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<document id="C6563716AC17176C9AC84CB388A97A58" ID-DOI="10.5281/zenodo.4525415" ID-ISSN="1638-9387" ID-RefBankID="E114FFC7FFB8FFD32C2E585BCB79FFF4" ID-Zenodo-Dep="4525415" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612878832874" checkinUser="felipe" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2006" docId="03AE8781FF94FFF4F19BFA60926AFAC2" docLanguage="en" docName="Zoosystema.28.4.887-915.pdf" docOrigin="Zoosystema 28 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Hyleoglomeris reducta Golovatch &amp; Geoffroy &amp; Mauriès 2006, n. sp." docType="treatment" docVersion="3" lastPageNumber="901" masterDocId="FF97FFF9FF98FFFAF17FFFAC9009FF85" masterDocTitle="Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia" masterLastPageNumber="915" masterPageNumber="887" pageNumber="899" updateTime="1698868940497" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2349FF39BE87FC94F547629DC1D81BA5">Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia</mods:title>
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<mods:name id="A1B5D964C1290D3435026CDF5A251D8C" type="personal">
<mods:role id="DA4A9C79AB4EE26AD05C0703CE27725E">
<mods:roleTerm id="F62DAC42A3E345B5C2D2C38CFC6F96B5">Author</mods:roleTerm>
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<mods:namePart id="F337E9D0719A537C99B442BE56C96665">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="10792BBED272E32152D486EDE6B8EA88">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:roleTerm id="9783058168D168E693394D1E5259861F">Author</mods:roleTerm>
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<mods:namePart id="DF9C604CDD9BCB55DAB437FEE9DC7352">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="86929890DE5A80CE710C6C287DCA4FAA">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, EA 3778 - USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="14EF067F9E25490C62962CFA2D291A46" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="79FFF31CF35E049D8BF7F36CD340EF95">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="0FFAB6AD1592E48CB247F731E0E5F6D6">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:typeOfResource id="F9C26E839619E680034DFE5197408F18">text</mods:typeOfResource>
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<mods:title id="05F9C9DECA01A353A130BF2489E672C4">Zoosystema</mods:title>
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<mods:part id="529445F3A5848A2A2CB69525BE6CF870">
<mods:date id="32A7A9869DE78C84329A57EA6C11652C">2006</mods:date>
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<subSubSection id="C31D651CFF94FFF6F19BFA60921DFA63" box="[228,532,1484,1510]" pageId="12" pageNumber="899" type="nomenclature">
<paragraph id="8BB83697FF94FFF6F19BFA60921DFA63" blockId="12.[228,532,1484,1541]" box="[228,532,1484,1510]" pageId="12" pageNumber="899">
<heading id="D0F081FBFF94FFF6F19BFA60921DFA63" box="[228,532,1484,1510]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="899" reason="2">
<taxonomicName id="4C074D14FF94FFF6F19BFA6091DBFA63" ID-CoL="3NF2C" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[228,466,1484,1510]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="12" pageNumber="899" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF94FFF6F19BFA6091DBFA63" bold="true" box="[228,466,1484,1510]" italics="true" pageId="12" pageNumber="899">Hyleoglomeris reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF94FFF6F0A6FA60921DFA63" box="[473,532,1484,1510]" pageId="12" pageNumber="899" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C31D651CFF94FFF6F02BFA4791ADF980" box="[340,420,1515,1541]" pageId="12" pageNumber="899" type="description">
<paragraph id="8BB83697FF94FFF6F02BFA4791ADF980" blockId="12.[228,532,1484,1541]" box="[340,420,1515,1541]" pageId="12" pageNumber="899">
(
<figureCitation id="133C2A12FF94FFF6F023FA479192F980" box="[348,411,1515,1541]" captionStart="FIG" captionStartId="13.[113,124,831,848]" captionTargetBox="[126,1197,217,787]" captionTargetId="figure-14@13.[112,1210,214,790]" captionTargetPageId="13" captionText="FIG. 7. — Hyleoglomeris reducta n. sp., ♂ paratype: A, lateral side of thoracic shield; B, leg 17; C, leg 18; D, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="12" pageNumber="899">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF94FFF6F10EF98E911FF926" pageId="12" pageNumber="899" type="materials_examined">
<paragraph id="8BB83697FF94FFF6F10EF98E911FF926" blockId="12.[113,648,1570,1699]" pageId="12" pageNumber="899">
<materialsCitation id="3B6F3CCAFF94FFF6F10EF98E90ECF90D" collectingDate="1989-01-12" collectionCode="TYPE, MATERIAL, NMNHS" collectorName="P. Beron" country="China" county="Jianshui County" location="Yan Dong Cave" pageId="12" pageNumber="899" specimenCount="33" specimenCount-female="12" specimenCount-male="21" stateProvince="Yunnan" typeStatus="holotype">
<collectionCode id="ED16AE52FF94FFF6F10EF98E910FF9BD" box="[113,262,1570,1593]" pageId="12" pageNumber="899">TYPE MATERIAL</collectionCode>
. —
<emphasis id="B973EA85FF94FFF6F049F98E9173F9BC" bold="true" box="[310,378,1570,1593]" pageId="12" pageNumber="899">
<collectingCountry id="F3107607FF94FFF6F049F98E917FF9BC" box="[310,374,1570,1593]" name="China" pageId="12" pageNumber="899">China</collectingCountry>
.
</emphasis>
<collectingRegion id="49C3F875FF94FFF6F0FEF98E9225F9BC" box="[385,556,1570,1593]" country="China" name="Yunnan" pageId="12" pageNumber="899">Yunnan Province</collectingRegion>
,
<collectingCounty id="62D94E1BFF94FFF6F348F98E90CAF9D6" pageId="12" pageNumber="899">Jianshui County</collectingCounty>
,
<location id="8ED8604CFF94FFF6F1B0F9909171F9D6" LSID="urn:lsid:plazi:treatment:03AE8781FF94FFF4F19BFA60926AFAC2:8ED8604CFF94FFF6F1B0F9909171F9D6" box="[207,376,1596,1619]" country="China" county="Jianshui County" name="Yan Dong Cave" pageId="12" pageNumber="899" stateProvince="Yunnan">Yan Dong Cave</location>
,
<date id="FFB91057FF94FFF6F0FAF99091E7F9D6" box="[389,494,1596,1619]" pageId="12" pageNumber="899" value="1989-01-12">
<collectingDate id="EFFDE9BFFF94FFF6F0FAF99091E7F9D6" box="[389,494,1596,1619]" pageId="12" pageNumber="899" value="1989-01-12">12.I.1989</collectingDate>
</date>
, leg.
<collectorName id="26F25341FF94FFF6F356F990928DF9D6" box="[553,644,1596,1619]" pageId="12" pageNumber="899">P. Beron</collectorName>
,
<typeStatus id="54BC8835FF94FFF6F10CF9FB90C7F9EB" box="[115,206,1623,1646]" pageId="12" pageNumber="899" type="holotype">holotype</typeStatus>
<specimenCount id="9D01FD1EFF94FFF6F1AAF9FB90E0F9EB" box="[213,233,1623,1646]" pageId="12" pageNumber="899" type="male"></specimenCount>
(
<collectionCode id="ED16AE52FF94FFF6F185F9FB916BF9EB" box="[250,354,1623,1646]" pageId="12" pageNumber="899">NMNHS</collectionCode>
),
<typeStatus id="54BC8835FF94FFF6F007F9FB91D0F9EB" box="[376,473,1623,1646]" pageId="12" pageNumber="899" type="paratype">paratypes</typeStatus>
<specimenCount id="9D01FD1EFF94FFF6F09DF9FB9227F9EB" box="[482,558,1623,1646]" pageId="12" pageNumber="899" type="male">20 ♂♂</specimenCount>
,
<specimenCount id="9D01FD1EFF94FFF6F340F9FB928FF9EB" box="[575,646,1623,1646]" pageId="12" pageNumber="899" type="female">12 ♀♀</specimenCount>
(
<collectionCode id="ED16AE52FF94FFF6F106F9DD90D7F90D" box="[121,222,1649,1672]" pageId="12" pageNumber="899">NMNHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B6F3CCAFF94FFF6F18EF9DD923FF90D" box="[241,566,1649,1673]" collectionCode="MNHN" country="China" pageId="12" pageNumber="899" specimenCode="CC155" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="holotype">
<specimenCount id="9D01FD1EFF94FFF6F18EF9DD9124F90C" box="[241,301,1649,1673]" pageId="12" pageNumber="899" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D01FD1EFF94FFF6F043F9DD917DF90C" box="[316,372,1649,1673]" pageId="12" pageNumber="899" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED16AE52FF94FFF6F0FBF9DD91D0F90D" box="[388,473,1649,1672]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="899">MNHN</collectionCode>
<specimenCode id="DBA19EECFF94FFF6F09EF9DD9227F90D" box="[481,558,1649,1672]" collectionCode="CC" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="12" pageNumber="899">CC155</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B6F3CCAFF94FFF6F33DF9DD911BF926" collectionCode="ZMUM" country="China" pageId="12" pageNumber="899" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="holotype">
<specimenCount id="9D01FD1EFF94FFF6F33DF9DD9276F90C" box="[578,639,1649,1673]" pageId="12" pageNumber="899" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D01FD1EFF94FFF6F10EF92090AEF926" box="[113,167,1676,1699]" pageId="12" pageNumber="899" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED16AE52FF94FFF6F1CAF9209102F926" box="[181,267,1676,1699]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="12" pageNumber="899">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF94FFF6F3DAFCDD9360FC38" pageId="12" pageNumber="899" type="etymology">
<paragraph id="8BB83697FF94FFF6F3DAFCDD9360FC38" blockId="12.[675,1210,881,957]" pageId="12" pageNumber="899">ETYMOLOGY. — To emphasize the coloration, hyposchism and the number of the striae crossing the thoracic shield, all strongly reduced.</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF94FFF6F3DAFC7A93F5FBA7" pageId="12" pageNumber="899" type="diagnosis">
<paragraph id="8BB83697FF94FFF6F3DAFC7A93F5FBA7" blockId="12.[677,1210,982,1058]" pageId="12" pageNumber="899">DIAGNOSIS. — Differs from congeners by the strongly reduced coloration and hyposchism, combined with certain details of telopod structure.</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF94FFF7F3DAFBE391D3FA43" lastPageId="13" lastPageNumber="900" pageId="12" pageNumber="899" type="description">
<paragraph id="8BB83697FF94FFF6F3DAFBE3933CFBE2" blockId="12.[674,1212,1103,1700]" box="[677,821,1103,1129]" pageId="12" pageNumber="899">DESCRIPTION</paragraph>
<paragraph id="8BB83697FF94FFF6F3DAFBC393ABFB4C" blockId="12.[674,1212,1103,1700]" pageId="12" pageNumber="899">
Length 5.5-7.5 (
<specimenCount id="9D01FD1EFF94FFF6F21CFBDC9370FB0F" box="[867,889,1136,1162]" pageId="12" pageNumber="899" type="male"></specimenCount>
) or 6.0-
<quantity id="4CFF9B72FF94FFF6F2A3FBC3943EFB0C" box="[988,1079,1135,1161]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="899" unit="mm" value="8.5">8.5 mm</quantity>
(
<specimenCount id="9D01FD1EFF94FFF6F535FBDC9457FB0F" box="[1098,1118,1136,1162]" pageId="12" pageNumber="899" type="female"></specimenCount>
), width 2.8-3.6 (
<specimenCount id="9D01FD1EFF94FFF6F27DFB3C9311FB2F" box="[770,792,1168,1194]" pageId="12" pageNumber="899" type="male"></specimenCount>
) or 3.0-4.0 mm (
<specimenCount id="9D01FD1EFF94FFF6F2AEFB3C93EDFB2F" box="[977,996,1168,1194]" pageId="12" pageNumber="899" type="female"></specimenCount>
).
<typeStatus id="54BC8835FF94FFF6F286FB239456FB2C" box="[1017,1119,1167,1193]" pageId="12" pageNumber="899" type="holotype">Holotype</typeStatus>
<quantity id="4CFF9B72FF94FFF6F51AFB2394B0FB2C" box="[1125,1209,1167,1193]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="899" unit="mm" value="7.5">7.5 mm</quantity>
long and
<quantity id="4CFF9B72FF94FFF6F274FB039368FB4C" box="[779,865,1199,1225]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="12" pageNumber="899" unit="mm" value="3.7">3.7 mm</quantity>
wide.
</paragraph>
<paragraph id="8BB83697FF94FFF6F3C4FB62943FFA63" blockId="12.[674,1212,1103,1700]" pageId="12" pageNumber="899">
Coloration usually entirely pallid, only in few
<typeStatus id="54BC8835FF94FFF6F3D9FB429302FA8D" box="[678,779,1262,1288]" pageId="12" pageNumber="899" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D01FD1EFF94FFF6F264FB429353FA8C" box="[795,858,1262,1289]" pageId="12" pageNumber="899" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D01FD1EFF94FFF6F217FB429399FA8C" box="[872,912,1262,1289]" pageId="12" pageNumber="899" type="female">1 ♀</specimenCount>
) with remnants of pigment expressed as head being light red-brown above antennal sockets, and antennae brown; in one of these males a light red-brownish tergal pattern is also retained as large, transverse, apparently marbled, paramedian spots nearly in touch anteromedially on terga 3-11 and pygidium, and as increasingly small sublateral spotlets on terga 3-8.
</paragraph>
<paragraph id="8BB83697FF94FFF6F3C4FA479353F901" blockId="12.[674,1212,1103,1700]" pageId="12" pageNumber="899">Antennomere 6 about 2.5 times longer than wide; ocelli 6 + 1 to 7 + 1, unpigmented, convex, especially evident in pigmented specimens; Tömösvárys organ transverse-oval, only slightly wider than long.</paragraph>
<paragraph id="8BB83697FF94FFF6F3C4F9269427F921" blockId="12.[674,1212,1103,1700]" box="[699,1070,1674,1700]" pageId="12" pageNumber="899">Collum with two transverse striae.</paragraph>
<caption id="DF78661FFF95FFF7F10EFC9391FEFCE3" ID-DOI="http://doi.org/10.5281/zenodo.4525433" ID-Zenodo-Dep="4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="13" pageNumber="900" startId="13.[113,124,831,848]" targetBox="[126,1197,217,787]" targetPageId="13">
<paragraph id="8BB83697FF95FFF7F10EFC9391FEFCE3" blockId="13.[113,1211,831,870]" pageId="13" pageNumber="900">
FIG. 7. —
<taxonomicName id="4C074D14FF95FFF7F1BBFC939176FCD5" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[196,383,831,848]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF95FFF7F1BBFC939176FCD5" box="[196,383,831,848]" italics="true" pageId="13" pageNumber="900">Hyleoglomeris reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF95FFF7F0FAFC9391B1FCD5" box="[389,440,831,848]" pageId="13" pageNumber="900" rank="species">n. sp.</taxonomicNameLabel>
, ♂ paratype:
<emphasis id="B973EA85FF95FFF7F356FC93923FFCD5" bold="true" box="[553,566,831,848]" pageId="13" pageNumber="900">A</emphasis>
, lateral side of thoracic shield;
<emphasis id="B973EA85FF95FFF7F240FC939345FCD5" bold="true" box="[831,844,831,848]" pageId="13" pageNumber="900">B</emphasis>
, leg 17;
<emphasis id="B973EA85FF95FFF7F2E9FC9393ADFCD5" bold="true" box="[918,932,831,848]" pageId="13" pageNumber="900">C</emphasis>
, leg 18;
<emphasis id="B973EA85FF95FFF7F291FC9393F5FCD5" bold="true" box="[1006,1020,831,848]" pageId="13" pageNumber="900">D</emphasis>
, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BB83697FF95FFF7F1F7FC1D9211FBCF" blockId="13.[111,648,945,1478]" pageId="13" pageNumber="900">
Thoracic shield with a very small and narrow hyposchism not reaching the caudal tergal contour (
<figureCitation id="133C2A12FF95FFF7F108FC5C90C5FB8F" box="[119,204,1008,1034]" captionStart="FIG" captionStartId="13.[113,124,831,848]" captionTargetBox="[126,1197,217,787]" captionTargetId="figure-14@13.[112,1210,214,790]" captionTargetPageId="13" captionText="FIG. 7. — Hyleoglomeris reducta n. sp., ♂ paratype: A, lateral side of thoracic shield; B, leg 17; C, leg 18; D, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="13" pageNumber="900">Fig. 7A</figureCitation>
); 6 or 7 transverse striae, of which 4 or 5 start above the schism and (1)2-3 ones (never the first and last though) cross the dorsum.
</paragraph>
<paragraph id="8BB83697FF95FFF7F1F7FBE3911AFB0C" blockId="13.[111,648,945,1478]" pageId="13" pageNumber="900">Male pygidium virtually not sinuate medially at caudal margin.</paragraph>
<paragraph id="8BB83697FF95FFF7F1F7FB23917CFB4C" blockId="13.[111,648,945,1478]" pageId="13" pageNumber="900">
Male leg 17 (
<figureCitation id="133C2A12FF95FFF7F071FB239155FB2C" box="[270,348,1167,1193]" captionStart="FIG" captionStartId="13.[113,124,831,848]" captionTargetBox="[126,1197,217,787]" captionTargetId="figure-14@13.[112,1210,214,790]" captionTargetPageId="13" captionText="FIG. 7. — Hyleoglomeris reducta n. sp., ♂ paratype: A, lateral side of thoracic shield; B, leg 17; C, leg 18; D, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="13" pageNumber="900">Fig. 7B</figureCitation>
) with a low outer coxal lobe; telopodite 4-segmented.
</paragraph>
<paragraph id="8BB83697FF95FFF7F1F7FB629270FA8D" blockId="13.[111,648,945,1478]" pageId="13" pageNumber="900">
Male leg 18 (
<figureCitation id="133C2A12FF95FFF7F073FB629153FB6D" box="[268,346,1230,1256]" captionStart="FIG" captionStartId="13.[113,124,831,848]" captionTargetBox="[126,1197,217,787]" captionTargetId="figure-14@13.[112,1210,214,790]" captionTargetPageId="13" captionText="FIG. 7. — Hyleoglomeris reducta n. sp., ♂ paratype: A, lateral side of thoracic shield; B, leg 17; C, leg 18; D, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="13" pageNumber="900">Fig.7C</figureCitation>
) with a more or less narrowly ogival syncoxital notch; telopodite 4-segmented.
</paragraph>
<paragraph id="8BB83697FF95FFF7F1F7FAA291D3FA43" blockId="13.[111,648,945,1478]" pageId="13" pageNumber="900">
Telopods (
<figureCitation id="133C2A12FF95FFF7F18DFAA2914AFAAD" box="[242,323,1294,1320]" captionStart="FIG" captionStartId="13.[113,124,831,848]" captionTargetBox="[126,1197,217,787]" captionTargetId="figure-14@13.[112,1210,214,790]" captionTargetPageId="13" captionText="FIG. 7. — Hyleoglomeris reducta n. sp., ♂ paratype: A, lateral side of thoracic shield; B, leg 17; C, leg 18; D, telopod, caudal view. Scale bar: A, drawn not to scale; B-D, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525433" httpUri="https://zenodo.org/record/4525433/files/figure.png" pageId="13" pageNumber="900">Fig.7D</figureCitation>
) with a rather high and roundly subtrapeziform syncoxital lobe flanked by two setose horns crowned with an apical setoid. Prefemur micropapillate laterally. Caudomedial outgrowth of tibia with a micropapillate tubercle at base. Tarsus rather modestly rounded apically.
</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF95FFF4F10EFA47926AFAC2" lastPageId="14" lastPageNumber="901" pageId="13" pageNumber="900" type="discussion">
<paragraph id="8BB83697FF95FFF7F10EFA4790DCF986" blockId="13.[111,646,1515,1700]" box="[113,213,1515,1541]" pageId="13" pageNumber="900">REMARKS</paragraph>
<paragraph id="8BB83697FF95FFF7F10EF9A79379FAE2" blockId="13.[111,646,1515,1700]" lastBlockId="13.[674,1212,945,1700]" pageId="13" pageNumber="900">
Only one cavernicolous congener has hitherto been described from
<collectingCountry id="F3107607FF95FFF7F06BF987915EF9C0" box="[276,343,1579,1605]" name="China" pageId="13" pageNumber="900">China</collectingCountry>
in general and from
<collectingRegion id="49C3F875FF95FFF7F34EF987928CF9C0" box="[561,645,1579,1605]" country="China" name="Yunnan" pageId="13" pageNumber="900">Yunnan</collectingRegion>
in particular:
<taxonomicName id="4C074D14FF95FFF7F184F9E7928BF9E1" authority="Zhang &amp; Zhang, 1995" authorityName="Zhang &amp; Zhang" authorityYear="1995" box="[251,642,1610,1637]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="albicorporis">
<emphasis id="B973EA85FF95FFF7F184F9E79186F9E0" box="[251,399,1611,1637]" italics="true" pageId="13" pageNumber="900">H. albicorporis</emphasis>
<bibRefCitation id="EF964B66FF95FFF7F0ECF9E6928BF9E1" author="ZHANG F. &amp; ZHANG C. Z." box="[403,642,1610,1636]" pageId="13" pageNumber="900" pagination="17 - 21" refId="ref18661" refString="ZHANG F. &amp; ZHANG C. Z. 1995. - [A new troglobitic species of glomerid millipeds from Yunnan (Diplopoda, Glomerida, Glomeridae)]. Zoological Research 16 (1): 17 - 21 (in Chinese, English summary)." type="journal article" year="1995">Zhang &amp; Zhang, 1995</bibRefCitation>
</taxonomicName>
, from Shihua Cave at Baoshan. Based only on three specimens, which were described quite superficially (
<bibRefCitation id="EF964B66FF95FFF7F3D3FC1D939CFC4E" author="ZHANG F. &amp; ZHANG C. Z." box="[684,917,945,971]" pageId="13" pageNumber="900" pagination="17 - 21" refId="ref18661" refString="ZHANG F. &amp; ZHANG C. Z. 1995. - [A new troglobitic species of glomerid millipeds from Yunnan (Diplopoda, Glomerida, Glomeridae)]. Zoological Research 16 (1): 17 - 21 (in Chinese, English summary)." type="journal article" year="1995">Zhang &amp; Zhang 1995</bibRefCitation>
), it is hard to decide whether
<taxonomicName id="4C074D14FF95FFF7F3DCFC7D933AFC6E" authorityName="Zhang &amp; Zhang" authorityYear="1995" box="[675,819,977,1003]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="albicorporis">
<emphasis id="B973EA85FF95FFF7F3DCFC7D933AFC6E" box="[675,819,977,1003]" italics="true" pageId="13" pageNumber="900">H.albicorporis</emphasis>
</taxonomicName>
is indeed devoid of pigmentation, and thus whether it differs from some other, especially endogean, congeners encountered in SE
<collectingCountry id="F3107607FF95FFF7F50BFBBC94B0FBAF" box="[1140,1209,1040,1066]" name="China" pageId="13" pageNumber="900">China</collectingCountry>
(see also below). It matches
<taxonomicName id="4C074D14FF95FFF7F2B1FB9C9435FBCF" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[974,1084,1072,1098]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF95FFF7F2B1FB9C9435FBCF" box="[974,1084,1072,1098]" italics="true" pageId="13" pageNumber="900">H. reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF95FFF7F53DFB9C9475FBCF" box="[1090,1148,1072,1098]" pageId="13" pageNumber="900" rank="species">n. sp.</taxonomicNameLabel>
quite closely, not only on geographical grounds, but also in terms of at least body size and the conformation of male legs 17-19. The only meaningful differences between these two species that can be drawn from the original description of
<taxonomicName id="4C074D14FF95FFF7F2D2FB629437FB6D" authorityName="Zhang &amp; Zhang" authorityYear="1995" box="[941,1086,1230,1256]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="albicorporis">
<emphasis id="B973EA85FF95FFF7F2D2FB629437FB6D" box="[941,1086,1230,1256]" italics="true" pageId="13" pageNumber="900">H. albicorporis</emphasis>
</taxonomicName>
concern the latters ordinary (i.e. small) hyposchism, a slenderer male telopodite 17, and the tip of the telopod syncoxital horns supplied with a subapical, rather than apical, setoid.
</paragraph>
<paragraph id="8BB83697FF95FFF4F3C4FAC191A4FB0C" blockId="13.[674,1212,945,1700]" lastBlockId="14.[111,648,977,1351]" lastPageId="14" lastPageNumber="901" pageId="13" pageNumber="900">
The fact that only a few (6 of 42, or
<emphasis id="B973EA85FF95FFF7F51EFAC1947AFA02" box="[1121,1139,1389,1415]" italics="true" pageId="13" pageNumber="900">c.</emphasis>
14%) specimens from the
<typeStatus id="54BC8835FF95FFF7F2E5FA2093C5FA23" box="[922,972,1420,1446]" pageId="13" pageNumber="900">type</typeStatus>
series of
<taxonomicName id="4C074D14FF95FFF7F540FA2094B0FA23" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[1087,1209,1420,1446]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF95FFF7F540FA2094B0FA23" box="[1087,1209,1420,1446]" italics="true" pageId="13" pageNumber="900">H. reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF95FFF7F3DAFA0092EBFA43" box="[677,738,1452,1478]" pageId="13" pageNumber="900" rank="species">n. sp.</taxonomicNameLabel>
still retain pigment, and only one shows some traces of a tergal pattern, is important, apparently reflecting an intermediate stage of evolution toward complete cavernicoly. Whether or not
<taxonomicName id="4C074D14FF95FFF7F38DF987936DF9C0" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[754,868,1579,1605]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="900" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF95FFF7F38DF987936DF9C0" box="[754,868,1579,1605]" italics="true" pageId="13" pageNumber="900">H. reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF95FFF7F212F98793A3F9C0" box="[877,938,1579,1605]" pageId="13" pageNumber="900" rank="species">n. sp.</taxonomicNameLabel>
is already a troglobite is impossible to decide based solely on the above morphological and geographical evidence. The colour pattern of the single particularly “atavis- tic” male in the sample strongly resembles that of
<taxonomicName id="4C074D14FF96FFF4F1EDFC5D9117FB8E" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[146,286,1009,1035]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="901" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B973EA85FF96FFF4F1EDFC5D9117FB8E" box="[146,286,1009,1035]" italics="true" pageId="14" pageNumber="901">H. maculata</emphasis>
</taxonomicName>
, but differs in the paramedian spots being nearly in touch anteromedially. In addition,
<taxonomicName id="4C074D14FF96FFF4F19FFB9C915DFBCF" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[224,340,1072,1098]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="901" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="B973EA85FF96FFF4F19FFB9C915DFBCF" box="[224,340,1072,1098]" italics="true" pageId="14" pageNumber="901">H. reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF96FFF4F022FB9C9192FBCF" box="[349,411,1072,1098]" pageId="14" pageNumber="901" rank="species">n. sp.</taxonomicNameLabel>
shows an even more strongly reduced hyposchism and less numerous striae on the thoracic shield.
</paragraph>
<caption id="DF78661FFF96FFF4F10EFCCE93FEFC0F" ID-DOI="http://doi.org/10.5281/zenodo.4525437" ID-Zenodo-Dep="4525437" httpUri="https://zenodo.org/record/4525437/files/figure.png" pageId="14" pageNumber="901" startId="14.[113,124,866,883]" targetBox="[120,1198,218,824]" targetPageId="14">
<paragraph id="8BB83697FF96FFF4F10EFCCE93FEFC0F" blockId="14.[113,1211,866,906]" pageId="14" pageNumber="901">
FIG. 8. —
<taxonomicName id="4C074D14FF96FFF4F1BBFCCF9186FCF1" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[196,399,867,884]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="901" phylum="Arthropoda" rank="species" species="eusulcata" status="sp. nov.">
<emphasis id="B973EA85FF96FFF4F1BBFCCF9186FCF1" box="[196,399,867,884]" italics="true" pageId="14" pageNumber="901">Hyleoglomeris eusulcata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF96FFF4F0EAFCCE91C1FCF6" box="[405,456,866,883]" pageId="14" pageNumber="901" rank="species">n. sp.</taxonomicNameLabel>
, ♂ paratype:
<emphasis id="B973EA85FF96FFF4F345FCCF924EFCF1" bold="true" box="[570,583,867,884]" pageId="14" pageNumber="901">A</emphasis>
, lateral side of thoracic shield;
<emphasis id="B973EA85FF96FFF4F230FCCF9355FCF1" bold="true" box="[847,860,867,884]" pageId="14" pageNumber="901">B</emphasis>
, leg 17;
<emphasis id="B973EA85FF96FFF4F2D9FCCF93BDFCF1" bold="true" box="[934,948,867,884]" pageId="14" pageNumber="901">C</emphasis>
, leg 18;
<emphasis id="B973EA85FF96FFF4F281FCCF9405FCF1" bold="true" box="[1022,1036,867,884]" pageId="14" pageNumber="901">D</emphasis>
, telopod, front view;
<emphasis id="B973EA85FF96FFF4F10EFCD59074FC0F" bold="true" box="[113,125,889,906]" pageId="14" pageNumber="901">E</emphasis>
, distal part of telopod, caudal view. Abbreviation:
<emphasis id="B973EA85FF96FFF4F361FCD59221FC0F" bold="true" box="[542,552,889,906]" pageId="14" pageNumber="901">s</emphasis>
, sulcus. Scale bar: A, drawn not to scale; B-E, 0.4 mm.
</paragraph>
</caption>
<paragraph id="8BB83697FF96FFF4F1F7FB23926AFAC2" blockId="14.[111,648,977,1351]" pageId="14" pageNumber="901">
In general,
<collectingRegion id="49C3F875FF96FFF4F07BFB239151FB2C" box="[260,344,1167,1193]" country="China" name="Yunnan" pageId="14" pageNumber="901">Yunnan</collectingRegion>
is among Chinas provinces particularly rich in karst caves that support numerous troglobitic animals (
<bibRefCitation id="EF964B66FF96FFF4F037FB62920FFB6D" author="CHEN Z. &amp; DECU V. &amp; JUBERTHIE C. &amp; UENO S. - I." box="[328,518,1230,1256]" pageId="14" pageNumber="901" pagination="1763 - 1781" refId="ref16716" refString="CHEN Z., DECU V., JUBERTHIE C. &amp; UENO S. - I. 2001. - Chine, in JUBERTHIE C. &amp; DECU V. (eds), Encyclopaedia Biospeologica 3. Societe internationale de Biospeologie, Moulis: 1763 - 1781." type="book chapter" year="2001">
Chen
<emphasis id="B973EA85FF96FFF4F0F0FB6291CAFB6D" box="[399,451,1230,1256]" italics="true" pageId="14" pageNumber="901">et al.</emphasis>
2001
</bibRefCitation>
). Yan Dong Cave is known to contain several troglobitic arthropods, including the millipede
<taxonomicName id="4C074D14FF96FFF4F0E0FAA29125FAC2" authority="Stoev &amp; Enghoff, 2005" authorityName="Stoev &amp; Enghoff" authorityYear="2005" class="Diplopoda" family="Caspiopetalidae" genus="Bollmania" kingdom="Animalia" order="Callipodida" pageId="14" pageNumber="901" phylum="Arthropoda" rank="species" species="beroni">
<emphasis id="B973EA85FF96FFF4F0E0FAA29243FAAD" box="[415,586,1294,1320]" italics="true" pageId="14" pageNumber="901">Bollmania beroni</emphasis>
<bibRefCitation id="EF964B66FF96FFF4F331FAA29125FAC2" author="STOEV P. &amp; ENGHOFF H." pageId="14" pageNumber="901" pagination="1875 - 1891" refId="ref18107" refString="STOEV P. &amp; ENGHOFF H. 2005. - A new cave-dwelling millipede of the genus Bollmania Silvestri, 1896 from Yunnan, China, with remarks on the reduction of the second female leg-pair (Diplopoda: Callipodida: Caspiopetalidae). Journal of Natural History 39: 1875 - 1891." type="journal article" year="2005">Stoev &amp; Enghoff, 2005</bibRefCitation>
</taxonomicName>
(cf.
<bibRefCitation id="EF964B66FF96FFF4F022FA81925FFAC2" author="STOEV P. &amp; ENGHOFF H." box="[349,598,1325,1351]" pageId="14" pageNumber="901" pagination="1875 - 1891" refId="ref18107" refString="STOEV P. &amp; ENGHOFF H. 2005. - A new cave-dwelling millipede of the genus Bollmania Silvestri, 1896 from Yunnan, China, with remarks on the reduction of the second female leg-pair (Diplopoda: Callipodida: Caspiopetalidae). Journal of Natural History 39: 1875 - 1891." type="journal article" year="2005">Stoev &amp; Enghoff 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>