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<document id="6A870679FC41689D5C0FBB40E0B99F85" ID="10.11646/zootaxa.4139.1.4" ID-DOI="10.11646/zootaxa.4139.1.4" ID-GBIF-Dataset="3cf8edb7-4be5-4ec3-803e-ca1f224d703b" ID-ISSN="1175-5326" ID-Zenodo-Dep="271999" ID-ZooBank="2744EC46-70A1-4267-AAB8-2131F5DC21DC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468909435473" checkinUser="plazi" docAuthor="Nunes, Rafael V. &amp; Vaz-De-Mello, Fernando Z." docDate="2016" docId="03C187AE4533FF890EC8FEF0FDB7F8B1" docLanguage="en" docName="zootaxa.4139.1.4.pdf" docOrigin="Zootaxa 4139 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Selenocopris) manni Nunes &amp; Vaz-De-Mello, 2016, new species" docType="treatment" docVersion="7" masterDocId="FFF8FFD6453BFF810E5FFF9FFF9CFFFA" masterDocTitle="New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus" updateTime="1698365276952" updateUser="plazi">
<mods:mods id="0AE1E0BB1F6BAB1E38CDDB89A6FC7405" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D8BFEACD28F5E03B29B421A2F070BA8B">
<mods:title id="132F6FFDE314E0368892A0306F0C2560">New brachypterous species of Dichotomius (Selenocopris) Burmeister (Coleoptera: Scarabaeidae: Scarabaeinae) with the definition of species groups and taxonomic notes in the subgenus</mods:title>
</mods:titleInfo>
<mods:name id="1061915889BBCE799ADC0F08EEE1516E" type="personal">
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<mods:namePart id="FE42E0AF10D4AB4A813DFB2D111667B3">Nunes, Rafael V.</mods:namePart>
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<mods:name id="260192FD5BFB0A4E96D42D5255CF9A2F" type="personal">
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<mods:roleTerm id="61AB4CCBC439F94285F1809A07B2360E">Author</mods:roleTerm>
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<mods:namePart id="393165AFBA07A775E90018E0A0FCD919">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="BEB386389E25C21A87DE48BEB3DA4EBC">text</mods:typeOfResource>
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<mods:title id="2D65A6E130890AFAB29F237EA116E4D8">Zootaxa</mods:title>
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<mods:part id="67EC947BA92E22AE29B8A284BB5EDA30">
<mods:date id="2CB9770C65E83FE7FDD008CDD9ED5D37">2016</mods:date>
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<mods:number id="D905D09E9D8A4B38D501C8FE39B80FB8">4139</mods:number>
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<mods:classification id="E3F3FB928305E06CBB3BD4555F1BA50F">journal volume</mods:classification>
<mods:identifier id="1D6F999FF59400EDC74C6E2923DE9E70" type="DOI">10.11646/zootaxa.4139.1.4</mods:identifier>
<mods:identifier id="61EAF93BC594259ED9381DFC46DC97C0" type="GBIF-Dataset">3cf8edb7-4be5-4ec3-803e-ca1f224d703b</mods:identifier>
<mods:identifier id="2D6D0834E740F2102F2EBEFBAD390484" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C187AE4533FF890EC8FEF0FDB7F8B1" ID-DOI="http://doi.org/10.5281/zenodo.6090773" ID-GBIF-Taxon="123198330" ID-Zenodo-Dep="6090773" LSID="urn:lsid:plazi:treatment:03C187AE4533FF890EC8FEF0FDB7F8B1" httpUri="http://treatment.plazi.org/id/03C187AE4533FF890EC8FEF0FDB7F8B1" pageId="8">
<subSubSection id="C37265334533FF890EC8FEF0FF77FE51" pageId="8" type="nomenclature">
<paragraph id="8BD736B84533FF890EC8FEF0FD4FFE70" blockId="8.[151,723,367,427]" box="[151,723,367,394]" pageId="8">
<heading id="D09F81D44533FF890EC8FEF0FD4FFE70" bold="true" box="[151,723,367,394]" fontSize="11" level="1" pageId="8" reason="1">
<emphasis id="B91CEAAA4533FF890EC8FEF0FD4FFE70" bold="true" box="[151,723,367,394]" pageId="8">
<taxonomicName id="4C684D3B4533FF890EC8FEF0FDA1FE73" ID-CoL="6D3RL" box="[151,573,367,393]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="manni" status="sp. nov." subGenus="Selenocopris">
<emphasis id="B91CEAAA4533FF890EC8FEF0FDA1FE73" bold="true" box="[151,573,367,393]" italics="true" pageId="8">Dichotomius (Selenocopris) manni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57D14533FF890C1BFEEFFD4FFE70" box="[580,723,368,394]" pageId="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD736B84533FF890EC8FE0DFF77FE51" blockId="8.[151,723,367,427]" box="[151,235,402,427]" pageId="8">
(
<figureCitation id="13532A3D4533FF890EC0FE0DFF7EFE51" box="[159,226,402,427]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="8">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890EC8FE45FBD0FDED" pageId="8" type="materials_examined">
<paragraph id="8BD736B84533FF890EC8FE45FBD0FDED" blockId="8.[151,1437,474,1867]" pageId="8">
<emphasis id="B91CEAAA4533FF890EC8FE45FE74FE09" bold="true" box="[151,488,474,499]" pageId="8">
Material studied.
<typeStatus id="54D3881A4533FF890F2DFE45FE7FFE09" box="[370,483,474,499]" pageId="8" type="holotype">Holotype</typeStatus>
:
</emphasis>
Ƌ Serra da Canastra, M.G
<collectingCountry id="F37F76284533FF890D71FE44FC0EFE09" box="[814,914,475,499]" name="Brazil" pageId="8">BRASIL</collectingCountry>
/ OUMNH-2003-051 W.D.Hamilton Colln. /
<taxonomicName id="4C684D3B4533FF890EC8FE61FEEFFDEC" box="[151,371,510,534]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="manni" status="sp. nov.">
<emphasis id="B91CEAAA4533FF890EC8FE61FEEFFDEC" box="[151,371,510,534]" italics="true" pageId="8">Dichotomius manni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57D14533FF890F25FE60FE29FDEC" box="[378,437,511,534]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
,
<typeStatus id="54D3881A4533FF890F9BFE61FDB6FDED" box="[452,554,510,535]" pageId="8" type="holotype">holotype</typeStatus>
. RVNunes &amp; FZVaz-de-Mello, (det) [
<collectionCode id="ED79AE7D4533FF890D90FE60FBDEFDED" box="[975,1090,511,535]" pageId="8">OUMNH</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890E98FDBDFC41FD31" pageId="8" type="diagnosis">
<paragraph id="8BD736B84533FF890E98FDBDFC41FD31" blockId="8.[151,1437,474,1867]" pageId="8">
<emphasis id="B91CEAAA4533FF890E98FDBDFEDDFDC1" bold="true" box="[199,321,546,571]" pageId="8">Diagnosis.</emphasis>
In the group,
<taxonomicName id="4C684D3B4533FF890FB4FDBCFDCBFDC0" box="[491,599,547,570]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="manni" status="sp. nov.">
<emphasis id="B91CEAAA4533FF890FB4FDBCFDCBFDC0" box="[491,599,547,570]" italics="true" pageId="8">D. manni</emphasis>
</taxonomicName>
<emphasis id="B91CEAAA4533FF890C3EFDBDFD3FFDC1" bold="true" box="[609,675,546,571]" pageId="8">
<taxonomicNameLabel id="A22F57D14533FF890C3EFDBDFD3FFDC1" box="[609,675,546,571]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the largest of the five recognized brachypterous species and is separated from them by the following combined characters: cephalic carina higher than wide bearing a feeble emargination at the apex (
<figureCitation id="13532A3D4533FF890FE8FDF5FE65FD79" box="[439,505,618,643]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="8">Fig. 5</figureCitation>
A); anterior pronotal excavation bearing spaced fine punctures spaced by more than twice their diameter (
<figureCitation id="13532A3D4533FF890FDAFD11FE5BFD5D" box="[389,455,654,679]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="8">Fig. 5</figureCitation>
A); central curved lobe delimiting the pronotal disc from the anterior declivity (
<figureCitation id="13532A3D4533FF890B62FD11FAE2FD5D" box="[1341,1406,654,679]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="8">Fig. 5</figureCitation>
A) and paramera longitudinal excavation taking the apical one third (
<figureCitation id="13532A3D4533FF890D27FD2DFC27FD31" box="[888,955,690,715]" captionStart="FIGURE 5" captionStartId="9.[151,250,1708,1730]" captionTargetBox="[341,1244,193,1685]" captionTargetId="figure@9.[341,1245,193,1686]" captionTargetPageId="9" captionText="FIGURE 5. Holotype of Dichotomius (Selenocopris) manni n. sp.: (A) male dorsal habitus; (B) male paramera." httpUri="https://zenodo.org/record/272004/files/figure.png" pageId="8">Fig. 5</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890E98FD49FDC1F9B5" pageId="8" type="materials_examined">
<paragraph id="8BD736B84533FF890E98FD49FDC1F9B5" blockId="8.[151,1437,474,1867]" pageId="8">
<emphasis id="B91CEAAA4533FF890E98FD49FE1AFD15" bold="true" box="[199,390,726,751]" pageId="8">
<typeStatus id="54D3881A4533FF890E98FD49FEAAFD15" box="[199,310,726,751]" pageId="8" type="holotype">Holotype</typeStatus>
. male:
</emphasis>
length: 25.0 mm. Maximum width (pronotum):
<quantity id="4C909B5D4533FF890DEFFD48FB98FD14" box="[944,1028,727,751]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="8" unit="mm" value="14.0">14 mm</quantity>
. Black, shiny.
<emphasis id="B91CEAAA4533FF890AF3FD49FB69FD15" bold="true" box="[1196,1269,726,751]" pageId="8">Head:</emphasis>
dorsal surface smooth, bearing few ocellate punctures s close to the internal border of the eyes. Clypeogenal junction with obtuse angle. Margins of clypeal teeth bearing a row of scattered setae. Ventral clypeal process strongly bifurcated. Cephalic carina higher than wide and bearing a feeble emargination at the apex. Clypeo-genal suture distinct and producing a knob close to the posterior portion. Antennal club with light brown tumescence.
<emphasis id="B91CEAAA4533FF890A8FFCF9FACEFC85" bold="true" box="[1232,1362,870,895]" pageId="8">Pronotum:</emphasis>
wider than long and wider than elytra. Anterior excavation bearing fine punctures spaced by more than twice their diameter. Anterior portion of pronotal lacking lobes. Anterior angle acute.
<emphasis id="B91CEAAA4533FF890DA8FC31FB12FC3D" bold="true" box="[1015,1166,942,967]" pageId="8">Hypomeron:</emphasis>
anterior portion with a lateral band of ocellate setose punctures. Posterior portion almost glabrous with few sparse ocellate setose punctures.
<emphasis id="B91CEAAA4533FF890F47FC69FE2CFBF5" bold="true" box="[280,432,1014,1039]" pageId="8">Prosternum:</emphasis>
covered by ocellate setose punctures: setae do not reach one quarter the length of hypomeron setae.
<emphasis id="B91CEAAA4533FF890F31FB85FD80FBC9" bold="true" box="[366,540,1050,1075]" pageId="8">Mesosternum:</emphasis>
strongly narrowed, glabrous and opaque medially.
<emphasis id="B91CEAAA4533FF890ADEFB85FADFFBC9" bold="true" box="[1153,1347,1050,1075]" pageId="8">Mesepisternum:</emphasis>
having ocellate setose punctures. Posterior portion of upper margin ending in an acute angle, reaching pseudoepipleuron (
<figureCitation id="13532A3D4533FF890EC0FBFDFF73FB81" box="[159,239,1122,1147]" captionStart="FIGURE 1" captionStartId="3.[151,250,1738,1760]" captionTargetBox="[255,1318,202,1704]" captionTargetId="figure@3.[249,1332,187,1718]" captionTargetPageId="3" captionText="FIGURE 1. (A) Ventral habitus of a female of Dichotomius (Selenocopris) periotoi n. sp. The white arrow at the head indicates the position of the ventral clypeal process and the white at the abdomen indicates the modification at the medial portion of the 6 th sternite; (B) lateral view of the same specimen. The white arrow indicating the position of the mesepisternum.; (C) detail of the mesepisternum of the same specimen. The white arrow indicates the posterior portion of the upper margin of the mesepisternum ending in an acute angle, which is typical of Selenocopris; (D) elytral striae and interstriae of D. (S.) fissus (Gillet); (E) elytral striae and interstriae of D. (S.) fissiceps (Felsche) (quadraticeps group); (F) head and clypeal teeth of D. (S.) bicuspis (Germar); (G) head and clypeal teeth of D. (S.) fissiceps (quadraticeps group); (H) 6 th abdominal sternite of the female of D. (S.) spadiceus (Luederwaldt); (I) 6 th abdominal sternite of the female of D. (S.) punctulatipennis (Luederwaldt) (bicuspis group); (J) 6 th abdominal sternite of the male of D. (S.) ascanius (Harold); (K) 6 th abdominal sternite of the female of D. (S.) ascanius (Luederwaldt); (L) 6 th abdominal sternite of the female of D. (S.) quadraticeps (Felsche)." httpUri="https://zenodo.org/record/272000/files/figure.png" pageId="8">Figs. 1</figureCitation>
BC).
<emphasis id="B91CEAAA4533FF890F1BFBFDFE73FB81" bold="true" box="[324,495,1122,1147]" pageId="8">Metasternum:</emphasis>
mesometasternal suture indistinct. Ocellate setose punctures present along the margins of the metasternal lobe. Metasternal disc with a feeble longitudinal sulcus producing a feeble posterior concavity.
<emphasis id="B91CEAAA4533FF890F4CFB35FEF4FB39" bold="true" box="[275,360,1194,1219]" pageId="8">Elytra:</emphasis>
shinny, lacking humeral callus. Points of the striae separated by twice their diameter. Elytral interstriae feebly convex.
<emphasis id="B91CEAAA4533FF890FE7FB51FDD4FB1D" bold="true" box="[440,584,1230,1255]" pageId="8">Hind wings:</emphasis>
length:
<quantity id="4C909B5D4533FF890CFCFB50FD68FB1C" box="[675,756,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" unit="mm" value="11.0">11 mm</quantity>
. Reduced but articulated at the folding region. Basal sclerites glabrous. Subcostal vein intersecting costal axis at its basal quarter. Medial vein extending to folding articulation. Distinct setae present on the basal third of costal axis and at folding articulation. Cubital vein present, anal vein absent. Small developed membranous portion apical to folding articulation with incomplete or weakly-marked venations.
<emphasis id="B91CEAAA4533FF890F4BFAC1FEC9FA8D" bold="true" box="[276,341,1374,1399]" pageId="8">Legs:</emphasis>
anterior tibial spur strongly bent downward at apical fifth. Anterior tibiae bearing four lateral teeth. Ventral surface of anterior femur smooth and glabrous. Length of the fifth metatarsomere twice the length of the fourth metatarsomere.
<emphasis id="B91CEAAA4533FF890F98FA39FDD9FA45" bold="true" box="[455,581,1446,1471]" pageId="8">Abdomen:</emphasis>
sternites 1 to 6 glabrous and having ocellate punctures along the anterior border.
<emphasis id="B91CEAAA4533FF890EABFA55FEECFA19" bold="true" box="[244,368,1482,1507]" pageId="8">Pygidium:</emphasis>
shinny, smooth, wider than long, lacking punctures.
<emphasis id="B91CEAAA4533FF890DA6FA55FBE6FA19" bold="true" box="[1017,1146,1482,1507]" pageId="8">Paramera:</emphasis>
having rounded/circular excavations near the basis (dorsal view) and longitudinal excavation toward the apex (lateral and dorsal view). Anterior border of circular/rounded excavation carinates. Longitudinal excavation taking 1/3 the length of paramera. Apex divergent (dorsal view).
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890E98F9C5FA07F989" box="[199,1435,1626,1651]" pageId="8" type="distribution">
<paragraph id="8BD736B84533FF890E98F9C5FA07F989" blockId="8.[151,1437,474,1867]" box="[199,1435,1626,1651]" pageId="8">
<emphasis id="B91CEAAA4533FF890E98F9C5FEFCF989" bold="true" box="[199,352,1626,1651]" pageId="8">Distribution.</emphasis>
the
<typeStatus id="54D3881A4533FF890FD0F9C5FE6DF989" box="[399,497,1626,1651]" pageId="8" type="holotype">holotype</typeStatus>
is known from Serra da Canastra, south of the state of
<collectingRegion id="49ACF85A4533FF890A0BF9C5FB70F989" box="[1108,1260,1626,1651]" country="Brazil" name="Minas Gerais" pageId="8">Minas Gerais</collectingRegion>
,
<collectingCountry id="F37F76284533FF890AA8F9C5FAA7F989" box="[1271,1339,1626,1651]" name="Brazil" pageId="8">Brazil</collectingCountry>
(
<figureCitation id="13532A3D4533FF890B15F9C5FA12F989" box="[1354,1422,1626,1651]" captionStart="FIGURE 8" captionStartId="13.[151,250,1913,1935]" captionTargetBox="[235,1353,676,1876]" captionTargetId="figure@13.[222,1365,661,1892]" captionTargetPageId="13" captionText="FIGURE 8. Distribution of the brachypterous species of Selenocopris." httpUri="https://zenodo.org/record/272007/files/figure.png" pageId="8">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890E98F9E1FCE2F941" pageId="8" type="etymology">
<paragraph id="8BD736B84533FF890E98F9E1FCE2F941" blockId="8.[151,1437,474,1867]" pageId="8">
<emphasis id="B91CEAAA4533FF890E98F9E1FED2F96D" bold="true" box="[199,334,1662,1687]" pageId="8">Etymology.</emphasis>
Named after Dr. Darren J. Mann, Head of life collections at Hope Entomological Collections of OUMNH and that kindly landed to us the
<typeStatus id="54D3881A4533FF890C31F93DFD4CF941" box="[622,720,1698,1723]" pageId="8" type="holotype">holotype</typeStatus>
of this species.
</paragraph>
</subSubSection>
<subSubSection id="C37265334533FF890E98F959FDB7F8B1" pageId="8" type="discussion">
<paragraph id="8BD736B84533FF890E98F959FDB7F8B1" blockId="8.[151,1437,474,1867]" pageId="8">
<emphasis id="B91CEAAA4533FF890E98F959FEA7F925" bold="true" box="[199,315,1734,1759]" pageId="8">Remarks.</emphasis>
<taxonomicName id="4C684D3B4533FF890F1CF958FE30F924" box="[323,428,1735,1758]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="manni" status="sp. nov.">
<emphasis id="B91CEAAA4533FF890F1CF958FE30F924" box="[323,428,1735,1758]" italics="true" pageId="8">D. manni</emphasis>
</taxonomicName>
<emphasis id="B91CEAAA4533FF890FEBF959FE68F925" bold="true" box="[436,500,1734,1759]" pageId="8">
<taxonomicNameLabel id="A22F57D14533FF890FEBF959FE68F925" box="[436,500,1734,1759]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the largest brachypterpus species in
<taxonomicName id="4C684D3B4533FF890DECF959FBDFF924" box="[947,1091,1734,1758]" class="Insecta" family="Scarabaeidae" genus="Selenocopris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="B91CEAAA4533FF890DECF959FBDFF924" box="[947,1091,1734,1758]" italics="true" pageId="8">Selenocopris</emphasis>
</taxonomicName>
. It is similar to
<taxonomicName id="4C684D3B4533FF890AA3F958FAE4F924" box="[1276,1400,1735,1758]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="periotoi">
<emphasis id="B91CEAAA4533FF890AA3F958FAE4F924" box="[1276,1400,1735,1758]" italics="true" pageId="8">D. periotoi</emphasis>
</taxonomicName>
by the pronotum and the clypeo-frontal carina. However, the short longitudinal excavation of the paramera are similar to those found in
<taxonomicName id="4C684D3B4533FF890F07F890FE23F8DC" box="[344,447,1807,1830]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="paresi" status="sp. nov.">
<emphasis id="B91CEAAA4533FF890F07F890FE23F8DC" box="[344,447,1807,1830]" italics="true" pageId="8">D. paresi</emphasis>
</taxonomicName>
<emphasis id="B91CEAAA4533FF890F9AF891FD9FF8DD" bold="true" box="[453,515,1806,1831]" pageId="8">
<taxonomicNameLabel id="A22F57D14533FF890F9AF891FD9FF8DD" box="[453,515,1806,1831]" pageId="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The label of the
<typeStatus id="54D3881A4533FF890CE1F891FCBCF8DD" box="[702,800,1806,1831]" pageId="8" type="holotype">holotype</typeStatus>
does not bring additional information concerning habitat or other natural history information.
</paragraph>
</subSubSection>
</treatment>
</document>