treatments-xml/data/03/B4/87/03B4878D181DFFEAB6C6FCD0FD4FFAAE.xml

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<mods:title>Caridina simoni Bouvier, 1904 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) and the synonymy by Johnson, 1963</mods:title>
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<mods:namePart>Richard, Jasmine</mods:namePart>
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<mods:namePart>Clark, Paul F.</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2014</mods:date>
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<mods:number>2014-07-28</mods:number>
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<mods:number>3841</mods:number>
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<taxonomicName ID-CoL="8VK4V" ID-ENA="382064" authority="De Man, 1908" authorityName="De Man" authorityYear="1908" box="[151,616,870,896]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis bold="true" box="[151,616,870,896]" pageId="7" pageNumber="308">
<emphasis bold="true" box="[151,435,870,896]" italics="true" pageId="7" pageNumber="308">Caridina brachydactyla</emphasis>
<bibRefCitation author="De Man" box="[442,616,870,896]" firstAuthor="De Man" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="308" pagination="219 - 283" part="2" refId="ref24738" refString="Man, J. G. de (1908 a) The fauna of brackish ponds at port Canning, Lower Bengal. Records of the Indian Museum, 2, 219 - 283." title="The fauna of brackish ponds at port Canning, Lower Bengal" type="journal article" year="1908">
De Man, 
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(
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Fig. 
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)
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<taxonomicName ID-CoL="8VKH6" authority="De Man, 1892: 386" authorityName="De Man" authorityPageNumber="386" authorityYear="1892" box="[151,516,977,999]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="wyckii">
<emphasis box="[151,314,977,999]" italics="true" pageId="7" pageNumber="308">Caridina wyckii</emphasis>
<bibRefCitation author="De Man" box="[320,516,977,999]" firstAuthor="De Man" page="386" pageId="7" pageNumber="308" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. In: Weber, M. Zoologische Ergebnisse einer Reise in Netherlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">
De 
<treatmentCitation author="Man" box="[355,516,977,999]" page="386" pageId="7" pageNumber="308" year="1892">
Man, 
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: 
<number box="[478,516,977,999]" pageId="7" pageNumber="308" value="386.0">386</number>
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(partim), pl. 
<number box="[650,678,977,999]" pageId="7" pageNumber="308" value="24.0">24</number>
, fig. 
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<number box="[729,756,977,999]" pageId="7" pageNumber="308" value="29.0">29</number>
f.
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<paragraph blockId="7.[151,1436,977,1275]" box="[151,1142,1007,1029]" pageId="7" pageNumber="308">
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<taxonomicName ID-CoL="8V757" authority="De Man, 1908: 269" authorityName="De Man" authorityPageNumber="269" authorityYear="1908" box="[151,727,1007,1029]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[151,326,1007,1029]" italics="true" pageId="7" pageNumber="308">Caridina nilotica</emphasis>
var. 
<emphasis box="[376,521,1007,1029]" italics="true" pageId="7" pageNumber="308">brachydactyla</emphasis>
De Man, 
<number box="[622,676,1007,1029]" pageId="7" pageNumber="308" value="1908.0">1908</number>
: 
<number box="[686,727,1007,1029]" pageId="7" pageNumber="308" value="269.0">269</number>
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, pl. 
<number box="[769,797,1007,1029]" pageId="7" pageNumber="308" value="20.0">20</number>
, fig. 
<number box="[847,859,1007,1029]" pageId="7" pageNumber="308" value="8.0">8</number>
a–c;? 
<treatmentCitation author="Johnson" box="[926,1138,1007,1029]" page="123" pageId="7" pageNumber="308" year="1961">
<bibRefCitation author="Johnson" box="[926,1138,1007,1029]" firstAuthor="Johnson" page="123" pageId="7" pageNumber="308" refString="Johnson, D. S. (1961 a) Notes on the freshwater Crustacea of Malaya, I. The Atyidae. Bulletin of the Raffles Museum, 26, 120 - 153." type="journal article" year="1961" yearSuffix="a">
Johnson, 
<number box="[1022,1076,1007,1029]" pageId="7" pageNumber="308" value="1961.0">1961</number>
a: 
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.
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<paragraph blockId="7.[151,1436,977,1275]" box="[151,744,1038,1060]" pageId="7" pageNumber="308">
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<taxonomicName ID-CoL="8V757" box="[151,504,1038,1060]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[151,326,1038,1060]" italics="true" pageId="7" pageNumber="308">Caridina nilotica</emphasis>
var. 
<emphasis box="[376,504,1038,1060]" italics="true" pageId="7" pageNumber="308">meridionalis</emphasis>
</taxonomicName>
. 
<treatmentCitation author="Reik" box="[516,674,1038,1060]" page="117" pageId="7" pageNumber="308" year="1953">
<bibRefCitation author="Reik" box="[516,674,1038,1060]" firstAuthor="Reik" page="117" pageId="7" pageNumber="308" refString="Reik, E. F. (1953) The Australian freshwater prawns of the family Atyidae. Records of the Australian Museum, 23, 111 - 121. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.23.1953.625" type="journal article" year="1953">
Reik 
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: 
<number box="[635,674,1038,1060]" pageId="7" pageNumber="308" value="117.0">117</number>
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, fig. 
<number box="[725,740,1038,1060]" pageId="7" pageNumber="308" value="6.0">6</number>
.
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<paragraph blockId="7.[151,1436,977,1275]" pageId="7" pageNumber="308">
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<taxonomicName ID-CoL="8VK4V" authority="De Man, 1908" box="[151,398,1069,1091]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[151,398,1069,1091]" italics="true" pageId="7" pageNumber="308">Caridina brachydactyla</emphasis>
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.? 
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<bibRefCitation author="Johnson" box="[438,660,1069,1091]" firstAuthor="Johnson" page="178" pageId="7" pageNumber="308" refString="Johnson, D. S. (1960 a) Some aspects of the distribution of freshwater organisms in the Indo-Pacific Area, and their relevance to the validity of the concept of an oriental region on zoogeography. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 170 - 181." type="journal article" year="1960" yearSuffix="a">
Johnson, 
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a: 
<number box="[619,660,1069,1091]" pageId="7" pageNumber="308" value="178.0">178</number>
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; 
<bibRefCitation author="Johnson" box="[675,798,1069,1091]" firstAuthor="Johnson" journalOrPublisher="Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore" page="266" pageId="7" pageNumber="308" pagination="261 - 267" part="1958" refId="ref24457" refString="Johnson, D. S. (1960 b) Sub-specific and infra specific variation in some freshwater prawns of the Indo-Pacific region. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 261 - 267." title="Sub-specific and infra specific variation in some freshwater prawns of the Indo-Pacific region" type="journal article" year="1960" yearSuffix="b">
<number box="[675,729,1069,1091]" pageId="7" pageNumber="308" value="1960.0">1960</number>
b: 
<number box="[757,798,1069,1091]" pageId="7" pageNumber="308" value="266.0">266</number>
</bibRefCitation>
; 
<bibRefCitation author="Johnson" box="[813,910,1069,1091]" firstAuthor="Johnson" journalOrPublisher="Bulletin of the National Museum of Singapore" page="21" pageId="7" pageNumber="308" pagination="5 - 30" part="32" refId="ref24584" refString="Johnson, D. S. (1963) Distribution and other notes on some freshwater prawns (Atyidae and Palaemonidae) mainly from the Indo-west Pacific region. Bulletin of the National Museum of Singapore, 32, 5 - 30." title="Distribution and other notes on some freshwater prawns (Atyidae and Palaemonidae) mainly from the Indo-west Pacific region" type="journal article" year="1963">
<number box="[813,867,1069,1091]" pageId="7" pageNumber="308" value="1963.0">1963</number>
: 
<number box="[882,910,1069,1091]" pageId="7" pageNumber="308" value="21.0">21</number>
</bibRefCitation>
; 
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<bibRefCitation author="Richard &amp; Clark" box="[925,1231,1069,1091]" firstAuthor="Richard" page="317" pageId="7" pageNumber="308" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark, 
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: 
<number box="[1191,1231,1069,1091]" pageId="7" pageNumber="308" value="317.0">317</number>
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, figs. 
<number box="[1302,1340,1069,1091]" pageId="7" pageNumber="308" value="7.5" valueMax="8.0" valueMin="7.0">7–8</number>
(for full synonymy).
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</paragraph>
<paragraph blockId="7.[151,1436,977,1275]" box="[151,684,1130,1152]" pageId="7" pageNumber="308">
<treatmentCitationGroup box="[151,684,1130,1152]" pageId="7" pageNumber="308">
<taxonomicName ID-CoL="8VKBR" authority="J. Roux, 1926b: 246" authorityName="J. Roux" authorityPageNumber="246" authorityYear="1926" box="[151,587,1130,1152]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[151,377,1130,1152]" italics="true" pageId="7" pageNumber="308">Caridina meridionalis</emphasis>
<treatmentCitation author="Roux" box="[383,587,1130,1152]" page="246" pageId="7" pageNumber="308" year="1926">
<bibRefCitation author="Roux" box="[383,539,1130,1152]" pageId="7" pageNumber="308" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">
J. 
<bibRefCitation author="Roux" box="[405,587,1130,1152]" firstAuthor="Roux" page="246" pageId="7" pageNumber="308" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">
Roux, 
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b: 
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(partim).
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</paragraph>
<paragraph blockId="7.[151,1436,977,1275]" box="[151,865,1161,1183]" pageId="7" pageNumber="308">
<treatmentCitationGroup box="[151,865,1161,1183]" pageId="7" pageNumber="308">
<taxonomicName ID-CoL="8VK4V" authority="Tiwari &amp; Pillai, 1971:80" authorityName="Tiwari &amp; Pillai" authorityPageNumber="80" authorityYear="1971" box="[151,801,1161,1183]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="subSpecies" species="brachydactyla" subSpecies="brachydactyla">
<emphasis box="[151,545,1161,1183]" italics="true" pageId="7" pageNumber="308">Caridina brachydactyla brachydactyla</emphasis>
<treatmentCitation author="Tiwari" box="[551,801,1161,1183]" page="80" pageId="7" pageNumber="308" year="1971">
<bibRefCitation author="Tiwari &amp; Pillai" box="[551,801,1161,1183]" firstAuthor="Tiwari" page="80" pageId="7" pageNumber="308" refString="Tiwari, K. K. &amp; Pillai, R. S. (1971) Atyid shrimps of the genus Caridina H. Milne Edwards, 1837, from the Andaman Islands (Decapoda, Caridea). Crustaceana, 21, 79 - 91. http: // dx. doi. org / 10.1163 / 156854071 x 00247" type="journal article" year="1971">
Tiwari &amp; Pillai, 
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: 
<number box="[773,801,1161,1183]" pageId="7" pageNumber="308" value="80.0">80</number>
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</taxonomicName>
, fig. 
<number box="[852,861,1161,1183]" pageId="7" pageNumber="308" value="1.0">1</number>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[151,1436,977,1275]" box="[151,1406,1191,1213]" pageId="7" pageNumber="308">
<treatmentCitationGroup box="[151,1406,1191,1213]" pageId="7" pageNumber="308">
Nec 
<taxonomicName ID-CoL="8VKH6" authority="De Man, 1892: 386" authorityName="De Man" authorityPageNumber="386" authorityYear="1892" box="[198,563,1191,1213]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="wyckii">
<emphasis box="[198,361,1191,1213]" italics="true" pageId="7" pageNumber="308">Caridina wyckii</emphasis>
<bibRefCitation author="De Man" box="[367,563,1191,1213]" firstAuthor="De Man" page="386" pageId="7" pageNumber="308" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. In: Weber, M. Zoologische Ergebnisse einer Reise in Netherlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">
De 
<treatmentCitation author="Man" box="[403,563,1191,1213]" page="386" pageId="7" pageNumber="308" year="1892">
Man, 
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: 
<number box="[525,563,1191,1213]" pageId="7" pageNumber="308" value="386.0">386</number>
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</taxonomicName>
(partim), fig. 
<quantity box="[706,744,1191,1213]" metricMagnitude="-2" metricUnit="kg" metricValue="2.9" pageId="7" pageNumber="308" unit="g" value="29.0">29g</quantity>
(= undescribed species A, see 
<bibRefCitation author="Richard &amp; Clark" box="[1057,1293,1191,1213]" firstAuthor="Richard" pageId="7" pageNumber="308" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark, 
<number box="[1238,1293,1191,1213]" pageId="7" pageNumber="308" value="2010.0">2010</number>
</bibRefCitation>
: fig. 
<number box="[1343,1356,1191,1213]" pageId="7" pageNumber="308" value="9.0">9</number>
a–c).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="7.[151,1436,977,1275]" pageId="7" pageNumber="308">
<treatmentCitationGroup pageId="7" pageNumber="308">
Nec 
<taxonomicName ID-CoL="8VKH6" authority="De Man, 1892: 386" authorityName="De Man" authorityPageNumber="386" authorityYear="1892" box="[198,564,1222,1244]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="wyckii">
<emphasis box="[198,361,1222,1244]" italics="true" pageId="7" pageNumber="308">Caridina wyckii</emphasis>
<bibRefCitation author="De Man" box="[367,564,1222,1244]" firstAuthor="De Man" page="386" pageId="7" pageNumber="308" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. In: Weber, M. Zoologische Ergebnisse einer Reise in Netherlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">
De 
<treatmentCitation author="Man" box="[403,564,1222,1244]" page="386" pageId="7" pageNumber="308" year="1892">
Man, 
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: 
<number box="[526,564,1222,1244]" pageId="7" pageNumber="308" value="386.0">386</number>
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(partim), pl. 
<number box="[698,726,1222,1244]" pageId="7" pageNumber="308" value="24.0">24</number>
, fig. 
<date box="[777,844,1222,1244]" pageId="7" pageNumber="308" value="1929-01" valueMax="1929-02-31" valueMin="1929-01-29">
<number box="[777,803,1222,1244]" pageId="7" pageNumber="308" value="29.0">29</number>
i–ii.
</date>
(= undescribed species B, see 
<bibRefCitation author="Richard &amp; Clark" box="[1155,1392,1222,1244]" firstAuthor="Richard" pageId="7" pageNumber="308" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark, 
<number box="[1337,1392,1222,1244]" pageId="7" pageNumber="308" value="2010.0">2010</number>
</bibRefCitation>
: fig. 
<number box="[199,212,1253,1275]" pageId="7" pageNumber="308" value="9.0">9</number>
d–e).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="308" type="materials_examined">
<paragraph blockId="7.[151,1437,1319,2031]" pageId="7" pageNumber="308">
<emphasis bold="true" box="[151,385,1319,1344]" pageId="7" pageNumber="308">Material examined.</emphasis>
<typeStatus box="[392,457,1320,1344]" pageId="7" pageNumber="308">Types</typeStatus>
: 
<materialsCitation ID-GBIF-Occurrence="4029644313" pageId="7" pageNumber="308">
<taxonomicName box="[474,594,1319,1344]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[474,493,1320,1343]" italics="true" pageId="7" pageNumber="308">C</emphasis>
. 
<emphasis box="[508,594,1319,1344]" italics="true" pageId="7" pageNumber="308">nilotica</emphasis>
</taxonomicName>
<taxonomicName box="[601,810,1319,1344]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="variety" species="brachydactyla" variety="brachydactyla">
var. 
<emphasis box="[650,810,1319,1344]" italics="true" pageId="7" pageNumber="308">brachydactyla</emphasis>
</taxonomicName>
, 
<collectingCountry box="[825,941,1319,1344]" name="Indonesia" pageId="7" pageNumber="308">
<emphasis bold="true" box="[825,941,1319,1344]" pageId="7" pageNumber="308">Indonesia</emphasis>
</collectingCountry>
. 
<typeStatus box="[956,1072,1320,1344]" pageId="7" pageNumber="308">Lectotype</typeStatus>
. 
<specimenCount box="[1082,1117,1320,1344]" pageId="7" pageNumber="308" type="generic">
<number box="[1082,1093,1320,1344]" pageId="7" pageNumber="308" value="1.0">1</number>
♂
</specimenCount>
(with rostral formula (
<number box="[1376,1391,1320,1344]" pageId="7" pageNumber="308" value="2.0">2</number>
) 
<number box="[1408,1436,1320,1344]" pageId="7" pageNumber="308" value="19.0">19</number>
+ 
<number box="[173,183,1356,1380]" pageId="7" pageNumber="308" value="1.0">1</number>
/ 
<number box="[193,225,1356,1380]" pageId="7" pageNumber="308" value="23.0">23</number>
),
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4029644318" box="[246,1048,1355,1380]" pageId="7" pageNumber="308">
<typeStatus box="[246,413,1355,1380]" pageId="7" pageNumber="308">Paralectotypes</typeStatus>
. 
<specimenCount box="[422,459,1356,1380]" pageId="7" pageNumber="308" type="generic">
<number box="[422,438,1356,1380]" pageId="7" pageNumber="308" value="4.0">4</number>
♂
</specimenCount>
, 
<number box="[471,485,1356,1380]" pageId="7" pageNumber="308" value="1.0">1</number>
♀ ovig., by Reo, 
<collectingCountry box="[676,749,1355,1380]" name="Indonesia" pageId="7" pageNumber="308">Flores</collectingCountry>
, 
<collectionCode LSID="urn:lsid:biocol.org:col:34992" box="[759,843,1356,1379]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="308">RMNH</collectionCode>
reg. Coll NR 
<number box="[999,1044,1356,1380]" pageId="7" pageNumber="308" value="977.0">977</number>
;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="4029644309" pageId="7" pageNumber="308">
<number box="[1055,1073,1356,1380]" pageId="7" pageNumber="308" value="2.0">2</number>
♀ ovig., river near Palopa Luwu, Celebes (Sulawesi), coll. Max Webber, 
<collectionCode LSID="urn:lsid:biocol.org:col:34992" box="[594,678,1392,1415]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="308">RMNH</collectionCode>
reg. Coll NR 
<number box="[836,896,1391,1416]" pageId="7" pageNumber="308" value="2552.0">2552</number>
.
</materialsCitation>
</paragraph>
<paragraph blockId="7.[151,1437,1319,2031]" pageId="7" pageNumber="308">
Non 
<typeStatus box="[261,320,1428,1452]" pageId="7" pageNumber="308">types</typeStatus>
:
<materialsCitation ID-GBIF-Occurrence="4029644314" pageId="7" pageNumber="308">
? 
<collectingCountry box="[370,516,1427,1452]" name="Madagascar" pageId="7" pageNumber="308">
<emphasis bold="true" box="[370,516,1427,1452]" pageId="7" pageNumber="308">Madagascar</emphasis>
</collectingCountry>
. det. E.L. Bouvier 
<number box="[760,819,1428,1452]" pageId="7" pageNumber="308" value="1918.0">1918</number>
, received from E.L. Bouvier 
<date box="[1184,1301,1428,1452]" pageId="7" pageNumber="308" value="1920-01">
Jan. 
<number box="[1243,1301,1428,1452]" pageId="7" pageNumber="308" value="1920.0">1920</number>
</date>
, NHM not registered, 
<specimenCount box="[275,313,1464,1488]" pageId="7" pageNumber="308" type="generic">
<number box="[275,290,1464,1488]" pageId="7" pageNumber="308" value="2.0">2</number>
♂
</specimenCount>
, 
<number box="[324,341,1463,1488]" pageId="7" pageNumber="308" value="5.0">5</number>
♀ ovig., 
<number box="[437,455,1463,1488]" pageId="7" pageNumber="308" value="5.0">5</number>
♀.
</materialsCitation>
</paragraph>
<paragraph blockId="7.[151,1437,1319,2031]" pageId="7" pageNumber="308">
Other material: 
<materialsCitation ID-GBIF-Occurrence="4029644312" pageId="7" pageNumber="308">
<taxonomicName box="[380,574,1499,1524]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[380,574,1499,1524]" italics="true" pageId="7" pageNumber="308">Caridina nilotica</emphasis>
</taxonomicName>
<taxonomicName box="[584,775,1499,1524]" pageId="7" pageNumber="308" rank="variety" variety="meridionalis">
var. 
<emphasis box="[634,775,1499,1524]" italics="true" pageId="7" pageNumber="308">meridionalis</emphasis>
</taxonomicName>
. 
<typeStatus box="[791,858,1500,1524]" pageId="7" pageNumber="308">Types</typeStatus>
: Cotype. 
<emphasis bold="true" box="[971,1090,1499,1524]" pageId="7" pageNumber="308">
<collectingCountry box="[971,1085,1499,1524]" name="Australia" pageId="7" pageNumber="308">Australia</collectingCountry>
.
</emphasis>
Cari Creek at Cook Town, N. Queensland; exch. with Sydney; coll. A.R. McCulloch, 
<date box="[782,900,1536,1560]" pageId="7" pageNumber="308" value="1918-05">
May 
<number box="[841,900,1536,1560]" pageId="7" pageNumber="308" value="1918.0">1918</number>
</date>
; originally P. 
<number box="[1049,1108,1535,1560]" pageId="7" pageNumber="308" value="4296.0">4296</number>
, now 
<collectionCode box="[1176,1240,1536,1559]" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="308">NMB</collectionCode>
reg. 
<date box="[1298,1374,1536,1560]" pageId="7" pageNumber="308">
<number box="[1298,1313,1536,1560]" pageId="7" pageNumber="308" value="4.0">4</number>
.VIII.
</date>
a 
<specimenCount box="[1394,1431,1536,1560]" pageId="7" pageNumber="308" type="generic">
<number box="[1394,1405,1536,1560]" pageId="7" pageNumber="308" value="1.0">1</number>
♂
</specimenCount>
, 
<number box="[151,165,1572,1596]" pageId="7" pageNumber="308" value="1.0">1</number>
♀ ovig., 
<number box="[264,283,1572,1596]" pageId="7" pageNumber="308" value="2.0">2</number>
♀.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[199,1380,1607,1632]" pageId="7" pageNumber="308" type="distribution">
<paragraph blockId="7.[151,1437,1319,2031]" box="[199,1380,1607,1632]" pageId="7" pageNumber="308">
<emphasis bold="true" box="[199,352,1607,1632]" pageId="7" pageNumber="308">Distribution.</emphasis>
Celebes (Sulawesi), 
<collectingCountry box="[589,701,1607,1632]" name="Indonesia" pageId="7" pageNumber="308">Indonesia</collectingCountry>
; Andaman Islands; 
<collectingCountry box="[922,1068,1607,1632]" name="South Africa" pageId="7" pageNumber="308">South Africa</collectingCountry>
, 
<collectingCountry box="[1080,1215,1607,1632]" name="Madagascar" pageId="7" pageNumber="308">Madagascar</collectingCountry>
and 
<collectingCountry box="[1270,1376,1607,1632]" name="Australia" pageId="7" pageNumber="308">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,925,1643,1668]" pageId="7" pageNumber="308" type="materials_examined">
<paragraph blockId="7.[151,1437,1319,2031]" box="[199,925,1643,1668]" pageId="7" pageNumber="308">
<emphasis bold="true" box="[199,357,1643,1668]" pageId="7" pageNumber="308">
<typeStatus box="[199,258,1643,1668]" pageId="7" pageNumber="308">Type</typeStatus>
locality.
</emphasis>
Celebes (Sulawesi) Selayar and 
<collectingCountry box="[725,922,1643,1668]" name="Indonesia" pageId="7" pageNumber="308">Flores, Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308" type="discussion">
<paragraph blockId="7.[151,1437,1319,2031]" pageId="7" pageNumber="308">
<emphasis bold="true" box="[199,308,1679,1704]" pageId="7" pageNumber="308">Remarks</emphasis>
. While studying 
<taxonomicName box="[531,742,1679,1704]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="niliotica">
<emphasis box="[531,742,1679,1704]" italics="true" pageId="7" pageNumber="308">Caridina niliotica</emphasis>
</taxonomicName>
and its varieties, 
<bibRefCitation author="De Man" box="[980,1172,1680,1704]" firstAuthor="De Man" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="308" pagination="219 - 283" part="2" refId="ref24738" refString="Man, J. G. de (1908 a) The fauna of brackish ponds at port Canning, Lower Bengal. Records of the Indian Museum, 2, 219 - 283." title="The fauna of brackish ponds at port Canning, Lower Bengal" type="journal article" year="1908">
De Man (
<number box="[1106,1164,1680,1704]" pageId="7" pageNumber="308" value="1908.0">1908</number>
)
</bibRefCitation>
described 
<emphasis box="[1314,1377,1680,1704]" italics="true" pageId="7" pageNumber="308">C. n.</emphasis>
<taxonomicName class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="variety" species="brachydactyla" variety="brachydactyla">
var. 
<emphasis box="[151,311,1715,1740]" italics="true" pageId="7" pageNumber="308">brachydactyla</emphasis>
</taxonomicName>
from specimens that he earlier considered as the typical form of 
<taxonomicName authority="Hickson" authorityName="Hickson" box="[1071,1379,1715,1740]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="wyckii">
<emphasis box="[1071,1253,1715,1740]" italics="true" pageId="7" pageNumber="308">Caridina wyckii</emphasis>
(Hickson)
</taxonomicName>
. The new variety was characterised by the slender carpus of the first pereiopod and the stout dactylus (
<number box="[1308,1318,1754,1779]" pageId="7" pageNumber="308" value="1.0">1</number>
/ 
<number box="[1329,1344,1754,1779]" pageId="7" pageNumber="308" value="6.0">6</number>
th of the propodus) of the three posterior legs. Later, 
<bibRefCitation author="Tiwari and Pillai" box="[642,910,1790,1815]" firstAuthor="Tiwari" pageId="7" pageNumber="308" refString="Tiwari, K. K. &amp; Pillai, R. S. (1971) Atyid shrimps of the genus Caridina H. Milne Edwards, 1837, from the Andaman Islands (Decapoda, Caridea). Crustaceana, 21, 79 - 91. http: // dx. doi. org / 10.1163 / 156854071 x 00247" type="journal article" year="1971">
Tiwari and Pillai (
<number box="[844,902,1791,1815]" pageId="7" pageNumber="308" value="1971.0">1971</number>
)
</bibRefCitation>
considered 
<taxonomicName box="[1045,1265,1790,1815]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="subSpecies" species="brachydactyla" subSpecies="brachydactyla">
<emphasis box="[1045,1265,1790,1815]" italics="true" pageId="7" pageNumber="308">C. n. brachydactyla</emphasis>
</taxonomicName>
to be a distinct species thereby rejecting the decision of 
<bibRefCitation author="Johnson" box="[607,780,1826,1851]" firstAuthor="Johnson" pageId="7" pageNumber="308" refString="Johnson, D. S. (1963) Distribution and other notes on some freshwater prawns (Atyidae and Palaemonidae) mainly from the Indo-west Pacific region. Bulletin of the National Museum of Singapore, 32, 5 - 30." type="journal article" year="1963">
Johnson (
<number box="[714,772,1826,1851]" pageId="7" pageNumber="308" value="1963.0">1963</number>
)
</bibRefCitation>
to make this subspecies a junior synonym of 
<emphasis box="[1291,1406,1827,1850]" italics="true" pageId="7" pageNumber="308">
<taxonomicName box="[1291,1402,1827,1850]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="simoni">C. simoni</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="7.[151,1437,1319,2031]" pageId="7" pageNumber="308">
<bibRefCitation author="Richard &amp; Clark" box="[199,484,1862,1887]" firstAuthor="Richard" pageId="7" pageNumber="308" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark (
<number box="[415,476,1863,1887]" pageId="7" pageNumber="308" value="2010.0">2010</number>
)
</bibRefCitation>
, who reported and described 
<taxonomicName box="[836,1034,1862,1887]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[836,1034,1862,1887]" italics="true" pageId="7" pageNumber="308">C. brachydactyla</emphasis>
</taxonomicName>
from 
<collectingCountry box="[1112,1264,1862,1887]" name="South Africa" pageId="7" pageNumber="308">South Africa</collectingCountry>
, supported the decision of 
<bibRefCitation author="Tiwari &amp; Pillai" box="[281,529,1898,1923]" firstAuthor="Tiwari" pageId="7" pageNumber="308" refString="Tiwari, K. K. &amp; Pillai, R. S. (1971) Atyid shrimps of the genus Caridina H. Milne Edwards, 1837, from the Andaman Islands (Decapoda, Caridea). Crustaceana, 21, 79 - 91. http: // dx. doi. org / 10.1163 / 156854071 x 00247" type="journal article" year="1971">
Tiwari &amp; Pillai (
<number box="[463,521,1899,1923]" pageId="7" pageNumber="308" value="1971.0">1971</number>
)
</bibRefCitation>
and considered this species to be distinct from 
<taxonomicName box="[1056,1164,1899,1922]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[1056,1164,1899,1922]" italics="true" pageId="7" pageNumber="308">C. simoni</emphasis>
</taxonomicName>
. They (
<bibRefCitation author="Richard &amp; Clark" firstAuthor="Richard" pageId="7" pageNumber="308" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark 
<number box="[151,209,1935,1959]" pageId="7" pageNumber="308" value="2010.0">2010</number>
</bibRefCitation>
) synonymised the following taxa as junior synonyms of 
<taxonomicName box="[854,1048,1934,1959]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[854,1048,1934,1959]" italics="true" pageId="7" pageNumber="308">C. brachydactyla</emphasis>
</taxonomicName>
: 
<taxonomicName box="[1064,1169,1934,1959]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="wyckii">
<emphasis box="[1064,1169,1934,1959]" italics="true" pageId="7" pageNumber="308">C. wyckii</emphasis>
</taxonomicName>
of 
<bibRefCitation author="De Man" box="[1209,1368,1935,1959]" firstAuthor="De Man" pageId="7" pageNumber="308" refString="Man, J. G. de (1892) Decapoden des Indischen Archipels. In: Weber, M. Zoologische Ergebnisse einer Reise in Netherlandisch Ost-Indien, 2, 265 - 527." type="journal article" year="1892">
De Man 
<number box="[1309,1368,1935,1959]" pageId="7" pageNumber="308" value="1892.0">1892</number>
</bibRefCitation>
, 
<emphasis box="[1381,1436,1935,1958]" italics="true" pageId="7" pageNumber="308">C. n.</emphasis>
<taxonomicName box="[151,363,1970,1995]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="variety" species="brachydactyla" variety="brachydactyla">
var. 
<emphasis box="[203,363,1970,1995]" italics="true" pageId="7" pageNumber="308">brachydactyla</emphasis>
</taxonomicName>
of 
<bibRefCitation author="De Man" box="[408,573,1971,1995]" firstAuthor="De Man" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="308" pagination="219 - 283" part="2" refId="ref24738" refString="Man, J. G. de (1908 a) The fauna of brackish ponds at port Canning, Lower Bengal. Records of the Indian Museum, 2, 219 - 283." title="The fauna of brackish ponds at port Canning, Lower Bengal" type="journal article" year="1908">
De Man 
<number box="[514,573,1971,1995]" pageId="7" pageNumber="308" value="1908.0">1908</number>
</bibRefCitation>
, 
<emphasis box="[588,646,1971,1994]" italics="true" pageId="7" pageNumber="308">C. n.</emphasis>
<taxonomicName box="[657,869,1970,1995]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="variety" species="brachydactyla" variety="brachydactyla">
var. 
<emphasis box="[709,869,1970,1995]" italics="true" pageId="7" pageNumber="308">brachydactyla</emphasis>
</taxonomicName>
of 
<bibRefCitation author="Johnson" box="[915,1088,1970,1995]" firstAuthor="Johnson" pageId="7" pageNumber="308" refString="Johnson, D. S. (1961 a) Notes on the freshwater Crustacea of Malaya, I. The Atyidae. Bulletin of the Raffles Museum, 26, 120 - 153." type="journal article" year="1961" yearSuffix="a">
Johnson 
<number box="[1017,1075,1970,1995]" pageId="7" pageNumber="308" value="1961.0">1961</number>
a
</bibRefCitation>
, 
<taxonomicName box="[1104,1300,1970,1995]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1104,1300,1970,1995]" italics="true" pageId="7" pageNumber="308">C. brachydactyla</emphasis>
</taxonomicName>
of 
<bibRefCitation author="Bouvier" firstAuthor="Bouvier" pageId="7" pageNumber="308" refString="Bouvier, E. L. (1913 a) Les caridines des Seychelles, The Percy Sladen Trust Expedition, 28. Transactions of the Linnean Society, London, 15, 447 - 472." type="journal article" year="1913" yearSuffix="a">
Bouvier 
<number box="[151,209,2007,2031]" pageId="7" pageNumber="308" value="1913.0">1913</number>
a
</bibRefCitation>
, 
<taxonomicName box="[233,426,2006,2031]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[233,426,2006,2031]" italics="true" pageId="7" pageNumber="308">C. brachydactyla</emphasis>
</taxonomicName>
of 
<bibRefCitation author="Johnson" box="[463,633,2006,2031]" firstAuthor="Johnson" pageId="7" pageNumber="308" refString="Johnson, D. S. (1960 a) Some aspects of the distribution of freshwater organisms in the Indo-Pacific Area, and their relevance to the validity of the concept of an oriental region on zoogeography. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 170 - 181." type="journal article" year="1960" yearSuffix="a">
Johnson 
<number box="[562,620,2006,2031]" pageId="7" pageNumber="308" value="1960.0">1960</number>
a
</bibRefCitation>
, 
<bibRefCitation author="Johnson" box="[645,659,2006,2031]" pageId="7" pageNumber="308" refString="Johnson, D. S. (1960 b) Sub-specific and infra specific variation in some freshwater prawns of the Indo-Pacific region. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 261 - 267." type="journal article" year="1960" yearSuffix="b">b</bibRefCitation>
and 
<emphasis box="[713,767,2007,2030]" italics="true" pageId="7" pageNumber="308">C. n.</emphasis>
<taxonomicName box="[773,936,2006,2031]" pageId="7" pageNumber="308" rank="variety" variety="natalensis">
var. 
<emphasis box="[822,936,2006,2031]" italics="true" pageId="7" pageNumber="308">natalensis</emphasis>
</taxonomicName>
of 
<bibRefCitation author="De Man" box="[973,1137,2007,2031]" firstAuthor="De Man" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="308" pagination="219 - 283" part="2" refId="ref24738" refString="Man, J. G. de (1908 a) The fauna of brackish ponds at port Canning, Lower Bengal. Records of the Indian Museum, 2, 219 - 283." title="The fauna of brackish ponds at port Canning, Lower Bengal" type="journal article" year="1908">
De Man, 
<number box="[1078,1137,2007,2031]" pageId="7" pageNumber="308" value="1908.0">1908</number>
</bibRefCitation>
.
</paragraph>
<caption httpUri="https://zenodo.org/record/228247/files/figure.png" pageId="8" pageNumber="309" targetBox="[167,1432,221,1051]" targetPageId="8">
<paragraph blockId="8.[151,1436,1099,1152]" pageId="8" pageNumber="309">
<emphasis bold="true" box="[151,276,1099,1121]" pageId="8" pageNumber="309">FIGURE 3.</emphasis>
<taxonomicName authority="De Man, 1908" authorityName="De Man" authorityYear="1908" box="[284,687,1099,1121]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[284,528,1099,1121]" italics="true" pageId="8" pageNumber="309">Caridina brachydactyla</emphasis>
<bibRefCitation author="De Man" box="[536,687,1099,1121]" firstAuthor="De Man" journalOrPublisher="Records of the Indian Museum" pageId="8" pageNumber="309" pagination="219 - 283" part="2" refId="ref24738" refString="Man, J. G. de (1908 a) The fauna of brackish ponds at port Canning, Lower Bengal. Records of the Indian Museum, 2, 219 - 283." title="The fauna of brackish ponds at port Canning, Lower Bengal" type="journal article" year="1908">De Man, 1908</bibRefCitation>
</taxonomicName>
, (was part of the type series of 
<taxonomicName box="[1015,1192,1099,1121]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[1015,1192,1099,1121]" italics="true" pageId="8" pageNumber="309">Caridina nilotica</emphasis>
</taxonomicName>
<taxonomicName authority="Roux 1926 Cotype" authorityName="Roux 1926 Cotype" pageId="8" pageNumber="309" rank="variety" variety="meridionalis">
var. 
<emphasis box="[1245,1373,1099,1121]" italics="true" pageId="8" pageNumber="309">meridionalis</emphasis>
<bibRefCitation author="Roux" editor="Sarasin, F. &amp; Roux, J." firstAuthor="Roux" pageId="8" pageNumber="309" pagination="181 - 240" part="4" refId="ref25389" refString="Roux, J. (1926 a) Crustaces decapodes d'eau douce de la Nouvelle-Caledonie. In: Sarasin, F. &amp; Roux, J. (Eds.), Nova Caledonia, Wiesbaden, A. Zoologie, 4, 181 - 240. [Footnote of J. Roux 1926 b: 237 states &quot; Since completion of this paper (Roux 1926 b Australian Atyidae) the author (J. Roux) has finished and published a work on Decapoda of New Caledonia &quot;]" title="Crustaces decapodes d'eau douce de la Nouvelle-Caledonie" type="book chapter" volumeTitle="Nova Caledonia, Wiesbaden, A. Zoologie" year="1926">Roux 1926</bibRefCitation>
Cotype
</taxonomicName>
) NMB 4.VIII.a ♀ovig.: a) Anterior region of the cepahalothorax; b) Posterior margin of telson; c) Egg.
</paragraph>
</caption>
<paragraph blockId="8.[151,1437,1191,2008]" pageId="8" pageNumber="309">
<taxonomicName box="[199,474,1191,1216]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[199,474,1191,1216]" italics="true" pageId="8" pageNumber="309">Caridina brachydactyla</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName box="[813,927,1192,1215]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[813,927,1192,1215]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
by possessing a bifid tipped rostrum (vs. terminating in a pointed tip in 
<taxonomicName box="[523,636,1228,1251]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[523,636,1228,1251]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
), teeth placed proximally leaving 
<number box="[1047,1145,1227,1252]" pageId="8" pageNumber="309" value="0.525" valueMax="0.75" valueMin="0.3">0.3–0.75</number>
of dorsal rostral margin unarmed distally (vs. 
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unarmed distally or interrupted by 
<number box="[936,979,1263,1287]" pageId="8" pageNumber="309" value="2.5" valueMax="4.0" valueMin="1.0">1–4</number>
teeth in 
<taxonomicName box="[1091,1204,1264,1287]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[1091,1204,1264,1287]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
), 
<number box="[1232,1302,1263,1288]" pageId="8" pageNumber="309" value="18.5" valueMax="25.0" valueMin="12.0">12–25</number>
teeth from proximal end either up to the tip or with a short unarmed end distally (vs. 
<number box="[987,1043,1299,1324]" pageId="8" pageNumber="309" value="9.5" valueMax="14.0" valueMin="5.0">5–14</number>
teeth proximally leaving the distal ventral margin always unarmed in 
<taxonomicName box="[538,645,1336,1359]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[538,645,1336,1359]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
), posterior margin of telson triangular and with a median protuberance (vs. rounded posterior margin mostly devoid of a median protuberance in 
<taxonomicName box="[971,1078,1372,1395]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[971,1078,1372,1395]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
) and in carrying ca. 
<specimenCount box="[1307,1406,1371,1396]" pageId="8" pageNumber="309" type="egg">
<number box="[1307,1349,1371,1396]" pageId="8" pageNumber="309" value="850.0">850</number>
eggs
</specimenCount>
of 
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× 
<quantity box="[277,428,1407,1432]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="8" pageNumber="309" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.2–0.23 mm</quantity>
size (vs. ca. 
<number box="[572,656,1407,1432]" pageId="8" pageNumber="309" value="105.0" valueMax="160.0" valueMin="50.0">
50– 
<specimenCount box="[615,656,1407,1432]" pageId="8" pageNumber="309" type="egg">160</specimenCount>
</number>
eggs of 
<number box="[752,801,1407,1432]" pageId="8" pageNumber="309" value="0.65">0.65</number>
–1.0× 
<quantity box="[864,1014,1407,1432]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="8" pageNumber="309" unit="mm" value="0.525" valueMax="0.6" valueMin="0.45">0.45–0.6 mm</quantity>
size in 
<taxonomicName box="[1100,1207,1408,1431]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="simoni">
<emphasis box="[1100,1207,1408,1431]" italics="true" pageId="8" pageNumber="309">C. simoni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="8.[151,1437,1191,2008]" lastBlockId="9.[151,1437,151,1760]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
Jean 
<bibRefCitation author="Roux" box="[259,411,1443,1468]" firstAuthor="Roux" pageId="8" pageNumber="309" refString="Roux, J. (1926 a) Crustaces decapodes d'eau douce de la Nouvelle-Caledonie. In: Sarasin, F. &amp; Roux, J. (Eds.), Nova Caledonia, Wiesbaden, A. Zoologie, 4, 181 - 240. [Footnote of J. Roux 1926 b: 237 states &quot; Since completion of this paper (Roux 1926 b Australian Atyidae) the author (J. Roux) has finished and published a work on Decapoda of New Caledonia &quot;]" type="journal article" year="1926" yearSuffix="a">
Roux (
<number box="[339,398,1443,1468]" pageId="8" pageNumber="309" value="1926.0">1926</number>
a
</bibRefCitation>
, 
<bibRefCitation author="Roux" box="[426,441,1443,1468]" pageId="8" pageNumber="309" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">b</bibRefCitation>
) described 
<taxonomicName box="[578,832,1443,1468]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[578,832,1443,1468]" italics="true" pageId="8" pageNumber="309">Caridina meridionalis</emphasis>
</taxonomicName>
from 
<collectingCountry box="[907,1085,1443,1468]" name="New Caledonia" pageId="8" pageNumber="309">New Caledonia</collectingCountry>
and 
<collectingCountry box="[1146,1248,1443,1468]" name="Australia" pageId="8" pageNumber="309">Australia</collectingCountry>
and the present study examined the 
<typeStatus box="[380,428,1480,1504]" pageId="8" pageNumber="309">type</typeStatus>
specimens. As rightly pointed out by 
<bibRefCitation author="Roux" box="[865,1011,1479,1504]" firstAuthor="Roux" pageId="8" pageNumber="309" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">
Roux, 
<number box="[941,997,1479,1504]" pageId="8" pageNumber="309" value="1926.0">1926</number>
b
</bibRefCitation>
the specimens from a Creek at Cook Town, Queensland, 
<collectingCountry box="[374,477,1515,1540]" name="Australia" pageId="8" pageNumber="309">Australia</collectingCountry>
have a “rostrum mostly without teeth on the distal part of the upper border” and “It is usually devoid of teeth on a longer or shorter distal part of the upper border (about 
<number box="[1087,1097,1551,1575]" pageId="8" pageNumber="309" value="1.0">1</number>
/ 
<number box="[1109,1123,1551,1575]" pageId="8" pageNumber="309" value="3.0">3</number>
)” and the ovigerous female carries “numerous eggs”. The size of eggs differs fractionally from measurement of 
<bibRefCitation author="Roux" box="[1212,1378,1587,1612]" firstAuthor="Roux" pageId="8" pageNumber="309" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">
Roux (
<number box="[1297,1355,1587,1612]" pageId="8" pageNumber="309" value="1926.0">1926</number>
b)
</bibRefCitation>
i.e., 
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× 
<quantity box="[279,444,1623,1648]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="8" pageNumber="309" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.21–0.23 mm</quantity>
being 
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× 
<quantity box="[638,790,1623,1648]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="8" pageNumber="309" unit="mm" value="0.225" valueMax="0.25" valueMin="0.2">0.2–0.25 mm</quantity>
size. The specimens described by 
<bibRefCitation author="Roux" box="[1192,1350,1623,1648]" firstAuthor="Roux" pageId="8" pageNumber="309" refString="Roux, J. (1926 b) An account of Australian Atyidae. Records of the Australian Museum, 15, 237 - 254." type="journal article" year="1926" yearSuffix="b">
Roux (
<number box="[1270,1328,1623,1648]" pageId="8" pageNumber="309" value="1926.0">1926</number>
b)
</bibRefCitation>
from a creek at Cook Town have the following characters: rostrum reaching the end of the antennal scale or slightly longer; 
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teeth on the proximal dorsal margin leaving 
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of the dorsal margin unarmed distally; 
<number box="[1422,1436,1695,1719]" pageId="8" pageNumber="309" value="2.0">2</number>
postorbital teeth; tip bifid; 
<number box="[458,527,1731,1756]" pageId="8" pageNumber="309" value="13.5" valueMax="16.0" valueMin="11.0">11–16</number>
teeth on the ventral margin either up to the tip or with a short unarmed margin distally; first pereiopod dactylus 
<number box="[518,600,1767,1792]" pageId="8" pageNumber="309" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5</number>
×palm of propodus; chela 
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×long as broad and carpus 
<number box="[1256,1289,1767,1792]" pageId="8" pageNumber="309" value="1.8">1.8</number>
–2.0×long as broad, with the anterior excavation; second pereiopod-dactylus 
<number box="[930,1014,1803,1828]" pageId="8" pageNumber="309" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4</number>
×long as palm of propodus, chela 
<number box="[151,236,1839,1864]" pageId="8" pageNumber="309" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8</number>
×long as broad and carpus 5.0– 
<number box="[604,639,1839,1864]" pageId="8" pageNumber="309" value="5.6">5.6</number>
×long as broad; third pereiopod-dactylus 
<number box="[1118,1204,1839,1864]" pageId="8" pageNumber="309" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6</number>
×long as broad, 
<number box="[1394,1436,1839,1864]" pageId="8" pageNumber="309" value="7.0" valueMax="8.0" valueMin="6.0">6–8</number>
spines on dactylus (including terminal spines), propodus 
<number box="[788,873,1875,1900]" pageId="8" pageNumber="309" value="6.1" valueMax="6.5" valueMin="5.7">5.7–6.5</number>
×long as dactylus and 
<number box="[1122,1212,1875,1900]" pageId="8" pageNumber="309" value="12.25" valueMax="14.0" valueMin="10.5">10.5–14</number>
×long as broad with 
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spines arranged along inner margin, carpus 
<number box="[739,824,1911,1936]" pageId="8" pageNumber="309" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6</number>
×long as propodus, with 
<number box="[1113,1127,1911,1935]" pageId="8" pageNumber="309" value="1.0">1</number>
large spine and 
<number box="[1323,1366,1911,1935]" pageId="8" pageNumber="309" value="3.0" valueMax="4.0" valueMin="2.0">2–4</number>
small spines on inner margin, merus 
<number box="[518,551,1947,1971]" pageId="8" pageNumber="309" value="1.9">1.9</number>
–2.0×carpus length, merus with 
<number box="[932,946,1947,1971]" pageId="8" pageNumber="309" value="3.0">3</number>
large spines along posterior margin; fifth pereiopod, dactylus 
<number box="[386,421,1983,2008]" pageId="8" pageNumber="309" value="3.6">3.6</number>
–4.0×long as broad with 
<number box="[718,788,1983,2008]" pageId="8" pageNumber="309" value="67.5" valueMax="75.0" valueMin="60.0">60–75</number>
spines arranged in comb-like fashion on inner margin, propodus 
<number box="[260,325,151,176]" pageId="9" pageNumber="310" value="14.5" valueMax="15.0" valueMin="14.0">14–15</number>
×long as broad and 
<number box="[554,589,151,176]" pageId="9" pageNumber="310" value="4.5">4.5</number>
–5.0×long as dactylus with 
<number box="[906,976,151,176]" pageId="9" pageNumber="310" value="17.5" valueMax="19.0" valueMin="16.0">16–19</number>
spines arranged along posterior margin, carpus 
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×propodus length and with 
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large spines and 
<number box="[902,943,188,213]" pageId="9" pageNumber="310" value="4.5" valueMax="5.0" valueMin="4.0">4–5</number>
minute spines along inner margin, merus 
<number box="[151,233,223,248]" pageId="9" pageNumber="310" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6</number>
×carpus length, with 
<number box="[511,525,224,248]" pageId="9" pageNumber="310" value="2.0">2</number>
large spines along posterior margin; first male pleopod with endopod 
<number box="[151,249,260,285]" pageId="9" pageNumber="310" value="0.225" valueMax="0.25" valueMin="0.2">0.2–0.25</number>
×exopod length, without appendix interna; numerous eggs of 
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× 
<quantity box="[1103,1259,260,285]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="9" pageNumber="310" unit="mm" value="0.225" valueMax="0.25" valueMin="0.2">0.2–0.25 mm</quantity>
size; posterior margin with triangular median process, bearing 
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pair of long lateral spines and 
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pairs intermediate spines that are shorter than lateral spines; the inner pair of spines longer than the outer and 
<number box="[1063,1133,332,357]" pageId="9" pageNumber="310" value="13.5" valueMax="15.0" valueMin="12.0">12–15</number>
uropod diaeresis spinules present. Based on the structure of the rostrum, posterior margin of the telson and the presence of numerous eggs, the specimens from the creek at Cook Town, are here identified as 
<taxonomicName box="[899,1092,404,429]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[899,918,405,428]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[932,1092,404,429]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="310">
<taxonomicName box="[199,454,439,464]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis box="[199,454,439,464]" italics="true" pageId="9" pageNumber="310">Caridina meridionalis</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[619,817,439,464]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[619,817,439,464]" italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Richard &amp; Clark" box="[887,1156,439,464]" firstAuthor="Richard" pageId="9" pageNumber="310" refString="Richard, J. &amp; Clark, P. F. (2010) Caridina H. Milne Edwards, 1837 (Crustacea: Decapoda: Caridea: Atyoidea: Atyidae) freshwater shrimps from eastern and southern Africa. Zootaxa, 2372, 305 - 337." type="journal article" year="2010">
Richard &amp; Clark 
<number box="[1098,1156,440,464]" pageId="9" pageNumber="310" value="2010.0">2010</number>
</bibRefCitation>
) by the morphology of rostrum that is equal to antennular peduncle or slightly longer (vs. equal to longer than the antennal scale in 
<taxonomicName class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
) 
<number box="[331,401,512,536]" pageId="9" pageNumber="310" value="23.0" valueMax="27.0" valueMin="19.0">19–27</number>
teeth on the dorsal margin up to the tip of the rostrum of which the proximal teeth are compact and the distal 
<number box="[413,455,548,573]" pageId="9" pageNumber="310" value="3.0" valueMax="5.0" valueMin="1.0">1–5</number>
teeth spaced (vs. 
<number box="[659,729,548,573]" pageId="9" pageNumber="310" value="18.5" valueMax="25.0" valueMin="12.0">12–25</number>
teeth proximally leaving distal dorsal margin always unarmed in 
<taxonomicName box="[180,372,583,608]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[180,372,583,608]" italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
), 
<number box="[396,452,584,608]" pageId="9" pageNumber="310" value="7.0" valueMax="10.0" valueMin="4.0">4–10</number>
teeth on the ventral margin up to the tip or with a short unarmed end distally (vs. 
<number box="[1366,1436,583,608]" pageId="9" pageNumber="310" value="18.5" valueMax="25.0" valueMin="12.0">12–25</number>
teeth either up to the tip or with a short unarmed end in 
<taxonomicName box="[824,1020,620,645]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[824,1020,620,645]" italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
) and in possessing ca. 
<specimenCount box="[1298,1402,620,645]" pageId="9" pageNumber="310" type="egg">
<number box="[1298,1340,620,644]" pageId="9" pageNumber="310" value="300.0">300</number>
eggs
</specimenCount>
of 
<number box="[151,249,655,680]" pageId="9" pageNumber="310" value="0.625" valueMax="0.65" valueMin="0.6">0.6–0.65</number>
× 
<quantity box="[264,428,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="9" pageNumber="310" unit="mm" value="0.37" valueMax="0.39" valueMin="0.35">0.35–0.39 mm</quantity>
size (vs. ca. 
<specimenCount box="[572,672,655,680]" pageId="9" pageNumber="310" type="egg">
<number box="[572,614,655,680]" pageId="9" pageNumber="310" value="850.0">850</number>
eggs
</specimenCount>
of 
<number box="[710,822,655,680]" pageId="9" pageNumber="310" value="0.38" valueMax="0.41" valueMin="0.35">0.35–0.41</number>
× 
<quantity box="[836,986,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="9" pageNumber="310" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.2–0.23 mm</quantity>
size in 
<taxonomicName box="[1072,1265,655,680]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1072,1265,655,680]" italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="9.[151,1437,151,1760]" pageId="9" pageNumber="310">
The references by Johnson to 
<taxonomicName box="[545,666,692,717]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[545,666,692,717]" italics="true" pageId="9" pageNumber="310">C. nilotica</emphasis>
</taxonomicName>
<taxonomicName box="[675,885,692,717]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="variety" species="brachydactyla" variety="brachydactyla">
var. 
<emphasis box="[725,885,692,717]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Johnson" box="[948,1119,692,717]" firstAuthor="Johnson" pageId="9" pageNumber="310" refString="Johnson, D. S. (1961 a) Notes on the freshwater Crustacea of Malaya, I. The Atyidae. Bulletin of the Raffles Museum, 26, 120 - 153." type="journal article" year="1961" yearSuffix="a">
Johnson 
<number box="[1050,1107,692,717]" pageId="9" pageNumber="310" value="1961.0">1961</number>
a
</bibRefCitation>
) and 
<taxonomicName box="[1187,1381,692,717]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1187,1206,693,716]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1221,1381,692,717]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
(see 
<bibRefCitation author="Johnson" box="[151,324,727,752]" firstAuthor="Johnson" pageId="9" pageNumber="310" refString="Johnson, D. S. (1960 a) Some aspects of the distribution of freshwater organisms in the Indo-Pacific Area, and their relevance to the validity of the concept of an oriental region on zoogeography. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 170 - 181." type="journal article" year="1960" yearSuffix="a">
Johnson 
<number box="[253,311,727,752]" pageId="9" pageNumber="310" value="1960.0">1960</number>
a
</bibRefCitation>
, 
<bibRefCitation author="Johnson" box="[338,355,727,752]" pageId="9" pageNumber="310" refString="Johnson, D. S. (1960 b) Sub-specific and infra specific variation in some freshwater prawns of the Indo-Pacific region. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 261 - 267." type="journal article" year="1960" yearSuffix="b">b</bibRefCitation>
; 
<bibRefCitation author="Johnson" box="[370,473,727,752]" firstAuthor="Johnson" page="21" pageId="9" pageNumber="310" refString="Johnson, D. S. (1963) Distribution and other notes on some freshwater prawns (Atyidae and Palaemonidae) mainly from the Indo-west Pacific region. Bulletin of the National Museum of Singapore, 32, 5 - 30." type="journal article" year="1963">
<number box="[370,430,727,752]" pageId="9" pageNumber="310" value="1963.0">1963</number>
: 
<number box="[444,473,728,752]" pageId="9" pageNumber="310" value="21.0">21</number>
</bibRefCitation>
) are questioned in the synonymy presented above because he (
<bibRefCitation author="Johnson" box="[1209,1428,727,752]" firstAuthor="Johnson" page="23" pageId="9" pageNumber="310" refString="Johnson, D. S. (1961 a) Notes on the freshwater Crustacea of Malaya, I. The Atyidae. Bulletin of the Raffles Museum, 26, 120 - 153." type="journal article" year="1961" yearSuffix="a">
Johnson 
<number box="[1313,1372,727,752]" pageId="9" pageNumber="310" value="1961.0">1961</number>
a: 
<number box="[1399,1428,728,752]" pageId="9" pageNumber="310" value="23.0">23</number>
</bibRefCitation>
) synonymises 
<emphasis box="[302,321,765,788]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[335,349,764,789]" italics="true" pageId="9" pageNumber="310">b</emphasis>
. subsp. 
<taxonomicName box="[441,580,764,789]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis box="[441,580,764,789]" italics="true" pageId="9" pageNumber="310">peninsularis</emphasis>
</taxonomicName>
with 
<taxonomicName box="[644,836,764,789]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[644,663,765,788]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[676,836,764,789]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
, a decision rejected by the present study as these two species are considered as distinct. Therefore, the reference of 
<bibRefCitation author="Johnson" box="[855,1042,799,824]" firstAuthor="Johnson" pageId="9" pageNumber="310" refString="Johnson, D. S. (1961 a) Notes on the freshwater Crustacea of Malaya, I. The Atyidae. Bulletin of the Raffles Museum, 26, 120 - 153." type="journal article" year="1961" yearSuffix="a">
Johnson (
<number box="[964,1021,799,824]" pageId="9" pageNumber="310" value="1961.0">1961</number>
a)
</bibRefCitation>
could have been to either of these two species. Moreover, 
<bibRefCitation author="Johnson" box="[431,684,836,861]" firstAuthor="Johnson" page="266" pageId="9" pageNumber="310" refString="Johnson, D. S. (1960 b) Sub-specific and infra specific variation in some freshwater prawns of the Indo-Pacific region. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 261 - 267." type="journal article" year="1960" yearSuffix="b">
Johnson (
<number box="[542,602,836,861]" pageId="9" pageNumber="310" value="1960.0">1960</number>
b: 
<number box="[632,676,836,861]" pageId="9" pageNumber="310" value="266.0">266</number>
)
</bibRefCitation>
states “If however attention is confined to limited geographical areas, it is found that these are inhabited by one or two quiet distinct forms which show little infra-specific variation. The majority of these local forms would appear to be true geographical races of the single species 
<taxonomicName class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis italics="true" pageId="9" pageNumber="310">C. brachydactyla</emphasis>
</taxonomicName>
. This species is readily distinguishable from true 
<taxonomicName box="[904,1027,943,968]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[904,1027,943,968]" italics="true" pageId="9" pageNumber="310">C. nilotica</emphasis>
</taxonomicName>
by a number of features of which detailed armature of the 
<number box="[433,445,980,1005]" pageId="9" pageNumber="310" value="3.0">3</number>
rd thoracic leg is the most characteristic and constant”. For the present study there are many features that distinguish 
<taxonomicName box="[510,631,1015,1040]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[510,529,1016,1039]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[546,631,1015,1040]" italics="true" pageId="9" pageNumber="310">nilotica</emphasis>
</taxonomicName>
from 
<taxonomicName box="[708,904,1015,1040]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[708,727,1016,1039]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[744,904,1015,1040]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
including posterior margin of the telson, and number and size of eggs. However, the detailed armature of the 
<number box="[877,889,1052,1077]" pageId="9" pageNumber="310" value="3.0">3</number>
rd pereiopod is not regarded as a good character for separating these two species. Consequently the identification by Johnson of 
<taxonomicName box="[1044,1162,1087,1112]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="nilotica">
<emphasis box="[1044,1063,1088,1111]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1077,1162,1087,1112]" italics="true" pageId="9" pageNumber="310">nilotica</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1217,1410,1087,1112]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1217,1236,1088,1111]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1250,1410,1087,1112]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
is considered doubtful. With respect to the latter species 
<bibRefCitation author="Johnson" box="[761,977,1124,1149]" firstAuthor="Johnson" page="23" pageId="9" pageNumber="310" refString="Johnson, D. S. (1963) Distribution and other notes on some freshwater prawns (Atyidae and Palaemonidae) mainly from the Indo-west Pacific region. Bulletin of the National Museum of Singapore, 32, 5 - 30." type="journal article" year="1963">
Johnson (
<number box="[869,930,1124,1149]" pageId="9" pageNumber="310" value="1963.0">1963</number>
: 
<number box="[941,970,1124,1148]" pageId="9" pageNumber="310" value="23.0">23</number>
)
</bibRefCitation>
writes “In the British Museum there is a specimen from 
<collectingCountry box="[331,466,1159,1184]" name="Madagascar" pageId="9" pageNumber="310">Madagascar</collectingCountry>
which Bouvier has identified as belonging to 
<taxonomicName box="[1005,1201,1159,1184]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1005,1024,1160,1183]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1041,1201,1159,1184]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
. I am very doubtful about this identification”. Johnson then discusses various characters of this specimen including the rostrum which he considers “differs strikingly from any rostral forms found in authentic specimens of 
<taxonomicName box="[1162,1357,1231,1256]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1162,1181,1232,1255]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1197,1357,1231,1256]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
, being much longer and with a very long unarmed portion of the dorsal margin”. Specimens of 
<taxonomicName box="[1230,1429,1268,1293]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1230,1249,1269,1292]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[1269,1429,1268,1293]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
, determined by E.L. Bouvier 
<number box="[473,532,1304,1328]" pageId="9" pageNumber="310" value="1918.0">1918</number>
, received from him in 
<date box="[788,938,1304,1329]" pageId="9" pageNumber="310" value="1920-01">
January 
<number box="[882,938,1304,1328]" pageId="9" pageNumber="310" value="1920.0">1920</number>
</date>
are extant in the NHM reference collection. There are no locality details on the label, the material has not been registered and comprises 
<specimenCount box="[1178,1216,1340,1364]" pageId="9" pageNumber="310" type="generic">
<number box="[1178,1194,1340,1364]" pageId="9" pageNumber="310" value="2.0">2</number>
♂
</specimenCount>
, 
<number box="[1227,1244,1340,1365]" pageId="9" pageNumber="310" value="5.0">5</number>
♀ ovig., 
<number box="[1339,1357,1340,1365]" pageId="9" pageNumber="310" value="5.0">5</number>
♀. Two of these specimens were placed in individual tubes, a female and an ovigerous specimen. This material was examined and the posterior margin of the telson had been cleanly removed from every specimen. The ovigerous female placed in a separate tube seemed to fit the rostral description of Johnston (
<number box="[1060,1118,1447,1472]" pageId="9" pageNumber="310" value="1963.0">1963</number>
) and the unarmed portion of the dorsal margin is 
<number box="[398,451,1484,1509]" pageId="9" pageNumber="310" value="0.52">0.52</number>
. This portion is considered to be “long” and falls within the range for the species. Furthermore, the eggs of another female were measured and counted. These were small, measuring 
<number box="[151,249,1556,1581]" pageId="9" pageNumber="310" value="0.375" valueMax="0.4" valueMin="0.35">0.35–0.4</number>
× 
<quantity box="[263,406,1556,1581]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="9" pageNumber="310" unit="mm" value="0.215" valueMax="0.23" valueMin="0.2">0.2–0.23mm</quantity>
in size and 
<quantity box="[576,657,1556,1581]" metricMagnitude="1" metricUnit="m" metricValue="2.2479" pageId="9" pageNumber="310" unit="in" value="885.0">885 in</quantity>
number. The eggs too were within the range expected for 
<taxonomicName class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1410,1429,1557,1580]" italics="true" pageId="9" pageNumber="310">C</emphasis>
. 
<emphasis box="[151,311,1591,1616]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
. Furthermore with reference to distribution, 
<bibRefCitation author="Johnson" box="[820,1065,1591,1616]" firstAuthor="Johnson" page="178" pageId="9" pageNumber="310" refString="Johnson, D. S. (1960 a) Some aspects of the distribution of freshwater organisms in the Indo-Pacific Area, and their relevance to the validity of the concept of an oriental region on zoogeography. Proceedings of the Centenary and Bicentenary Congress of Biology, Singapore, 1958, 170 - 181." type="journal article" year="1960" yearSuffix="a">
Johnson (
<number box="[929,990,1591,1616]" pageId="9" pageNumber="310" value="1960.0">1960</number>
a: 
<number box="[1015,1058,1592,1616]" pageId="9" pageNumber="310" value="178.0">178</number>
)
</bibRefCitation>
says “An interesting member of the group is 
<taxonomicName box="[289,556,1628,1653]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[289,556,1628,1653]" italics="true" pageId="9" pageNumber="310">Caridina brachydactyla</emphasis>
</taxonomicName>
which, though comparatively intolerant of salt water and normally confined to freshwaters, is nevertheless widely distributed from Papuasia and N. 
<collectingCountry box="[941,1044,1663,1688]" name="Australia" pageId="9" pageNumber="310">Australia</collectingCountry>
to 
<collectingCountry box="[1083,1144,1663,1688]" name="Thailand" pageId="9" pageNumber="310">Siam</collectingCountry>
; it does not seem to have reached 
<collectingCountry box="[250,311,1700,1725]" name="India" pageId="9" pageNumber="310">India</collectingCountry>
. ….”. Although the distribution data presented by Johnson is correct for C. 
<taxonomicName box="[1224,1384,1700,1725]" class="Malacostraca" family="Atyidae" genus="Caridina" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="brachydactyla">
<emphasis box="[1224,1384,1700,1725]" italics="true" pageId="9" pageNumber="310">brachydactyla</emphasis>
</taxonomicName>
, his identification of the species could be questionable.
</paragraph>
</subSubSection>
</treatment>
</document>