="9" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FDABFEF0FD18" box="[223,343,636,658]" gridcol="0" gridrow="9" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE91FDAAFE97FD18" bold="true" box="[263,304,637,658]" pageId="10" pageNumber="71">PW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFDABFDE5FD18" box="[508,578,636,658]" gridcol="1" gridrow="9" pageId="10" pageNumber="71">0.2739</td>
<td id="76895DE7AD320068FD63FDABFCE0FD18" box="[757,839,636,658]" gridcol="2" gridrow="9" pageId="10" pageNumber="71">–0.0537</td>
<td id="76895DE7AD320068FC62FDABFBE0FD18" box="[1012,1095,636,658]" gridcol="3" gridrow="9" pageId="10" pageNumber="71">0.1803</td>
<td id="76895DE7AD320068FB65FDABFAE1FD18" box="[1267,1350,636,658]" gridcol="4" gridrow="9" pageId="10" pageNumber="71">–0.1468</td>
</tr>
<tr id="3558349BAD320068FF49FD7FFAE1FD34" box="[223,1350,680,702]" gridrow="10" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FD7FFEF0FD34" box="[223,343,680,702]" gridcol="0" gridrow="10" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE96FD7EFE90FD34" bold="true" box="[256,311,681,702]" pageId="10" pageNumber="71">PpW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFD7FFDE5FD34" box="[508,578,680,702]" gridcol="1" gridrow="10" pageId="10" pageNumber="71">0.2758</td>
<td id="76895DE7AD320068FD63FD7FFCE0FD34" box="[757,839,680,702]" gridcol="2" gridrow="10" pageId="10" pageNumber="71">0.0473</td>
<td id="76895DE7AD320068FC62FD7FFBE0FD34" box="[1012,1095,680,702]" gridcol="3" gridrow="10" pageId="10" pageNumber="71">0.0436</td>
<td id="76895DE7AD320068FB65FD7FFAE1FD34" box="[1267,1350,680,702]" gridcol="4" gridrow="10" pageId="10" pageNumber="71">0.1971</td>
</tr>
<tr id="3558349BAD320068FF49FD03FAE1FD60" box="[223,1350,724,746]" gridrow="11" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FD03FEF0FD60" box="[223,343,724,746]" gridcol="0" gridrow="11" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE97FD02FE92FD60" bold="true" box="[257,309,725,746]" pageId="10" pageNumber="71">PrW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFD03FDE5FD60" box="[508,578,724,746]" gridcol="1" gridrow="11" pageId="10" pageNumber="71">0.2767</td>
<td id="76895DE7AD320068FD63FD03FCE0FD60" box="[757,839,724,746]" gridcol="2" gridrow="11" pageId="10" pageNumber="71">0.0585</td>
<td id="76895DE7AD320068FC62FD03FBE0FD60" box="[1012,1095,724,746]" gridcol="3" gridrow="11" pageId="10" pageNumber="71">–0.0261</td>
<td id="76895DE7AD320068FB65FD03FAE1FD60" box="[1267,1350,724,746]" gridcol="4" gridrow="11" pageId="10" pageNumber="71">0.0450</td>
</tr>
<tr id="3558349BAD320068FF49FCD7FAE1FC9D" box="[223,1350,768,791]" gridrow="12" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FCD7FEF0FC9D" box="[223,343,768,791]" gridcol="0" gridrow="12" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE91FCD7FE97FC9D" bold="true" box="[263,304,768,791]" pageId="10" pageNumber="71">TiL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFCD7FDE5FC9D" box="[508,578,768,791]" gridcol="1" gridrow="12" pageId="10" pageNumber="71">0.2440</td>
<td id="76895DE7AD320068FD63FCD7FCE0FC9D" box="[757,839,768,791]" gridcol="2" gridrow="12" pageId="10" pageNumber="71">–0.1402</td>
<td id="76895DE7AD320068FC62FCD7FBE0FC9D" box="[1012,1095,768,791]" gridcol="3" gridrow="12" pageId="10" pageNumber="71">–0.8193</td>
<td id="76895DE7AD320068FB65FCD7FAE1FC9D" box="[1267,1350,768,791]" gridcol="4" gridrow="12" pageId="10" pageNumber="71">–0.4051</td>
</tr>
<tr id="3558349BAD320068FF49FCFBFAE1FCC9" box="[223,1350,812,835]" gridrow="13" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FCFBFEF0FCC9" box="[223,343,812,835]" gridcol="0" gridrow="13" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFE95FCFBFE93FCC9" bold="true" box="[259,308,812,835]" pageId="10" pageNumber="71">TiW</emphasis>
</th>
<td id="76895DE7AD320068FE6AFCFBFDE5FCC9" box="[508,578,812,835]" gridcol="1" gridrow="13" pageId="10" pageNumber="71">0.2672</td>
<td id="76895DE7AD320068FD63FCFBFCE0FCC9" box="[757,839,812,835]" gridcol="2" gridrow="13" pageId="10" pageNumber="71">–0.0193</td>
<td id="76895DE7AD320068FC62FCFBFBE0FCC9" box="[1012,1095,812,835]" gridcol="3" gridrow="13" pageId="10" pageNumber="71">–0.2126</td>
<td id="76895DE7AD320068FB65FCFBFAE1FCC9" box="[1267,1350,812,835]" gridcol="4" gridrow="13" pageId="10" pageNumber="71">0.5268</td>
</tr>
<tr id="3558349BAD320068FF49FC8FFAE1FCE4" box="[223,1350,856,878]" gridrow="14" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FC8FFEF0FCE4" box="[223,343,856,878]" gridcol="0" gridrow="14" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF69FC8EFE90FCE4" bold="true" box="[255,311,857,878]" pageId="10" pageNumber="71">WbL</emphasis>
</th>
<td id="76895DE7AD320068FE6AFC8FFDE5FCE4" box="[508,578,856,878]" gridcol="1" gridrow="14" pageId="10" pageNumber="71">0.2752</td>
<td id="76895DE7AD320068FD63FC8FFCE0FCE4" box="[757,839,856,878]" gridcol="2" gridrow="14" pageId="10" pageNumber="71">–0.0823</td>
<td id="76895DE7AD320068FC62FC8FFBE0FCE4" box="[1012,1095,856,878]" gridcol="3" gridrow="14" pageId="10" pageNumber="71">0.0967</td>
<td id="76895DE7AD320068FB65FC8FFAE1FCE4" box="[1267,1350,856,878]" gridcol="4" gridrow="14" pageId="10" pageNumber="71">–0.1464</td>
</tr>
<tr id="3558349BAD320068FF49FC67FAE1FC4D" box="[223,1350,944,967]" gridrow="15" pageId="10" pageNumber="71">
<th id="76895DE7AD320068FF49FC67FEF0FC4D" box="[223,343,944,967]" gridcol="0" gridrow="15" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF49FC67FEF0FC4D" bold="true" box="[223,343,944,967]" pageId="10" pageNumber="71">Eigenvalue</emphasis>
</th>
<td id="76895DE7AD320068FE6AFC67FDE5FC4D" box="[508,578,944,967]" gridcol="1" gridrow="15" pageId="10" pageNumber="71">0.9154</td>
<td id="76895DE7AD320068FD63FC67FCE0FC4D" box="[757,839,944,967]" gridcol="2" gridrow="15" pageId="10" pageNumber="71">0.0334</td>
<td id="76895DE7AD320068FC62FC67FBE0FC4D" box="[1012,1095,944,967]" gridcol="3" gridrow="15" pageId="10" pageNumber="71">0.0211</td>
<td id="76895DE7AD320068FB65FC67FAE1FC4D" box="[1267,1350,944,967]" gridcol="4" gridrow="15" pageId="10" pageNumber="71">0.0112</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD736D9AD32FF9DFF51FBDFFC63FAEE" blockId="10.[151,1437,1032,1992]" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF51FBDFFE96FBAA" box="[199,305,1032,1056]" italics="true" pageId="10" pageNumber="71">Sculpture</emphasis>
. The strength and depth of all sculpture patterns is accentuated in larger sizes. Collections from 
<collectingCountry id="F37F7649AD32FF9DFF01FBFBFEADFBCE" box="[151,266,1068,1092]" name="Nicaragua" pageId="10" pageNumber="71">Nicaragua</collectingCountry>
also tend to present more accentuated sculpture patterns. The head dorsum is usually smooth and shining, except for the area below eyes, which presents longitudinal carinae. The head vertex is covered with transverse carinulae. The lateral surface of the mandible is smooth and shining except for longitudinal superficial striae on the side that vary in depth. The antennal scape is shagreened and superficially areolate. The surface of the ventrolateral part of the pronotum varies strongly across specimens, from smooth and shining to strongly striate, or carinulate. The dorsum of the mesosoma has concentric carinulae and sometimes is slightly punctate. The mesopleuron is smooth and shining except for punctuations and areolae on the ventral margin. The propodeal declivity is smooth with fine transverse striae. The petiole and postpetiole are dorso-laterally strigulate. The gaster is mostly smooth and shiny but sometimes finely and sparsely strigulate.
</paragraph>
</subSubSection>
<subSubSection id="C3726552AD32FF9BFF51FAA7FB36F810" lastPageId="12" lastPageNumber="73" pageId="10" pageNumber="71" type="distribution">
<paragraph id="8BD736D9AD32FF9DFF51FAA7FC02F9B6" blockId="10.[151,1437,1032,1992]" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF51FAA7FEF7FA02" box="[199,336,1392,1416]" italics="true" pageId="10" pageNumber="71">Distribution</emphasis>
. The genus 
<taxonomicName id="4C684D5AAD32FF9DFE77FAA7FDE2FA02" box="[481,581,1392,1416]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFE77FAA7FDE2FA02" box="[481,581,1392,1416]" italics="true" pageId="10" pageNumber="71">Tatuidris</emphasis>
</taxonomicName>
is restricted to the Neotropics, but it has an ample distribution that spans from northern 
<collectingCountry id="F37F7649AD32FF9DFEABFA43FE35FA26" box="[317,402,1428,1452]" name="Mexico" pageId="10" pageNumber="71">Mexico</collectingCountry>
to central 
<collectingCountry id="F37F7649AD32FF9DFD98FA43FDF2FA26" box="[526,597,1428,1452]" name="Brazil" pageId="10" pageNumber="71">Brazil</collectingCountry>
, 
<collectingCountry id="F37F7649AD32FF9DFDF4FA43FCAEFA26" box="[610,777,1428,1452]" name="French Guiana" pageId="10" pageNumber="71">French Guiana</collectingCountry>
(
<bibRefCitation id="EFF94B28AD32FF9DFC8FFA43FC43FA26" author="Lacau" box="[793,996,1428,1452]" pageId="10" pageNumber="71" refString="Lacau, S, Groc, S., Dejean, A. de Oliveira, M. L. &amp; Delabie J. C. H. (2012) Tatuidris kapasi sp. nov.: a new armadillo ant from French Guiana (Formicidae: Agroecomyrmecinae). Psyche, 2012, Article ID 926089, 6 pages, doi: 10.1155 / 2012 / 926089" type="book" year="2012">
Lacau 
<emphasis id="B91CEACBAD32FF9DFCFEFA42FC05FA26" box="[872,930,1428,1452]" italics="true" pageId="10" pageNumber="71">et al.</emphasis>
2012
</bibRefCitation>
) and Amazonian 
<collectingCountry id="F37F7649AD32FF9DFB38FA42FB46FA26" box="[1198,1249,1429,1452]" name="Peru" pageId="10" pageNumber="71">Peru</collectingCountry>
(
<figureCitation id="13532A5CAD32FF9DFB67FA43FAC3FA26" box="[1265,1380,1428,1452]" captionStart="FIGURE 13" captionStartId="17.[151,250,1289,1312]" captionTargetBox="[156,1430,321,1248]" captionTargetId="figure@17.[151,1436,301,1268]" captionTargetPageId="17" captionText="FIGURE 13. Map of localities of specimens included in this study. Black diamonds represent localities from which specimens included in the CO 1 DNA barcodes analysis were obtained. Circled stars represent type localities from the two previously know species (e. g. Tatuidris tatusia Brown and Kempf, and Tatuidris kapasi Lacau and Groc syn. nov.)." httpUri="https://zenodo.org/record/282492/files/figure.png" pageId="10" pageNumber="71">Figure 13</figureCitation>
). No collections are known from the Caribbean, Galápagos, or other islands. Most specimens and collections are currently known to occur in localities west of the Andes, with more collections tending to occur towards Central 
<collectingCountry id="F37F7649AD32FF9DFF01F9D7FF50F992" box="[151,247,1536,1560]" name="United States of America" pageId="10" pageNumber="71">America</collectingCountry>
and 
<collectingCountry id="F37F7649AD32FF9DFEA5F9D7FE2BF992" box="[307,396,1536,1560]" name="Mexico" pageId="10" pageNumber="71">Mexico</collectingCountry>
. Most collections come from mountainside (pre-montane) areas at mid elevations (usually 
<quantity id="4C909B3CAD32FF9DFF01F9F3FEBBF9B6" box="[151,284,1572,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="10" pageNumber="71" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">800–1200m</quantity>
of altitude). Collections from lowland Amazonia are few.
</paragraph>
<paragraph id="8BD736D9AD32FF9DFF51F99FFE5CF842" blockId="10.[151,1437,1032,1992]" pageId="10" pageNumber="71">
<emphasis id="B91CEACBAD32FF9DFF51F99FFEDFF9EA" box="[199,376,1608,1632]" italics="true" pageId="10" pageNumber="71">Natural history.</emphasis>
Little is known about the biology of the ant genus 
<taxonomicName id="4C684D5AAD32FF9DFC3AF99FFBB7F9EA" box="[940,1040,1608,1632]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFC3AF99FFBB7F9EA" box="[940,1040,1608,1632]" italics="true" pageId="10" pageNumber="71">Tatuidris</emphasis>
</taxonomicName>
and, until recently, no observations of live specimens were registered. Details of a first collection event of a small live colony (
<specimenCount id="9D6EFD50AD32FF9DFB00F9BBFAA2F90E" box="[1174,1285,1644,1668]" pageId="10" pageNumber="71" type="worker">3 workers</specimenCount>
and 4 gynes) by Dr. Thibaut Delsinne (pers. comm.) in a mid-elevation forest in southeastern 
<collectingCountry id="F37F7649AD32FF9DFB85F947FBD7F922" box="[1043,1136,1680,1704]" name="Ecuador" pageId="10" pageNumber="71">Ecuador</collectingCountry>
suggest that 
<taxonomicName id="4C684D5AAD32FF9DFA97F947FAC2F922" box="[1281,1381,1680,1704]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFA97F947FAC2F922" box="[1281,1381,1680,1704]" italics="true" pageId="10" pageNumber="71">Tatuidris</emphasis>
</taxonomicName>
may well be a highly specialized predator, as colonies kept in captivity did not accept any food item offered to them. Food items rejected by the ants included live and dead termites, millipedes, mites, various insect parts, sugar water, tuna, biscuits, live and dead fruit flies (
<taxonomicName id="4C684D5AAD32FF9DFDCDF92BFD7EF89E" box="[603,729,1788,1812]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFDCDF92BFD7EF89E" box="[603,729,1788,1812]" italics="true" pageId="10" pageNumber="71">Drosophila</emphasis>
</taxonomicName>
), live springtails, live myriapods (Chilopoda and Diplopoda), live and dead Diplura, small live spiders, small live pseudoscorpions, one small snail, uncooked hen egg (i.e. piece of cotton wool soaked with fresh whisked hen egg; 
<bibRefCitation id="EFF94B28AD32FF9DFD61F892FC2EF8D6" author="Brown" box="[759,905,1860,1884]" pageId="10" pageNumber="71" refString="Brown, W. L. (1977) An aberrant new genus of myrmicine ant from Madagascar. Psyche, 84, 218 - 224." type="journal article" year="1977">Brown 1977</bibRefCitation>
), ant larvae (
<taxonomicName id="4C684D5AAD32FF9DFBB3F892FB6AF8D6" box="[1061,1229,1861,1884]" class="Insecta" family="Formicidae" genus="Gnamptogenys" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFBB3F892FB6AF8D6" box="[1061,1229,1861,1884]" italics="true" pageId="10" pageNumber="71">Gnamptogenys</emphasis>
</taxonomicName>
sp.), and live ant workers (
<taxonomicName id="4C684D5AAD32FF9DFE95F8BFFE02F80A" box="[259,421,1896,1920]" class="Insecta" family="Formicidae" genus="Cyphomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFE95F8BFFE02F80A" box="[259,421,1896,1920]" italics="true" pageId="10" pageNumber="71">Cyphomyrmex</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C684D5AAD32FF9DFE4BF8BFFD20F80A" box="[477,647,1896,1920]" class="Insecta" family="Formicidae" genus="Brachymyrmex" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFE4BF8BFFD20F80A" box="[477,647,1896,1920]" italics="true" pageId="10" pageNumber="71">Brachymyrmex</emphasis>
</taxonomicName>
sp.). Potential food items (arthropods) for 
<taxonomicName id="4C684D5AAD32FF9DFBE2F8BFFB7FF80A" box="[1140,1240,1896,1920]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFBE2F8BFFB7FF80A" box="[1140,1240,1896,1920]" italics="true" pageId="10" pageNumber="71">Tatuidris</emphasis>
</taxonomicName>
were taken from soil samples and Winkler samples (following 
<bibRefCitation id="EFF94B28AD32FF9DFD0BF85BFC17F82E" author="Silva" box="[669,944,1932,1956]" pageId="10" pageNumber="71" refString="Silva, R. R. &amp; Brandao, C. R. F. (2010) Morphological patterns and community organization in leaf-litter ant assemblages. Ecological Monographs, 80, 107 - 124" type="journal article" year="2010">Silva and Brandão 2010</bibRefCitation>
) collected at the site where 
<taxonomicName id="4C684D5AAD32FF9DFB78F85BFAF5F82E" box="[1262,1362,1932,1956]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD32FF9DFB78F85BFAF5F82E" box="[1262,1362,1932,1956]" italics="true" pageId="10" pageNumber="71">Tatuidris</emphasis>
</taxonomicName>
was 
<emphasis id="B91CEACBAD32FF9DFA18F85AFF7FF842" italics="true" pageId="10" pageNumber="71">a priori</emphasis>
determined to be present.
</paragraph>
<caption id="DF176651AD33FF9CFF01FA18FD4BF9CB" httpUri="https://zenodo.org/record/282485/files/figure.png" pageId="11" pageNumber="72" targetBox="[151,1435,183,1475]" targetPageId="11">
<paragraph id="8BD736D9AD33FF9CFF01FA18FD4BF9CB" blockId="11.[151,1436,1487,1601]" pageId="11" pageNumber="72">
<emphasis id="B91CEACBAD33FF9CFF01FA18FEB3FA6E" bold="true" box="[151,276,1487,1510]" pageId="11" pageNumber="72">FIGURE 6.</emphasis>
<taxonomicName id="4C684D5AAD33FF9CFE8AFA18FE65FA6F" box="[284,450,1487,1509]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFE8AFA18FE65FA6F" box="[284,450,1487,1509]" italics="true" pageId="11" pageNumber="72">Tatuidris tatusia</emphasis>
</taxonomicName>
and specimens showing pilosity pattern A. A–C) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD33FF9CFAABFA07FA3BFA6F" box="[1341,1436,1488,1509]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFAABFA07FA3BFA6F" box="[1341,1436,1488,1509]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
from Matagalpa, Nicaragua; D–F) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD33FF9CFC17FA38FC46F98E" box="[897,993,1519,1540]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFC17FA38FC46F98E" box="[897,993,1519,1540]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
from Otongachi, Ecuador; G–I) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD33FF9CFE3DF9D9FDACF9A9" box="[427,523,1550,1571]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFE3DF9D9FDACF9A9" box="[427,523,1550,1571]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
from Maquipucuna, Ecuador; J–L) Full-face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD33FF9CFA94F9D9FAC5F9A9" box="[1282,1378,1550,1571]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFA94F9D9FAC5F9A9" box="[1282,1378,1550,1571]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
from Cuzco, Peru. Images A–C and J–L courtesy of antweb.org.
</paragraph>
</caption>
<paragraph id="8BD736D9AD33FF9CFF51F9B9FC94F82C" blockId="11.[151,1437,1646,2030]" pageId="11" pageNumber="72">
Further observations by T. Delsinne suggest that 
<taxonomicName id="4C684D5AAD33FF9CFD6CF9B8FCC4F90C" box="[762,867,1647,1670]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFD6CF9B8FCC4F90C" box="[762,867,1647,1670]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
may be a sit-and-wait predator, as “both workers and gynes moved very slowly and were very clumsy. They often remained motionless during several tens of seconds or even several minutes when disturbed (either by my handling or by the contact with another arthropod). It is difficult to see them as powerful predators!” (pers. comm.). These observations were mainly performed at night, suggesting that 
<taxonomicName id="4C684D5AAD33FF9CFE18F928FE53F89C" box="[398,500,1791,1814]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFE18F928FE53F89C" box="[398,500,1791,1814]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
may be nocturnal, a hypothesis also supported by collection patterns. For example, in the Río Toachi forest of 
<collectingCountry id="F37F7649AD33FF9CFE76F8F5FD9AF8B0" box="[480,573,1826,1850]" name="Ecuador" pageId="11" pageNumber="72">Ecuador</collectingCountry>
<taxonomicName id="4C684D5AAD33FF9CFDDFF8F4FD12F8B0" box="[585,693,1827,1850]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="72" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD33FF9CFDDFF8F4FD12F8B0" box="[585,693,1827,1850]" italics="true" pageId="11" pageNumber="72">T. tatusia</emphasis>
</taxonomicName>
specimens tend to fall in pitfall traps, instead of Winkler sacs (Donoso and Ramón 2009). Because pitfall traps usually work 24-h, but Winkler sacs generally uses litter sifted during the day, then ants with nocturnal habits may be underrepresented in Winkler samples. The small eyes of 
<emphasis id="B91CEACBAD33FF9CFF01F859FF56F82C" box="[151,241,1934,1958]" italics="true" pageId="11" pageNumber="72">Tatudris</emphasis>
species provide further support for this hypothesis.
</paragraph>
<paragraph id="8BD736D9AD33FF9BFF51F865FB78FE0D" blockId="11.[151,1437,1646,2030]" lastBlockId="12.[151,1436,151,392]" lastPageId="12" lastPageNumber="73" pageId="11" pageNumber="72">
<emphasis id="B91CEACBAD33FF9CFF51F865FEAAF840" box="[199,269,1970,1994]" italics="true" pageId="11" pageNumber="72">Other.</emphasis>
The relative position of eyes is highly variable within the species. For example, eye location ranges from being completely within the antennal scrobes to completely outside the scrobes (
<figureCitation id="13532A5CAD33FF9CFBE5F801FB75F864" box="[1139,1234,2006,2030]" captionStart="FIGURE 1" captionStartId="1.[151,250,1499,1522]" captionTargetBox="[151,1436,555,1474]" captionTargetId="figure@1.[151,1436,555,1475]" captionTargetPageId="1" captionText="FIGURE 1. SEM images of Tatuidris tatusia external morphology. A) Head in (partial) full-face view; B) lateral view of the body; C) mandibular setae; D) close-up of mandible setae." httpUri="https://zenodo.org/record/282480/files/figure.png" pageId="11" pageNumber="72">Figure 1</figureCitation>
b). In some cases (specimen J.Longino#2088-S) the eye itself is located outside the antennal scrobe, but the eye’s fossa is well marked and confluent with the antennal scrobe. In most specimens, the antennal carina bifurcates from the antennal scrobes and lies straight above the eyes. However, in specimens from 
<collectingCountry id="F37F7649AD34FF9BFC31FF08FBBDFF7D" box="[935,1050,223,247]" name="Nicaragua" pageId="12" pageNumber="73">Nicaragua</collectingCountry>
(MGB#1179 colony collection), a strongly impressed antennal carina is present. In these specimens about 40% of the eye’s area lies within the antennal scrobes. In the gyne, only ~1/6 of the eye lies within the antennal scrobes. A depression sometimes forms in the integument in the sides of the propodeum, below the propodeal spiracle and above the metapleural gland. The depth of this depression varies among specimens and tends to be deepest in larger specimens.
</paragraph>
<caption id="DF176651AD34FF9BFF01FA46FB8AFA61" httpUri="https://zenodo.org/record/282486/files/figure.png" pageId="12" pageNumber="73" targetBox="[151,1435,437,1404]" targetPageId="12">
<paragraph id="8BD736D9AD34FF9BFF01FA46FB8AFA61" blockId="12.[151,1436,1425,1515]" pageId="12" pageNumber="73">
<emphasis id="B91CEACBAD34FF9BFF01FA46FEB5FA2D" bold="true" box="[151,274,1425,1448]" pageId="12" pageNumber="73">FIGURE 7.</emphasis>
<emphasis id="B91CEACBAD34FF9BFE8EFA45FE6DFA22" box="[280,458,1426,1448]" italics="true" pageId="12" pageNumber="73">
<taxonomicName id="4C684D5AAD34FF9BFE8EFA45FE19FA22" box="[280,446,1426,1448]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tatusia">Tatuidris tatusia</taxonomicName>
.
</emphasis>
Specimens showing pilosity pattern B. A–C) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD34FF9BFA91FA44FAC3FA22" box="[1287,1380,1427,1448]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD34FF9BFA91FA44FAC3FA22" box="[1287,1380,1427,1448]" italics="true" pageId="12" pageNumber="73">T. tatusia</emphasis>
</taxonomicName>
from Caves Branch, Belize; D–F) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD34FF9BFCA6FA63FC29FA43" box="[816,910,1460,1481]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD34FF9BFCA6FA63FC29FA43" box="[816,910,1460,1481]" italics="true" pageId="12" pageNumber="73">T. tatusia</emphasis>
</taxonomicName>
from Belmopan, Belize; G–I) Full-face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD34FF9BFEB8FA01FE2BFA61" box="[302,396,1494,1515]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD34FF9BFEB8FA01FE2BFA61" box="[302,396,1494,1515]" italics="true" pageId="12" pageNumber="73">T. tatusia</emphasis>
</taxonomicName>
from Maquipucuna, Ecuador. Images G–I courtesy of antweb.org.
</paragraph>
</caption>
<paragraph id="8BD736D9AD34FF9BFF51F9CDFB36F810" blockId="12.[151,1436,1562,2018]" pageId="12" pageNumber="73">
<emphasis id="B91CEACBAD34FF9BFF51F9CDFDB8F9B8" box="[199,543,1562,1586]" italics="true" pageId="12" pageNumber="73">Justification for the synonymy</emphasis>
. A comprehensive analysis of the description of 
<taxonomicName id="4C684D5AAD34FF9BFBF0F9CCFB69F9B8" box="[1126,1230,1562,1586]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFBF0F9CCFB69F9B8" box="[1126,1230,1562,1586]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
suggests that the morphological characters proposed by 
<bibRefCitation id="EFF94B28AD34FF9BFDDBF9E8FCF1F9DC" author="Lacau" box="[589,854,1598,1622]" pageId="12" pageNumber="73" refString="Lacau, S, Groc, S., Dejean, A. de Oliveira, M. L. &amp; Delabie J. C. H. (2012) Tatuidris kapasi sp. nov.: a new armadillo ant from French Guiana (Formicidae: Agroecomyrmecinae). Psyche, 2012, Article ID 926089, 6 pages, doi: 10.1155 / 2012 / 926089" type="book" year="2012">Lacau and Groc (2012)</bibRefCitation>
to separate 
<taxonomicName id="4C684D5AAD34FF9BFC75F9E8FBE1F9DC" box="[995,1094,1598,1622]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFC75F9E8FBE1F9DC" box="[995,1094,1598,1622]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C684D5AAD34FF9BFB05F9E8FB5CF9DC" box="[1171,1275,1599,1622]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD34FF9BFB05F9E8FB5CF9DC" box="[1171,1275,1599,1622]" italics="true" pageId="12" pageNumber="73">T. tatusia</emphasis>
</taxonomicName>
lie within the continuous variability encountered in 
<taxonomicName id="4C684D5AAD34FF9BFDA7F9B5FD35F9F0" box="[561,658,1634,1658]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD34FF9BFDA7F9B5FD35F9F0" box="[561,658,1634,1658]" italics="true" pageId="12" pageNumber="73">Tatuidris</emphasis>
</taxonomicName>
specimens across the Neotropics. Two considerations were taken. First, the description of 
<taxonomicName id="4C684D5AAD34FF9BFEF6F950FE64F914" box="[352,451,1670,1694]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFEF6F950FE64F914" box="[352,451,1670,1694]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
relies on a relatively large specimen where sculpture, body profile and body proportions are expected to vary the most. Diagnostic characters of 
<taxonomicName id="4C684D5AAD34FF9BFD60F97CFCFFF948" box="[758,856,1706,1730]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFD60F97CFCFFF948" box="[758,856,1706,1730]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
(i.e. occipital border more concave, clypeus with the free margin nearly straight and laterally concave, dorsum of mesosoma mostly sculptured with concentric rugulae and carinulae, and dorsum of the petiolar and postpetiolar nodes with superficial concentric rugulae and carinulae) are all present to some degree in larger specimens among the collections reviewed in the present study (
<figureCitation id="13532A5CAD34FF9BFA92F8C1FA37F8A4" box="[1284,1424,1814,1838]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="11.[151,250,1487,1510]" captionStartId-1="12.[151,250,1425,1448]" captionStartId-2="13.[151,250,849,872]" captionStartId-3="13.[151,250,1937,1960]" captionTargetBox-0="[151,1435,183,1475]" captionTargetBox-1="[151,1435,437,1404]" captionTargetBox-2="[151,1435,337,828]" captionTargetBox-3="[151,1435,952,1916]" captionTargetId-0="figure@11.[151,1436,183,1476]" captionTargetId-1="figure@12.[151,1436,437,1405]" captionTargetId-2="figure@13.[151,1436,337,829]" captionTargetId-3="figure@13.[151,1436,952,1916]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="13" captionText-0="FIGURE 6. Tatuidris tatusia and specimens showing pilosity pattern A. A – C) Full face, lateral and dorsal view of T. tatusia from Matagalpa, Nicaragua; D – F) Full face, lateral and dorsal view of T. tatusia from Otongachi, Ecuador; G – I) Full face, lateral and dorsal view of T. tatusia from Maquipucuna, Ecuador; J – L) Full-face, lateral and dorsal view of T. tatusia from Cuzco, Peru. Images A – C and J – L courtesy of antweb. org." captionText-1="FIGURE 7. Tatuidris tatusia. Specimens showing pilosity pattern B. A – C) Full face, lateral and dorsal view of T. tatusia from Caves Branch, Belize; D – F) Full face, lateral and dorsal view of T. tatusia from Belmopan, Belize; G – I) Full-face, lateral and dorsal view of T. tatusia from Maquipucuna, Ecuador. Images G – I courtesy of antweb. org." captionText-2="FIGURE 8. Tatuidris tatusia. Specimens showing pilosity pattern C. A) Full face and B) lateral view of T. tatusia from Puntarenas, Costa Rica." captionText-3="FIGURE 9. Tatuidris tatusia. Specimens showing pilosity pattern D. A – C) Full face, lateral and dorsal view of T. tatusia from Los Tuxtlas, Mexico; D – F) Full face, lateral and dorsal view of T. tatusia from Chiapas, Mexico; G – I) Full face, lateral and dorsal view of T. tatusia from Magdalena, Colombia." httpUri-0="https://zenodo.org/record/282485/files/figure.png" httpUri-1="https://zenodo.org/record/282486/files/figure.png" httpUri-2="https://zenodo.org/record/282487/files/figure.png" httpUri-3="https://zenodo.org/record/282488/files/figure.png" pageId="12" pageNumber="73">Figures 6–9</figureCitation>
). Another diagnostic character of 
<taxonomicName id="4C684D5AAD34FF9BFD91F8ECFDC9F8D8" box="[519,622,1850,1874]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFD91F8ECFDC9F8D8" box="[519,622,1850,1874]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
was denser pilosity without long setae (similar to Morph D), but setal patterns did not correlate with any morphology- or molecular-based grouping in this study. Second, the 
<typeStatus id="54D3887BAD34FF9BFB6FF888FA8FF8FD" box="[1273,1320,1887,1911]" pageId="12" pageNumber="73">type</typeStatus>
locality of 
<taxonomicName id="4C684D5AAD34FF9BFF01F854FF5EF810" box="[151,249,1922,1946]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="kapasi">
<emphasis id="B91CEACBAD34FF9BFF01F854FF5EF810" box="[151,249,1922,1946]" italics="true" pageId="12" pageNumber="73">T. kapasi</emphasis>
</taxonomicName>
lies at a range limit of the genus, where morphological extremes might be expected.
</paragraph>
</subSubSection>
<subSubSection id="C3726552AD34FF87FF51F871FBEBFE45" lastPageId="16" lastPageNumber="77" pageId="12" pageNumber="73" type="description">
<paragraph id="8BD736D9AD34FF9AFF51F871FDCCFE96" blockId="12.[151,1436,1562,2018]" lastBlockId="13.[151,1436,151,284]" lastPageId="13" lastPageNumber="74" pageId="12" pageNumber="73">
<emphasis id="B91CEACBAD34FF9BFF51F871FD0BF834" box="[199,684,1958,1982]" italics="true" pageId="12" pageNumber="73">Worker measurements (in mm) and indices:</emphasis>
[average (min–max) of 10– 
<specimenCount id="9D6EFD50AD34FF9BFC75F871FC58F834" box="[995,1023,1958,1982]" pageId="12" pageNumber="73" type="generic">58</specimenCount>
specimens]: AScL 0.46 (0.31, 0.67); AScW 0.24 (0.18, 0.36); CIx 129.03 (117.07, 137.93); EL 0.05 (0.03, 0.08); FL 0.43 (0.31, 0.70); FW 0.11 (0.08, 0.17); HL 0.59 (0.43, 0.88); HW 0.76 (0.56, 1.10); IAD 0.36 (0.25, 0.54); PL 0.16 (0.10, 0.24); PpL 0.16 (0.10, 0.25); PW 0.25 (0.18, 0.37); PpW 0.36 (0.26, 0.53); PPpIx 68.92 (58.14, 80.00); PrW 0.52 (0.38, 0.77); TiL 0.35 (0.27, 0.52); TiW 0.10 (0.06, 0.17); WbL 0.62 (0.45, 0.89); SL 0.42 (0.32, 0.51); SW 0.13 (0.11, 0.17); SI 329.82 (300.00, 360.00); REL 9.02 (5.41, 11.48).
</paragraph>
<caption id="DF176651AD35FF9AFF01FC86FE2FFC0C" httpUri="https://zenodo.org/record/282487/files/figure.png" pageId="13" pageNumber="74" targetBox="[151,1435,337,828]" targetPageId="13">
<paragraph id="8BD736D9AD35FF9AFF01FC86FE2FFC0C" blockId="13.[151,1436,849,902]" pageId="13" pageNumber="74">
<emphasis id="B91CEACBAD35FF9AFF01FC86FEB1FCED" bold="true" box="[151,278,849,872]" pageId="13" pageNumber="74">FIGURE 8.</emphasis>
<emphasis id="B91CEACBAD35FF9AFEB6FC85FE7EFCE2" box="[288,473,850,872]" italics="true" pageId="13" pageNumber="74">
<taxonomicName id="4C684D5AAD35FF9AFEB6FC85FE6EFCE2" box="[288,457,850,872]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">Tatuidris tatusia</taxonomicName>
.
</emphasis>
Specimens showing pilosity pattern C. A) Full face and B) lateral view of 
<taxonomicName id="4C684D5AAD35FF9AFA96FC84FAC6FCE2" box="[1280,1377,851,872]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD35FF9AFA96FC84FAC6FCE2" box="[1280,1377,851,872]" italics="true" pageId="13" pageNumber="74">T. tatusia</emphasis>
</taxonomicName>
from Puntarenas, Costa Rica.
</paragraph>
</caption>
<caption id="DF176651AD35FF9AFF01F846FD15F866" httpUri="https://zenodo.org/record/282488/files/figure.png" pageId="13" pageNumber="74" targetBox="[151,1435,952,1916]" targetPageId="13">
<paragraph id="8BD736D9AD35FF9AFF01F846FD15F866" blockId="13.[151,1437,1937,2028]" pageId="13" pageNumber="74">
<emphasis id="B91CEACBAD35FF9AFF01F846FEB5F82D" bold="true" box="[151,274,1937,1960]" pageId="13" pageNumber="74">FIGURE 9.</emphasis>
<emphasis id="B91CEACBAD35FF9AFE8EF845FE6DF822" box="[280,458,1938,1960]" italics="true" pageId="13" pageNumber="74">
<taxonomicName id="4C684D5AAD35FF9AFE8EF845FE19F822" box="[280,446,1938,1960]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">Tatuidris tatusia</taxonomicName>
.
</emphasis>
Specimens showing pilosity pattern D. A–C) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD35FF9AFA91F844FAC2F822" box="[1287,1381,1939,1960]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD35FF9AFA91F844FAC2F822" box="[1287,1381,1939,1960]" italics="true" pageId="13" pageNumber="74">T. tatusia</emphasis>
</taxonomicName>
from Los Tuxtlas, Mexico; D–F) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD35FF9AFCA4F863FC36F843" box="[818,913,1972,1993]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD35FF9AFCA4F863FC36F843" box="[818,913,1972,1993]" italics="true" pageId="13" pageNumber="74">T. tatusia</emphasis>
</taxonomicName>
from Chiapas, Mexico; G–I) Full face, lateral and dorsal view of 
<taxonomicName id="4C684D5AAD35FF9AFEB8F800FE2BF866" box="[302,396,2007,2028]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="74" phylum="Arthropoda" rank="species" species="tatusia">
<emphasis id="B91CEACBAD35FF9AFEB8F800FE2BF866" box="[302,396,2007,2028]" italics="true" pageId="13" pageNumber="74">T. tatusia</emphasis>
</taxonomicName>
from Magdalena, Colombia.
</paragraph>
</caption>
<caption id="DF176651AD36FF99FF01FBA4FD99FAC8" httpUri="https://zenodo.org/record/282489/files/figure.png" pageId="14" pageNumber="75" targetBox="[242,1345,195,1073]" targetPageId="14">
<paragraph id="8BD736D9AD36FF99FF01FBA4FD99FAC8" blockId="14.[151,1437,1139,1346]" pageId="14" pageNumber="75">
<emphasis id="B91CEACBAD36FF99FF01FBA4FE86FB03" bold="true" box="[151,289,1139,1162]" pageId="14" pageNumber="75">FIGURE 10.</emphasis>
Neighbor joining tree based on K2P distances for 28 
<taxonomicName id="4C684D5AAD36FF99FCC7FBA3FC0CFB00" box="[849,939,1140,1162]" class="Insecta" family="Formicidae" genus="Tatuidris" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD36FF99FCC7FBA3FC0CFB00" box="[849,939,1140,1162]" italics="true" pageId="14" pageNumber="75">Tatuidris</emphasis>
</taxonomicName>
specimens and four other ant taxa (
<taxonomicName id="4C684D5AAD36FF99FAB4FBA2FF45FB23" class="Insecta" family="Formicidae" genus="Paraponera" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="B91CEACBAD36FF99FAB4FBA2FF45FB23" italics="true" pageId="14" pageNumber="75">Paraponera clavata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C684D5AAD36FF99FF65FB44FE32FB23" box="[243,405,1171,1193]" class="Insecta" family="Formicidae" genus="Probolomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEACBAD36FF99FF65FB44FE32FB23" box="[243,405,1171,1193]" italics="true" pageId="14" pageNumber="75">Probolomyrmex</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C684D5AAD36FF99FE59FB43FD30FB23" box="[463,663,1172,1193]" class="Insecta" family="Formicidae" genus="Proceratium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="avium">
<emphasis id="B91CEACBAD36FF99FE59FB43FD30FB23" box="[463,663,1172,1193]" italics="true" pageId="14" pageNumber="75">Proceratium avium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C684D5AAD36FF99FD44FB44FC3EFB23" box="[722,921,1171,1193]" class="Insecta" family="Formicidae" genus="Strumigenys" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="coveri">
<emphasis id="B91CEACBAD36FF99FD44FB44FC3EFB23" box="[722,921,1171,1193]" italics="true" pageId="14" pageNumber="75">Strumigenys coveri</emphasis>
</taxonomicName>
) as outgroups. Labels consist of countries, first division, localities, and specimen IDs. Specimens with no color present a pilosity pattern “A”, similar to the type series. Specimens with pilosity pattern “B” are highlighted in light blue. Specimens with pilosity pattern “D” are highlighted in yellow. No specimen with pilosity pattern “C” was available for sequencing. Asterisks above nodes represent nodes with&gt;70% bootstrap support (999 repetitions). The tree is drawn to scale, with branch lengths in the same units as those of the genetic divergence inferred using the K2P model.
</paragraph>
</caption>
<paragraph id="8BD736D9AD36FF99FF51FABDFEFEFA43" blockId="14.[151,1436,1385,2022]" pageId="14" pageNumber="75">
<emphasis id="B91CEACBAD36FF99FF51FABDFD01FA0B" box="[199,678,1385,1409]" italics="true" pageId="14" pageNumber="75">Gyne measurements (in mm) and indices:</emphasis>
(n=1) AScL 0.47; AScW 0.15; EL 0.20; FFS 0.09; FL 0.77; FW 0.20; HL 0.88; HW 1.28; IAD 0.56; Pl 0.22; PPL 0.28; PPW 0.72; PW 0.45; TL 0.69; TW 0.20; WingL 4.60; WbL 1.53; REL 22.63.
</paragraph>
<paragraph id="8BD736D9AD36FF99FF51FA02FDDDF99B" blockId="14.[151,1436,1385,2022]" pageId="14" pageNumber="75">
<emphasis id="B91CEACBAD36FF99FF51FA02FD3BFA67" box="[199,668,1493,1518]" italics="true" pageId="14" pageNumber="75">Male measurements (in mm) and indices:</emphasis>
(n=1) EL 0.32; FFS 0.13; HL 0.66; HW 0.88; IAD 0.21; ScL 0.11; WingL 3.6; WbL 1.22; FL 0.9; REL 48.44.
</paragraph>
<paragraph id="8BD736D9AD36FF99FF51F9C9FB3DF9D3" blockId="14.[151,1436,1385,2022]" pageId="14" pageNumber="75">
<emphasis id="B91CEACBAD36FF99FF51F9C9FE1FF9BF" box="[199,440,1565,1589]" italics="true" pageId="14" pageNumber="75">Specimens examined:</emphasis>
Elements in this list follow this order: COUNTRY, 
<emphasis id="B91CEACBAD36FF99FC61F9C9FB1FF9BC" box="[1015,1208,1566,1590]" italics="true" pageId="14" pageNumber="75">State or Province</emphasis>
, Locality, elevation, coordinates, collector (collection code, habitat, method) [specimen code], [REPOSITORY].
</paragraph>
<paragraph id="8BD736D9AD36FF87FF51F9B1FBEBFE45" blockId="14.[151,1436,1385,2022]" lastBlockId="16.[151,1436,151,463]" lastPageId="16" lastPageNumber="77" pageId="14" pageNumber="75">
<collectingCountry id="F37F7649AD36FF99FF51F9B1FE8DF9F7" box="[199,298,1638,1661]" name="Belize" pageId="14" pageNumber="75">BELIZE</collectingCountry>
, 2.5 millas S Belmopan, S&amp;J Peck (B-242, Limestone forest, ex: Berlese) [MCZC]. Caves Branch, S&amp;J Peck (B-248, Hi Canopi Forest, ex: Berlese [MCZC]. 
<collectingCountry id="F37F7649AD36FF99FCFAF95DFC73F928" box="[876,980,1674,1698]" name="Brazil" pageId="14" pageNumber="75">BRAZIL</collectingCountry>
, 
<emphasis id="B91CEACBAD36FF99FC70F95EFBFEF92B" box="[998,1113,1673,1697]" italics="true" pageId="14" pageNumber="75">Amazonas</emphasis>
, Manaus, Universidade do Amazonas, 
<geoCoordinate id="EE5C501EAD36FF99FE8BF97AFE39F94F" box="[285,414,1709,1733]" direction="south" orientation="latitude" pageId="14" pageNumber="75" precision="15" value="-3.0933332">03°05'36''S</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD36FF99FE3DF97AFD93F94F" box="[427,564,1709,1733]" direction="west" orientation="longitude" pageId="14" pageNumber="75" precision="15" value="-59.964447">59°57'52''W</geoCoordinate>
, E Pereira Franken (Terra Firme: PLATO, ex: Pitfall) [INPA]. 
<collectingCountry id="F37F7649AD36FF99FA96F97AFA3FF94F" box="[1280,1432,1709,1733]" name="Colombia" pageId="14" pageNumber="75">COLOMBIA</collectingCountry>
, 
<emphasis id="B91CEACBAD36FF99FF01F906FEB0F963" box="[151,279,1745,1769]" italics="true" pageId="14" pageNumber="75">Magdalena</emphasis>
, El Campano, 
<quantity id="4C909B3CAD36FF99FE54F906FDB5F963" box="[450,530,1745,1769]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="75" unit="m" value="1300.0">1300m</quantity>
, 
<geoCoordinate id="EE5C501EAD36FF99FD89F906FDD9F963" box="[543,638,1745,1769]" direction="north" orientation="latitude" pageId="14" pageNumber="75" precision="925" value="11.116667">11°07'N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD36FF99FD1DF906FD49F963" box="[651,750,1745,1769]" direction="west" orientation="longitude" pageId="14" pageNumber="75" precision="925" value="-74.1">74°06'W</geoCoordinate>
, Phillip Ward (#7891-9, montane rainforest, ex: Sifted litter, leaf mold, rotten log) [QCAZ]. 
<collectingCountry id="F37F7649AD36FF99FD9CF922FD13F887" box="[522,692,1781,1805]" name="Costa Rica" pageId="14" pageNumber="75">COSTA RICA</collectingCountry>
, Rincon de la Vieja, Pailas, 
<quantity id="4C909B3CAD36FF99FB94F922FBF5F887" box="[1026,1106,1781,1805]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="75" unit="m" value="1000.0">1000m</quantity>
, 10.7887, –85.349, A Smith (10COSTA-0629 and 10COSTA-0630, Primary Forest) [AlexSmith]. 
<emphasis id="B91CEACBAD36FF99FC23F8CEFBB3F8BB" box="[949,1044,1817,1841]" italics="true" pageId="14" pageNumber="75">Alajuela</emphasis>
, Casa Eladio, Rio Peñas Blancas, 
<quantity id="4C909B3CAD36FF99FF01F8EAFF7EF8DF" box="[151,217,1853,1877]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="75" unit="m" value="800.0">800m</quantity>
, 
<geoCoordinate id="EE5C501EAD36FF99FF63F8EAFECBF8DF" box="[245,364,1853,1877]" direction="north" orientation="latitude" pageId="14" pageNumber="75" precision="925" value="10.316667">10°19' N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD36FF99FE1FF8EAFDA3F8DF" box="[393,516,1853,1877]" direction="west" orientation="longitude" pageId="14" pageNumber="75" precision="925" value="-84.71667">84°43' W</geoCoordinate>
, J Longino (Wet forest, ex: Sifted leaf litter) [INBIOCRI002280066, INBIOCRI002280068 and INBIOCRI002280065] [JTLC]. 
<emphasis id="B91CEACBAD36FF99FCD7F8B6FC07F8F3" box="[833,928,1889,1913]" italics="true" pageId="14" pageNumber="75">Alajuela</emphasis>
, El Aleman, Rio Peñas Blancas, 6,5 Km E Monteverde, 
<quantity id="4C909B3CAD36FF99FEABF852FE27F817" box="[317,384,1925,1949]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="14" pageNumber="75" unit="m" value="940.0">940m</quantity>
, 
<geoCoordinate id="EE5C501EAD36FF99FE0AF852FDB4F817" box="[412,531,1925,1949]" direction="north" orientation="latitude" pageId="14" pageNumber="75" precision="925" value="10.3">10°18' N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD36FF99FDA6F852FD0BF817" box="[560,684,1925,1949]" direction="west" orientation="longitude" pageId="14" pageNumber="75" precision="925" value="-84.75">84°45' W</geoCoordinate>
, J Longino (#0861-s, Wet forest, ex: Sifted leaf litter) [INBIOCRI002280059] [JTLC]. 
<emphasis id="B91CEACBAD36FF99FD83F87EFDD7F84B" box="[533,624,1961,1985]" italics="true" pageId="14" pageNumber="75">Heredia</emphasis>
, 
<quantity id="4C909B3CAD36FF99FD14F87EFD65F84B" box="[642,706,1961,1985]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="75" unit="km" value="16.0">16km</quantity>
SSE La Virgen, 
<quantity id="4C909B3CAD36FF99FC04F87EFC46F84B" box="[914,993,1961,1985]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="75" unit="m" value="1100.0">1100m</quantity>
, 
<geoCoordinate id="EE5C501EAD36FF99FC67F87EFBF6F84B" box="[1009,1105,1961,1985]" direction="north" orientation="latitude" pageId="14" pageNumber="75" precision="925" value="10.266666">10°16'N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD36FF99FBF7F87EFB64F84B" box="[1121,1219,1961,1985]" direction="west" orientation="longitude" pageId="14" pageNumber="75" precision="925" value="-84.083336">84°05'W</geoCoordinate>
[INB0003214087, INB0003214014, INB0003214104, INB0003214131, INB0003214232 and INB0003212497] [INBC]. 
<emphasis id="B91CEACBAD36FF99FA80F819FAD7F86C" box="[1302,1392,1998,2022]" italics="true" pageId="14" pageNumber="75">Heredia</emphasis>
, La Selva Biological Station-1, 50m, M Molina and H Garcia (AMI-1-W-006-05) [JTLC]. 
<emphasis id="B91CEACBAD37FF98FB29FBB9FABDFB0C" box="[1215,1306,1134,1158]" italics="true" pageId="15" pageNumber="76">Heredia</emphasis>
, La Selva Biological Station-2, 50m, 10.433, –84.0167, J Longino (All ants from Louisa Stark (OTS 89-1 student) study, ex: Litter and ground-nesting) [INBIOCRI001281194 and INBIOCRI001281181] [JTLC]. 
<emphasis id="B91CEACBAD37FF98FB2FFB61FAB3FB44" box="[1209,1300,1206,1230]" italics="true" pageId="15" pageNumber="76">Heredia</emphasis>
, La Selva Biological Station-2, 
<geoCoordinate id="EE5C501EAD37FF98FE11FB0DFE49FB78" box="[391,494,1242,1266]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="925" value="9.15">09°09' N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FE6CFB0DFDD4FB78" box="[506,627,1242,1266]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="925" value="-79.85">079°51' W</geoCoordinate>
, Michael Kaspari (45196.01 and 45000.6) [MEK]. 
<emphasis id="B91CEACBAD37FF98FB22FB0CFA93FB78" box="[1204,1332,1243,1266]" italics="true" pageId="15" pageNumber="76">Puntarenas</emphasis>
, Rio San Luis, 
<quantity id="4C909B3CAD37FF98FF41FB29FEBDFA9C" box="[215,282,1278,1302]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="15" pageNumber="76" unit="m" value="850.0">850m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FEB0FB29FE2AFA9C" box="[294,397,1278,1302]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="925" value="10.283334">10°17' N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FE0CFB29FDA2FA9C" box="[410,517,1278,1302]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="925" value="-84.8">84°48' W</geoCoordinate>
, J. Longino (Moist forest, ex: Sifted leaf litter on ground) [INBIOCRI002280062 and INBIOCRI002280063] [JTLC]. 
<collectingCountry id="F37F7649AD37FF98FDADFAF5FD61FAB0" box="[571,710,1314,1338]" name="Ecuador" pageId="15" pageNumber="76">ECUADOR</collectingCountry>
, 
<emphasis id="B91CEACBAD37FF98FD42FAF4FC9DFAB0" box="[724,826,1315,1338]" italics="true" pageId="15" pageNumber="76">Cotopaxi</emphasis>
, 
<quantity id="4C909B3CAD37FF98FCDDFAF5FC32FAB0" box="[843,917,1314,1338]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="76" unit="km" value="19.0">19 km</quantity>
ENE La Maná, 
<quantity id="4C909B3CAD37FF98FBCEFAF5FB0FFAB0" box="[1112,1192,1314,1338]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="76" unit="m" value="1100.0">1100m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FB20FAF5FABCFAB0" box="[1206,1307,1314,1338]" direction="south" orientation="latitude" pageId="15" pageNumber="76" precision="925" value="-0.8833333">00°53' S</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FABCFAF5FA3FFAB0" box="[1322,1432,1314,1338]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="925" value="-79.05">79°03' W</geoCoordinate>
, Phillip Ward (#11418-6, Second-Grown Rainforest, ex: Sifted leaf litter and logs) [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FB75FA91FAF2FAD4" box="[1251,1365,1350,1374]" italics="true" pageId="15" pageNumber="76">Pichincha</emphasis>
, R.B. Maquipucuna, 
<quantity id="4C909B3CAD37FF98FED6FABDFE37FA08" box="[320,400,1386,1410]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="76" unit="m" value="1200.0">1200m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FE0BFABDFD84FA08" box="[413,547,1386,1410]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="15" value="0.11666667">00°07'00''N</geoCoordinate>
; 78°38'06W, R Anderson (#99-208-6 and #99-208, Montane evergreen forest) [CASENT0423526 and CASENT0001968] [CASC]. 
<emphasis id="B91CEACBAD37FF98FD65FA59FCC2FA2C" box="[755,869,1422,1446]" italics="true" pageId="15" pageNumber="76">Pichincha</emphasis>
, Unión del Toachi-Otongachi, 
<quantity id="4C909B3CAD37FF98FB50FA59FAAFFA2C" box="[1222,1288,1422,1446]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="15" pageNumber="76" unit="m" value="850.0">850m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FA83FA59FA30FA2C" box="[1301,1431,1422,1446]" direction="south" orientation="latitude" pageId="15" pageNumber="76" precision="15" value="-0.35138887">00°21'05&quot;S</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FF01FA65FE86FA40" box="[151,289,1458,1482]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="15" value="-78.952774">78°57'10&quot;W</geoCoordinate>
, Donoso &amp; Vieira (Bosque Secundario, ex: Pitfall) [
<specimenCount id="9D6EFD50AD37FF98FCEBFA65FC59FA40" box="[893,1022,1458,1482]" pageId="15" pageNumber="76" type="worker">18 workers</specimenCount>
and 1 gyne, MEKOU12079-84 and MEKOU12103-4] [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FE7EFA00FDE7FA64" box="[488,576,1495,1518]" italics="true" pageId="15" pageNumber="76">Zamora</emphasis>
, Bombuscaro Numex, 
<quantity id="4C909B3CAD37FF98FCCBFA01FC07FA64" box="[861,928,1494,1518]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="76" unit="m" value="950.0">950m</quantity>
, – 
<geoCoordinate id="EE5C501EAD37FF98FC57FA01FBEDFA64" box="[961,1098,1494,1518]" direction="south" orientation="latitude" pageId="15" pageNumber="76" precision="1" value="-4.114972">4.114972 S</geoCoordinate>
; – 
<geoCoordinate id="EE5C501EAD37FF98FBFBFA01FAA7FA64" box="[1133,1280,1494,1518]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="1" value="-78.96794">78.96794 W</geoCoordinate>
, M Leponce (SPM_33796, Evergreen premontane rainforest, ex: Winkler sample) [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FBCFFA2CFB15F998" box="[1113,1202,1531,1554]" italics="true" pageId="15" pageNumber="76">Zamora</emphasis>
, Copalinga Private Reserve, 
<quantity id="4C909B3CAD37FF98FE91F9C9FEF0F9BC" box="[263,343,1566,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="76" unit="m" value="1000.0">1000m</quantity>
, – 
<geoCoordinate id="EE5C501EAD37FF98FEEFF9C9FDB7F9BC" box="[377,528,1566,1590]" direction="south" orientation="latitude" pageId="15" pageNumber="76" precision="1" value="-4.0912223">4.0912222 S</geoCoordinate>
; – 
<geoCoordinate id="EE5C501EAD37FF98FDA5F9C9FD62F9BC" box="[563,709,1566,1590]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="1" value="-78.96069">78.96069 W</geoCoordinate>
, T Delsinne and T. Arias-Penna (SPM_4130219, Secondary evergreen premontane rainforest, ex: Winkler sample) [QCAZ]. 
<collectingCountry id="F37F7649AD37FF98FCFEF994FB84F9D0" box="[872,1059,1602,1626]" name="El Salvador" pageId="15" pageNumber="76">EL SALVADOR</collectingCountry>
[
<typeStatus id="54D3887BAD37FF98FBA7F994FB31F9D1" box="[1073,1174,1603,1627]" pageId="15" pageNumber="76" type="paratype">Paratype</typeStatus>
and 
<typeStatus id="54D3887BAD37FF98FB5AF995FA9DF9D0" box="[1228,1338,1602,1626]" pageId="15" pageNumber="76" type="holotype">Holotype</typeStatus>
], 
<quantity id="4C909B3CAD37FF98FADAF995FA25F9D0" box="[1356,1410,1602,1626]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="15" pageNumber="76" unit="mi" value="2.0">2 mi</quantity>
. S Quetzaltepec, ME Erwin [MCZ 
<typeStatus id="54D3887BAD37FF98FD97F9B0FDC4F9F5" box="[513,611,1639,1663]" pageId="15" pageNumber="76" type="paratype">Paratype</typeStatus>
31577] [MCZC]. 
<collectingCountry id="F37F7649AD37FF98FCA7F9B1FC45F9F4" box="[817,994,1638,1662]" name="Guatemala" pageId="15" pageNumber="76">GUATEMALA</collectingCountry>
, 
<emphasis id="B91CEACBAD37FF98FC78F9B0FB8AF9F4" box="[1006,1069,1639,1662]" italics="true" pageId="15" pageNumber="76">Peten</emphasis>
, Parq. Nac. Tikal, 
<quantity id="4C909B3CAD37FF98FA97F9B1FAE3F9F4" box="[1281,1348,1638,1662]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="76" unit="m" value="270.0">270m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FAD9F9B1FF09F928" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="55" value="17.232">17.232 N</geoCoordinate>
; – 
<geoCoordinate id="EE5C501EAD37FF98FF5AF95DFE9AF928" box="[204,317,1674,1698]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="55" value="-89.623">89.623 W</geoCoordinate>
(Tropical Moist Forest) [CASC]. 
<collectingCountry id="F37F7649AD37FF98FD5BF95DFCCBF928" box="[717,876,1674,1698]" name="Honduras" pageId="15" pageNumber="76">HONDURAS</collectingCountry>
, 
<emphasis id="B91CEACBAD37FF98FCECF95CFBA6F928" box="[890,1025,1675,1698]" italics="true" pageId="15" pageNumber="76">Comayagua</emphasis>
, PN Cerro Azul Meambar, 
<quantity id="4C909B3CAD37FF98FADEF95DFA32F928" box="[1352,1429,1674,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="15" pageNumber="76" unit="m" value="1120.0">1120m</quantity>
, 14.887136; –87.89974 (LLAMA#Wa-C- 
<date id="FFD61019AD37FF98FDF8F979FD1BF94C" box="[622,700,1710,1734]" pageId="15" pageNumber="76" value="1915-01-04">04-1-15</date>
, Ridgetop cloud forest, ex: Sifted leaf litter) [CASENT 0615399] [CASC]. 
<collectingCountry id="F37F7649AD37FF98FE93F905FED1F960" box="[261,374,1746,1770]" name="Mexico" pageId="15" pageNumber="76">MEXICO</collectingCountry>
, 
<emphasis id="B91CEACBAD37FF98FE10F905FE44F960" box="[390,483,1746,1770]" italics="true" pageId="15" pageNumber="76">Chiapas</emphasis>
, 
<quantity id="4C909B3CAD37FF98FE60F905FD9FF960" box="[502,568,1746,1770]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="15" pageNumber="76" unit="m" value="860.0">860m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FDDEF905FD10F960" box="[584,695,1746,1770]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="55" value="16.98">16.980 N</geoCoordinate>
; – 
<geoCoordinate id="EE5C501EAD37FF98FD40F905FCEEF960" box="[726,841,1746,1770]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="55" value="-91.586">91.586 W</geoCoordinate>
, MG Branstetter (#856, Mesophil forest) [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FF01F921FF53F884" box="[151,244,1782,1806]" italics="true" pageId="15" pageNumber="76">Chiapas</emphasis>
, 
<quantity id="4C909B3CAD37FF98FE95F921FEE3F884" box="[259,324,1782,1806]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="15" pageNumber="76" unit="mi" value="12.0">12 mi</quantity>
NW Ocozocoautla, 
<quantity id="4C909B3CAD37FF98FDBCF921FDCAF884" box="[554,621,1782,1806]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="76" unit="m" value="400.0">400m</quantity>
, A Newton (ex: Berlese) [MCZC]. 
<emphasis id="B91CEACBAD37FF98FC68F921FBFCF884" box="[1022,1115,1782,1806]" italics="true" pageId="15" pageNumber="76">Chiapas</emphasis>
, 6kmSW Ocosingo, 
<quantity id="4C909B3CAD37FF98FAD1F921FA30F884" box="[1351,1431,1782,1806]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="76" unit="m" value="1400.0">1400m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FF01F8CDFE88F8B8" box="[151,303,1818,1842]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="1" value="16.867222">16.867221 N</geoCoordinate>
; – 
<geoCoordinate id="EE5C501EAD37FF98FEDBF8CDFE50F8B8" box="[333,503,1818,1842]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="1" value="-92.07871">92.0787132 W</geoCoordinate>
, R Anderson (# 91–116, Forest litter, ex: Berlese) [CASENT 0603397] [INBC]. 
<emphasis id="B91CEACBAD37FF98FF01F8E9FF53F8DC" box="[151,244,1854,1878]" italics="true" pageId="15" pageNumber="76">Chiapas</emphasis>
, Lago Metzabok, 
<quantity id="4C909B3CAD37FF98FE41F8E9FDBEF8DC" box="[471,537,1854,1878]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="15" pageNumber="76" unit="m" value="575.0">575m</quantity>
, 17.124, –91.636, MG Branstetter (Lowland wet forest) [CASC]. 
<emphasis id="B91CEACBAD37FF98FAA8F8E8FA32F8DC" box="[1342,1429,1855,1878]" italics="true" pageId="15" pageNumber="76">Oaxaca</emphasis>
, Mirador Grande, 
<quantity id="4C909B3CAD37FF98FEF5F8B5FE01F8F0" box="[355,422,1890,1914]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="15" pageNumber="76" unit="m" value="990.0">990m</quantity>
, 17.89844, –96.36253, MG Branstetter (#1405) [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FBD8F8B5FB77F8F0" box="[1102,1232,1890,1914]" italics="true" pageId="15" pageNumber="76">Tamaulipas</emphasis>
, El Cielo, 
<quantity id="4C909B3CAD37FF98FAC3F8B5FA30F8F0" box="[1365,1431,1890,1914]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="15" pageNumber="76" unit="m" value="870.0">870m</quantity>
, 23.276, –99.276, MG Branstetter (#1465) [QCAZ]. 
<emphasis id="B91CEACBAD37FF98FD4AF850FC9BF814" box="[732,828,1927,1950]" italics="true" pageId="15" pageNumber="76">Ve r a c r u z</emphasis>
, Los Tuxtlas, Ejido-López Mateos, 
<quantity id="4C909B3CAD37FF98FB44F851FAA1F814" box="[1234,1286,1926,1950]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="76" unit="m" value="50.0">50m</quantity>
, 
<geoCoordinate id="EE5C501EAD37FF98FA84F851FA30F814" box="[1298,1431,1926,1950]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="15" value="18.415556">18°24'56''N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FF01F87DFE87F848" box="[151,288,1962,1986]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="15" value="-94.94806">94°56'53''W</geoCoordinate>
, P Rojas (LM7S-H1, LM42S-H4, LM42S-H5 and LM21A-H3, Selva alta perennifolia, ex: Winkler) [IEXM]. 
<emphasis id="B91CEACBAD37FF98FE97F818FEC5F86C" box="[257,354,1999,2022]" italics="true" pageId="15" pageNumber="76">Veracruz</emphasis>
, Los Tuxtlas, Ejido-López Mateos, Ejido 2, 50m, 
<geoCoordinate id="EE5C501EAD37FF98FC3EF819FB89F86C" box="[936,1070,1998,2022]" direction="north" orientation="latitude" pageId="15" pageNumber="76" precision="15" value="18.415556">18°24'56''N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD37FF98FBAAF819FB63F86C" box="[1084,1220,1998,2022]" direction="west" orientation="longitude" pageId="15" pageNumber="76" precision="15" value="-94.94806">94°56'53''W</geoCoordinate>
, P Rojas (LM42S- H3, Selva alta perennifolia, ex: Winkler) [IEXM]. 
<emphasis id="B91CEACBAD28FF87FC86FF4EFCD6FF3A" box="[784,881,153,176]" italics="true" pageId="16" pageNumber="77">Veracruz</emphasis>
, Los Tuxtlas, Volcán S. M. Pajapan, 
<quantity id="4C909B3CAD28FF87FAC3FF40FA3FFF25" box="[1365,1432,151,175]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="16" pageNumber="77" unit="m" value="510.0">510m</quantity>
, 
<geoCoordinate id="EE5C501EAD28FF87FF01FF6BFEBBFF5E" box="[151,284,188,212]" direction="north" orientation="latitude" pageId="16" pageNumber="77" precision="15" value="18.266666">18°16'00''N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD28FF87FEBDFF6BFE14FF5E" box="[299,435,188,212]" direction="west" orientation="longitude" pageId="16" pageNumber="77" precision="15" value="-94.78639">94°46'71''W</geoCoordinate>
, A Cartas (847d, Selva 
<taxonomicName id="4C684D5AAD28FF87FD5FFF6BFC8FFF5E" box="[713,808,188,212]" class="Insecta" family="Drosophilidae" genus="Drosophila" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="77" phylum="Arthropoda" rank="species" species="mediana">mediana</taxonomicName>
subperennifolia, ex: Berlese) [IEXM]. 
<collectingCountry id="F37F7649AD28FF87FB7BFF6BFA3FFF5E" box="[1261,1432,188,212]" name="Nicaragua" pageId="16" pageNumber="77">NICARAGUA</collectingCountry>
, 
<emphasis id="B91CEACBAD28FF87FF01FF37FEB5FF72" box="[151,274,224,248]" italics="true" pageId="16" pageNumber="77">Matagalpa</emphasis>
, RN El Musún, 
<quantity id="4C909B3CAD28FF87FE5EFF08FDA8FF7D" box="[456,527,223,247]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="77" unit="km" value="4.8">4.8km</quantity>
NNW Rio Blanco, 
<quantity id="4C909B3CAD28FF87FD7AFF08FC9CFF7D" box="[748,827,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="16" pageNumber="77" unit="m" value="1170.0">1170m</quantity>
, 
<geoCoordinate id="EE5C501EAD28FF87FCD0FF08FC1DFF7D" box="[838,954,223,247]" direction="north" orientation="latitude" pageId="16" pageNumber="77" precision="92" value="12.973333">12°58.4'N</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD28FF87FC50FF08FB90FF7D" box="[966,1079,223,247]" direction="west" orientation="longitude" pageId="16" pageNumber="77" precision="925" value="-85.23333">085°14'W</geoCoordinate>
, M.G.Branstetter (#1179, Mesic forest, ex: Sifted leaf litter) [QCAZ]. 
<collectingCountry id="F37F7649AD28FF87FDACFED3FD13FE96" box="[570,692,260,284]" name="Panama" pageId="16" pageNumber="77">PANAMA</collectingCountry>
, 
<emphasis id="B91CEACBAD28FF87FD28FED3FCBAFE96" box="[702,797,260,284]" italics="true" pageId="16" pageNumber="77">Chiriqui</emphasis>
, 20,4 Km North San Felix, R. Anderson (#17768_1, Wet mountain forest, ex: Litter sample) [WEMC]. 
<emphasis id="B91CEACBAD28FF87FD3CFEFFFCAEFECA" box="[682,777,296,320]" italics="true" pageId="16" pageNumber="77">Chiriqui</emphasis>
, Alto Lino, 
<quantity id="4C909B3CAD28FF87FC0CFEF0FC4CFEB5" box="[922,1003,295,319]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="77" unit="m" value="3800.0">3800m</quantity>
, HG Real [Male, CASENT 0102681] [CASC]. 
<emphasis id="B91CEACBAD28FF87FE97FE9BFEC7FEEE" box="[257,352,332,356]" italics="true" pageId="16" pageNumber="77">Chiriqui</emphasis>
, La 
<collectingCountry id="F37F7649AD28FF87FE02FE9AFE49FEEE" box="[404,494,333,356]" name="Wallis and Futuna" pageId="16" pageNumber="77">Fortuna</collectingCountry>
, Finca La 
<collectingCountry id="F37F7649AD28FF87FDF3FE9BFD08FEEE" box="[613,687,332,356]" name="Switzerland" pageId="16" pageNumber="77">Suisse</collectingCountry>
, 
<quantity id="4C909B3CAD28FF87FD2DFE9BFCACFEEE" box="[699,779,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="77" unit="m" value="1200.0">1200m</quantity>
, R. Anderson (#17838, #17787, #17788, #17789, #17790, #17838 and #17839 [WEMC]. 
<collectingCountry id="F37F7649AD28FF87FD91FEA7FDF6FE0D" box="[519,593,368,391]" name="Peru" pageId="16" pageNumber="77">PERU</collectingCountry>
, 
<emphasis id="B91CEACBAD28FF87FDF7FEA7FD00FE0D" box="[609,679,368,391]" italics="true" pageId="16" pageNumber="77">Cuzco</emphasis>
, La convención Province, 
<quantity id="4C909B3CAD28FF87FC70FEB8FB83FE0D" box="[998,1060,367,391]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="77" unit="km" value="4.0">4 km</quantity>
S Camisea River. Campamento Cashiriari-2, Plot 
<quantity id="4C909B3CAD28FF87FEC9FE43FE19FE26" box="[351,446,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.579" pageId="16" pageNumber="77" unit="m" value="1579.0">1, 579m</quantity>
, 
<geoCoordinate id="EE5C501EAD28FF87FE5FFE43FDF9FE26" box="[457,606,404,428]" direction="south" orientation="latitude" pageId="16" pageNumber="77" precision="1" value="-11.86425">11°51'51.3''S</geoCoordinate>
; 
<geoCoordinate id="EE5C501EAD28FF87FDFCFE43FCAFFE26" box="[618,776,404,428]" direction="west" orientation="longitude" pageId="16" pageNumber="77" precision="1" value="-72.779335">72°46'45.6''W</geoCoordinate>
, J Santisteban et al. (#38, Primary Rainforest, hilly terrain, ex: Winkler Trap) [MUSM-ENT 0201599/ANTWEB-CASENT 0178882] [MUSM].
</paragraph>
</subSubSection>
</treatment>
</document>