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<document id="D3DB3E7554EF0E27CE677A28781FC1D3" ID-DOI="10.1649/0010-065X-67.2.61" ID-Zenodo-Dep="5368429" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630260625427" checkinUser="carolina" docAuthor="Anderson, Robert S. &amp; Marvaldi, Adriana E." docDate="2013" docId="03AC87C00932FF8539B5EB1C045247CC" docLanguage="en" docName="0010-065X-67.2.61.pdf" docOrigin="The Coleopterists Bulletin 67 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-67.2.61" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Archicorynus kuscheli Anderson and Marvaldi 2013, new species" docType="treatment" docUuid="22DAAC8F-D832-4C9D-9141-838C0ED90385" docUuidSource="ZooBank" docVersion="5" lastPageNumber="71" masterDocId="FF95FFB80938FF8E3922EE640511465D" masterDocTitle="Finding Unexpected Beetles in Odd Places: Archicorynus kuscheliAnderson and Marvaldi, a New Genus and Species Representing a New Tribe, Archicorynini, of Oxycoryninae (Coleoptera: Belidae) from Nicaragua" masterLastPageNumber="71" masterPageNumber="61" pageNumber="61" updateTime="1699201200614" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1DC70A004FE6493A0B4453074D794DD5">Finding Unexpected Beetles in Odd Places: Archicorynus kuscheliAnderson and Marvaldi, a New Genus and Species Representing a New Tribe, Archicorynini, of Oxycoryninae (Coleoptera: Belidae) from Nicaragua</mods:title>
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<mods:name id="179BE7E22593D29C892C9D00FEAC4B6A" type="personal">
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</mods:role>
<mods:namePart id="E1CB73594C51521C88BD878C336575F6">Anderson, Robert S.</mods:namePart>
</mods:name>
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<mods:role id="9609A704E1ECA3790ADE43FA3D56F4FC">
<mods:roleTerm id="FF24B147F688435CE4239A9C30B852E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37A12FFDE39CE8D3C0BE8B71441D0C9B">Marvaldi, Adriana E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="80FD47C81548A19405E0EEEA398B0A3B">text</mods:typeOfResource>
<mods:relatedItem id="12C89206B71749E919100ACBC3F6F176" type="host">
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<mods:title id="4FF9A27AEC4705BE9183F7688E525D46">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="6A1F1455234FC0E4226A8C745FF2E4C4">
<mods:date id="38DCB07B3B37A01717C9A48D18381C22">2013</mods:date>
<mods:detail id="779A310BDDBECB099C9F08B75755D6EC" type="pubDate">
<mods:number id="B4B04F2215B1AA9C077A56F2639ACBB7">2013-06-30</mods:number>
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<mods:number id="095D4AAE21975A9FA1CD6FA6289C0D5C">67</mods:number>
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<mods:number id="2E0991C030E55C881C1A472D23D5AC10">2</mods:number>
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<mods:classification id="4D3DD80C893B3D6C32D840CDE51043AA">journal article</mods:classification>
<mods:identifier id="82B08E854FF21A744441B3C652B252C0" type="DOI">10.1649/0010-065X-67.2.61</mods:identifier>
<mods:identifier id="9A12868498DA07D9D46C963B5BAB64AB" type="Zenodo-Dep">5368429</mods:identifier>
<mods:identifier id="1BEA7B50BB64F99D3CC3F58EE4290364" type="ZooBank">F9908407-F3AF-43F4-B9D2-EA892143231D</mods:identifier>
</mods:mods>
<treatment id="03AC87C00932FF8539B5EB1C045247CC" LSID="urn:lsid:zoobank.org:act:22DAAC8F-D832-4C9D-9141-838C0ED90385" httpUri="http://treatment.plazi.org/id/03AC87C00932FF8539B5EB1C045247CC" lastPageId="11" pageId="10">
<subSubSection id="C31F655D0932FF8439B5EB1C04D443F5" pageId="10" type="nomenclature">
<paragraph id="8BBA36D60932FF8439B5EB1C04D443F5" blockId="10.[151,638,1400,1475]" pageId="10">
<heading id="D0F281BA0932FF8439B5EB1C04D443F5" centered="true" fontSize="10" level="2" pageId="10" reason="2">
<emphasis id="B971EAC40932FF8439B5EB1C04D443F5" bold="true" pageId="10">
<taxonomicName id="4C054D550932FF8439B5EB1C076843D0" authority="Anderson and Marvaldi" authorityName="Anderson and Marvaldi" authorityYear="2013" box="[151,633,1400,1422]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="kuscheli" status="sp. nov.">
<emphasis id="B971EAC40932FF8439B5EB1C046443D3" bold="true" box="[151,373,1400,1422]" italics="true" pageId="10">Archicorynus kuscheli</emphasis>
Anderson and Marvaldi
</taxonomicName>
,
<taxonomicNameLabel id="A24257BF0932FF84386DEBF704D443F5" box="[335,453,1427,1448]" pageId="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF84386DEBC904D7439F" box="[335,454,1453,1475]" pageId="10" type="description">
<paragraph id="8BBA36D60932FF84386DEBC904D7439F" blockId="10.[151,638,1400,1475]" box="[335,454,1453,1475]" pageId="10">
(
<figureCitation id="133E2A530932FF843874EBC904AE439F" box="[342,447,1453,1475]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="7.[197,238,1686,1705]" captionStartId-1="7.[731,771,666,685]" captionStartId-2="7.[731,771,1662,1681]" captionStartId-3="8.[175,215,1313,1332]" captionTargetBox-0="[173,659,1248,1663]" captionTargetBox-1="[723,1188,193,644]" captionTargetBox-2="[721,1144,1063,1639]" captionTargetBox-3="[157,1166,190,1291]" captionTargetId-0="figure-388@7.[173,659,1248,1663]" captionTargetId-1="figure-404@7.[707,1193,183,654]" captionTargetId-2="figure-425@7.[707,1193,1047,1672]" captionTargetId-3="figure-167@8.[151,1171,183,1307]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figs. 12. Archicorynus kuscheli, male. 1) Dorsal habitus; 2) Lateral habitus." captionText-1="Figs. 34. Archicorynus kuscheli, female. 3) Ventral view; 4) Legs: A) Front, B) Middle, C) Hind." captionText-2="Figs. 57. Archicorynus kuscheli, genitalia. 5) Female; 6) Aedeagus and tegmen, dorsal view; 7) Aedeagus and tegmen, lateral view." captionText-3="Figs. 813. Archicorynus kuscheli. 8) Male, ventral view; 9) Ventral surface of elytra, male (PLR = pocket-like cavity); 10) Female genitalia (BC = bursa copulatrix; HLE = hook-like extensions of T VIII; HS = gonocoxites; S = spermatheca; SD = spermathecal duct; SG = spermathecal gland; S VIII = sternite VIII; T VIII = tergite VIII); 11) Aedeagus, lateral view (PN = penis; PE = pedon; TC = tectum; TG = tegmen); 12) Penis (a) and tegmen (b), separated; 13) Hindwing, male." figureDoi-0="http://doi.org/10.5281/zenodo.5368435" figureDoi-1="http://doi.org/10.5281/zenodo.5368431" figureDoi-2="http://doi.org/10.5281/zenodo.5368433" figureDoi-3="http://doi.org/10.5281/zenodo.5368437" httpUri-0="https://zenodo.org/record/5368435/files/figure.png" httpUri-1="https://zenodo.org/record/5368431/files/figure.png" httpUri-2="https://zenodo.org/record/5368433/files/figure.png" httpUri-3="https://zenodo.org/record/5368437/files/figure.png" pageId="10">Figs. 113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF84398DEBBC062946AF" pageId="10" type="materials_examined">
<paragraph id="8BBA36D60932FF84398DEBBC076C409C" blockId="10.[151,639,1496,1729]" pageId="10">
<emphasis id="B971EAC40932FF84398DEBBC046C43B0" bold="true" box="[175,381,1496,1517]" pageId="10">Material Examined.</emphasis>
<materialsCitation id="3B6D3C8B0932FF8438A9EBBC05E04061" collectingDate="2011-05-03" collectionCode="LLAMA" collectorName="RN Cerro Musun" country="Nicaragua" elevation="0" latitude="12.96966" location="Types" longLatPrecision="419" longitude="-85.23341" pageId="10" specimenCount="1" stateProvince="Matagalpa" typeStatus="holotype">
<location id="8EDA600D0932FF8438A9EBBC04DD43B0" LSID="urn:lsid:plazi:treatment:03AC87C00932FF8539B5EB1C045247CC:8EDA600D0932FF8438A9EBBC04DD43B0" box="[395,460,1496,1517]" country="Nicaragua" latitude="12.96966" longLatPrecision="419" longitude="-85.23341" name="Types" pageId="10" stateProvince="Matagalpa">Types</location>
:
<typeStatus id="54BE88740932FF8438F8EBBC072943B0" box="[474,568,1496,1517]" pageId="10" type="holotype">holotype</typeStatus>
, male,
<collectingCountry id="F31276460932FF8439B5EB960420405A" box="[151,305,1522,1543]" name="Nicaragua" pageId="10">NICARAGUA</collectingCountry>
,
<collectingRegion id="49C1F8340932FF843862EB9604A1405A" box="[320,432,1522,1543]" country="Nicaragua" name="Matagalpa" pageId="10">Matagalpa</collectingRegion>
,
<collectorName id="26F053000932FF84389CEB96076B405A" box="[446,634,1522,1543]" pageId="10">RN Cerro Musún</collectorName>
,
<geoCoordinate id="EE3150110932FF8439B5E86905FE407F" box="[151,239,1549,1570]" degrees="12.96966" direction="north" orientation="latitude" pageId="10" precision="300" value="12.96966">12.96966</geoCoordinate>
<geoCoordinate id="EE3150110932FF8439DAE8690449407F" box="[248,344,1549,1570]" degrees="85.23341" direction="west" orientation="longitude" pageId="10" precision="300" value="-85.23341">-85.23341</geoCoordinate>
±
<elevation id="0028D1E50932FF84384FE86904E4407F" box="[365,501,1549,1570]" isEnumeration="true" pageId="10">
<quantity id="4CFD9B330932FF84384FE86904B4407F" box="[365,421,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" unit="m" value="300.0">300m</quantity>
,
<quantity id="4CFD9B330932FF843893E86904E4407F" box="[433,501,1549,1570]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="10" unit="m" value="1090.0">1090m</quantity>
</elevation>
,
<date id="FFBB10160932FF843B23E869076B407F" box="[513,634,1549,1570]" pageId="10" value="2011-05-03">
<collectingDate id="EFFFE9FE0932FF843B23E869076B407F" box="[513,634,1549,1570]" pageId="10" value="2011-05-03">3 May 2011</collectingDate>
</date>
,
<collectionCode id="ED14AE130932FF8439B5E84305E04061" box="[151,241,1575,1596]" pageId="10">LLAMA</collectionCode>
</materialsCitation>
#
<materialsCitation id="3B6D3C8B0932FF8439DCE843042B40D1" collectingDate="2011-05-03" collectionCode="CMNC, LLAMA" collectorName="RN Cerro Musun" country="Nicaragua" elevation="1" latitude="12.96966" location="Types" longLatPrecision="419" longitude="-85.23341" pageId="10" specimenCount="2" stateProvince="Matagalpa" typeStatus="paratype">
Go-D-01-1-01, montane wet forest, beating vegetation (
<collectionCode id="ED14AE130932FF843813E826046C400A" box="[305,381,1602,1623]" pageId="10">CMNC</collectionCode>
);
<typeStatus id="54BE88740932FF8438ADE82604FA400A" box="[399,491,1602,1623]" pageId="10" type="paratype">paratypes</typeStatus>
(6), same data as holotype
<quantity id="4CFD9B330932FF84382BE8390438402F" box="[265,297,1629,1650]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1.0">
<elevation id="0028D1E50932FF84382BE8390438402F" box="[265,297,1629,1650]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1.0">1m</elevation>
</quantity>
, 2f (
<collectionCode id="ED14AE130932FF843877E839048F402F" box="[341,414,1629,1650]" pageId="10">CMNC</collectionCode>
), same data as
<typeStatus id="54BE88740932FF843B0FE839076F402F" box="[557,638,1629,1650]" pageId="10" type="holotype">holotype</typeStatus>
except
<collectionCode id="ED14AE130932FF8439FCE813042B40D1" box="[222,314,1655,1676]" pageId="10">LLAMA</collectionCode>
</materialsCitation>
#
<materialsCitation id="3B6D3C8B0932FF843865E813075B409C" collectingDate="2011-05-01" collectionCode="CMNC, LLAMA" collectorName="Same" country="Nicaragua" elevation="1" latitude="12.96094" location="Types" longLatPrecision="14" longitude="-85.23268" pageId="10" specimenCount="1" stateProvince="Matagalpa" typeStatus="paratype">
Go-D-01-1-02,
<quantity id="4CFD9B330932FF8438FAE81304E840D1" box="[472,505,1655,1676]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1.0">
<elevation id="0028D1E50932FF8438FAE81304E840D1" box="[472,505,1655,1676]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1.0">1m</elevation>
</quantity>
, 1f (
<collectionCode id="ED14AE130932FF843B0AE813076540D1" box="[552,628,1655,1676]" pageId="10">CMNC</collectionCode>
).
<collectorName id="26F053000932FF8439B5E8F605C040FA" box="[151,209,1682,1703]" pageId="10">Same</collectorName>
data as holotype except
<date id="FFBB10160932FF8438D2E8F6076B40FA" box="[496,634,1682,1703]" pageId="10" value="2011-05-01">
<collectingDate id="EFFFE9FE0932FF8438D2E8F6076B40FA" box="[496,634,1682,1703]" pageId="10" value="2011-05-01">1 May 2011</collectingDate>
</date>
,
<geoCoordinate id="EE3150110932FF8439B5E8C805E2409C" box="[151,243,1708,1729]" degrees="12.96094" direction="north" orientation="latitude" pageId="10" precision="10" value="12.96094">12.96094</geoCoordinate>
<geoCoordinate id="EE3150110932FF8439D9E8C80471409C" box="[251,352,1708,1729]" degrees="85.23268" direction="west" orientation="longitude" pageId="10" precision="10" value="-85.23268">-85.23268</geoCoordinate>
±
<elevation id="0028D1E50932FF843854E8C804F9409C" box="[374,488,1708,1729]" isEnumeration="true" pageId="10">
<quantity id="4CFD9B330932FF843854E8C804B3409C" box="[374,418,1708,1729]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="10.0">10m</quantity>
,
<quantity id="4CFD9B330932FF84388CE8C804F9409C" box="[430,488,1708,1729]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="10" unit="m" value="750.0">750m</quantity>
</elevation>
,
<collectionCode id="ED14AE130932FF8438D6E8C8075B409C" box="[500,586,1708,1729]" pageId="10">LLAMA</collectionCode>
</materialsCitation>
#Wa-
</paragraph>
<paragraph id="8BBA36D60932FF843B8FEEA6062946AF" blockId="10.[685,1173,194,1729]" pageId="10">
D-01-1-15, tropical wet forest, ex sifted leaf litter,
<quantity id="4CFD9B330932FF843B8FEEB907DD46AF" box="[685,716,221,242]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1.0">1m</quantity>
(
<collectionCode id="ED14AE130932FF843BFAEEB9063C46AF" box="[728,813,221,242]" pageId="10">IADIZA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF843BE7EE93015E4701" pageId="10" type="etymology">
<paragraph id="8BBA36D60932FF843BE7EE93015E4701" blockId="10.[685,1173,194,1729]" pageId="10">
<emphasis id="B971EAC40932FF843BE7EE93062D4751" bold="true" box="[709,828,247,268]" pageId="10">Etymology.</emphasis>
This species is named after Guillermo (Willy) Kuschel, the outstanding expert on Curculionoidea, for his work on the ancient lineages including
<taxonomicName id="4C054D550932FF843A9DEF23015D4701" box="[959,1100,327,348]" class="Insecta" family="Belidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="subFamily" subFamily="Oxycoryninae">Oxycoryninae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF843BE7EF0506DD47BC" pageId="10" type="diagnosis">
<paragraph id="8BBA36D60932FF843BE7EF0506DD47BC" blockId="10.[685,1173,194,1729]" pageId="10">
<emphasis id="B971EAC40932FF843BE7EF05063F472B" bold="true" box="[709,814,353,374]" pageId="10">Diagnosis.</emphasis>
This species is easily recognized by the generic characters, especially by the carinate form of elytral interstriae 3 and 5 and the raised pronotal sculpturing. The fine, erect hairs fringing the elytra are also distinctive.
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF843BE7EF82067945E2" pageId="10" type="description">
<paragraph id="8BBA36D60932FF843BE7EF8201214558" blockId="10.[685,1173,194,1729]" pageId="10">
<emphasis id="B971EAC40932FF843BE7EF82065047A6" bold="true" box="[709,833,486,507]" pageId="10">Description.</emphasis>
Male, length
<quantity id="4CFD9B330932FF843AF7EF82014147A6" box="[981,1104,486,508]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.8" metricValueMin="3.1" pageId="10" unit="mm" value="3.45" valueMax="3.8" valueMin="3.1">3.13.8 mm</quantity>
, width
<quantity id="4CFD9B330932FF843B8FEC65063A444B" box="[685,811,512,534]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="10" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
; female, length 4.0
<quantity id="4CFD9B330932FF843ADCEC65015F444B" box="[1022,1102,513,534]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="10" unit="mm" value="4.6">4.6 mm</quantity>
, width
<quantity id="4CFD9B330932FF843B8FEC7F0639446D" box="[685,808,539,561]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="10" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
. Cuticle brown, mostly obscured by dense, fine, appressed to erect pilosity (and trapped dirt and debris). Head concave between eyes (sulcus covered by dense vestiture), eyes widely separated by distance subequal to basal width of rostrum. Rostrum short and robust, straight, approximately twice longer than wide in dorsal view in male, approximately 34 times in female, finely densely pilose throughout length.
</paragraph>
<paragraph id="8BBA36D60932FF843BE7ED6E067945E2" blockId="10.[685,1173,194,1729]" pageId="10">
<emphasis id="B971EAC40932FF843BE7ED6E067C4542" bold="true" box="[709,877,778,799]" pageId="10">Natural History.</emphasis>
Six of the
<specimenCount id="9D03FD5F0932FF843AC9ED6E01854542" box="[1003,1172,778,799]" pageId="10" type="generic">seven specimens</specimenCount>
were collected by breaking open dead twigs in an upper montane, wet, evergreen forest at around
<quantity id="4CFD9B330932FF843B8FED3E06104532" box="[685,769,858,879]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" unit="m" value="1000.0">1,000 m</quantity>
elevation. One specimen was collected in a Winkler-processed sample of leaf litter at a slightly lower elevation (
<quantity id="4CFD9B330932FF843A84EDEB06F845F9" box="[934,1001,911,932]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="10" unit="m" value="750.0">750 m</quantity>
) but in the same general forest
<typeStatus id="54BE88740932FF843A1AEDCE067545E2" box="[824,868,938,959]" pageId="10">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF843BE7EDA0016340D1" pageId="10" type="distribution">
<paragraph id="8BBA36D60932FF843BE7EDA0016340D1" blockId="10.[685,1173,194,1729]" pageId="10">
<emphasis id="B971EAC40932FF843BE7EDA006594584" bold="true" box="[709,840,964,985]" pageId="10">Distribution.</emphasis>
<taxonomicName id="4C054D550932FF843A77EDA1013E4587" box="[853,1071,965,986]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B971EAC40932FF843A77EDA1013E4587" box="[853,1071,965,986]" italics="true" pageId="10">Archicorynus kuscheli</emphasis>
</taxonomicName>
is known only from Reserva Natural Cerro Musún (
<geoCoordinate id="EE3150110932FF843D1DEDBB018145A9" box="[1087,1168,991,1012]" degrees="12.96" direction="north" orientation="latitude" pageId="10" precision="555" value="12.96">12.96°N</geoCoordinate>
,
<geoCoordinate id="EE3150110932FF843B8FED9E06194252" box="[685,776,1018,1039]" degrees="85.23" direction="west" orientation="longitude" pageId="10" precision="555" value="-85.23">85.23°W</geoCoordinate>
). Cerro Musún is an isolated mountain located in central
<collectingCountry id="F31276460932FF843A5FEA7006F84274" box="[893,1001,1044,1065]" name="Nicaragua" pageId="10">Nicaragua</collectingCountry>
(
<collectingRegion id="49C1F8340932FF843ADFEA7006074219" country="Nicaragua" name="Matagalpa" pageId="10">Departamento Matagalpa</collectingRegion>
) east of
<collectingRegion id="49C1F8340932FF843A5BEA4B06F14219" box="[889,992,1071,1092]" country="Nicaragua" name="Matagalpa" pageId="10">Matagalpa</collectingRegion>
and northeast of
<collectingRegion id="49C1F8340932FF843B8FEA2D061C4203" box="[685,781,1097,1118]" country="Nicaragua" name="Managua" pageId="10">Managua</collectingRegion>
. The entrance to the reserve is from the small town of Río Blanco (46 hours drive from
<collectingRegion id="49C1F8340932FF843BC8EA1B065C42C9" box="[746,845,1151,1172]" country="Nicaragua" name="Managua" pageId="10">Managua</collectingRegion>
). The official reserve includes 4,142 ha of protected area with good secondary and primary forest starting at about
<quantity id="4CFD9B330932FF843D30EAD001404294" box="[1042,1105,1204,1225]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="10" unit="m" value="600.0">600 m</quantity>
elevation and continuing up to the peak at
<quantity id="4CFD9B330932FF843D18EAAA018042BE" box="[1082,1169,1230,1251]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="10" unit="m" value="1438.0">1,438 m</quantity>
. The land at the base of Musún has been converted mostly to pasture and agricultural fields. Going up the mountain, habitats include humid tropical forest, riparian forest, montane wet forest, cloud forest, and dwarf forest (nebliselva) at the peak. The park is administered by the Ministerio del Ambiente y los Recursos Naturales in
<collectingRegion id="49C1F8340932FF843D08EBEC018043C0" box="[1066,1169,1416,1437]" country="Nicaragua" name="Matagalpa" pageId="10">Matagalpa</collectingRegion>
, which maintains a small station at the edge of the forest. We used this station as our base of operations during our stay. From the station there are two main trails, the “Sendero El Mojón” and the “Sendero Golondrina.” The Mojón trail runs from the station to the peak of Musún (23 hours to reach the peak). The Golondrina trail is a loop trail that borders one of the main streams near the station and passes by a series of picturesque waterfalls.
</paragraph>
</subSubSection>
<subSubSection id="C31F655D0932FF853BE7E8F6045247CC" lastPageId="11" pageId="10" type="discussion">
<paragraph id="8BBA36D60932FF853BE7E8F6045247CC" blockId="10.[685,1173,194,1729]" lastBlockId="11.[173,660,194,401]" lastPageId="11" pageId="10">
<emphasis id="B971EAC40932FF843BE7E8F6063840FA" bold="true" box="[709,809,1682,1703]" pageId="10">Remarks.</emphasis>
As the title of this paper suggests, one is often surprised by unexpected findings in places where we would never had searched for them. This is certainly the case for
<taxonomicName id="4C054D550933FF853B2CEEB9079D46AF" authorityName="kuscheli Anderson and Marvaldi, a" authorityYear="2013" box="[526,652,221,242]" class="Insecta" family="Belidae" genus="Archicorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
<emphasis id="B971EAC40933FF853B2CEEB9079D46AF" box="[526,652,221,242]" italics="true" pageId="11">Archicorynus</emphasis>
</taxonomicName>
. It underscores the importance of continued field collections and to extend this fieldwork into areas that may not have previously been sampled. Certainly this is more difficult logistically (and sometimes physically), but the rewards are worth the extra effort.
</paragraph>
</subSubSection>
</treatment>
</document>