260 lines
29 KiB
XML
260 lines
29 KiB
XML
<document id="DBDE56CB7BA5E0210B8064012CBE3E7A" ID-DOI="10.5281/zenodo.176960" ID-GBIF-Dataset="a67acf3d-f43a-4d16-b437-aacf9f0d09ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="176960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459844304451" checkinUser="plazi" docAuthor="Ziegler, Thomas, Hendrix, Ralf, Thanh, Vu Ngoc, Vogt, Martina, Forster, Bernhard & Kien, Dang Ngoc" docDate="2007" docId="03AC87D0B11FFF92FF469004FD688AB5" docLanguage="en" docName="zt01493p040.pdf" docOrigin="Zootaxa 1493" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Lycodon ruhstrati Fischer 1886" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="FF95FFA8B115FF9EFFD19607FFD28827" masterDocTitle="The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key" masterLastPageNumber="40" masterPageNumber="1" pageNumber="11" updateTime="1698224724941" updateUser="plazi">
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<mods:title id="D3E014B7DFFB11C2EE78DD703D8E9A90">The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key</mods:title>
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<mods:name id="8A2C6A3B14DD6CD6AB77E062CFA07CCC" type="personal">
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<mods:namePart id="ED9A643A23BD82D31F8EAA73256F5503">Ziegler, Thomas</mods:namePart>
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<mods:roleTerm id="A0392FC54E156EA97473ED06A6298B63">Author</mods:roleTerm>
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<mods:namePart id="BB828B915E1A2C827DE49F32398BD21C">Hendrix, Ralf</mods:namePart>
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<mods:name id="A8B1D366C2C81509AA370EEC9918091D" type="personal">
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<mods:roleTerm id="3BA433B11C0B8EBEDEA0F0B85F9E72D0">Author</mods:roleTerm>
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<mods:namePart id="05B2B5E860436108E15912CAEA5B86B6">Thanh, Vu Ngoc</mods:namePart>
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<mods:namePart id="3502421AD0F505F79850CC84E047247D">Vogt, Martina</mods:namePart>
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<mods:roleTerm id="E95189FB33DE66DB0B7E39B14FC874FE">Author</mods:roleTerm>
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<mods:namePart id="85620F06576015DE25A88347AB3453AC">Forster, Bernhard</mods:namePart>
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<mods:roleTerm id="622B72AEBEB6BAF413A5B8B2AEBB744A">Author</mods:roleTerm>
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<mods:namePart id="6D3258DB83119AE883B8FE0AFF0975D6">Kien, Dang Ngoc</mods:namePart>
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<mods:typeOfResource id="2DA058A4C9FFFFC5782C23090432387B">text</mods:typeOfResource>
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<mods:title id="B9A0A772F0DFA2826C475547FD40F1A0">Zootaxa</mods:title>
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<mods:part id="3F070B99C839B02F36E142F8A31202E6">
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<mods:date id="724A3535A39D080AAD0DDD20E781A1F6">2007</mods:date>
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<mods:detail id="68C9BEFD8C31C85098CA5A2938B59EAB" type="volume">
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<mods:number id="DFFD4D9B2CD1A9F82587D56D0B67878A">1493</mods:number>
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<mods:classification id="041BF96FD109041C540F024FB6B42FAB">journal article</mods:classification>
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<mods:identifier id="F9ABD6DF0F204A0A034E81F0849673CB" type="DOI">10.5281/zenodo.176960</mods:identifier>
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<mods:identifier id="F76AFB42500B2686CBDBB1F5A91A9AC0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="33A6D5A361AFB7E451D53B9A24860079" type="Zenodo-Dep">176960</mods:identifier>
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<treatment id="03AC87D0B11FFF92FF469004FD688AB5" ID-DOI="http://doi.org/10.5281/zenodo.6247457" ID-GBIF-Taxon="119361338" ID-Zenodo-Dep="6247457" LSID="urn:lsid:plazi:treatment:03AC87D0B11FFF92FF469004FD688AB5" httpUri="http://treatment.plazi.org/id/03AC87D0B11FFF92FF469004FD688AB5" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection id="C31F654DB11FFF95FF469004FAE58975" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BBA36C6B11FFF94FF469004FBA28E3A" blockId="10.[151,1136,1539,1565]" box="[151,1136,1539,1565]" pageId="10" pageNumber="11">
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<heading id="D0F281AAB11FFF94FF469004FBA28E3A" bold="true" box="[151,1136,1539,1565]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B971EAD4B11FFF94FF469004FBA28E3A" bold="true" box="[151,1136,1539,1565]" pageId="10" pageNumber="11">
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<taxonomicName id="4C054D45B11FFF94FF469004FD858E3A" authority="Fischer, 1886" authorityName="Fischer" authorityYear="1886" box="[151,599,1539,1565]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="ruhstrati">
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<emphasis id="B971EAD4B11FFF94FF469004FF2C8E3A" bold="true" box="[151,254,1539,1565]" italics="true" pageId="10" pageNumber="11">Lycodon</emphasis>
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cf.
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<emphasis id="B971EAD4B11FFF94FEFA9004FE478E3A" bold="true" box="[299,405,1539,1565]" italics="true" pageId="10" pageNumber="11">ruhstrati</emphasis>
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(
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<bibRefCitation id="EF944B37B11FFF94FE759004FD9C8E3A" author="Fischer" box="[420,590,1539,1565]" pageId="10" pageNumber="38" refString="Fischer, J. G. (1886) Herpetologische Notizen. Abh. Naturw. Ver. Hamburg, 9, 1 - 19." type="journal article" year="1886">Fischer, 1886</bibRefCitation>
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)
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</taxonomicName>
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: Ruhstrat's wolf snake / Ran khuyet rutrat
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36C6B11FFF95FF469054FAE58975" blockId="10.[151,1437,1619,2005]" lastBlockId="11.[151,1437,152,338]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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A female specimen (ZFMK 86451, SVL 466, TaL 142, TL
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<quantity id="4CFD9B23B11FFF94FC9C9054FC628E4A" box="[845,944,1619,1645]" metricMagnitude="-1" metricUnit="m" metricValue="6.08" pageId="10" pageNumber="11" unit="mm" value="608.0">608 mm</quantity>
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) was discovered at night of
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<date id="FFBB1006B11FFF94FB289054FA4A8E4A" box="[1273,1432,1619,1645]" pageId="10" pageNumber="11" value="2006-06-20">20 June 2006</date>
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, during the dry season, in the primary forest of U Bo region (
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<figureCitation id="133E2A43B11FFF94FCBF907CFC348EB2" box="[878,998,1659,1685]" captionStart="FIGURE 12" captionStartId="11.[151,255,1127,1151]" captionTargetBox="[151,1434,378,1102]" captionTargetId="figure@11.[151,1436,376,1103]" captionTargetPageId="11" captionText="FIGURE 12. Portrait of the preserved specimen of Lycodon cf. ruhstrati from Phong Nha - Ke Bang (ZFMK 86451). Photo: Thomas Ziegler" httpUri="https://zenodo.org/record/176972/files/figure.png" pageId="10" pageNumber="11">Figure 12</figureCitation>
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). The specimen was crawling on the forest ground near a steep cascade at an altitude of about
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<quantity id="4CFD9B23B11FFF94FCE890A4FC518E9A" box="[825,899,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="10" pageNumber="11" unit="m" value="520.0">520 m</quantity>
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above sea level. Scalation features are as follows: Eight supralabials, of which third to fifth supralabials are in contact with eye; ten infralabials; one loreal, not in contact with eye; one preocular, not in contact with frontal; prefrontal not entering eye; two postoculars; two anterior temporals, followed by three posterior temporals; 17 dorsal scale rows at midbody, feebly keeled except for outermost rows; vertebral scales not enlarged; two undivided preventrals and 229 ventrals; ventrals distinctly angulate laterally; anal entire; 104 divided subcaudals. The specimen had 19 dark transversal crossbands on its body dorsum. These dark crossbands are largest anteriorly and become shorter towards the cloaca. The 18 interspaces of the dark body crossbands are creamy white in the neck region, but then become distinctly darker. After the first body third the light interspaces consist of a greyish-brown bar that is edged by white and contains in its centre a darker transversal bar. Such a pattern continues on the tail dorsum, which bears ten dark and 11–12 light crossbars, including the tail tip. The body venter is completely yellowish to cream in its anterior fourth, then in part interspersed with a brownish-grey marbling that becomes more intensive in the last body half. The dark dorsal crossbands continue on the underside of the tail.
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</paragraph>
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</subSubSection>
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<caption id="DF7A664EB11EFF95FF469260FE5E8C86" httpUri="https://zenodo.org/record/176972/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1434,378,1102]" targetPageId="11">
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<paragraph id="8BBA36C6B11EFF95FF469260FE5E8C86" blockId="11.[151,1437,1127,1185]" pageId="11" pageNumber="12">
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<emphasis id="B971EAD4B11EFF95FF469260FEFA8C58" bold="true" box="[151,296,1127,1151]" pageId="11" pageNumber="12">FIGURE 12.</emphasis>
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Portrait of the preserved specimen of
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<taxonomicName id="4C054D45B11EFF95FD1D9260FC7D8C59" box="[716,943,1127,1150]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">
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<emphasis id="B971EAD4B11EFF95FD1D9260FCF58C59" box="[716,807,1127,1150]" italics="true" pageId="11" pageNumber="12">Lycodon</emphasis>
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cf.
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<emphasis id="B971EAD4B11EFF95FC839260FC7D8C59" box="[850,943,1127,1150]" italics="true" pageId="11" pageNumber="12">ruhstrati</emphasis>
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</taxonomicName>
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from Phong Nha - Ke Bang (ZFMK 86451). Photo: Thomas Ziegler
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</paragraph>
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</caption>
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<subSubSection id="C31F654DB11EFF92FF1792D2FD688AB5" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
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<paragraph id="8BBA36C6B11EFF95FF1792D2FE198EE8" blockId="11.[151,1437,1237,2023]" pageId="11" pageNumber="12">
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The specimen clearly is a member of the genus
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<taxonomicName id="4C054D45B11EFF95FD2A92D2FC738CC8" authority="Boie" authorityName="Boie" box="[763,929,1237,1263]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
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<emphasis id="B971EAD4B11EFF95FD2A92D2FC8D8CC9" box="[763,863,1237,1262]" italics="true" pageId="11" pageNumber="12">Lycodon</emphasis>
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Boie
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</taxonomicName>
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, due to its pholidosis, a vertically elliptical pupil, and because of its arched maxillary bone, that is bent inwards anteriorly, with 6 anterior fang-like teeth increasing in size and separated by a toothless interspace of about 0.9 mm width at left side from 6 or 7 posterior teeth, the last two ones of which being enlarged (
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<bibRefCitation id="EF944B37B11EFF95FCC1934AFC728D40" author="Lanza" box="[784,928,1357,1383]" pageId="11" pageNumber="39" refString="Lanza, B. (1999) A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropic. Zool., 12, 89 - 104." type="journal article" year="1999">Lanza 1999</bibRefCitation>
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). In the key provided by
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<bibRefCitation id="EF944B37B11EFF95FB1B934AFAB98D40" author="Lanza" box="[1226,1387,1357,1383]" pageId="11" pageNumber="39" refString="Lanza, B. (1999) A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropic. Zool., 12, 89 - 104." type="journal article" year="1999">Lanza (1999)</bibRefCitation>
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, the specimen from Phong Nha - Ke Bang, with some of its dorsal scale rows keeled along the whole body, clearly keys out as
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<taxonomicName id="4C054D45B11EFF95FECE939AFE768D91" box="[287,420,1437,1462]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">
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<emphasis id="B971EAD4B11EFF95FECE939AFE768D91" box="[287,420,1437,1462]" italics="true" pageId="11" pageNumber="12">L. ruhstrati</emphasis>
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</taxonomicName>
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at point nine. From the similar
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<taxonomicName id="4C054D45B11EFF95FCC2939AFC488D91" box="[787,922,1437,1462]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fasciatus">
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<emphasis id="B971EAD4B11EFF95FCC2939AFC488D91" box="[787,922,1437,1462]" italics="true" pageId="11" pageNumber="12">L. fasciatus</emphasis>
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</taxonomicName>
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, which is a common snake around the limestone outcrops and caves in the karst forest of Phong Nha - Ke Bang,
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<taxonomicName id="4C054D45B11EFF95FC1493C2FB998DF9" box="[965,1099,1477,1502]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">
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<emphasis id="B971EAD4B11EFF95FC1493C2FB998DF9" box="[965,1099,1477,1502]" italics="true" pageId="11" pageNumber="12">L. ruhstrati</emphasis>
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</taxonomicName>
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differs e. g. by the loreal not touching the eye, having slightly higher ventral (193–237 vs. 186–220) and subcaudal counts (77–119 vs. 66– 90), shorter posterior chinshields than the anterior ones (which is vice versa in the species
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<taxonomicName id="4C054D45B11EFF95FB169012FAFD8E09" box="[1223,1327,1557,1582]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fasciatus">
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<emphasis id="B971EAD4B11EFF95FB169012FAFD8E09" box="[1223,1327,1557,1582]" italics="true" pageId="11" pageNumber="12">fasciatus</emphasis>
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</taxonomicName>
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), as well as in lacking dark bands that completely encircle the body, as it is the case in
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<taxonomicName id="4C054D45B11EFF95FB86903AFB338E71" box="[1111,1249,1597,1622]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fasciatus">
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<emphasis id="B971EAD4B11EFF95FB86903AFB338E71" box="[1111,1249,1597,1622]" italics="true" pageId="11" pageNumber="12">L. fasciatus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EF944B37B11EFF95FB25903AFA458E70" author="Bourret" box="[1268,1431,1597,1623]" pageId="11" pageNumber="38" refString="Bourret, R. (1936) Les serpents de l'Indochine. II. Catalogue systematique descriptif. Henri Basuyau et Cie, Toulouse, 505 pp." type="book" year="1936">Bourret 1936</bibRefCitation>
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,
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<bibRefCitation id="EF944B37B11EFF95FF469062FEF78E58" author="Lanza" box="[151,293,1637,1663]" pageId="11" pageNumber="39" refString="Lanza, B. (1999) A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropic. Zool., 12, 89 - 104." type="journal article" year="1999">Lanza 1999</bibRefCitation>
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||
).
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<bibRefCitation id="EF944B37B11EFF95FEE89062FE048E58" author="Lanza" box="[313,470,1637,1663]" pageId="11" pageNumber="39" refString="Lanza, B. (1999) A new species of Lycodon from the Philippines, with a key to the genus (Reptilia Serpentes Colubridae). Tropic. Zool., 12, 89 - 104." type="journal article" year="1999">Lanza (1999)</bibRefCitation>
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further distinguishes in his key between the subspecies
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<taxonomicName id="4C054D45B11EFF95FBBB9062FA498E59" box="[1130,1435,1637,1662]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="ruhstrati" subSpecies="multifasciatus">
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<emphasis id="B971EAD4B11EFF95FBBB9062FA498E59" box="[1130,1435,1637,1662]" italics="true" pageId="11" pageNumber="12">L. ruhstrati multifasciatus</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F3127656B11EFF95FF09908AFEC98E80" box="[216,283,1677,1703]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
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and the nominate form from southern
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<collectingCountry id="F3127656B11EFF95FD36908AFCFF8E80" box="[743,813,1677,1703]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
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and
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<collectingCountry id="F3127656B11EFF95FCB9908AFC138E80" box="[872,961,1677,1703]" name="Taiwan" pageId="11" pageNumber="12">Taiwan</collectingCountry>
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, with which our specimen appears to be conspecific at first glance.
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</paragraph>
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<paragraph id="8BBA36C6B11EFF92FF1790DAFE458975" blockId="11.[151,1437,1237,2023]" lastBlockId="12.[151,1437,152,658]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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However,
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<bibRefCitation id="EF944B37B11EFF95FE9390DAFE168ED0" author="Ota" box="[322,452,1757,1783]" pageId="11" pageNumber="39" refString="Ota, H. (1988) Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. J. Taiwan Mus., 41, 85 - 91." type="journal article" year="1988">Ota (1988)</bibRefCitation>
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additionally found distinct meristic differences between the Taiwanese and continental populations of
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<emphasis id="B971EAD4B11EFF95FE4D9102FDEF8F39" box="[412,573,1797,1822]" italics="true" pageId="11" pageNumber="12">
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L. r.
|
||
<taxonomicName id="4C054D45B11EFF95FE079102FDEF8F39" box="[470,573,1797,1822]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">ruhstrati</taxonomicName>
|
||
</emphasis>
|
||
. These results might support the validity of
|
||
<taxonomicName id="4C054D45B11EFF95FB879102FA4A8F38" authority="Pope" authorityName="Pope" box="[1110,1432,1797,1823]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="futsingensis">
|
||
<emphasis id="B971EAD4B11EFF95FB879102FA828F39" box="[1110,1360,1797,1822]" italics="true" pageId="11" pageNumber="12">Dinodon futsingensis</emphasis>
|
||
Pope
|
||
</taxonomicName>
|
||
, that was described from "Futsing Hsien" in Fujian Province, continental
|
||
<collectingCountry id="F3127656B11EFF95FBDD912AFB878F60" box="[1036,1109,1837,1863]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
|
||
, and subsequently synonymized by
|
||
<bibRefCitation id="EF944B37B11EFF95FEC39152FE748F48" author="Pope" box="[274,422,1877,1903]" pageId="11" pageNumber="39" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of central Asia. Vol. X. American Museum Natural History, New York, 604 p." type="book" year="1935">Pope (1935)</bibRefCitation>
|
||
with
|
||
<taxonomicName id="4C054D45B11EFF95FE219152FD228F48" authority="Fischer" authorityName="Fischer" box="[496,752,1877,1903]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B11EFF95FE219152FDAB8F49" box="[496,633,1877,1902]" italics="true" pageId="11" pageNumber="12">L. ruhstrati</emphasis>
|
||
(Fischer)
|
||
</taxonomicName>
|
||
, that was originally described as
|
||
<taxonomicName id="4C054D45B11EFF95FB5A9152FA8A8F49" box="[1163,1368,1877,1902]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B11EFF95FB5A9152FA8A8F49" box="[1163,1368,1877,1902]" italics="true" pageId="11" pageNumber="12">Ophites ruhstrati</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F3127656B11EFF95FED4917AFE898FB0" box="[261,347,1917,1943]" name="Taiwan" pageId="11" pageNumber="12">Taiwan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF944B37B11EFF95FEBC917AFDDC8FB0" author="Fischer" box="[365,526,1917,1943]" pageId="11" pageNumber="38" refString="Fischer, J. G. (1886) Herpetologische Notizen. Abh. Naturw. Ver. Hamburg, 9, 1 - 19." type="journal article" year="1886">Fischer 1886</bibRefCitation>
|
||
). However, it is noteworthy, that
|
||
<bibRefCitation id="EF944B37B11EFF95FC73917AFB868FB0" author="Bourret" box="[930,1108,1917,1943]" pageId="11" pageNumber="38" refString="Bourret, R. (1936) Les serpents de l'Indochine. II. Catalogue systematique descriptif. Henri Basuyau et Cie, Toulouse, 505 pp." type="book" year="1936">Bourret (1936)</bibRefCitation>
|
||
states for "
|
||
<taxonomicName id="4C054D45B11EFF95FB30917AFF088F99" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="futsingense">
|
||
<emphasis id="B971EAD4B11EFF95FB30917AFF088F99" italics="true" pageId="11" pageNumber="12">Dinodon futsingense</emphasis>
|
||
</taxonomicName>
|
||
", that the dorsal scales are smooth and not keeled as it is the case in our recent finding from Phong Nha - Ke Bang. Pope's (1928)
|
||
<typeStatus id="54BE8864B11EFF95FE1391CAFE268FC0" box="[450,500,1997,2023]" pageId="11" pageNumber="12">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C054D45B11EFF95FD5E91CAFC558FC1" box="[655,903,1997,2022]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="futsingensis">
|
||
<emphasis id="B971EAD4B11EFF95FD5E91CAFC558FC1" box="[655,903,1997,2022]" italics="true" pageId="11" pageNumber="12">Dinodon futsingensis</emphasis>
|
||
</taxonomicName>
|
||
had smooth scales and also another continental specimen that was studied by us (ZFMK 23363 from Fujian) as well as a specimen from the Vietnamese province Ha Tinh, listed as
|
||
<taxonomicName id="4C054D45B119FF92FE0B96C7FD7E88FE" box="[474,684,192,217]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B119FF92FE0B96C7FD7E88FE" box="[474,684,192,217]" italics="true" pageId="12" pageNumber="13">Lycodon ruhstrati</emphasis>
|
||
</taxonomicName>
|
||
in Ziegler (2002). However,
|
||
<bibRefCitation id="EF944B37B119FF92FBD396C7FB4088FD" author="Pope" box="[1026,1170,192,218]" pageId="12" pageNumber="39" refString="Pope, C. H. (1929) Notes on reptiles from Fukien and other Chinese provinces. Bull. Amer. Mus. Nat. Hist., 58, 335 - 487, Pl. 17 - 20." type="journal article" year="1929">Pope (1929)</bibRefCitation>
|
||
reported also on specimens from Fujian and
|
||
<bibRefCitation id="EF944B37B119FF92FE7A96EFFDEA8925" box="[427,568,232,258]" pageId="12" pageNumber="39" refString="Mell, R. (1922) Beitrage zur Fauna sinica. I Die Vertebraten Suchinas; Feldlisten und Feldnoten der Sauger, Vogel, Reptilien, Batrachier. Arch. Naturgesch. Berlin, (A), 80, 1 - 143." type="journal article">Mell (1922)</bibRefCitation>
|
||
on a specimen from Guangdong, that bear dorsal keels. Taiwanese specimens known so far always had keeled dorsal scales (
|
||
<bibRefCitation id="EF944B37B119FF92FCD09717FC72890D" author="Fischer" box="[769,928,272,298]" pageId="12" pageNumber="38" refString="Fischer, J. G. (1886) Herpetologische Notizen. Abh. Naturw. Ver. Hamburg, 9, 1 - 19." type="journal article" year="1886">Fischer 1886</bibRefCitation>
|
||
:
|
||
<typeStatus id="54BE8864B119FF92FC7A9717FC0F890D" box="[939,989,272,298]" pageId="12" pageNumber="13">type</typeStatus>
|
||
series
|
||
<taxonomicName id="4C054D45B119FF92FBFE9717FA99890D" authority="Maki 1931" authorityName="Maki" authorityYear="1931" box="[1071,1355,272,298]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B119FF92FBFE9717FB68890E" box="[1071,1210,272,297]" italics="true" pageId="12" pageNumber="13">
|
||
<collectingCountry id="F3127656B119FF92FBFE9717FB95890E" box="[1071,1095,272,297]" name="Iceland" pageId="12" pageNumber="13">O</collectingCountry>
|
||
. ruhstrati
|
||
</emphasis>
|
||
,
|
||
<bibRefCitation id="EF944B37B119FF92FB189717FA99890D" author="Maki" box="[1225,1355,272,298]" pageId="12" pageNumber="39" refString="Maki, M. (1931) A monograph of the snakes of Japan. Dai-Ichi Shobo, Tokyo, 240 pp." type="book" year="1931">Maki 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
, Vogel & Brachtel accepted).
|
||
</paragraph>
|
||
<paragraph id="8BBA36C6B119FF92FF179767FD688AB5" blockId="12.[151,1437,152,658]" pageId="12" pageNumber="13">
|
||
Thus, future studies must clarify, whether the coexistence of continental representatives both with keeled and smooth dorsal scales is due to individual variation or rather indicating that different taxa are involved. It is further noteworthy, that female
|
||
<taxonomicName id="4C054D45B119FF92FDDB97B7FD4289EE" box="[522,656,432,457]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B119FF92FDDB97B7FD4289EE" box="[522,656,432,457]" italics="true" pageId="12" pageNumber="13">L. ruhstrati</emphasis>
|
||
</taxonomicName>
|
||
from continental
|
||
<collectingCountry id="F3127656B119FF92FCB197B7FC7489ED" box="[864,934,432,458]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
|
||
that were studied by
|
||
<bibRefCitation id="EF944B37B119FF92FB7197B7FAF289ED" author="Ota" box="[1184,1312,432,458]" pageId="12" pageNumber="39" refString="Ota, H. (1988) Taxonomic notes on Lycodon ruhstrati (Colubridae: Ophidia) from East Asia. J. Taiwan Mus., 41, 85 - 91." type="journal article" year="1988">Ota (1988)</bibRefCitation>
|
||
had lower ventral counts (197–227), lower subcaudal counts (77–95), and relatively higher transverse body and tail bands (22–38, 12–18) compared to our recent finding from Phong Nha - Ke Bang.
|
||
<bibRefCitation id="EF944B37B119FF92FBA69407FABD8A3D" box="[1143,1391,512,538]" pageId="12" pageNumber="39" refString="Nguyen, V. S., Ho, T. C. & Nguyen, Q. T. (2005) A checklist of amphibians and reptiles of Vietnam. Nha xuat ban nong nghiep, Hanoi, 180 pp." type="book">
|
||
Nguyen
|
||
<emphasis id="B971EAD4B119FF92FB0D9407FACA8A3E" box="[1244,1304,512,537]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
list both
|
||
<taxonomicName id="4C054D45B119FF92FF03942FFE1B8A66" box="[210,457,552,577]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="futsingensis">
|
||
<emphasis id="B971EAD4B119FF92FF03942FFE1B8A66" box="[210,457,552,577]" italics="true" pageId="12" pageNumber="13">Dinodon futsingensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C054D45B119FF92FDD3942FFD068A66" box="[514,724,552,577]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B119FF92FDD3942FFD068A66" box="[514,724,552,577]" italics="true" pageId="12" pageNumber="13">Lycodon ruhstrati</emphasis>
|
||
</taxonomicName>
|
||
as occurring in
|
||
<collectingCountry id="F3127656B119FF92FC5E942FFC258A65" box="[911,1015,552,578]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C054D45B119FF92FBD3942FFB7D8A66" box="[1026,1199,552,577]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="futsingensis">
|
||
<emphasis id="B971EAD4B119FF92FBD3942FFB7D8A66" box="[1026,1199,552,577]" italics="true" pageId="12" pageNumber="13">D. futsingensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3127656B119FF92FB27942FFAF68A65" box="[1270,1316,552,578]" name="Laos" pageId="12" pageNumber="13">Lao</collectingCountry>
|
||
Cai Province in the north southwards to Hoa Binh Province, and
|
||
<taxonomicName id="4C054D45B119FF92FC929457FBCA8A4E" box="[835,1048,592,617]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ruhstrati">
|
||
<emphasis id="B971EAD4B119FF92FC929457FBCA8A4E" box="[835,1048,592,617]" italics="true" pageId="12" pageNumber="13">Lycodon ruhstrati</emphasis>
|
||
</taxonomicName>
|
||
from Cao Bang Province in the north to Da Nang Province in central
|
||
<collectingCountry id="F3127656B119FF92FD9E947FFD648AB5" box="[591,694,632,658]" name="Vietnam" pageId="12" pageNumber="13">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |