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<document id="D6DD9A555103CBD55E0C9F479CB2BE90" ID-DOI="10.5281/zenodo.183072" ID-GBIF-Dataset="38ff6ff6-74c9-4f36-ba62-37a58323edbe" ID-ISSN="1175-5326" ID-Zenodo-Dep="183072" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459890232028" checkinUser="plazi" docAuthor="Turan, Davut, Ekmekçï, Güler &amp; Ïlhan, Alï" docDate="2008" docId="03AC87CDFF96FF84BC9AFF3CF1EEF83E" docLanguage="en" docName="zt01824p044.pdf" docOrigin="Zootaxa 1824" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Luciobarbus kottelati Turan, Ekmekçï &amp; Ïlhan, 2008, sp. nov." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FF95FFB5FF93FF83BC0DFFABF316FFDC" masterDocTitle="Luciobarbus kottelati, a new species of barbel (Teleostei: Cyprinidae) from the Büyük Menderes River, Turkey, with rediagnose of L. lydianus" masterLastPageNumber="44" masterPageNumber="35" pageNumber="40" updateTime="1698227955931" updateUser="plazi">
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<mods:title id="68CD05713B232EAF4A4B743BC6EC764E">Luciobarbus kottelati, a new species of barbel (Teleostei: Cyprinidae) from the Büyük Menderes River, Turkey, with rediagnose of L. lydianus</mods:title>
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<mods:namePart id="FFEEB5104D0869115912108B85C160B2">Turan, Davut</mods:namePart>
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<mods:namePart id="E378BF68389488FB8B37E602614C3578">Ekmekçï, Güler</mods:namePart>
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<mods:namePart id="BCD0DA05F7B030BF48A2DA8CECBCC863">Ïlhan, Alï</mods:namePart>
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<mods:date id="F4EFDD196FA711B067EB6D50A32401DB">2008</mods:date>
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<treatment id="03AC87CDFF96FF84BC9AFF3CF1EEF83E" ID-DOI="http://doi.org/10.5281/zenodo.6233281" ID-GBIF-Taxon="119371835" ID-Zenodo-Dep="6233281" LSID="urn:lsid:plazi:treatment:03AC87CDFF96FF84BC9AFF3CF1EEF83E" httpUri="http://treatment.plazi.org/id/03AC87CDFF96FF84BC9AFF3CF1EEF83E" lastPageId="7" lastPageNumber="42" pageId="5" pageNumber="40">
<subSubSection id="C31F6550FF96FF86BC9AFF3CF3F8FF08" pageId="5" pageNumber="40" type="nomenclature">
<paragraph id="8BBA36DBFF96FF86BC9AFF3CF113FF6D" blockId="5.[151,517,151,212]" box="[151,517,151,177]" pageId="5" pageNumber="40">
<heading id="D0F281B7FF96FF86BC9AFF3CF113FF6D" bold="true" box="[151,517,151,177]" fontSize="11" level="1" pageId="5" pageNumber="40" reason="1">
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<taxonomicName id="4C054D58FF96FF86BC9AFF3CF28FFF6D" ID-CoL="3WCKN" box="[151,409,151,177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="kottelati" status="sp. nov.">
<emphasis id="B971EAC9FF96FF86BC9AFF3CF28FFF6D" bold="true" box="[151,409,151,177]" italics="true" pageId="5" pageNumber="40">Luciobarbus kottelati</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24257B2FF96FF86BDAAFF3CF113FF6D" box="[423,517,151,177]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBA36DBFF96FF86BC9AFF11F3F8FF08" blockId="5.[151,517,151,212]" box="[151,238,186,212]" pageId="5" pageNumber="40">
(
<figureCitation id="133E2A5EFF96FF86BC92FF11F3F3FF08" box="[159,229,186,212]" captionStart="FIGURE 5" captionStartId="5.[151,255,1843,1867]" captionTargetBox="[151,1433,1350,1819]" captionTargetId="figure@5.[151,1436,1349,1820]" captionTargetPageId="5" captionText="FIGURE 5. Luciobarbus kottelati, holotype, FFR 454, 155 mm SL; Turkey: Büyük Menderes drainage." httpUri="https://zenodo.org/record/183077/files/figure.png" pageId="5" pageNumber="40">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF96FF86BC9AFEA1F0D5FE40" pageId="5" pageNumber="40" type="materials_examined">
<paragraph id="8BBA36DBFF96FF86BC9AFEA1F76FFE90" blockId="5.[151,1437,266,1292]" pageId="5" pageNumber="40">
<emphasis id="B971EAC9FF96FF86BC9AFEA1F206FEF8" bold="true" box="[151,272,266,292]" pageId="5" pageNumber="40">
<typeStatus id="54BE8879FF96FF86BC9AFEA1F21DFEF8" box="[151,267,266,292]" pageId="5" pageNumber="40" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED14AE1EFF96FF86BD15FEA1F261FEF8" box="[280,375,266,292]" pageId="5" pageNumber="40">IUSHM</collectionCode>
<quantity id="4CFD9B3EFF96FF86BD72FEA1F166FEF8" box="[383,624,266,292]" metricMagnitude="2" metricUnit="m" metricValue="4.532324999999999" metricValueMax="8.79165" metricValueMin="0.273" pageId="5" pageNumber="40" unit="mm" value="453232.5" valueMax="879165.0" valueMin="27300.0">27300-879, 165 mm</quantity>
<collectionCode id="ED14AE1EFF96FF86BE75FEA1F18BFEF8" box="[632,669,266,292]" pageId="5" pageNumber="40">SL</collectionCode>
,
<collectingCountry id="F312764BFF96FF86BEA4FEA1F017FEF8" box="[681,769,266,292]" name="Turkey" pageId="5" pageNumber="40">Turkey</collectingCountry>
: Aydın Prov.: River Büyük Menderes, Dandalas Stream, Karacasu,
<quantity id="4CFD9B3EFF96FF86BD1FFE99F24AFE90" box="[274,348,306,332]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="40" unit="km" value="90.0">90 km</quantity>
south of Aydın; D. Turan, S. Engin &amp; M. Gölükçetin,
<date id="FFBB101BFF96FF86BFD1FE99F762FE90" box="[988,1140,306,332]" pageId="5" pageNumber="40" value="2005-04-06">6 April 2005</date>
.
</paragraph>
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<emphasis id="B971EAC9FF96FF86BCC8FEF1F25CFEA8" bold="true" box="[197,330,346,372]" pageId="5" pageNumber="40">
<typeStatus id="54BE8879FF96FF86BCC8FEF1F252FEA8" box="[197,324,346,372]" pageId="5" pageNumber="40" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED14AE1EFF96FF86BD5FFEF1F2A6FEA8" box="[338,432,346,372]" pageId="5" pageNumber="40">IUSHM</collectionCode>
27300-878, 4, 120
<quantity id="4CFD9B3EFF96FF86BEADFEF1F016FEA8" box="[672,768,346,372]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="40" unit="mm" value="145.0">145 mm</quantity>
<collectionCode id="ED14AE1EFF96FF86BF0AFEF1F03BFEA8" box="[775,813,346,372]" pageId="5" pageNumber="40">SL</collectionCode>
; CMK 18592, 4, 111
<quantity id="4CFD9B3EFF96FF86B838FEF1F782FEA8" box="[1077,1172,346,372]" metricMagnitude="-1" metricUnit="m" metricValue="1.51" pageId="5" pageNumber="40" unit="mm" value="151.0">151 mm</quantity>
<collectionCode id="ED14AE1EFF96FF86B891FEF1F7D7FEA8" box="[1180,1217,346,372]" pageId="5" pageNumber="40">SL</collectionCode>
;
<collectionCode id="ED14AE1EFF96FF86B8C3FEF1F613FEA8" box="[1230,1285,346,372]" httpUri="http://grbio.org/cool/146d-d288" name="Food Science Australia, Ryde" pageId="5" pageNumber="40">FRR</collectionCode>
454, 10,
<quantity id="4CFD9B3EFF96FF86B978FEF1F3EEFE40" metricMagnitude="-1" metricUnit="m" metricValue="1.855" metricValueMax="2.8" metricValueMin="0.9099999999999999" pageId="5" pageNumber="40" unit="mm" value="185.5" valueMax="280.0" valueMin="91.0">91- 280 mm</quantity>
<collectionCode id="ED14AE1EFF96FF86BCF2FE29F232FE40" box="[255,292,386,412]" pageId="5" pageNumber="40">SL</collectionCode>
;
<collectionCode id="ED14AE1EFF96FF86BD3CFE29F27AFE40" LSID="urn:lsid:biocol.org:col:14489" box="[305,364,386,412]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14489" name="Cambridge University" pageId="5" pageNumber="40">CGE</collectionCode>
uncat., 2, 124
<quantity id="4CFD9B3EFF96FF86BE2CFE29F197FE40" box="[545,641,386,412]" metricMagnitude="-1" metricUnit="m" metricValue="1.43" pageId="5" pageNumber="40" unit="mm" value="143.0">143 mm</quantity>
<collectionCode id="ED14AE1EFF96FF86BE85FE29F1BBFE40" box="[648,685,386,412]" pageId="5" pageNumber="40">SL</collectionCode>
; same data as
<typeStatus id="54BE8879FF96FF86BF58FE29F0A9FE40" box="[853,959,386,412]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF96FF86BCCBFE01F70AFD68" pageId="5" pageNumber="40" type="diagnosis">
<paragraph id="8BBA36DBFF96FF86BCCBFE01F110FDE0" blockId="5.[151,1437,266,1292]" pageId="5" pageNumber="40">
<emphasis id="B971EAC9FF96FF86BCCBFE01F250FE18" bold="true" box="[198,326,426,452]" pageId="5" pageNumber="40">Diagnosis.</emphasis>
<taxonomicName id="4C054D58FF96FF86BD42FE01F142FE1F" box="[335,596,426,451]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="kotellatii">
<emphasis id="B971EAC9FF96FF86BD42FE01F142FE1F" box="[335,596,426,451]" italics="true" pageId="5" pageNumber="40">Luciobarbus kotellatii</emphasis>
</taxonomicName>
is distinguished from all other species of
<taxonomicName id="4C054D58FF96FF86B845FE01F7CAFE1F" box="[1096,1244,426,451]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF96FF86B845FE01F7CAFE1F" box="[1096,1244,426,451]" italics="true" pageId="5" pageNumber="40">Luciobarbus</emphasis>
</taxonomicName>
in
<collectingCountry id="F312764BFF96FF86B909FE01F67FFE18" box="[1284,1385,426,452]" name="Turkey" pageId="5" pageNumber="40">Anatolia</collectingCountry>
and Europe by the position of the dorsal-fin origin conspicuously behind the pelvic-fin origin (vs. directly above or very slightly behind) and by having 4346 total lateral line scales (vs.
<quantity id="4CFD9B3EFF96FF86B80CFE51F77CFDC8" box="[1025,1130,506,532]" metricMagnitude="0" metricUnit="m" metricValue="1.778" metricValueMax="2.286" metricValueMin="1.27" pageId="5" pageNumber="40" unit="in" value="70.0" valueMax="90.0" valueMin="50.0">5090 in</quantity>
all others, except
<taxonomicName id="4C054D58FF96FF86B94BFE51F3D9FDE7" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF96FF86B94BFE51F3D9FDE7" italics="true" pageId="5" pageNumber="40">L. lydianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF96FF86BD0AFD89F295FDE7" box="[263,387,546,571]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF96FF86BD0AFD89F295FDE7" box="[263,387,546,571]" italics="true" pageId="5" pageNumber="40">L. graecus</emphasis>
</taxonomicName>
[
<tableCitation id="C6870360FF96FF86BD9DFD89F2F8FDE0" box="[400,494,546,572]" captionStart="TABLE 2" captionStartId="6.[151,239,517,541]" captionText="TABLE 2. Frequency distribution of meristic features of Luciobarbus lydianus, L. graecus, L. kottelati and L. pectoralis." pageId="5" pageNumber="40">Table 2</tableCitation>
]).
</paragraph>
<paragraph id="8BBA36DBFF96FF86BC9AFDE1F70AFD68" blockId="5.[151,1437,266,1292]" pageId="5" pageNumber="40">
Other characters useful for identification are: 1415, modally 15, gill rakers on the first gill arch; lips thin; lower lip with a swollen median pad margined by a shallow groove in some specimens; last simple ray of the dorsal fin well ossified, with moderate serrae along posterior edge (
<figureCitation id="133E2A5EFF96FF86BFA0FD31F0E2FD68" box="[941,1012,666,692]" captionStart="FIGURE 2" captionStartId="2.[151,255,1476,1500]" captionTargetBox="[588,998,705,1452]" captionTargetId="figure@2.[588,1000,704,1452]" captionTargetPageId="2" captionText="FIGURE 2. Last simple dorsal-fin ray of: a, Luciobarbus lydianus, FFR 454, 157 mm SL; b, L. kottelati, FFR 452, 142 mm SL." httpUri="https://zenodo.org/record/183074/files/figure.png" pageId="5" pageNumber="40">Fig. 2</figureCitation>
-b).
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF96FF85BCCBFD69F68AFF06" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40" type="description">
<paragraph id="8BBA36DBFF96FF86BCCBFD69F1EBFC78" blockId="5.[151,1437,266,1292]" pageId="5" pageNumber="40">
<emphasis id="B971EAC9FF96FF86BCCBFD69F248FD00" bold="true" box="[198,350,706,732]" pageId="5" pageNumber="40">Description.</emphasis>
General appearance of body is shown in
<figureCitation id="133E2A5EFF96FF86BF47FD69F0BBFD00" box="[842,941,706,732]" captionStart="FIGURE 5" captionStartId="5.[151,255,1843,1867]" captionTargetBox="[151,1433,1350,1819]" captionTargetId="figure@5.[151,1436,1349,1820]" captionTargetPageId="5" captionText="FIGURE 5. Luciobarbus kottelati, holotype, FFR 454, 155 mm SL; Turkey: Büyük Menderes drainage." httpUri="https://zenodo.org/record/183077/files/figure.png" pageId="5" pageNumber="40">Figure 5</figureCitation>
and head from below in
<figureCitation id="133E2A5EFF96FF86B8DAFD69F62DFD00" box="[1239,1339,706,732]" captionStart="FIGURE 3" captionStartId="4.[151,255,1015,1039]" captionTargetBox="[151,1425,194,990]" captionTargetId="figure@4.[151,1436,194,991]" captionTargetPageId="4" captionText="FIGURE 3. Ventral view of head of: a, Luciobarbus lydianus, FFR 452, 187 mm SL; b, L. kottelati, FFR 454, 152 mm SL." httpUri="https://zenodo.org/record/183075/files/figure.png" pageId="5" pageNumber="40">Figure 3</figureCitation>
-b; morphometric and meristic data are given in Tables 1 and 2. Body moderately high and slightly compressed laterally, caudal peduncle deep. Dorsal profile of body slightly arched, ventral profile more or less similar. Dorsal head profile slightly convex. Upper lip thin; lower lip thicker than upper lip, with a median swollen pad margined by a shallow groove in some specimens (
<figureCitation id="133E2A5EFF96FF86BEDCFCC9F00FFCA0" box="[721,793,866,892]" captionStart="FIGURE 3" captionStartId="4.[151,255,1015,1039]" captionTargetBox="[151,1425,194,990]" captionTargetId="figure@4.[151,1436,194,991]" captionTargetPageId="4" captionText="FIGURE 3. Ventral view of head of: a, Luciobarbus lydianus, FFR 452, 187 mm SL; b, L. kottelati, FFR 454, 152 mm SL." httpUri="https://zenodo.org/record/183075/files/figure.png" pageId="5" pageNumber="40">Fig. 3</figureCitation>
-b). Maxillary barbel reaching anterior margin of eye. Mandibular barbel reaching posterior margin of eye.
</paragraph>
<paragraph id="8BBA36DBFF96FF86BCCBFC19F181FAD0" blockId="5.[151,1437,266,1292]" pageId="5" pageNumber="40">
Dorsal fin with 4 simple and 8 branched rays, outer margin slightly concave, its origin markedly behind vertical through pelvic-fin origin, last simple ray thick and strongly ossified (
<figureCitation id="133E2A5EFF96FF86B810FC71F772FC28" box="[1053,1124,986,1012]" captionStart="FIGURE 2" captionStartId="2.[151,255,1476,1500]" captionTargetBox="[588,998,705,1452]" captionTargetId="figure@2.[588,1000,704,1452]" captionTargetPageId="2" captionText="FIGURE 2. Last simple dorsal-fin ray of: a, Luciobarbus lydianus, FFR 454, 157 mm SL; b, L. kottelati, FFR 452, 142 mm SL." httpUri="https://zenodo.org/record/183074/files/figure.png" pageId="5" pageNumber="40">Fig. 2</figureCitation>
a). Pectoral fin with 1719 branched rays, outer margin convex. Pelvic fin with 1 simple and 8 branched rays, outer margin convex. Anal fin with 3 simple and 5 branched rays, outer margin convex. Caudal fin deeply forked, with lobes slightly rounded, lower lobe longer than upper lobe. Lateral line with 4346 pored scales on body and anterior part of caudal; 78 scale rows between lateral line and dorsal-fin origin; 56 scales between lateral line and anal-fin origin. Gill rakers 4+911= 1415 on outer side of fist gill arch; 4143 (modally 42) total vertebrae. Pharyngeal teeth 5.3.22.3.4, usually four on right and five on left in main row. Sexual dimorphism is present, male with tubercles on snout in spawning period.
</paragraph>
<caption id="DF7A6653FF96FF86BC9AF898F7E3F897" box="[151,1269,1843,1867]" httpUri="https://zenodo.org/record/183077/files/figure.png" pageId="5" pageNumber="40" targetBox="[151,1433,1350,1819]" targetPageId="5">
<paragraph id="8BBA36DBFF96FF86BC9AF898F7E3F897" blockId="5.[151,1269,1843,1867]" box="[151,1269,1843,1867]" pageId="5" pageNumber="40">
<emphasis id="B971EAC9FF96FF86BC9AF898F20CF897" bold="true" box="[151,282,1843,1867]" pageId="5" pageNumber="40">FIGURE 5.</emphasis>
<emphasis id="B971EAC9FF96FF86BD2BF89FF104F897" box="[294,530,1844,1867]" italics="true" pageId="5" pageNumber="40">
<taxonomicName id="4C054D58FF96FF86BD2BF89FF118F897" box="[294,526,1844,1867]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="40" phylum="Chordata" rank="species" species="kottelati">Luciobarbus kottelati</taxonomicName>
,
</emphasis>
holotype, FFR 454, 155 mm SL; Turkey: Büyük Menderes drainage.
</paragraph>
</caption>
<paragraph id="8BBA36DBFF96FF85BCC8F8D5F68AFF06" blockId="5.[151,1437,1918,2025]" lastBlockId="6.[151,1436,152,458]" lastPageId="6" lastPageNumber="41" pageId="5" pageNumber="40">
<emphasis id="B971EAC9FF96FF86BCC8F8D5F242F844" bold="true" box="[197,340,1918,1944]" pageId="5" pageNumber="40">Coloration.</emphasis>
Formalin preserved specimens, grey on back and brown on flanks, and yellowish white on belly. Dorsal and caudal fins greyish, pectoral, anal and pelvic fins yellowish. Flank scales with a faintly but distinctly marked dark brown edge along posterior margin; dark spot on scale pocket faintly marked. In life, dark brown on back and flanks, yellowish white on belly. Dorsal and caudal fins brown, pectoral, anal and pelvic fins yellowish. Each scale margined by a band of black pigments, forming a regular reticulated pattern.
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF95FF85BCC8FF4CF165FEA6" pageId="6" pageNumber="41" type="distribution">
<paragraph id="8BBA36DBFF95FF85BCC8FF4CF165FEA6" blockId="6.[151,1436,152,458]" pageId="6" pageNumber="41">
<emphasis id="B971EAC9FF95FF85BCC8FF4CF270FEDD" bold="true" box="[197,358,231,257]" pageId="6" pageNumber="41">Distribution.</emphasis>
<taxonomicName id="4C054D58FF95FF85BD7DFF43F178FEDD" box="[368,622,232,257]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BD7DFF43F178FEDD" box="[368,622,232,257]" italics="true" pageId="6" pageNumber="41">Luciobarbus kottelati</emphasis>
</taxonomicName>
is presently known only from the River Büyük Menderes drainage (
<figureCitation id="133E2A5EFF95FF85BC92FEBBF3FDFEF6" box="[159,235,272,298]" captionStart="FIGURE 4" captionStartId="4.[151,255,1849,1873]" captionTargetBox="[153,1434,1198,1825]" captionTargetId="figure@4.[151,1436,1197,1825]" captionTargetPageId="4" captionText="FIGURE 4. Distribution of some species of Luciobarbus in Turkey" httpUri="https://zenodo.org/record/183076/files/figure.png" pageId="6" pageNumber="41">Fig. 4</figureCitation>
). The Büyük Menderes originates from the Denizli and Aydın Mountains and it discharges into the Aegean Sea in Aydın.
<taxonomicName id="4C054D58FF95FF85BD96FE93F180FE8D" box="[411,662,312,337]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BD96FE93F180FE8D" box="[411,662,312,337]" italics="true" pageId="6" pageNumber="41">Luciobarbus kottelati</emphasis>
</taxonomicName>
was collected in clean and very slowly running waters, in habitats with coarse sand and fine gravel bottom.
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF95FF84BCCBFE2CF7A3FBFE" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41" type="etymology">
<paragraph id="8BBA36DBFF95FF85BCCBFE2CF2DCFE16" blockId="6.[151,1436,152,458]" pageId="6" pageNumber="41">
<emphasis id="B971EAC9FF95FF85BCCBFE2CF245FE7D" bold="true" box="[198,339,391,417]" pageId="6" pageNumber="41">Etymology.</emphasis>
The species is named for
<collectingCountry id="F312764BFF95FF85BE9FFE23F1E2FE7E" box="[658,756,392,418]" name="Mauritius" pageId="6" pageNumber="41">Maurice</collectingCountry>
<taxonomicName id="4C054D58FF95FF85BEF2FE23F077FE7E" box="[767,865,392,418]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="kottelat">Kottelat</taxonomicName>
, who contributed to the knowledge of the fish fauna of Europe and Asia.
</paragraph>
<caption id="DF7A6653FF95FF85BC9AFDAEF68BFDC1" box="[151,1437,517,541]" pageId="6" pageNumber="41">
<paragraph id="8BBA36DBFF95FF85BC9AFDAEF68BFDC1" blockId="6.[151,1437,517,850]" box="[151,1437,517,541]" pageId="6" pageNumber="41">
<emphasis id="B971EAC9FF95FF85BC9AFDAEF21EFDC1" bold="true" box="[151,264,517,541]" pageId="6" pageNumber="41">TABLE 2.</emphasis>
Frequency distribution of meristic features of
<emphasis id="B971EAC9FF95FF85BEFAFDADF7CDFDC1" box="[759,1243,518,541]" italics="true" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BEFAFDADF0F7FDC1" box="[759,993,518,541]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">Luciobarbus lydianus</taxonomicName>
,
<taxonomicName id="4C054D58FF95FF85BFE1FDADF748FDC1" box="[1004,1118,518,541]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="graecus">L. graecus</taxonomicName>
,
<taxonomicName id="4C054D58FF95FF85B864FDADF7CDFDC1" box="[1129,1243,518,541]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">L. kottelati</taxonomicName>
</emphasis>
and
<taxonomicName id="4C054D58FF95FF85B902FDADF68EFDC1" box="[1295,1432,518,541]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">L. pectoralis</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BBA36DBFF95FF85BDAAFD83F142FDE1" blockId="6.[151,1437,517,850]" box="[423,596,552,573]" pageId="6" pageNumber="41">Lateral line scales</paragraph>
<paragraph id="8BBA36DBFF95FF85BC92FDF9F64CFCDE" blockId="6.[151,1437,517,850]" pageId="6" pageNumber="41">
Species
<geoCoordinate id="EE31501CFF95FF85BD54FDF9F150FDBB" box="[345,582,594,615]" direction="north" orientation="latitude" pageId="6" pageNumber="41" precision="15" value="43.745834">N 43 44 45</geoCoordinate>
46 47 48 49 50 51 52 53 54 Mean
<taxonomicName id="4C054D58FF95FF85BC92FDD3F21FFD51" box="[159,265,632,653]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BC92FDD3F21FFD51" box="[159,265,632,653]" italics="true" pageId="6" pageNumber="41">L. lydianus</emphasis>
</taxonomicName>
14 - - 1 8 4 1 - - - - - -
<specimenCount id="9D03FD52FF95FF85B94FFDDCF3A5FD6E" pageId="6" pageNumber="41" type="generic">
46
<taxonomicName id="4C054D58FF95FF85BC92FD36F3A5FD6E" box="[159,179,669,690]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF95FF85BC92FD36F3A5FD6E" box="[159,179,669,690]" italics="true" pageId="6" pageNumber="41">L.</emphasis>
</taxonomicName>
</specimenCount>
graecus 4 - - 3 1 - - - - - - - -
<specimenCount id="9D03FD52FF95FF85B94FFD36F3A5FD0B" pageId="6" pageNumber="41" type="generic">
45
<taxonomicName id="4C054D58FF95FF85BC92FD69F3A5FD0B" box="[159,179,706,727]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BC92FD69F3A5FD0B" box="[159,179,706,727]" italics="true" pageId="6" pageNumber="41">L.</emphasis>
</taxonomicName>
</specimenCount>
kottelati
<date id="FFBB101BFF95FF85BD54FD69F2F8FD0B" box="[345,494,706,727]" pageId="6" pageNumber="41" value="1915-02-01">15 1 2</date>
4 8 - - - - - - -
<specimenCount id="9D03FD52FF95FF85B94FFD69F3A5FCDE" pageId="6" pageNumber="41" type="generic">
45
<taxonomicName id="4C054D58FF95FF85BC92FD46F3A5FCDE" box="[159,179,749,770]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BC92FD46F3A5FCDE" box="[159,179,749,770]" italics="true" pageId="6" pageNumber="41">L.</emphasis>
</taxonomicName>
</specimenCount>
pectoralis 11 - - - - - - 1 1 1 2
<date id="FFBB101BFF95FF85B8A8FD46F64CFCDE" box="[1189,1370,749,770]" pageId="6" pageNumber="41" value="1953-02-04">4 2 53</date>
</paragraph>
<paragraph id="8BBA36DBFF95FF85BDAAFCB9F16EFCFB" blockId="6.[151,1437,517,850]" box="[423,632,786,807]" pageId="6" pageNumber="41">Transverse line scales</paragraph>
<paragraph id="8BBA36DBFF95FF85BDAAFC96F772FC8E" blockId="6.[151,1437,517,850]" box="[423,1124,829,850]" pageId="6" pageNumber="41">Above lateral line Below lateral line</paragraph>
<paragraph id="8BBA36DBFF95FF85BD54FCCCF616FBFB" pageId="6" pageNumber="41">
<table id="F905C47BFF95007CBC9AFCCCF68BFBFB" box="[151,1437,871,1063]" gridcols="12" gridrows="5" pageId="6" pageNumber="41">
<tr id="35353499FF95007CBC9AFCCCF68BFCA0" box="[151,1437,871,892]" gridrow="0" pageId="6" pageNumber="41" rowspan-0="1">
<th id="76E45DE5FF95007CBD54FCCCF267FCA0" box="[345,369,871,892]" gridcol="1" gridrow="0" pageId="6" pageNumber="41">N</th>
<th id="76E45DE5FF95007CBDAAFCCCF2ECFCA0" box="[423,506,871,892]" gridcol="2" gridrow="0" pageId="6" pageNumber="41">7 8</th>
<th id="76E45DE5FF95007CBE23FCCCF12CFCA0" box="[558,570,871,892]" gridcol="3" gridrow="0" pageId="6" pageNumber="41">9</th>
<th id="76E45DE5FF95007CBE76FCCCF185FCA0" box="[635,659,871,892]" gridcol="4" gridrow="0" pageId="6" pageNumber="41">10</th>
<th id="76E45DE5FF95007CBECAFCCCF1C8FCA0" box="[711,734,871,892]" gridcol="5" gridrow="0" pageId="6" pageNumber="41">11</th>
<th id="76E45DE5FF95007CBF12FCCCF040FCA0" box="[799,854,871,892]" gridcol="6" gridrow="0" pageId="6" pageNumber="41">Mean</th>
<th id="76E45DE5FF95007CBFB7FCCCF0D0FCA0" box="[954,966,871,892]" gridcol="7" gridrow="0" pageId="6" pageNumber="41">5</th>
<th id="76E45DE5FF95007CB805FCCCF736FCA0" box="[1032,1056,871,892]" gridcol="8" gridrow="0" pageId="6" pageNumber="41">6</th>
<th id="76E45DE5FF95007CB85AFCCCF775FCA0" box="[1111,1123,871,892]" gridcol="9" gridrow="0" pageId="6" pageNumber="41">7</th>
<th id="76E45DE5FF95007CB8A8FCCCF7A7FCA0" box="[1189,1201,871,892]" gridcol="10" gridrow="0" pageId="6" pageNumber="41">8</th>
<th id="76E45DE5FF95007CB8F9FCCCF68BFCA0" box="[1268,1437,871,892]" gridcol="11" gridrow="0" pageId="6" pageNumber="41">Mean</th>
</tr>
<tr id="35353499FF95007CBC9AFC39F68BFC7B" box="[151,1437,914,935]" gridrow="1" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFC39F20EFC7B" box="[151,280,914,935]" gridcol="0" gridrow="1" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FC39F21FFC7B" box="[159,265,914,935]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BC92FC39F21FFC7B" box="[159,265,914,935]" italics="true" pageId="6" pageNumber="41">L. lydianus</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FC39F267FC7B" box="[345,369,914,935]" gridcol="1" gridrow="1" pageId="6" pageNumber="41">14</td>
<td id="76E45DE5FF95007CBDAAFC39F2ECFC7B" box="[423,506,914,935]" gridcol="2" gridrow="1" pageId="6" pageNumber="41">- 14</td>
<td id="76E45DE5FF95007CBE23FC39F12CFC7B" box="[558,570,914,935]" gridcol="3" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBE76FC39F185FC7B" box="[635,659,914,935]" gridcol="4" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBECAFC39F1C8FC7B" box="[711,734,914,935]" gridcol="5" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF12FC39F040FC7B" box="[799,854,914,935]" gridcol="6" gridrow="1" pageId="6" pageNumber="41">8</td>
<td id="76E45DE5FF95007CBFB7FC39F0D0FC7B" box="[954,966,914,935]" gridcol="7" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FC39F736FC7B" box="[1032,1056,914,935]" gridcol="8" gridrow="1" pageId="6" pageNumber="41">14</td>
<td id="76E45DE5FF95007CB85AFC39F775FC7B" box="[1111,1123,914,935]" gridcol="9" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FC39F7A7FC7B" box="[1189,1201,914,935]" gridcol="10" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FC39F68BFC7B" box="[1268,1437,914,935]" gridcol="11" gridrow="1" pageId="6" pageNumber="41">6</td>
</tr>
<tr id="35353499FF95007CBC9AFC16F68BFC0E" box="[151,1437,957,978]" gridrow="2" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFC16F20EFC0E" box="[151,280,957,978]" gridcol="0" gridrow="2" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FC16F212FC0E" box="[159,260,957,978]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF95FF85BC92FC16F212FC0E" box="[159,260,957,978]" italics="true" pageId="6" pageNumber="41">L. graecus</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FC16F267FC0E" box="[345,369,957,978]" gridcol="1" gridrow="2" pageId="6" pageNumber="41">4</td>
<td id="76E45DE5FF95007CBDAAFC16F2ECFC0E" box="[423,506,957,978]" gridcol="2" gridrow="2" pageId="6" pageNumber="41">- 4</td>
<td id="76E45DE5FF95007CBE23FC16F12CFC0E" box="[558,570,957,978]" gridcol="3" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBE76FC16F185FC0E" box="[635,659,957,978]" gridcol="4" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBECAFC16F1C8FC0E" box="[711,734,957,978]" gridcol="5" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF12FC16F040FC0E" box="[799,854,957,978]" gridcol="6" gridrow="2" pageId="6" pageNumber="41">8</td>
<td id="76E45DE5FF95007CBFB7FC16F0D0FC0E" box="[954,966,957,978]" gridcol="7" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FC16F736FC0E" box="[1032,1056,957,978]" gridcol="8" gridrow="2" pageId="6" pageNumber="41">4</td>
<td id="76E45DE5FF95007CB85AFC16F775FC0E" box="[1111,1123,957,978]" gridcol="9" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FC16F7A7FC0E" box="[1189,1201,957,978]" gridcol="10" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FC16F68BFC0E" box="[1268,1437,957,978]" gridcol="11" gridrow="2" pageId="6" pageNumber="41">6</td>
</tr>
<tr id="35353499FF95007CBC9AFC4CF68BFC20" box="[151,1437,999,1020]" gridrow="3" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFC4CF20EFC20" box="[151,280,999,1020]" gridcol="0" gridrow="3" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FC4CF211FC20" box="[159,263,999,1020]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BC92FC4CF211FC20" box="[159,263,999,1020]" italics="true" pageId="6" pageNumber="41">L. kottelati</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FC4CF267FC20" box="[345,369,999,1020]" gridcol="1" gridrow="3" pageId="6" pageNumber="41">15</td>
<td id="76E45DE5FF95007CBDAAFC4CF2ECFC20" box="[423,506,999,1020]" gridcol="2" gridrow="3" pageId="6" pageNumber="41">3 12</td>
<td id="76E45DE5FF95007CBE23FC4CF12CFC20" box="[558,570,999,1020]" gridcol="3" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBE76FC4CF185FC20" box="[635,659,999,1020]" gridcol="4" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBECAFC4CF1C8FC20" box="[711,734,999,1020]" gridcol="5" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF12FC4CF040FC20" box="[799,854,999,1020]" gridcol="6" gridrow="3" pageId="6" pageNumber="41">8</td>
<td id="76E45DE5FF95007CBFB7FC4CF0D0FC20" box="[954,966,999,1020]" gridcol="7" gridrow="3" pageId="6" pageNumber="41">4</td>
<td id="76E45DE5FF95007CB805FC4CF736FC20" box="[1032,1056,999,1020]" gridcol="8" gridrow="3" pageId="6" pageNumber="41">11</td>
<td id="76E45DE5FF95007CB85AFC4CF775FC20" box="[1111,1123,999,1020]" gridcol="9" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FC4CF7A7FC20" box="[1189,1201,999,1020]" gridcol="10" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FC4CF68BFC20" box="[1268,1437,999,1020]" gridcol="11" gridrow="3" pageId="6" pageNumber="41">6</td>
</tr>
<tr id="35353499FF95007CBC9AFBB9F68BFBFB" box="[151,1437,1042,1063]" gridrow="4" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFBB9F20EFBFB" box="[151,280,1042,1063]" gridcol="0" gridrow="4" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FBB9F20EFBFB" box="[159,280,1042,1063]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BC92FBB9F20EFBFB" box="[159,280,1042,1063]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FBB9F267FBFB" box="[345,369,1042,1063]" gridcol="1" gridrow="4" pageId="6" pageNumber="41">11</td>
<td id="76E45DE5FF95007CBDAAFBB9F2ECFBFB" box="[423,506,1042,1063]" gridcol="2" gridrow="4" pageId="6" pageNumber="41">- -</td>
<td id="76E45DE5FF95007CBE23FBB9F12CFBFB" box="[558,570,1042,1063]" gridcol="3" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBE76FBB9F185FBFB" box="[635,659,1042,1063]" gridcol="4" gridrow="4" pageId="6" pageNumber="41">5</td>
<td id="76E45DE5FF95007CBECAFBB9F1C8FBFB" box="[711,734,1042,1063]" gridcol="5" gridrow="4" pageId="6" pageNumber="41">6</td>
<td id="76E45DE5FF95007CBF12FBB9F040FBFB" box="[799,854,1042,1063]" gridcol="6" gridrow="4" pageId="6" pageNumber="41">11</td>
<td id="76E45DE5FF95007CBFB7FBB9F0D0FBFB" box="[954,966,1042,1063]" gridcol="7" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FBB9F736FBFB" box="[1032,1056,1042,1063]" gridcol="8" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB85AFBB9F775FBFB" box="[1111,1123,1042,1063]" gridcol="9" gridrow="4" pageId="6" pageNumber="41">9</td>
<td id="76E45DE5FF95007CB8A8FBB9F7A7FBFB" box="[1189,1201,1042,1063]" gridcol="10" gridrow="4" pageId="6" pageNumber="41">2</td>
<td id="76E45DE5FF95007CB8F9FBB9F68BFBFB" box="[1268,1437,1042,1063]" gridcol="11" gridrow="4" pageId="6" pageNumber="41">7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBA36DBFF95FF85BDAAFB96F11EF8FD" blockId="6.[423,902,1085,1106]" lastBlockId="6.[151,1436,1479,2025]" pageId="6" pageNumber="41">
Pectoral rays Gill rakers The species of
<taxonomicName id="4C054D58FF95FF85BD4AFA63F2CDFA3D" box="[327,475,1480,1505]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF95FF85BD4AFA63F2CDFA3D" box="[327,475,1480,1505]" italics="true" pageId="6" pageNumber="41">Luciobarbus</emphasis>
</taxonomicName>
from western
<collectingCountry id="F312764BFF95FF85BE89FA6CF1FFFA3D" box="[644,745,1479,1505]" name="Turkey" pageId="6" pageNumber="41">Anatolia</collectingCountry>
(
<taxonomicName id="4C054D58FF95FF85BEF7FA63F06AFA3D" box="[762,892,1480,1505]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BEF7FA63F06AFA3D" box="[762,892,1480,1505]" italics="true" pageId="6" pageNumber="41">L. lydianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF95FF85BFB9FA63F725FA3D" box="[948,1075,1480,1505]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BFB9FA63F725FA3D" box="[948,1075,1480,1505]" italics="true" pageId="6" pageNumber="41">L. kottelati</emphasis>
</taxonomicName>
) had been identified as
<taxonomicName id="4C054D58FF95FF85B946FA63F3F2F9D5" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85B946FA63F3F2F9D5" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF944B2AFF95FF85BD1EFA44F2DDF9D5" author="Ladiges" box="[275,459,1519,1545]" pageId="6" pageNumber="43" refString="Ladiges, W. (1960). Susswasserfische der Turkei, Teil 1 Cyprinidae. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 58, 105 - 150." type="journal article" year="1960">Ladiges (1960)</bibRefCitation>
.
<taxonomicName id="4C054D58FF95FF85BDD5FA5BF1A8F9D5" box="[472,702,1520,1545]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF95FF85BDD5FA5BF1A8F9D5" box="[472,702,1520,1545]" italics="true" pageId="6" pageNumber="41">Luciobarbus capito</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF95FF85BEF6FA5BF087F9D5" box="[763,913,1520,1545]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BEF6FA5BF087F9D5" box="[763,913,1520,1545]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
have been considered conspecific by
<bibRefCitation id="EF944B2AFF95FF85B954FA44F234F9ED" author="Karaman" pageId="6" pageNumber="43" refString="Karaman, L. S. (1971) Susswasserfische der Turkei, 8. Teil, Revision der Barben Europas, Vorderasiens und Nordafrikas. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 67: 175 - 254." type="journal article" year="1971">Karaman (1971)</bibRefCitation>
, followed by most Turkish authors (e.g.,
<bibRefCitation id="EF944B2AFF95FF85BF05F9BCF701F9ED" author="Geldiay" box="[776,1047,1559,1585]" pageId="6" pageNumber="43" refString="Geldiay, R. &amp; Balik, S. (1999) Turkiye Tatlisu Baliklari, Ege Universitesi Su Urunleri Fakultesi Yayinlari, No: 46, Ders Kitabi Dizini, No: 16, Izmir, 532 pp. [in Turkish]." type="book" year="1999">Geldiay &amp; Balık, 1999</bibRefCitation>
;
<bibRefCitation id="EF944B2AFF95FF85B829F9BCF7A6F9ED" author="Kuru" box="[1060,1200,1559,1585]" pageId="6" pageNumber="43" refString="Kuru, M. (2004). Recent systematic status of inland fishes of Turkey. Gazi Universitesi, Gazi E g itim Fakultesi Dergisi, 24, 3, 1 - 21." type="journal article" year="2004">Kuru, 2004</bibRefCitation>
). This point of view was not followed by
<bibRefCitation id="EF944B2AFF95FF85BDACF994F159F985" author="Almaca" box="[417,591,1599,1625]" pageId="6" pageNumber="43" refString="Almaca, C. (1982) Remarks on some Heckel's species of Barbus from Western Asia. Arquivos do Museu Bocage, Serie B, 2: 95 - 102." type="journal article" year="1982">Almaça (1982</bibRefCitation>
,
<bibRefCitation id="EF944B2AFF95FF85BE6CF994F188F985" author="Almaca" box="[609,670,1599,1625]" pageId="6" pageNumber="43" refString="Almaca, C. (1984) Form Relationships among western Palaearctic Species of Barbus (Cyprinidae, Pisces). Arquivos do Museu Bocage, Serie A, 2: 207 - 248." type="journal article" year="1984">1984</bibRefCitation>
,
<bibRefCitation id="EF944B2AFF95FF85BEBDF994F1F8F985" author="Almaca" box="[688,750,1599,1625]" pageId="6" pageNumber="43" refString="Almaca, C. (1986) On some Barbus Species from Western Asia (Cyprinidae, Pisces). Annalen des Naturhistorischen Museums in Wien, 87: 5 - 30." type="journal article" year="1986">1986</bibRefCitation>
),
<bibRefCitation id="EF944B2AFF95FF85BF05F994F0D3F985" author="Doadrio" box="[776,965,1599,1625]" pageId="6" pageNumber="43" refString="Doadrio, I. (1990) Phylogenetic relationships and classification of western Palearctic species of the genus Barbus (Osteichthyes, Cyprinidae). Aquatic Living Resources, 3: 265 - 282." type="journal article" year="1990">Doadrio (1990)</bibRefCitation>
,
<bibRefCitation id="EF944B2AFF95FF85BFDBF994F673F985" author="Banarescu" box="[982,1381,1599,1625]" pageId="6" pageNumber="43" refString="Banarescu, P. &amp; Bogustkaya, N. G. (2003) Barbus Cuvier, 1816. In: Banarescu, P. M. &amp; Bogustkaya, N. G. (Eds) The freshwater fishes of Europe. 5. Cyprinidae 2. Part II. Barbus. Aula Wiesbaden, 1 - 10" type="journal article" year="2003">Banarescu &amp; Bogutskaya (2003)</bibRefCitation>
and
<bibRefCitation id="EF944B2AFF95FF85BC9AF9CCF2EEF95D" author="Bogutskaya" box="[151,504,1639,1665]" pageId="6" pageNumber="43" refString="Bogutskaya, N. G. &amp; Naseka, A. M. (2004) Catalogue of agnathans and fishes of fresh and brackish waters of Russia with comments on nomenclature and taxonomy. Moscow: KMK Scientific Press. 389 pp. (in Russian)." type="book" year="2004">Bogutskaya &amp; Naseka (2004)</bibRefCitation>
, they considered
<taxonomicName id="4C054D58FF95FF85BECAF9C3F04AF95D" box="[711,860,1640,1665]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BECAF9C3F04AF95D" box="[711,860,1640,1665]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
as a valid species.
<taxonomicName id="4C054D58FF95FF85B84FF9C3F631F95D" box="[1090,1319,1640,1665]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF95FF85B84FF9C3F631F95D" box="[1090,1319,1640,1665]" italics="true" pageId="6" pageNumber="41">Luciobarbus capito</emphasis>
</taxonomicName>
is known from the Caspian basin and
<taxonomicName id="4C054D58FF95FF85BDE5F93BF168F975" box="[488,638,1680,1705]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BDE5F93BF168F975" box="[488,638,1680,1705]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
from the Orontes basin (
<bibRefCitation id="EF944B2AFF95FF85BFA6F924F758F975" author="Almaca" box="[939,1102,1679,1705]" pageId="6" pageNumber="43" refString="Almaca, C. (1986) On some Barbus Species from Western Asia (Cyprinidae, Pisces). Annalen des Naturhistorischen Museums in Wien, 87: 5 - 30." type="journal article" year="1986">Almaça,1986</bibRefCitation>
;
<bibRefCitation id="EF944B2AFF95FF85B856F924F699F975" author="Kottelat" box="[1115,1423,1679,1705]" pageId="6" pageNumber="43" refString="Kottelat, M. &amp; Freyhof, J. (2007). Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Freyhof, Berlin, Germany, xiv + 646 pp." type="book" year="2007">
<taxonomicName id="4C054D58FF95FF85B856F924F7AFF975" box="[1115,1209,1679,1705]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="kottelat">Kottelat</taxonomicName>
&amp; Freyhof, 2007
</bibRefCitation>
). Their ranges are widely disjunct (
<figureCitation id="133E2A5EFF95FF85BE28F91CF17AF90D" box="[549,620,1719,1745]" captionStart="FIGURE 4" captionStartId="4.[151,255,1849,1873]" captionTargetBox="[153,1434,1198,1825]" captionTargetId="figure@4.[151,1436,1197,1825]" captionTargetPageId="4" captionText="FIGURE 4. Distribution of some species of Luciobarbus in Turkey" httpUri="https://zenodo.org/record/183076/files/figure.png" pageId="6" pageNumber="41">Fig. 4</figureCitation>
) and they are readily distinguished by differences in total lateral line scale counts (5765 total scales in
<taxonomicName id="4C054D58FF95FF85BE21F94BF182F925" box="[556,660,1760,1785]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF95FF85BE21F94BF182F925" box="[556,660,1760,1785]" italics="true" pageId="6" pageNumber="41">L. capito</emphasis>
</taxonomicName>
vs.
<quantity id="4CFD9B3EFF95FF85BECEF974F03CF925" box="[707,810,1759,1785]" metricMagnitude="0" metricUnit="m" metricValue="1.3335" metricValueMax="1.3969999999999998" metricValueMin="1.27" pageId="6" pageNumber="41" unit="in" value="52.5" valueMax="55.0" valueMin="50.0">5055 in</quantity>
<taxonomicName id="4C054D58FF95FF85BF3CF94BF0D3F925" box="[817,965,1760,1785]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BF3CF94BF0D3F925" box="[817,965,1760,1785]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
) and gill raker counts (1719 gill rakers on the first gill arch vs. 1718).
</paragraph>
<paragraph id="8BBA36DBFF95FF85BD54FBCCF20DFA4A" pageId="6" pageNumber="41">
<table id="F905C47BFF95007CBC9AFBCCF68BFA4A" box="[151,1437,1127,1430]" gridcols="12" gridrows="6" pageId="6" pageNumber="41">
<tr id="35353499FF95007CBC9AFBCCF68BFBA0" box="[151,1437,1127,1148]" gridrow="0" pageId="6" pageNumber="41" rowspan-0="1">
<th id="76E45DE5FF95007CBD54FBCCF267FBA0" box="[345,369,1127,1148]" gridcol="1" gridrow="0" pageId="6" pageNumber="41">N</th>
<th id="76E45DE5FF95007CBDAAFBCCF2ECFBA0" box="[423,506,1127,1148]" gridcol="2" gridrow="0" pageId="6" pageNumber="41">17 18</th>
<th id="76E45DE5FF95007CBE23FBCCF150FBA0" box="[558,582,1127,1148]" gridcol="3" gridrow="0" pageId="6" pageNumber="41">19</th>
<th id="76E45DE5FF95007CBE76FBCCF1A4FBA0" box="[635,690,1127,1148]" gridcol="4" gridrow="0" pageId="6" pageNumber="41">Mean</th>
<th id="76E45DE5FF95007CBF12FBCCF021FBA0" box="[799,823,1127,1148]" gridcol="5" gridrow="0" pageId="6" pageNumber="41">14</th>
<th id="76E45DE5FF95007CBF63FBCCF090FBA0" box="[878,902,1127,1148]" gridcol="6" gridrow="0" pageId="6" pageNumber="41">15</th>
<th id="76E45DE5FF95007CBFB7FBCCF0C4FBA0" box="[954,978,1127,1148]" gridcol="7" gridrow="0" pageId="6" pageNumber="41">16</th>
<th id="76E45DE5FF95007CB805FBCCF736FBA0" box="[1032,1056,1127,1148]" gridcol="8" gridrow="0" pageId="6" pageNumber="41">17</th>
<th id="76E45DE5FF95007CB85AFBCCF779FBA0" box="[1111,1135,1127,1148]" gridcol="9" gridrow="0" pageId="6" pageNumber="41">18</th>
<th id="76E45DE5FF95007CB8A8FBCCF7ABFBA0" box="[1189,1213,1127,1148]" gridcol="10" gridrow="0" pageId="6" pageNumber="41">19</th>
<th id="76E45DE5FF95007CB8F9FBCCF68BFBA0" box="[1268,1437,1127,1148]" gridcol="11" gridrow="0" pageId="6" pageNumber="41">Mean</th>
</tr>
<tr id="35353499FF95007CBC9AFB39F68BFB7B" box="[151,1437,1170,1191]" gridrow="1" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFB39F20DFB7B" box="[151,283,1170,1191]" gridcol="0" gridrow="1" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FB39F21FFB7B" box="[159,265,1170,1191]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BC92FB39F21FFB7B" box="[159,265,1170,1191]" italics="true" pageId="6" pageNumber="41">L. lydianus</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FB39F267FB7B" box="[345,369,1170,1191]" gridcol="1" gridrow="1" pageId="6" pageNumber="41">14</td>
<td id="76E45DE5FF95007CBDAAFB39F2ECFB7B" box="[423,506,1170,1191]" gridcol="2" gridrow="1" pageId="6" pageNumber="41">10 3</td>
<td id="76E45DE5FF95007CBE23FB39F150FB7B" box="[558,582,1170,1191]" gridcol="3" gridrow="1" pageId="6" pageNumber="41">1</td>
<td id="76E45DE5FF95007CBE76FB39F1A4FB7B" box="[635,690,1170,1191]" gridcol="4" gridrow="1" pageId="6" pageNumber="41">17</td>
<td id="76E45DE5FF95007CBF12FB39F021FB7B" box="[799,823,1170,1191]" gridcol="5" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF63FB39F090FB7B" box="[878,902,1170,1191]" gridcol="6" gridrow="1" pageId="6" pageNumber="41">3</td>
<td id="76E45DE5FF95007CBFB7FB39F0C4FB7B" box="[954,978,1170,1191]" gridcol="7" gridrow="1" pageId="6" pageNumber="41">9</td>
<td id="76E45DE5FF95007CB805FB39F736FB7B" box="[1032,1056,1170,1191]" gridcol="8" gridrow="1" pageId="6" pageNumber="41">2</td>
<td id="76E45DE5FF95007CB85AFB39F779FB7B" box="[1111,1135,1170,1191]" gridcol="9" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FB39F7ABFB7B" box="[1189,1213,1170,1191]" gridcol="10" gridrow="1" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FB39F68BFB7B" box="[1268,1437,1170,1191]" gridcol="11" gridrow="1" pageId="6" pageNumber="41">16</td>
</tr>
<tr id="35353499FF95007CBC9AFB16F68BFB0E" box="[151,1437,1213,1234]" gridrow="2" pageId="6" pageNumber="41" rowspan-11="1">
<th id="76E45DE5FF95007CBC9AFB16F20DFB0E" box="[151,283,1213,1234]" gridcol="0" gridrow="2" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FB16F212FB0E" box="[159,260,1213,1234]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF95FF85BC92FB16F212FB0E" box="[159,260,1213,1234]" italics="true" pageId="6" pageNumber="41">L. graecus</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FB16F267FB0E" box="[345,369,1213,1234]" gridcol="1" gridrow="2" pageId="6" pageNumber="41">4</td>
<td id="76E45DE5FF95007CBDAAFB16F2ECFB0E" box="[423,506,1213,1234]" gridcol="2" gridrow="2" pageId="6" pageNumber="41">3 1</td>
<td id="76E45DE5FF95007CBE23FB16F150FB0E" box="[558,582,1213,1234]" gridcol="3" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBE76FB16F1A4FB0E" box="[635,690,1213,1234]" gridcol="4" gridrow="2" pageId="6" pageNumber="41">17</td>
<td id="76E45DE5FF95007CBF12FB16F021FB0E" box="[799,823,1213,1234]" gridcol="5" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF63FB16F090FB0E" box="[878,902,1213,1234]" gridcol="6" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBFB7FB16F0C4FB0E" box="[954,978,1213,1234]" gridcol="7" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FB16F736FB0E" box="[1032,1056,1213,1234]" gridcol="8" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB85AFB16F779FB0E" box="[1111,1135,1213,1234]" gridcol="9" gridrow="2" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FB16F7ABFB0E" box="[1189,1213,1213,1234]" gridcol="10" gridrow="2" pageId="6" pageNumber="41">-</td>
</tr>
<tr id="35353499FF95007CBC9AFB4CF68BFB20" box="[151,1437,1255,1276]" gridrow="3" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFB4CF20DFB20" box="[151,283,1255,1276]" gridcol="0" gridrow="3" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FB4CF211FB20" box="[159,263,1255,1276]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85BC92FB4CF211FB20" box="[159,263,1255,1276]" italics="true" pageId="6" pageNumber="41">L. kottelati</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FB4CF267FB20" box="[345,369,1255,1276]" gridcol="1" gridrow="3" pageId="6" pageNumber="41">15</td>
<td id="76E45DE5FF95007CBDAAFB4CF2ECFB20" box="[423,506,1255,1276]" gridcol="2" gridrow="3" pageId="6" pageNumber="41">4 9</td>
<td id="76E45DE5FF95007CBE23FB4CF150FB20" box="[558,582,1255,1276]" gridcol="3" gridrow="3" pageId="6" pageNumber="41">2</td>
<td id="76E45DE5FF95007CBE76FB4CF1A4FB20" box="[635,690,1255,1276]" gridcol="4" gridrow="3" pageId="6" pageNumber="41">18</td>
<td id="76E45DE5FF95007CBF12FB4CF021FB20" box="[799,823,1255,1276]" gridcol="5" gridrow="3" pageId="6" pageNumber="41">3</td>
<td id="76E45DE5FF95007CBF63FB4CF090FB20" box="[878,902,1255,1276]" gridcol="6" gridrow="3" pageId="6" pageNumber="41">12</td>
<td id="76E45DE5FF95007CBFB7FB4CF0C4FB20" box="[954,978,1255,1276]" gridcol="7" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FB4CF736FB20" box="[1032,1056,1255,1276]" gridcol="8" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB85AFB4CF779FB20" box="[1111,1135,1255,1276]" gridcol="9" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8A8FB4CF7ABFB20" box="[1189,1213,1255,1276]" gridcol="10" gridrow="3" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FB4CF68BFB20" box="[1268,1437,1255,1276]" gridcol="11" gridrow="3" pageId="6" pageNumber="41">15</td>
</tr>
<tr id="35353499FF95007CBC9AFAB9F68BFAFB" box="[151,1437,1298,1319]" gridrow="4" pageId="6" pageNumber="41">
<th id="76E45DE5FF95007CBC9AFAB9F20DFAFB" box="[151,283,1298,1319]" gridcol="0" gridrow="4" pageId="6" pageNumber="41">
<taxonomicName id="4C054D58FF95FF85BC92FAB9F20EFAFB" box="[159,280,1298,1319]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BC92FAB9F20EFAFB" box="[159,280,1298,1319]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
</th>
<td id="76E45DE5FF95007CBD54FAB9F267FAFB" box="[345,369,1298,1319]" gridcol="1" gridrow="4" pageId="6" pageNumber="41">11</td>
<td id="76E45DE5FF95007CBDAAFAB9F2ECFAFB" box="[423,506,1298,1319]" gridcol="2" gridrow="4" pageId="6" pageNumber="41">2 6</td>
<td id="76E45DE5FF95007CBE23FAB9F150FAFB" box="[558,582,1298,1319]" gridcol="3" gridrow="4" pageId="6" pageNumber="41">3</td>
<td id="76E45DE5FF95007CBE76FAB9F1A4FAFB" box="[635,690,1298,1319]" gridcol="4" gridrow="4" pageId="6" pageNumber="41">18</td>
<td id="76E45DE5FF95007CBF12FAB9F021FAFB" box="[799,823,1298,1319]" gridcol="5" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBF63FAB9F090FAFB" box="[878,902,1298,1319]" gridcol="6" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CBFB7FAB9F0C4FAFB" box="[954,978,1298,1319]" gridcol="7" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB805FAB9F736FAFB" box="[1032,1056,1298,1319]" gridcol="8" gridrow="4" pageId="6" pageNumber="41">8</td>
<td id="76E45DE5FF95007CB85AFAB9F779FAFB" box="[1111,1135,1298,1319]" gridcol="9" gridrow="4" pageId="6" pageNumber="41">3</td>
<td id="76E45DE5FF95007CB8A8FAB9F7ABFAFB" box="[1189,1213,1298,1319]" gridcol="10" gridrow="4" pageId="6" pageNumber="41">-</td>
<td id="76E45DE5FF95007CB8F9FAB9F68BFAFB" box="[1268,1437,1298,1319]" gridcol="11" gridrow="4" pageId="6" pageNumber="41">17</td>
</tr>
<tr id="35353499FF95007CBC9AFAD7F68BFA4A" box="[151,1437,1404,1430]" gridrow="5" pageId="6" pageNumber="41" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76E45DE5FF95007CBC9AFAD7F20DFA4A" box="[151,283,1404,1430]" gridcol="0" gridrow="5" pageId="6" pageNumber="41">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="8BBA36DBFF95FF84BCCBF884F168FEF6" blockId="6.[151,1436,1479,2025]" lastBlockId="7.[151,1436,152,2018]" lastPageId="7" lastPageNumber="42" pageId="6" pageNumber="41">
The range of the
<taxonomicName id="4C054D58FF95FF85BD9DF89BF132F895" box="[400,548,1840,1865]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF95FF85BD9DF89BF132F895" box="[400,548,1840,1865]" italics="true" pageId="6" pageNumber="41">Luciobarbus</emphasis>
</taxonomicName>
species of western
<collectingCountry id="F312764BFF95FF85BF01F884F067F895" box="[780,881,1839,1865]" name="Turkey" pageId="6" pageNumber="41">Anatolia</collectingCountry>
(
<taxonomicName id="4C054D58FF95FF85BF8EF89BF713F895" box="[899,1029,1840,1865]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BF8EF89BF713F895" box="[899,1029,1840,1865]" italics="true" pageId="6" pageNumber="41">L. lydianus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C054D58FF95FF85B819F89BF782F895" box="[1044,1172,1840,1865]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85B819F89BF782F895" box="[1044,1172,1840,1865]" italics="true" pageId="6" pageNumber="41">L. kottelati</emphasis>
</taxonomicName>
) are very distant from those of
<taxonomicName id="4C054D58FF95FF85BCF1F8F3F272F8AD" box="[252,356,1880,1905]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF95FF85BCF1F8F3F272F8AD" box="[252,356,1880,1905]" italics="true" pageId="6" pageNumber="41">L. capito</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF95FF85BD93F8F3F125F8AD" box="[414,563,1880,1905]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF95FF85BD93F8F3F125F8AD" box="[414,563,1880,1905]" italics="true" pageId="6" pageNumber="41">L. pectoralis</emphasis>
</taxonomicName>
, separated by several river drainages and belong to conspicuously distinct zoogeographic regions; also they have very different appearance and it is difficult to understand why they had been considered conspecific. Further, they do not seem closely related. Molecular data of
<bibRefCitation id="EF944B2AFF95FF85B8BDF80CF3F3F835" pageId="6" pageNumber="44" refString="Tsigenopoulos, C. S., Durand, J. D., Unlu, E. &amp; Berrebi, P. (2003). Rapid radiation of the Mediterranean Luciobarbus species (Cyprinidae) after the Messinian salinity crisis of the Mediterranean Sea, inferred from mitochondrial phylogenetic analysis. Biological Journal of the Linnean Society, 80: 207 - 222." type="journal article">
Tsigenopoulos
<emphasis id="B971EAC9FF95FF85B96CF803F68DF81D" box="[1377,1435,1960,1985]" italics="true" pageId="6" pageNumber="41">et al.</emphasis>
(2003)
</bibRefCitation>
also show that they belong to different lineages.
<taxonomicName id="4C054D58FF95FF85BF40F87BF746F835" box="[845,1104,2000,2025]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF95FF85BF40F87BF746F835" box="[845,1104,2000,2025]" italics="true" pageId="6" pageNumber="41">Luciobarbus lydianus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF95FF85B89FF87BF603F835" box="[1170,1301,2000,2025]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF95FF85B89FF87BF603F835" box="[1170,1301,2000,2025]" italics="true" pageId="6" pageNumber="41">L. kottelati</emphasis>
</taxonomicName>
are distinguished from
<taxonomicName id="4C054D58FF94FF84BD34FF33F2B6FF6D" box="[313,416,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF94FF84BD34FF33F2B6FF6D" box="[313,416,152,177]" italics="true" pageId="7" pageNumber="42">L. capito</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF94FF84BDD4FF33F17BFF6D" box="[473,621,152,177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF94FF84BDD4FF33F17BFF6D" box="[473,621,152,177]" italics="true" pageId="7" pageNumber="42">L. pectoralis</emphasis>
</taxonomicName>
by having fewer total lateral line scales (4347 vs.
<quantity id="4CFD9B3EFF94FF84B8CBFF33F63BFF6E" box="[1222,1325,152,178]" metricMagnitude="0" metricUnit="m" metricValue="1.3335" metricValueMax="1.3969999999999998" metricValueMin="1.27" pageId="7" pageNumber="42" unit="in" value="52.5" valueMax="55.0" valueMin="50.0">5055 in</quantity>
<taxonomicName id="4C054D58FF94FF84B939FF33F3DBFF05" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF94FF84B939FF33F3DBFF05" italics="true" pageId="7" pageNumber="42">L. pectoralis</emphasis>
</taxonomicName>
and 5272,usually
<quantity id="4CFD9B3EFF94FF84BDB6FF6BF134FF06" box="[443,546,192,218]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" metricValueMax="1.6764" metricValueMin="1.524" pageId="7" pageNumber="42" unit="in" value="63.0" valueMax="66.0" valueMin="60.0">6066 in</quantity>
<taxonomicName id="4C054D58FF94FF84BE27FF6BF184FF05" box="[554,658,192,217]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF94FF84BE27FF6BF184FF05" box="[554,658,192,217]" italics="true" pageId="7" pageNumber="42">L. capito</emphasis>
</taxonomicName>
), fewer scale rows between dorsal-fin origin and lateral line (78, vs.
<quantity id="4CFD9B3EFF94FF84BCB2FF43F230FEDE" box="[191,294,232,258]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="7" pageNumber="42" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">1011 in</quantity>
<taxonomicName id="4C054D58FF94FF84BD20FF43F2D7FEDD" box="[301,449,232,258]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF94FF84BD20FF43F22BFEDD" box="[301,317,232,257]" italics="true" pageId="7" pageNumber="42">L</emphasis>
.
<emphasis id="B971EAC9FF94FF84BD41FF43F2D7FEDD" box="[332,449,232,257]" italics="true" pageId="7" pageNumber="42">pectoralis</emphasis>
</taxonomicName>
and
<quantity id="4CFD9B3EFF94FF84BDF7FF43F174FEDE" box="[506,610,232,258]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="7" pageNumber="42" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
<taxonomicName id="4C054D58FF94FF84BE64FF43F1C7FEDD" box="[617,721,232,257]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF94FF84BE64FF43F1C7FEDD" box="[617,721,232,257]" italics="true" pageId="7" pageNumber="42">L. capito</emphasis>
</taxonomicName>
) and fewer scale row between anal-fin origin and lateral line (6, vs.
<quantity id="4CFD9B3EFF94FF84BCEBFEBBF227FEF6" box="[230,305,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="7" pageNumber="42" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">78 in</quantity>
<taxonomicName id="4C054D58FF94FF84BD35FEBBF2DAFEF5" box="[312,460,272,297]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF94FF84BD35FEBBF2DAFEF5" box="[312,460,272,297]" italics="true" pageId="7" pageNumber="42">L. pectoralis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C054D58FF94FF84BE08FEBBF17BFEF5" box="[517,621,272,297]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF94FF84BE08FEBBF17BFEF5" box="[517,621,272,297]" italics="true" pageId="7" pageNumber="42">L. capito</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBA36DBFF94FF84BCCBFE93F08BFD9E" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
The populations of
<taxonomicName id="4C054D58FF94FF84BDBFFE93F150FE8D" box="[434,582,312,337]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF94FF84BDBFFE93F150FE8D" box="[434,582,312,337]" italics="true" pageId="7" pageNumber="42">Luciobarbus</emphasis>
</taxonomicName>
from western
<collectingCountry id="F312764BFF94FF84BEFBFE93F04DFE8E" box="[758,859,312,338]" name="Turkey" pageId="7" pageNumber="42">Anatolia</collectingCountry>
belong to two species, at once distinguished by the respective positions of the origin of the dorsal and pelvic fins.
<taxonomicName id="4C054D58FF94FF84BFACFECBF78BFEA5" box="[929,1181,352,377]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84BFACFECBF78BFEA5" box="[929,1181,352,377]" italics="true" pageId="7" pageNumber="42">Luciobarbus kottelati</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C054D58FF94FF84BC9AFE23F20CFE7D" box="[151,282,392,417]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BC9AFE23F20CFE7D" box="[151,282,392,417]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
by having the pelvic-fin origin markedly in front of the dorsal-fin origin (vs. originating directly below or slightly in front of the dorsal-fin), by having a greater predorsal distance (54.960.0 % SL, vs. 52.0
<quantity id="4CFD9B3EFF94FF84BC9AFE73F3FFFE2E" box="[151,233,472,498]" metricMagnitude="0" metricUnit="m" metricValue="1.39954" pageId="7" pageNumber="42" unit="in" value="55.1">55.1 in</quantity>
<taxonomicName id="4C054D58FF94FF84BCFCFE73F262FE2D" box="[241,372,472,497]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BCFCFE73F262FE2D" box="[241,372,472,497]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
) and by having fewer serrae along posterior edge of last simple dorsal-fin ray (2327, vs. 2933). The position of the pelvic-fin origin in front of the dorsal-fin origin in fact also distinguishes
<taxonomicName id="4C054D58FF94FF84B937FDABF3A8FD9D" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84B937FDABF3A8FD9D" italics="true" pageId="7" pageNumber="42">L. kottelati</emphasis>
</taxonomicName>
from all other species of
<taxonomicName id="4C054D58FF94FF84BDE4FD83F16BFD9D" box="[489,637,552,577]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF94FF84BDE4FD83F16BFD9D" box="[489,637,552,577]" italics="true" pageId="7" pageNumber="42">Luciobarbus</emphasis>
</taxonomicName>
in Europe and
<collectingCountry id="F312764BFF94FF84BF3DFD83F08FFD9E" box="[816,921,552,578]" name="Turkey" pageId="7" pageNumber="42">Anatolia</collectingCountry>
.
</paragraph>
<paragraph id="8BBA36DBFF94FF84BCCBFDFBF756FCD6" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
In
<taxonomicName id="4C054D58FF94FF84BCE8FDFBF272FDB5" box="[229,356,592,617]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84BCE8FDFBF272FDB5" box="[229,356,592,617]" italics="true" pageId="7" pageNumber="42">L. kottelati</emphasis>
</taxonomicName>
, the anterior-most dorsal pterygiophore is inserted between the neural spines of vertebrae 12 and 13, or 13 and 14, the dorsal-fin origin is above vertebrae 15 or in space between vertebrae 14 and 15, and the base of the anterior pelvic-fin ray is below vertebra 14. In
<taxonomicName id="4C054D58FF94FF84BF76FD0BF0E8FD65" box="[891,1022,672,697]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BF76FD0BF0E8FD65" box="[891,1022,672,697]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
, the anterior-most dorsal pterygiophore is inserted between the neural spines of vertebrae 12 and 13, the dorsal-fin origin is above vertebra 14, and the base of the anterior pelvic-fin ray below vertebra 14 or 15 (modally 15).
</paragraph>
<paragraph id="8BBA36DBFF94FF84BCC8FCB3F7A3FBFE" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
Besides the very important difference in the respective position of the dorsal and pelvic fins,
<taxonomicName id="4C054D58FF94FF84B911FCB3F68AFCED" box="[1308,1436,792,817]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84B911FCB3F68AFCED" box="[1308,1436,792,817]" italics="true" pageId="7" pageNumber="42">L. kottelati</emphasis>
</taxonomicName>
further differs from
<taxonomicName id="4C054D58FF94FF84BD8EFCEBF113FC85" box="[387,517,832,857]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BD8EFCEBF113FC85" box="[387,517,832,857]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
by having a well ossified last simple dorsal-fin ray in adults (vs. weakly ossified;
<figureCitation id="133E2A5EFF94FF84BCDEFCC3F20FFC5E" box="[211,281,872,898]" captionStart="FIGURE 2" captionStartId="2.[151,255,1476,1500]" captionTargetBox="[588,998,705,1452]" captionTargetId="figure@2.[588,1000,704,1452]" captionTargetPageId="2" captionText="FIGURE 2. Last simple dorsal-fin ray of: a, Luciobarbus lydianus, FFR 454, 157 mm SL; b, L. kottelati, FFR 452, 142 mm SL." httpUri="https://zenodo.org/record/183074/files/figure.png" pageId="7" pageNumber="42">Fig. 2</figureCitation>
a,b), somewhat fewer scales in lateral line (4346, vs. 4548), fewer gill rakers on the first gill arch (1415, mode 15, vs. 1517, mode 16), the lower caudal-fin lobe longer than the upper one (vs. equal), a deeper body (body depth at dorsal-fin origin 25.628.9 % SL, mean 27.4, vs. 22.726.7, mean 24.4), a deeper caudal peduncle (caudal peduncle depth 11.412.6 % SL, mean 12.1, vs. 10.611.8, mean 11.4), and a shallower head (head depth at occiput 63.068.3, mean 66.1 % HL vs. 58.764.6, mean 62.1).
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF94FF84BCC8FB9BF69DF8C6" pageId="7" pageNumber="42" type="reference_group">
<paragraph id="8BBA36DBFF94FF84BCC8FB9BF258FB36" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
<taxonomicName id="4C054D58FF94FF84BCC8FB9BF2D5FB95" ID-CoL="3WCKN" box="[197,451,1072,1097]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84BCC8FB9BF2D5FB95" box="[197,451,1072,1097]" italics="true" pageId="7" pageNumber="42">Luciobarbus kottelati</emphasis>
</taxonomicName>
is also distinguished from all other
<taxonomicName id="4C054D58FF94FF84BF78FB9BF71FFB95" ID-CoL="5H62" box="[885,1033,1072,1097]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF94FF84BF78FB9BF71FFB95" box="[885,1033,1072,1097]" italics="true" pageId="7" pageNumber="42">Luciobarbus</emphasis>
</taxonomicName>
species in Western Asia (all geographically very distant) by having fewer lateral line scales (4346, vs.
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<taxonomicName id="4C054D58FF94FF84B840FBF3F633FBAD" ID-CoL="3WCK5" box="[1101,1317,1112,1137]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B971EAC9FF94FF84B840FBF3F633FBAD" box="[1101,1317,1112,1137]" italics="true" pageId="7" pageNumber="42">L. brachycephalus</emphasis>
</taxonomicName>
,
<quantity id="4CFD9B3EFF94FF84B938FBF3F68AFBAE" box="[1333,1436,1112,1138]" metricMagnitude="0" metricUnit="m" metricValue="1.5748" metricValueMax="1.8288" metricValueMin="1.3208" pageId="7" pageNumber="42" unit="in" value="62.0" valueMax="72.0" valueMin="52.0">5272 in</quantity>
<taxonomicName id="4C054D58FF94FF84BC9AFB2BF216FB45" ID-CoL="3WCK8" box="[151,256,1152,1177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="capito">
<emphasis id="B971EAC9FF94FF84BC9AFB2BF216FB45" box="[151,256,1152,1177]" italics="true" pageId="7" pageNumber="42">L. capito</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C054D58FF94FF84BD83FB2BF2E1FB45" ID-CoL="3WCL2" box="[398,503,1152,1177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="mursa">
<emphasis id="B971EAC9FF94FF84BD83FB2BF2E1FB45" box="[398,503,1152,1177]" italics="true" pageId="7" pageNumber="42">L. mursa</emphasis>
</taxonomicName>
,
<quantity id="4CFD9B3EFF94FF84BE0AFB2BF152FB46" box="[519,580,1152,1178]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="7" pageNumber="42" unit="in" value="56.0">56 in</quantity>
<emphasis id="B971EAC9FF94FF84BE41FB2BF1ADFB45" box="[588,699,1152,1177]" italics="true" pageId="7" pageNumber="42">
<taxonomicName id="4C054D58FF94FF84BE41FB2BF1A0FB45" ID-CoL="3WCKL" box="[588,694,1152,1177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kersin">L. kersin</taxonomicName>
,
</emphasis>
<quantity id="4CFD9B3EFF94FF84BECFFB2BF03DFB46" box="[706,811,1152,1178]" metricMagnitude="0" metricUnit="m" metricValue="1.3589000000000002" metricValueMax="1.3969999999999998" metricValueMin="1.3208000000000002" pageId="7" pageNumber="42" unit="in" value="53.5" valueMax="55.0" valueMin="52.0">5255 in</quantity>
<taxonomicName id="4C054D58FF94FF84BF3EFB2BF0DDFB45" ID-CoL="3WCL3" box="[819,971,1152,1177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="mystaceus">
<emphasis id="B971EAC9FF94FF84BF3EFB2BF0DDFB45" box="[819,971,1152,1177]" italics="true" pageId="7" pageNumber="42">L. mystaceus</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C054D58FF94FF84B846FB2BF612FB45" ID-CoL="3WCLF" box="[1099,1284,1152,1177]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="xanthopterus">
<emphasis id="B971EAC9FF94FF84B846FB2BF612FB45" box="[1099,1284,1152,1177]" italics="true" pageId="7" pageNumber="42">L. xanthopterus</emphasis>
</taxonomicName>
,
<quantity id="4CFD9B3EFF94FF84B919FB2BF66AFB46" box="[1300,1404,1152,1178]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" metricValueMax="1.8034" metricValueMin="1.6002" pageId="7" pageNumber="42" unit="in" value="67.0" valueMax="71.0" valueMin="63.0">6371 in</quantity>
<taxonomicName id="4C054D58FF94FF84B989FB2BF3EAFB1D" ID-CoL="3WCKD" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="esocinus">
<emphasis id="B971EAC9FF94FF84B989FB2BF3EAFB1D" italics="true" pageId="7" pageNumber="42">L. esocinus</emphasis>
</taxonomicName>
,
<quantity id="4CFD9B3EFF94FF84BD07FB03F253FB1E" box="[266,325,1192,1218]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="7" pageNumber="42" unit="in" value="53.0">53 in</quantity>
<taxonomicName id="4C054D58FF94FF84BD40FB03F2D2FB1D" ID-CoL="3WCL9" box="[333,452,1192,1217]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="scheich">
<emphasis id="B971EAC9FF94FF84BD40FB03F2D2FB1D" box="[333,452,1192,1217]" italics="true" pageId="7" pageNumber="42">L. scheich</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C054D58FF94FF84BE4CFB03F1C2FB1D" ID-CoL="3WCL6" box="[577,724,1192,1217]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pectoralis">
<emphasis id="B971EAC9FF94FF84BE4CFB03F1C2FB1D" box="[577,724,1192,1217]" italics="true" pageId="7" pageNumber="42">L. pectoralis</emphasis>
</taxonomicName>
,
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<taxonomicName id="4C054D58FF94FF84BF5FFB03F0CFFB1D" ID-CoL="3WCK2" box="[850,985,1192,1217]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="barbulus">
<emphasis id="B971EAC9FF94FF84BF5FFB03F0CFFB1D" box="[850,985,1192,1217]" italics="true" pageId="7" pageNumber="42">L. barbulus</emphasis>
</taxonomicName>
) (data from
<bibRefCitation id="EF944B2AFF94FF84B861FB03F605FB1E" author="Almaca" box="[1132,1299,1192,1218]" pageId="7" pageNumber="43" refString="Almaca, C. (1986) On some Barbus Species from Western Asia (Cyprinidae, Pisces). Annalen des Naturhistorischen Museums in Wien, 87: 5 - 30." type="journal article" year="1986">Almaça, 1986</bibRefCitation>
;
<bibRefCitation id="EF944B2AFF94FF84B912FB03F257FB36" author="Kottelat" pageId="7" pageNumber="43" refString="Kottelat, M. &amp; Freyhof, J. (2007). Handbook of European Freshwater Fishes. Kottelat, Cornol, Switzerland and Freyhof, Berlin, Germany, xiv + 646 pp." type="book" year="2007">
<taxonomicName id="4C054D58FF94FF84B912FB03F66BFB1E" box="[1311,1405,1192,1218]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="38" phylum="Chordata" rank="species" species="kottelat">Kottelat</taxonomicName>
&amp; Freyhof, 2007
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBA36DBFF94FF84BCC8FB53F69DF8C6" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
<taxonomicName id="4C054D58FF94FF84BCC8FB53F2D2FACD" ID-CoL="6QQCK" box="[197,452,1272,1297]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BCC8FB53F2D2FACD" box="[197,452,1272,1297]" italics="true" pageId="7" pageNumber="42">Luciobarbus lydianus</emphasis>
</taxonomicName>
is similar in appearance to
<taxonomicName id="4C054D58FF94FF84BF05FB53F092FACD" ID-CoL="3WCKG" box="[776,900,1272,1297]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF94FF84BF05FB53F092FACD" box="[776,900,1272,1297]" italics="true" pageId="7" pageNumber="42">L. graecus</emphasis>
</taxonomicName>
from the Kifissos drainage of central
<collectingCountry id="F312764BFF94FF84B94FFB53F68FFACE" box="[1346,1433,1272,1298]" name="Greece" pageId="7" pageNumber="42">Greece</collectingCountry>
. Their morphometric and meristic datas are given the Tables 1 and 2. Molecular data of
<bibRefCitation id="EF944B2AFF94FF84B8A1FA8BF3F3FABE" pageId="7" pageNumber="44" refString="Tsigenopoulos, C. S., Durand, J. D., Unlu, E. &amp; Berrebi, P. (2003). Rapid radiation of the Mediterranean Luciobarbus species (Cyprinidae) after the Messinian salinity crisis of the Mediterranean Sea, inferred from mitochondrial phylogenetic analysis. Biological Journal of the Linnean Society, 80: 207 - 222." type="journal article">
Tsigenopoulos
<emphasis id="B971EAC9FF94FF84B96DFA8BF68AFAE5" box="[1376,1436,1312,1337]" italics="true" pageId="7" pageNumber="42">et al.</emphasis>
(2003)
</bibRefCitation>
suggest that there is close relationship between
<taxonomicName id="4C054D58FF94FF84BF11FAE3F088FABD" ID-CoL="6QQCK" box="[796,926,1352,1377]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BF11FAE3F088FABD" box="[796,926,1352,1377]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
from Gediz River (Western
<collectingCountry id="F312764BFF94FF84B8E0FAE3F655FABE" box="[1261,1347,1352,1378]" name="Turkey" pageId="7" pageNumber="42">Turkey</collectingCountry>
) and
<taxonomicName id="4C054D58FF94FF84B989FAE3F3E2FA55" ID-CoL="3WCKG" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF94FF84B989FAE3F3E2FA55" italics="true" pageId="7" pageNumber="42">L. graecus</emphasis>
</taxonomicName>
from Sperchios River (Central
<collectingCountry id="F312764BFF94FF84BE65FADBF1D6FA56" box="[616,704,1392,1418]" name="Greece" pageId="7" pageNumber="42">Greece</collectingCountry>
). Their ranges are separated by the Aegean Sea and by a large area where no other
<taxonomicName id="4C054D58FF94FF84BD81FA33F136FA6D" ID-CoL="5H62" box="[396,544,1432,1457]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="B971EAC9FF94FF84BD81FA33F136FA6D" box="[396,544,1432,1457]" italics="true" pageId="7" pageNumber="42">Luciobarbus</emphasis>
</taxonomicName>
is known (part of Central
<collectingCountry id="F312764BFF94FF84BF52FA33F0A3FA6E" box="[863,949,1432,1458]" name="Greece" pageId="7" pageNumber="42">Greece</collectingCountry>
, northern
<collectingCountry id="F312764BFF94FF84B822FA33F794FA6E" box="[1071,1154,1432,1458]" name="Greece" pageId="7" pageNumber="42">Greece</collectingCountry>
and European
<collectingCountry id="F312764BFF94FF84B93AFA33F686FA6E" box="[1335,1424,1432,1458]" name="Turkey" pageId="7" pageNumber="42">Turkey</collectingCountry>
). The two species can be distinguished by the well-developed black spot on each scale pocket in
<taxonomicName id="4C054D58FF94FF84B8E5FA6BF67FFA05" ID-CoL="6QQCK" box="[1256,1385,1472,1497]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84B8E5FA6BF67FFA05" box="[1256,1385,1472,1497]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
(vs. absent or inconspicuous in
<taxonomicName id="4C054D58FF94FF84BDD4FA43F140F9DD" ID-CoL="3WCKG" box="[473,598,1512,1537]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF94FF84BDD4FA43F140F9DD" box="[473,598,1512,1537]" italics="true" pageId="7" pageNumber="42">L. graecus</emphasis>
</taxonomicName>
), the faintly marked and narrow black margin along the posterior edge of scales (vs. well marked and wide), the more pointed snout (vs. somewhat blunt), the wider mouth (vs. narrower; not quantifiable), the lips fleshy (vs. thin), with the lower lip conspicuously thicker than the upper lip (vs. only slightly thicker), the middle of the lower lip with a median fleshy pad with a shallow groove posteriorly (vs. without median fleshy pad). Moreover
<taxonomicName id="4C054D58FF94FF84BEDAF923F04CF97D" ID-CoL="6QQCK" box="[727,858,1672,1697]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="lydianus">
<emphasis id="B971EAC9FF94FF84BEDAF923F04CF97D" box="[727,858,1672,1697]" italics="true" pageId="7" pageNumber="42">L. lydianus</emphasis>
</taxonomicName>
has a shorter pectoral fin than
<taxonomicName id="4C054D58FF94FF84B8DCF923F651F97D" ID-CoL="3WCKG" box="[1233,1351,1672,1697]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="graecus">
<emphasis id="B971EAC9FF94FF84B8DCF923F651F97D" box="[1233,1351,1672,1697]" italics="true" pageId="7" pageNumber="42">L graecus</emphasis>
</taxonomicName>
(14.0
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% SL, mean 17.7, vs.19.920.8, mean 20.4), a wider head at anterior margin of eye (39.743.5 % HL, mean 41.1, vs. 34.738.8, mean 37.4), a more slender head (depth at occiput 58.764.6 % HL, mean 6.12, vs. 67.371.4, mean 68.3), and a wider interorbital (33.536.6 % HL, mean 34.9, vs. 31.633.0, mean 32.3).
</paragraph>
</subSubSection>
<subSubSection id="C31F6550FF94FF84BCCBF883F1EEF83E" pageId="7" pageNumber="42" type="description">
<paragraph id="8BBA36DBFF94FF84BCCBF883F1EEF83E" blockId="7.[151,1436,152,2018]" pageId="7" pageNumber="42">
<taxonomicName id="4C054D58FF94FF84BCCBF883F2D7F89D" box="[198,449,1832,1857]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84BCCBF883F2D7F89D" box="[198,449,1832,1857]" italics="true" pageId="7" pageNumber="42">Luciobarbus kottelati</emphasis>
</taxonomicName>
occurs in sympatry with
<taxonomicName id="4C054D58FF94FF84BEE5F883F0E2F89D" box="[744,1012,1832,1857]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pergamonensis">
<emphasis id="B971EAC9FF94FF84BEE5F883F0E2F89D" box="[744,1012,1832,1857]" italics="true" pageId="7" pageNumber="42">Barbus pergamonensis</emphasis>
</taxonomicName>
in the Büyük Menderes River. They are easily distinguished, with
<taxonomicName id="4C054D58FF94FF84BDF1F8FBF16BF8B5" box="[508,637,1872,1897]" class="Actinopterygii" family="Cyprinidae" genus="Luciobarbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="kottelati">
<emphasis id="B971EAC9FF94FF84BDF1F8FBF16BF8B5" box="[508,637,1872,1897]" italics="true" pageId="7" pageNumber="42">L. kottelati</emphasis>
</taxonomicName>
having fewer scales in lateral line (43 46, vs.
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<taxonomicName id="4C054D58FF94FF84B921F8FBF212F84D" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="42" phylum="Chordata" rank="species" species="pergamonensis">
<emphasis id="B971EAC9FF94FF84B921F8FBF212F84D" italics="true" pageId="7" pageNumber="42">B. pergamonensis</emphasis>
</taxonomicName>
), more gill rakers on the first gill arch (1415, vs. 810), a strong, spinous last simple dorsal ray (vs. weak, flexible), a lower lip without median lobe (vs. thick, fleshy, with a median lobe) and by the absence of black spots and blotches on the body (vs. presence).
</paragraph>
</subSubSection>
</treatment>
</document>