279 lines
47 KiB
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279 lines
47 KiB
XML
<document id="8E45351A9CCDDC9E5C076C7AB5E4F502" ID-DOI="10.11646/zootaxa.4742.3.11" ID-GBIF-Dataset="a1b47413-2f76-404c-86e4-571b06c6a1cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677917" ID-ZooBank="B5B11009-059B-4DC1-85B6-831A67B72166" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272996897" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="03BE87E8FF9A7F199CF7FEC9545DFF1C" docLanguage="en" docName="zootaxa.4742.3.11.pdf" docOrigin="Zootaxa 4742 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Monilistylus) Kluge 2020, subgen. n." docType="treatment" docVersion="7" lastPageNumber="579" masterDocId="FF87FF90FF9F7F1F9C60FFB85021FFCB" masterDocTitle="New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon)" masterLastPageNumber="587" masterPageNumber="573" pageNumber="578" updateTime="1698720838785" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2A8D0079BCACC2F2C200DDC88DFA408A">New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon)</mods:title>
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<mods:namePart id="E2F373AEDA157C16C5C4D5FDCE10B520">Kluge, Nikita J.</mods:namePart>
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<mods:title id="1C093CBB037888AD3724B7F1773DE4CC">Zootaxa</mods:title>
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<mods:part id="2DDA508FA957E67FE1DEF3CAE84D6D97">
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<mods:date id="5FE03298DF06C14FBB0AB727D9DC6A64">2020</mods:date>
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<mods:number id="1210447CDB6744EF04CC025B89F55A71">2020-02-21</mods:number>
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<treatment id="03BE87E8FF9A7F199CF7FEC9545DFF1C" ID-DOI="http://doi.org/10.5281/zenodo.3684990" ID-GBIF-Taxon="162244917" ID-Zenodo-Dep="3684990" LSID="urn:lsid:plazi:treatment:03BE87E8FF9A7F199CF7FEC9545DFF1C" httpUri="http://treatment.plazi.org/id/03BE87E8FF9A7F199CF7FEC9545DFF1C" lastPageId="6" lastPageNumber="579" pageId="5" pageNumber="578">
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<subSubSection id="C30D6575FF9A7F1A9CF7FEC95192FE40" box="[151,435,369,396]" pageId="5" pageNumber="578" type="nomenclature">
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<paragraph id="8BA836FEFF9A7F1A9CF7FEC95192FE40" blockId="5.[151,435,369,431]" box="[151,435,369,396]" pageId="5" pageNumber="578">
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<heading id="D0E08192FF9A7F1A9CF7FEC95192FE40" bold="true" box="[151,435,369,396]" fontSize="11" level="1" pageId="5" pageNumber="578" reason="1">
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<emphasis id="B963EAECFF9A7F1A9CF7FEC95192FE40" bold="true" box="[151,435,369,396]" pageId="5" pageNumber="578">
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<taxonomicName id="4C174D7DFF9A7F1A9CF7FEC9510CFE47" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[151,301,369,396]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Monilistylus">
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<emphasis id="B963EAECFF9A7F1A9CF7FEC9510CFE47" bold="true" box="[151,301,369,396]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2505797FF9A7F1A9D54FEC95192FE40" box="[308,435,369,395]" pageId="5" pageNumber="578" rank="subGenus">subgen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D6575FF9A7F1A9CF7FE2D5138FE64" box="[151,281,404,431]" pageId="5" pageNumber="578" type="description">
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<paragraph id="8BA836FEFF9A7F1A9CF7FE2D5138FE64" blockId="5.[151,435,369,431]" box="[151,281,404,431]" pageId="5" pageNumber="578">
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(
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<figureCitation id="132C2A7BFF9A7F1A9CFFFE2D5130FE64" box="[159,273,404,431]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURES 9–25" captionStart-2="FIGURES 26–29" captionStart-3="FIGURES 30–32" captionStart-4="FIGURES 33–38" captionStart-5="FIGURES 39–46" captionStart-6="FIGURES 47–51" captionStart-7="FIGURES 52–54" captionStartId-0="2.[151,264,1897,1922]" captionStartId-1="3.[151,264,1705,1730]" captionStartId-2="6.[151,264,1811,1836]" captionStartId-3="8.[151,264,1781,1806]" captionStartId-4="9.[151,264,1848,1873]" captionStartId-5="10.[151,264,1885,1910]" captionStartId-6="11.[151,264,1579,1604]" captionStartId-7="12.[151,264,1768,1793]" captionTargetBox-0="[151,1436,270,1869]" captionTargetBox-1="[189,1402,191,1671]" captionTargetBox-2="[151,1436,236,1786]" captionTargetBox-3="[151,1436,182,1755]" captionTargetBox-4="[164,1423,273,1823]" captionTargetBox-5="[151,1436,181,1860]" captionTargetBox-6="[164,1423,181,1549]" captionTargetBox-7="[165,1436,188,1732]" captionTargetId-0="figure@2.[151,1436,270,1869]" captionTargetId-1="figure@3.[177,1411,181,1679]" captionTargetId-2="figure@6.[151,1436,236,1786]" captionTargetId-3="figure@8.[151,1436,181,1755]" captionTargetId-4="figure@9.[164,1423,273,1823]" captionTargetId-5="figure@10.[151,1436,181,1860]" captionTargetId-6="figure@11.[164,1423,181,1553]" captionTargetId-7="figure@12.[151,1436,181,1743]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionText-0="FIGURES 1–8. Larval exuviae. 1–2, Procloeon (Monilistylus) monilistylus: 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8, Procloeon (Monilistylus) ornatipennis sp. n.: 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged." captionText-1="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." captionText-2="FIGURES 26–29. Procloeon (Monilistylus) ornatipennis sp. n., exuviae of labium: 26, general view with focus on ventral side; 27, labial palp with focus on dorsal side; 28, glossa and paraglossa with focus on ventral side; 29, the same with focus on dorsal side (26, holotype)." captionText-3="FIGURES 30–32. Procloeon (Monilistylus) ornatipennis sp. n., legs: 30, foreleg in anterior view (holotype); 31, base of tibia in anterior view; 32, claw." captionText-4="FIGURES 33–38. Procloeon (Monilistylus) ornatipennis sp. n.: 33, larval exuviae of posterior margin of 9th abdominal sternum with protogonostyli; 34, subimaginal tissues with gonostyli, extracted from it; 35, exuviae of paraprocts and protogonostyli of another specimen; 36, exuviae of 5th abdominal sternum (Figs 33–36 under the same magnification); 37,middle of posterior margin of 9th abdominal tergum; 38, portion of mesonotum (magnification as in Fig. 37).Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle." captionText-5="FIGURES 39–46. Procloeon (Monilistylus) ornatipennis sp. n.: 39–40, male imagines; 41, male imaginal genitals; 42, exuviae of right half of subimaginal mesonotum; 43, exuviae of left half of subimaginal mesopleuron; 44, wing; 45, abdominal terga II–VII; 46, tibia and tarsus of middle leg (41–43 and 45–46, holotype)." captionText-6="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." captionText-7="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi-0="http://doi.org/10.5281/zenodo.3677921" figureDoi-1="http://doi.org/10.5281/zenodo.3677923" figureDoi-2="http://doi.org/10.5281/zenodo.3677925" figureDoi-3="http://doi.org/10.5281/zenodo.3677927" figureDoi-4="http://doi.org/10.5281/zenodo.3677929" figureDoi-5="http://doi.org/10.5281/zenodo.3677931" figureDoi-6="http://doi.org/10.5281/zenodo.3677933" figureDoi-7="http://doi.org/10.5281/zenodo.3677935" httpUri-0="https://zenodo.org/record/3677921/files/figure.png" httpUri-1="https://zenodo.org/record/3677923/files/figure.png" httpUri-2="https://zenodo.org/record/3677925/files/figure.png" httpUri-3="https://zenodo.org/record/3677927/files/figure.png" httpUri-4="https://zenodo.org/record/3677929/files/figure.png" httpUri-5="https://zenodo.org/record/3677931/files/figure.png" httpUri-6="https://zenodo.org/record/3677933/files/figure.png" httpUri-7="https://zenodo.org/record/3677935/files/figure.png" pageId="5" pageNumber="578">Figs 1–54</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D6575FF9A7F1A9CF7FE655458F870" pageId="5" pageNumber="578" type="type_taxon">
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<paragraph id="8BA836FEFF9A7F1A9CF7FE6553EDFE3C" blockId="5.[151,1437,476,2015]" box="[151,972,476,503]" pageId="5" pageNumber="578">
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<emphasis id="B963EAECFF9A7F1A9CF7FE655115FE3C" bold="true" box="[151,308,476,503]" pageId="5" pageNumber="578">
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<typeStatus id="54AC885CFF9A7F1A9CF7FE6550F3FE3C" box="[151,210,477,503]" pageId="5" pageNumber="578">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C174D7DFF9A7F1A9D5CFE6553E9FE3C" authority="Kluge, Tiunova & Novikova 2014" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[316,968,477,503]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="monilistylus">
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<emphasis id="B963EAECFF9A7F1A9D5CFE65521CFE3C" box="[316,573,477,503]" italics="true" pageId="5" pageNumber="578">Procloeon monilistylus</emphasis>
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<bibRefCitation id="EF864B0FFF9A7F1A9E24FE6553E9FE3C" author="Kluge, N. J. & Tiunova, T. M. & Novikova, E. A." box="[580,968,477,503]" pageId="5" pageNumber="578" pagination="483 - 491" refId="ref7118" refString="Kluge, N. J., Tiunova, T. M. & Novikova, E. A. (2014) A new species, Procloeon monilistylus sp. n. (Ephemeroptera, Baetidae), from the Russian Far East. Zootaxa, 3786 (4), 483 - 491. https: // doi. org / 10.11646 / zootaxa. 3786.4.6" type="journal article" year="2014">Kluge, Tiunova & Novikova 2014</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA836FEFF9A7F1A9CA7FDB8526AFDD0" blockId="5.[151,1437,476,2015]" box="[199,587,512,539]" pageId="5" pageNumber="578">
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<emphasis id="B963EAECFF9A7F1A9CA7FDB8526AFDD0" bold="true" box="[199,587,512,539]" pageId="5" pageNumber="578">
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Autapomorphies of
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<emphasis id="B963EAECFF9A7F1A9DD5FDB85265FDD0" bold="true" box="[437,580,512,539]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
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.
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</emphasis>
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</paragraph>
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<paragraph id="8BA836FEFF9A7F1A9CA7FD9C50E8FD4C" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
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<emphasis id="B963EAECFF9A7F1A9CA7FD9C50C6FDF4" bold="true" box="[199,231,548,575]" pageId="5" pageNumber="578">(1)</emphasis>
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In male imago, sterno-styligeral muscle (initially unpaired) has enlarged paired lateral portions, which are attached to the bases of unistyligers (
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<figureCitation id="132C2A7BFF9A7F1A9E59FDF152B1FDA8" box="[569,656,584,611]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="5" pageNumber="578">Figs 51</figureCitation>
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,
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<figureCitation id="132C2A7BFF9A7F1A9EFDFDF0529DFDA8" box="[669,700,584,611]" captionStart="FIGURES 52–54" captionStartId="12.[151,264,1768,1793]" captionTargetBox="[165,1436,188,1732]" captionTargetId="figure@12.[151,1436,181,1743]" captionTargetPageId="12" captionText="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677935" httpUri="https://zenodo.org/record/3677935/files/figure.png" pageId="5" pageNumber="578">53</figureCitation>
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), so that each lateral portion can serve as adductor of the unistyliger.
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</paragraph>
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<paragraph id="8BA836FEFF9A7F1A9CA7FD2952AFFC6C" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
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In our previous paper (Kluge
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<emphasis id="B963EAECFF9A7F1A9E77FD2A5268FD60" box="[535,585,657,683]" italics="true" pageId="5" pageNumber="578">et al</emphasis>
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. 2014) the lateral portions of the sterno-styligeral muscle were wrongly interpreted as «penial muscles» (Kluge
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<emphasis id="B963EAECFF9A7F1A9E5CFD0E524DFD04" box="[572,620,693,719]" italics="true" pageId="5" pageNumber="578">et al</emphasis>
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. 2014: legend to fig. 25) or as parts of the «penial muscles» (ibid: legend to fig. 31). This error was caused by the fact that in
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<taxonomicName id="4C174D7DFF9A7F1A9EBEFD62539CFD38" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[734,957,729,755]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="monilistylus" subGenus="Monilistylus">
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<emphasis id="B963EAECFF9A7F1A9EBEFD6252CEFD38" box="[734,751,730,755]" italics="true" pageId="5" pageNumber="578">P</emphasis>
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. (
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<emphasis id="B963EAECFF9A7F1A9F67FD62533FFD38" box="[775,798,730,755]" italics="true" pageId="5" pageNumber="578">M</emphasis>
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.)
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<emphasis id="B963EAECFF9A7F1A9F56FD61539CFD38" box="[822,957,729,755]" italics="true" pageId="5" pageNumber="578">monilistylus</emphasis>
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</taxonomicName>
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the lateral portions of the sterno-styligeral muscle run parallel to the gonovectal (penial) muscles and completely overlap them in ventral view, so that these two pairs of muscles look as one pair (compare left and right halves of
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<figureCitation id="132C2A7BFF9A7F1A9FD4FC995429FCF0" box="[948,1032,800,827]" captionStart="FIGURES 52–54" captionStartId="12.[151,264,1768,1793]" captionTargetBox="[165,1436,188,1732]" captionTargetId="figure@12.[151,1436,181,1743]" captionTargetPageId="12" captionText="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677935" httpUri="https://zenodo.org/record/3677935/files/figure.png" pageId="5" pageNumber="578">Fig. 52</figureCitation>
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). In the new species,
|
||
<taxonomicName id="4C174D7DFF9A7F1A9897FC9A50C1FC94" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="ornatipennis" status="sp. nov." subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9A7F1A9897FC9A5529FCF0" box="[1271,1288,802,827]" italics="true" pageId="5" pageNumber="578">P</emphasis>
|
||
. (
|
||
<emphasis id="B963EAECFF9A7F1A9940FC9A5516FCF0" box="[1312,1335,802,827]" italics="true" pageId="5" pageNumber="578">M</emphasis>
|
||
.)
|
||
<emphasis id="B963EAECFF9A7F1A992EFC9A50C1FC94" italics="true" pageId="5" pageNumber="578">ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2505797FF9A7F1A9C87FCFD5105FC94" box="[231,292,837,863]" pageId="5" pageNumber="578" rank="species">sp. n.</taxonomicNameLabel>
|
||
, the lateral portions of the sterno-styligeral muscle are divergent caudally (
|
||
<figureCitation id="132C2A7BFF9A7F1A9806FCFD5493FC94" box="[1126,1202,836,863]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="5" pageNumber="578">Fig 51</figureCitation>
|
||
), while the gonovectal (penial) muscles are convergent caudally (
|
||
<figureCitation id="132C2A7BFF9A7F1A9EF7FCD152CAFC48" box="[663,747,872,899]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="5" pageNumber="578">Fig. 50</figureCitation>
|
||
), so that in ventral view these two pairs of muscles cross one another, being well distinguishable (
|
||
<figureCitation id="132C2A7BFF9A7F1A9E4DFC3552A3FC6C" box="[557,642,909,935]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="5" pageNumber="578">Fig. 49</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7FC095259FA9C" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
|
||
Initially for
|
||
<taxonomicName id="4C174D7DFF9A7F1A9D2DFC0951DAFC00" ID-CoL="372" box="[333,507,945,971]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
|
||
, the sterno-styligeral muscle is unpaired and serves for rotation of the unpaired styliger as a whole to the ventral direction; when this muscle is contracted, the styliger becomes directed perpendicular to the body and the penis becomes protracted (
|
||
<bibRefCitation id="EF864B0FFF9A7F1A9ECEFC415318FBD8" author="Kluge, N. J." box="[686,825,1017,1043]" pageId="5" pageNumber="578" pagination="3 - 16" refId="ref6514" refString="Kluge, N. J. (2003) About evolution and homology of genital appendages of insects. [Trudy Russkogo Entomologicheskogo Obshestva] (Proceedings of the Russian Entomological Society), 74, 3 - 16. [in Russian]" type="journal article" year="2003">Kluge 2003</bibRefCitation>
|
||
: figs 3–7; 2004: figs 10–11). In
|
||
<taxonomicName id="4C174D7DFF9A7F1A98CCFC415530FBD8" ID-CoL="6ZG" authorityName="Leach" authorityYear="1815" box="[1196,1297,1017,1043]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
, the styliger is divided to a pair of unistyligers separated by a deep incision, and the sterno-styligeral muscle is attached to the integument of this incision (
|
||
<bibRefCitation id="EF864B0FFF9A7F1A9DB2FBF952CCFB90" author="Kluge, N. J. & Novikova, E. A." box="[466,749,1089,1115]" pageId="5" pageNumber="578" pagination="1 - 56" refId="ref6931" refString="Kluge, N. J. & Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56. https: // doi. org / 10.15298 / rusentj. 20.1.01" type="journal article" year="2011">Kluge & Novikova 2011</bibRefCitation>
|
||
: fig. 3); as a result of this, contraction of the sterno-styligeral muscle leads not to a rotation of the styliger ventrally, but to inclining the unistyligers toward one another. Primitively for
|
||
<taxonomicName id="4C174D7DFF9A7F1A9D65FB31514BFB68" ID-CoL="6ZG" authorityName="Leach" authorityYear="1815" box="[261,362,1161,1187]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
, the sterno-styligeral muscle is wide and strong, the place of its attachment between unistyligers is sclerotized, and its lateral parts touch the bases of unistyligers (
|
||
<bibRefCitation id="EF864B0FFF9A7F1A9F0EFB1553D7FB0C" author="Kluge, N. J." box="[878,1014,1197,1223]" pageId="5" pageNumber="578" pagination="115 - 128" refId="ref6766" refString="Kluge, N. J. (2018) A new Afrotropical genus Monocentroptilum gen. n. (Ephemeroptera: Baetidae: Protopatellata). Zootaxa, 4486 (2), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4486.2.2" type="journal article" year="2018">Kluge 2018</bibRefCitation>
|
||
: fig. 46). In many baetid taxa, the median sclerite between unistyligers is lost; the sterno-styligeral muscle is often weak and unable to move the unistyligers; in selected taxa this muscle is completely lost (e.g., in Labiobaetini
|
||
<bibRefCitation id="EF864B0FFF9A7F1A9FA9FB4D54FDFAC4" author="Kluge, N. J. & Novikova, E. A." box="[969,1244,1269,1295]" pageId="5" pageNumber="578" pagination="1 - 43" refId="ref7052" refString="Kluge, N. J. & Novikova, E. A. (2016) New tribe Labiobaetini tribus n., redefinition of Pseudopannota Waltz & McCafferty 1987 and descriptions of new and little known species from Zambia and Uganda. Zootaxa, 4169 (1), 1 - 43. https: // doi. org / 10.11646 / zootaxa. 4169.1.1" type="journal article" year="2016">Kluge & Novikova 2016</bibRefCitation>
|
||
and
|
||
<taxonomicName id="4C174D7DFF9A7F1A9970FB4D513DFAF8" authority="Jacob 2003" authorityName="Jacob" authorityYear="2003" class="Insecta" family="Baetidae" genus="Rhodobaetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9970FB4D55BDFAC4" box="[1296,1436,1269,1295]" italics="true" pageId="5" pageNumber="578">Rhodobaetis</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9A7F1A9CF7FAA1513DFAF8" author="Jacob, U." box="[151,284,1305,1331]" pageId="5" pageNumber="578" pagination="59 - 129" refId="ref6197" refString="Jacob, U. (2003) Baetis Leach 1815, sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimmungsschlusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
). In
|
||
<emphasis id="B963EAECFF9A7F1A9D2EFAA151F9FAF8" box="[334,472,1305,1331]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
|
||
, the sterno-styligeral muscle evolved in opposite direction, so that its lateral portions become strong and look as paired muscles.
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7FAD855B6FA54" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7FAD850C9FAB0" bold="true" box="[199,232,1376,1403]" pageId="5" pageNumber="578">(2)</emphasis>
|
||
The second segment of the gonostylus has a composite shape: its proximal 2/3 has a groove on its concave median side; the distal 1/3 has no groove and forms a blunt angle with the grooved proximal portion (
|
||
<figureCitation id="132C2A7BFF9A7F1A9968FA3D5541FA54" box="[1288,1376,1413,1439]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="5" pageNumber="578">Figs 49</figureCitation>
|
||
,
|
||
<figureCitation id="132C2A7BFF9A7F1A990BFA3C55ABFA54" box="[1387,1418,1412,1439]" captionStart="FIGURES 52–54" captionStartId="12.[151,264,1768,1793]" captionTargetBox="[165,1436,188,1732]" captionTargetId="figure@12.[151,1436,181,1743]" captionTargetPageId="12" captionText="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677935" httpUri="https://zenodo.org/record/3677935/files/figure.png" pageId="5" pageNumber="578">53</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7FA105353FA08" blockId="5.[151,1437,476,2015]" box="[199,882,1448,1475]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7FA105353FA08" bold="true" box="[199,882,1448,1475]" pageId="5" pageNumber="578">
|
||
Characters of
|
||
<emphasis id="B963EAECFF9A7F1A9D12FA105220FA08" bold="true" box="[370,513,1448,1475]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
|
||
of unclear phylogenetic status.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7FA745122F928" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7FA7450C9FA2C" bold="true" box="[199,232,1484,1511]" pageId="5" pageNumber="578">(3)</emphasis>
|
||
Mandibles of the «
|
||
<emphasis id="B963EAECFF9A7F1A9DA4FA755279FA2C" box="[452,600,1485,1511]" italics="true" pageId="5" pageNumber="578">Centroptilum</emphasis>
|
||
-
|
||
<typeStatus id="54AC885CFF9A7F1A9E01FA7552B6FA2C" box="[609,663,1485,1511]" pageId="5" pageNumber="578">type</typeStatus>
|
||
»: the kinetodontium is deeply separated from the incisor and rotated perpendicular to the plane of the mandible, so that its denticles hide one another in dorsal or ventral view; the left prostheca is bifurcate and rotated in the same manner (
|
||
<figureCitation id="132C2A7BFF9A7F1A9F63F9AD536DF9E4" box="[771,844,1557,1583]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1897,1922]" captionTargetBox="[151,1436,270,1869]" captionTargetId="figure@2.[151,1436,270,1869]" captionTargetPageId="2" captionText="FIGURES 1–8. Larval exuviae. 1–2, Procloeon (Monilistylus) monilistylus: 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8, Procloeon (Monilistylus) ornatipennis sp. n.: 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3677921" httpUri="https://zenodo.org/record/3677921/files/figure.png" pageId="5" pageNumber="578">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="132C2A7BFF9A7F1A9F39F9AC5382F9E4" box="[857,931,1556,1583]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="5" pageNumber="578">18–25</figureCitation>
|
||
). Among
|
||
<taxonomicName id="4C174D7DFF9A7F1A9871F9AD54ABF9E4" ID-CoL="63PRK" authorityName=": Kluge" authorityYear="2016" box="[1041,1162,1557,1583]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="587" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1, such «
|
||
<emphasis id="B963EAECFF9A7F1A989FF9AD55B2F9E4" box="[1279,1427,1557,1583]" italics="true" pageId="5" pageNumber="578">Centroptilum</emphasis>
|
||
-
|
||
<typeStatus id="54AC885CFF9A7F1A9CF7F98150E8F998" box="[151,201,1593,1619]" pageId="5" pageNumber="578">type</typeStatus>
|
||
» of mandibles, besides
|
||
<emphasis id="B963EAECFF9A7F1A9DB8F9815243F998" box="[472,610,1593,1619]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
|
||
, occurs in
|
||
<taxonomicName id="4C174D7DFF9A7F1A9EBFF98153F9F998" ID-CoL="6ZGW" box="[735,984,1593,1619]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9EBFF98153F9F998" box="[735,984,1593,1619]" italics="true" pageId="5" pageNumber="578">Pseudocentroptiloides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D7DFF9A7F1A9870F981545FF998" ID-CoL="7F34" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[1040,1150,1593,1619]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9870F981545FF998" box="[1040,1150,1593,1619]" italics="true" pageId="5" pageNumber="578">Securiops</emphasis>
|
||
</taxonomicName>
|
||
as well; it is also present in some other baetid taxa (Kluge, in press). In other taxa of
|
||
<taxonomicName id="4C174D7DFF9A7F1A9F5DF9E55397F9BC" ID-CoL="63PRK" authorityName=": Kluge" authorityYear="2016" box="[829,950,1629,1655]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="587" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1, the mandibles are of the «
|
||
<taxonomicName id="4C174D7DFF9A7F1A9967F9E6556CF9BC" ID-CoL="62FJ6" authorityName="Leach" authorityYear="1815" box="[1287,1357,1630,1655]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9967F9E6556CF9BC" box="[1287,1357,1630,1655]" italics="true" pageId="5" pageNumber="578">Baetis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54AC885CFF9A7F1A9935F9E555AAF9BC" box="[1365,1419,1629,1655]" pageId="5" pageNumber="578">type</typeStatus>
|
||
»: the kinetodontium is fused with the incisor; the left prostheca is wide, terminated by 3–6 blunt, more distal denticles and 2–4 pointed, more proximal denticles; all denticles of incisor, kinetodontium, prostheca and mola are visible in one view.
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7F95453A5F8E0" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7F95450C9F8CC" bold="true" box="[199,232,1772,1799]" pageId="5" pageNumber="578">(4)</emphasis>
|
||
Larval femur without stout apical setae on outer margin (
|
||
<figureCitation id="132C2A7BFF9A7F1A9F00F95553FEF8CC" box="[864,991,1773,1799]" captionStart="FIGURES 30–32" captionStartId="8.[151,264,1781,1806]" captionTargetBox="[151,1436,182,1755]" captionTargetId="figure@8.[151,1436,181,1755]" captionTargetPageId="8" captionText="FIGURES 30–32. Procloeon (Monilistylus) ornatipennis sp. n., legs: 30, foreleg in anterior view (holotype); 31, base of tibia in anterior view; 32, claw." figureDoi="http://doi.org/10.5281/zenodo.3677927" httpUri="https://zenodo.org/record/3677927/files/figure.png" pageId="5" pageNumber="578">Figs 30–31</figureCitation>
|
||
) (in contrast to
|
||
<taxonomicName id="4C174D7DFF9A7F1A98EBF9555534F8CC" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1163,1301,1773,1799]" genus="Oculogaster" pageId="5" pageNumber="587" rank="subGenus" status="subgen. nov." subGenus="Oculogaster">
|
||
<emphasis id="B963EAECFF9A7F1A98EBF9555534F8CC" box="[1163,1301,1773,1799]" italics="true" pageId="5" pageNumber="578">Oculogaster</emphasis>
|
||
</taxonomicName>
|
||
and various taxa not belonging to
|
||
<taxonomicName id="4C174D7DFF9A7F1A9DEBF8A951C0F8E0" ID-CoL="3R5Y" authorityName="Leach" authorityYear="1815" box="[395,481,1809,1835]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1, which have 2 stout apical setae).
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7F88C5458F870" blockId="5.[151,1437,476,2015]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7F88C521FF884" bold="true" box="[199,574,1844,1871]" pageId="5" pageNumber="578">
|
||
Plesiomorphies of
|
||
<emphasis id="B963EAECFF9A7F1A9DC8F88C5216F884" bold="true" box="[424,567,1844,1871]" italics="true" pageId="5" pageNumber="578">Monilistylus</emphasis>
|
||
.
|
||
</emphasis>
|
||
In contrast to
|
||
<taxonomicName id="4C174D7DFF9A7F1A9E8BF88D5354F884" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[747,885,1845,1871]" genus="Oculogaster" pageId="5" pageNumber="587" rank="subGenus" status="subgen. nov." subGenus="Oculogaster">
|
||
<emphasis id="B963EAECFF9A7F1A9E8BF88D5354F884" box="[747,885,1845,1871]" italics="true" pageId="5" pageNumber="578">Oculogaster</emphasis>
|
||
</taxonomicName>
|
||
, pterostigma with several cross veins (
|
||
<figureCitation id="132C2A7BFF9A7F1A9959F88D55AEF884" box="[1337,1423,1845,1871]" captionStart="FIGURES 39–46" captionStartId="10.[151,264,1885,1910]" captionTargetBox="[151,1436,181,1860]" captionTargetId="figure@10.[151,1436,181,1860]" captionTargetPageId="10" captionText="FIGURES 39–46. Procloeon (Monilistylus) ornatipennis sp. n.: 39–40, male imagines; 41, male imaginal genitals; 42, exuviae of right half of subimaginal mesonotum; 43, exuviae of left half of subimaginal mesopleuron; 44, wing; 45, abdominal terga II–VII; 46, tibia and tarsus of middle leg (41–43 and 45–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3677931" httpUri="https://zenodo.org/record/3677931/files/figure.png" pageId="5" pageNumber="578">Fig. 44</figureCitation>
|
||
), eggs with sculptured chorion (known for
|
||
<taxonomicName id="4C174D7DFF9A7F1A9E01F8E2531FF8B8" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[609,830,1881,1907]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="monilistylus" subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9A7F1A9E01F8E25253F8B8" box="[609,626,1882,1907]" italics="true" pageId="5" pageNumber="578">P</emphasis>
|
||
. (
|
||
<emphasis id="B963EAECFF9A7F1A9EE9F8E25281F8B8" box="[649,672,1882,1907]" italics="true" pageId="5" pageNumber="578">M</emphasis>
|
||
.)
|
||
<emphasis id="B963EAECFF9A7F1A9ED7F8E1531FF8B8" box="[695,830,1881,1907]" italics="true" pageId="5" pageNumber="578">monilistylus</emphasis>
|
||
</taxonomicName>
|
||
only). In contrast to
|
||
<taxonomicName id="4C174D7DFF9A7F1A9844F8E1553CF8B8" ID-CoL="6ZGW" box="[1060,1309,1881,1907]" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9844F8E1553CF8B8" box="[1060,1309,1881,1907]" italics="true" pageId="5" pageNumber="578">Pseudocentroptiloides</emphasis>
|
||
</taxonomicName>
|
||
, the maxillae are narrowed distally (
|
||
<figureCitation id="132C2A7BFF9A7F1A9DDDF8C5522EF85C" box="[445,527,1917,1943]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="5" pageNumber="578">Fig. 17</figureCitation>
|
||
), the labrum is not deeply emarginated (
|
||
<figureCitation id="132C2A7BFF9A7F1A9FB7F8C5540AF853" box="[983,1067,1917,1944]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="5" pageNumber="578">Fig. 16</figureCitation>
|
||
). In contrast to
|
||
<taxonomicName id="4C174D7DFF9A7F1A98BCF8C550FBF870" ID-CoL="6ZGW" class="Insecta" family="Baetidae" genus="Pseudocentroptiloides" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A98BCF8C550FBF870" italics="true" pageId="5" pageNumber="578">Pseudocentroptiloides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C174D7DFF9A7F1A9D70F819515FF870" ID-CoL="7F34" authorityName="Jacobus, McCafferty & Gattolliat" authorityYear="2006" box="[272,382,1953,1979]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9A7F1A9D70F819515FF870" box="[272,382,1953,1979]" italics="true" pageId="5" pageNumber="578">Securiops</emphasis>
|
||
</taxonomicName>
|
||
, the glossae are not much longer than the paraglossae (
|
||
<figureCitation id="132C2A7BFF9A7F1A9F89F819544CF870" box="[1001,1133,1953,1979]" captionStart="FIGURES 26–29" captionStartId="6.[151,264,1811,1836]" captionTargetBox="[151,1436,236,1786]" captionTargetId="figure@6.[151,1436,236,1786]" captionTargetPageId="6" captionText="FIGURES 26–29. Procloeon (Monilistylus) ornatipennis sp. n., exuviae of labium: 26, general view with focus on ventral side; 27, labial palp with focus on dorsal side; 28, glossa and paraglossa with focus on ventral side; 29, the same with focus on dorsal side (26, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3677925" httpUri="https://zenodo.org/record/3677925/files/figure.png" pageId="5" pageNumber="578">Figs 28–29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D6575FF9A7F199CA7F87C545DFF1C" lastPageId="6" lastPageNumber="579" pageId="5" pageNumber="578" type="distribution">
|
||
<paragraph id="8BA836FEFF9A7F1A9CA7F87C534AF814" blockId="5.[151,1437,476,2015]" box="[199,875,1988,2015]" pageId="5" pageNumber="578">
|
||
<emphasis id="B963EAECFF9A7F1A9CA7F87C5140F814" bold="true" box="[199,353,1988,2015]" pageId="5" pageNumber="578">Distribution.</emphasis>
|
||
Oriental Region and south-east of Palaearctic.
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF997F199CA7FF20545DFF1C" blockId="6.[151,1436,152,215]" pageId="6" pageNumber="579">
|
||
<emphasis id="B963EAECFF997F199CA7FF20519EFF78" bold="true" box="[199,447,152,179]" pageId="6" pageNumber="579">Species composition.</emphasis>
|
||
Two species:
|
||
<taxonomicName id="4C174D7DFF997F199E03FF2055BDFF78" authority="monilistylus Kluge, Tiunova & Novikova 2014" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[611,1436,152,179]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="monilistylus" subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF997F199E03FF2052F6FF79" box="[611,727,152,178]" italics="true" pageId="6" pageNumber="579">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF997F199E89FF205352FF79" box="[745,883,152,178]" italics="true" pageId="6" pageNumber="579">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF997F199FE5FF20542DFF79" box="[901,1036,152,178]" italics="true" pageId="6" pageNumber="579">monilistylus</emphasis>
|
||
<bibRefCitation id="EF864B0FFF997F199875FF2055BDFF78" author="Kluge, N. J. & Tiunova, T. M. & Novikova, E. A." box="[1045,1436,152,179]" pageId="6" pageNumber="579" pagination="483 - 491" refId="ref7118" refString="Kluge, N. J., Tiunova, T. M. & Novikova, E. A. (2014) A new species, Procloeon monilistylus sp. n. (Ephemeroptera, Baetidae), from the Russian Far East. Zootaxa, 3786 (4), 483 - 491. https: // doi. org / 10.11646 / zootaxa. 3786.4.6" type="journal article" year="2014">Kluge, Tiunova & Novikova 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(south of Russian Far East) and
|
||
<taxonomicName id="4C174D7DFF997F199D9EFF04538AFF1D" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[510,939,188,215]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="579" phylum="Arthropoda" rank="species" species="ornatipennis" status="sp. nov." subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF997F199D9EFF045253FF1D" box="[510,626,188,214]" italics="true" pageId="6" pageNumber="579">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF997F199EE2FF04532DFF1D" box="[642,780,188,214]" italics="true" pageId="6" pageNumber="579">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF997F199F7DFF05538AFF1D" box="[797,939,189,214]" italics="true" pageId="6" pageNumber="579">ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2505797FF997F199FD2FF0553CFFF1C" box="[946,1006,189,215]" pageId="6" pageNumber="579" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<collectingCountry id="F300766EFF997F199F9CFF045451FF1C" box="[1020,1136,188,215]" name="Indonesia" pageId="6" pageNumber="579">Indonesia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |