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<document id="9FD5B6F7C95F2BE06D3ED6E92C2F9174" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFEB0732564EF8FE7CC9F936" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Rathbunaja ursus Ng &amp; Forges 2015, n. sp." docType="treatment" docUuid="EA0364F5-146A-4288-929C-60D73C5D0A3B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="216" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="216" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B6DE78854BE9DB798CD2E44EB47D8322" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B94750C58834A98ED085587954747618">
<mods:title id="CEE56F36A409A4D01D746FD58396F2F0">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
</mods:titleInfo>
<mods:name id="FBAA72F9CB3CB23753743A36EC6F3511" type="personal">
<mods:role id="EAB9564EE3F3B236D36E0B1EA132893E">
<mods:roleTerm id="5673D442049AB91B5F685B448A39BF3E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D2AF6C3C2D2C10EC5803AAF1065BB51">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="8F764F8751E3CA8AD88D4AFA987B6644" type="personal">
<mods:role id="EA6E7A7EEE87763C322BD98D1FED754D">
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</mods:role>
<mods:namePart id="6E9BBF1C55C6B585C1F9B0425D5B4D81">Forges, Bertrand Richer De</mods:namePart>
</mods:name>
<mods:typeOfResource id="34CA17D8DCA2854D92D9B51FE0AC0644">text</mods:typeOfResource>
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<mods:titleInfo id="78598E9101BD7EDCC4DEE4BD2952C962">
<mods:title id="B5D406F726AFE221A536F5697476964F">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="F31ECE691E4AB8FDF0EB46BD57452816">
<mods:date id="0DCB39C7896DDBBE14F14C736244D3BA">2015</mods:date>
<mods:detail id="900C0B88A3105A4B9F0EDF0E1EC69C32" type="pubDate">
<mods:number id="87044CB82A7928C4A54082EEE3B7D55A">2015-05-29</mods:number>
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<mods:detail id="7804F9AB90FD6B22BB7615E736D9C7AB" type="volume">
<mods:number id="B5D464066FFDE5EB5271EE2401E9F8CC">63</mods:number>
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<mods:start id="0FEEDFE6DAC00902B9681C1EE54972F0">110</mods:start>
<mods:end id="346731B5B6F6E2568C22F780A5BBBA08">225</mods:end>
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<mods:classification id="826935DCC8A20646F6313F0ADBFE34FE">journal article</mods:classification>
<mods:identifier id="AE1561B82DDF6E0CBDC314FC3B72C371" type="DOI">10.5281/zenodo.5384590</mods:identifier>
<mods:identifier id="D4DD139C1C58C4E98CE4765CE4AA6EF1" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="9EC46EF78854E13FB96DC70BF6625663" type="Zenodo-Dep">5384590</mods:identifier>
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<treatment id="03A9654BFFEB0732564EF8FE7CC9F936" ID-DOI="http://doi.org/10.5281/zenodo.5466833" ID-Zenodo-Dep="5466833" LSID="urn:lsid:zoobank.org:act:EA0364F5-146A-4288-929C-60D73C5D0A3B" httpUri="http://treatment.plazi.org/id/03A9654BFFEB0732564EF8FE7CC9F936" lastPageNumber="216" pageId="106" pageNumber="216">
<subSubSection id="C31A87D6FFEB0732564EF8FE7B5AF836" box="[297,568,1917,1941]" pageId="106" pageNumber="216" type="nomenclature">
<paragraph id="8BBFD45DFFEB0732564EF8FE7B5AF836" blockId="106.[148,716,1917,1973]" box="[297,568,1917,1941]" pageId="106" pageNumber="216">
<heading id="D0F76331FFEB0732564EF8FE7B5AF836" bold="true" box="[297,568,1917,1941]" centered="true" fontSize="10" level="2" pageId="106" pageNumber="216" reason="2">
<emphasis id="B974084FFFEB0732564EF8FE7B5AF836" bold="true" box="[297,568,1917,1941]" pageId="106" pageNumber="216">
<taxonomicName id="4C00AFDEFFEB0732564EF8FE7893F836" ID-CoL="4RLXG" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[297,497,1917,1941]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFEB0732564EF8FE7893F836" bold="true" box="[297,497,1917,1941]" italics="true" pageId="106" pageNumber="216">Rathbunaja ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFEB0732569DF8FE7B5AF836" box="[506,568,1917,1941]" pageId="106" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFEB073257F3F81E7BAEF816" box="[148,716,1949,1973]" pageId="106" pageNumber="216" type="description">
<paragraph id="8BBFD45DFFEB073257F3F81E7BAEF816" blockId="106.[148,716,1917,1973]" box="[148,716,1949,1973]" pageId="106" pageNumber="216">
(
<figureCitation id="133BC8D8FFEB073257FBF81E7995F816" box="[156,247,1949,1973]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Figs. 60</figureCitation>
,
<figureCitation id="133BC8D8FFEB07325663F81E783AF816" box="[260,344,1949,1973]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="106" pageNumber="216">61G, H</figureCitation>
,
<figureCitation id="133BC8D8FFEB07325603F81E78D0F816" box="[356,434,1949,1973]" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="106" pageNumber="216">63E–H</figureCitation>
,
<figureCitation id="133BC8D8FFEB073256D8F81E7B68F816" box="[447,522,1949,1973]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">64E, F</figureCitation>
,
<figureCitation id="133BC8D8FFEB07325570F81E7B0AF816" box="[535,616,1949,1973]" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="106" pageNumber="216">65H–K</figureCitation>
,
<figureCitation id="133BC8D8FFEB07325512F81E7BA7F816" box="[629,709,1949,1973]" captionStart="Fig" captionStartId="105.[135,170,1959,1981]" captionTargetBox="[137,1474,266,1944]" captionTargetId="figure-8@105.[127,1495,262,1947]" captionTargetPageId="105" captionText="Fig. 67. Left G1s, Rathbunaja species. A–D, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; E–G, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea. A, E = 5.0 mm; B-G = 1.0 mm; F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384740" httpUri="https://zenodo.org/record/5384740/files/figure.png" pageId="106" pageNumber="216">67A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFEB07325716F85E7CCFFD96" pageId="106" pageNumber="216" type="materials_examined">
<paragraph id="8BBFD45DFFEB07325716F85E7CCFFD96" blockId="106.[113,752,2013,2101]" lastBlockId="106.[811,1453,189,566]" pageId="106" pageNumber="216">
<emphasis id="B974084FFFEB07325716F85E783EF856" bold="true" box="[113,348,2013,2037]" pageId="106" pageNumber="216">Material examined.</emphasis>
<materialsCitation id="3B68DE00FFEB07325600F85E7D65FF76" collectingDate="2004-03-02" collectionCode="NMCR" county="Balicasag Island" elevation="250" location="Philippines" municipality="Panglao" pageId="106" pageNumber="216" specimenCode="ZRC 2013.1275" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54BB6AFFFFEB07325600F85E78B1F856" box="[359,467,2013,2037]" pageId="106" pageNumber="216" type="holotype">Holotype</typeStatus>
: male (50.1 ×
<quantity id="4CF879B8FFEB073255E6F85E7B8AF856" box="[641,744,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" pageId="106" pageNumber="216" unit="mm" value="41.5">41.5 mm</quantity>
) (
<collectionCode id="ED114C98FFEB0732571FF87E79AEF7B6" box="[120,204,2045,2069]" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="106" pageNumber="216" type="Herbarium">NMCR</collectionCode>
, ex
<specimenCode id="DBA67C26FFEB07325799F87E78D7F7B6" box="[254,437,2045,2069]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="106" pageNumber="216">ZRC 2013.1275</specimenCode>
),
<collectingCounty id="62DEACD1FFEB073256ACF87E7BE1F7B6" box="[459,643,2045,2069]" pageId="106" pageNumber="216">Balicasag Island</collectingCounty>
,
<collectingMunicipality id="6BDB4E27FFEB073255F5F87E7B8EF7B6" box="[658,748,2045,2069]" pageId="106" pageNumber="216">Panglao</collectingMunicipality>
,
<collectingRegion id="49C41ABFFFEB07325716F79E79D7F796" box="[113,181,2077,2101]" country="Philippines" name="Bohol" pageId="106" pageNumber="216">Bohol</collectingRegion>
,
<location id="8EDF8286FFEB073257A6F79E785EF796" LSID="urn:lsid:plazi:treatment:03A9654BFFEB0732564EF8FE7CC9F936:8EDF8286FFEB073257A6F79E785EF796" box="[193,316,2077,2101]" county="Balicasag Island" municipality="Panglao" name="Philippines" pageId="106" pageNumber="216" stateProvince="Bohol">Philippines</location>
,
<quantity id="4CF879B8FFEB07325620F79E78A1F796" box="[327,451,2077,2101]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFEB07325620F79E78A1F796" box="[327,451,2077,2101]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200–300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFEB0732540FFF3E7D65FF76" box="[872,1031,189,213]" pageId="106" pageNumber="216" value="2004-03-02">
<collectingDate id="EFFA0B75FFEB0732540FFF3E7D65FF76" box="[872,1031,189,213]" pageId="106" pageNumber="216" value="2004-03-02">2 March 2004</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B68DE00FFEB07325374FF3D7DD0FE96" collectingDate="2001-11-28" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="106" pageNumber="216" specimenCode="ZRC 2001.0594" specimenCount="1" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="54BB6AFFFFEB07325374FF3D7D1DFF75" box="[1043,1151,190,214]" pageId="106" pageNumber="216" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B974084FFFEB073253EBFF3E7C69FF76" bold="true" box="[1164,1291,189,213]" pageId="106" pageNumber="216">
<collectingCountry id="F31794CDFFEB073253EBFF3E7C69FF76" box="[1164,1291,189,213]" name="Philippines" pageId="106" pageNumber="216">Philippines</collectingCountry>
</emphasis>
– 2 ovigerous females (46.8 ×
<quantity id="4CF879B8FFEB073254B4FF5E7D50FF56" box="[979,1074,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="106" pageNumber="216" unit="mm" value="36.6">36.6 mm</quantity>
, 45.0 × 36.0 mm) (
<specimenCode id="DBA67C26FFEB07325394FF5E7CC2FF56" box="[1267,1440,221,245]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="106" pageNumber="216">ZRC 2001.0594</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFEB0732544CFF7E7ABBFEB6" box="[811,985,253,277]" pageId="106" pageNumber="216">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFEB07325485FF7E7D5BFEB6" LSID="urn:lsid:plazi:treatment:03A9654BFFEB0732564EF8FE7CC9F936:8EDF8286FFEB07325485FF7E7D5BFEB6" box="[994,1081,253,277]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="106" pageNumber="216" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFEB07325325FF7E7DE7FEB6" box="[1090,1157,253,277]" country="Philippines" name="Bohol" pageId="106" pageNumber="216">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFEB073253E9FF7E7C64FEB6" box="[1166,1286,253,277]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFEB073253E9FF7E7C64FEB6" box="[1166,1286,253,277]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200–300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFEB07325486FE9E7DD0FE96" box="[993,1202,285,309]" pageId="106" pageNumber="216" value="2001-11-28">
<collectingDate id="EFFA0B75FFEB07325486FE9E7DD0FE96" box="[993,1202,285,309]" pageId="106" pageNumber="216" value="2001-11-28">28 November 2001</collectingDate>
</date>
</materialsCitation>
. —
<specimenCount id="9D061FD4FFEB073253BBFE9E7C46FE96" box="[1244,1316,285,309]" pageId="106" pageNumber="216" type="male">1 male</specimenCount>
(41.6 ×
<quantity id="4CF879B8FFEB0732521AFE9E7A3BFEF6" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="106" pageNumber="216" unit="mm" value="32.8">32.8 mm</quantity>
), 1 ovigerous female (45.6 ×
<quantity id="4CF879B8FFEB073253FFFEBE7D99FEF6" box="[1176,1275,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="3.57" pageId="106" pageNumber="216" unit="mm" value="35.7">35.7 mm</quantity>
),
<materialsCitation id="3B68DE00FFEB07325268FEBE7A04FE16" collectingDate="2002-06" collectionCode="ZRC" country="Philippines" elevation="250" location="Panglao" municipality="Balicasag Island" pageId="106" pageNumber="216" specimenCode="ZRC 2013.1276" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFEB07325268FEBE7C0FFEF6" box="[1295,1389,317,341]" pageId="106" pageNumber="216" type="female">1 female</specimenCount>
(47.2 ×
<quantity id="4CF879B8FFEB07325421FEDE7ACBFED6" box="[838,937,349,373]" metricMagnitude="-2" metricUnit="m" metricValue="3.76" pageId="106" pageNumber="216" unit="mm" value="37.6">37.6 mm</quantity>
) (
<specimenCode id="DBA67C26FFEB073254A6FEDE7D15FED6" box="[961,1143,349,373]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="106" pageNumber="216">ZRC 2013.1276</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFEB073253EBFEDE7C23FED6" box="[1164,1345,349,373]" pageId="106" pageNumber="216">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFEB07325228FEDE7CCAFED6" LSID="urn:lsid:plazi:treatment:03A9654BFFEB0732564EF8FE7CC9F936:8EDF8286FFEB07325228FEDE7CCAFED6" box="[1359,1448,349,373]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="106" pageNumber="216" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFEB0732544AFEFE7A13FE36" box="[813,881,381,405]" country="Philippines" name="Bohol" pageId="106" pageNumber="216">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFEB07325418FEFE7A9CFE36" box="[895,1022,381,405]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFEB07325418FEFE7A9CFE36" box="[895,1022,381,405]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="106" pageNumber="216" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200–300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFEB0732521DFEFD7A04FE16" pageId="106" pageNumber="216" value="2002-06">
<collectingDate id="EFFA0B75FFEB0732521DFEFD7A04FE16" pageId="106" pageNumber="216" value="2002-06">June 2002</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFEB073254FEFE1E7C0EFE56" collectingDate="2003-02" collectorName="H. Komatsu &amp; M. Takeda" country="Philippines" location="Panglao" municipality="Balicasag Island" pageId="106" pageNumber="216" specimenCount="1" specimenCount-female="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFEB073254FEFE1E7A9BFE16" box="[921,1017,413,437]" pageId="106" pageNumber="216" type="female">1 female</specimenCount>
(49.4 ×
<quantity id="4CF879B8FFEB07325307FE1E7DA6FE16" box="[1120,1220,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="3.84" pageId="106" pageNumber="216" unit="mm" value="38.4">38.4 mm</quantity>
) (NSMT-Cr 15386),
<collectingMunicipality id="6BDB4E27FFEB0732544AFE3E7ABFFE76" box="[813,989,445,469]" pageId="106" pageNumber="216">Balicasag Island</collectingMunicipality>
,
<location id="8EDF8286FFEB07325481FE3E7D5CFE76" LSID="urn:lsid:plazi:treatment:03A9654BFFEB0732564EF8FE7CC9F936:8EDF8286FFEB07325481FE3E7D5CFE76" box="[998,1086,445,469]" country="Philippines" municipality="Balicasag Island" name="Panglao" pageId="106" pageNumber="216" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFEB07325320FE3E7DE9FE76" box="[1095,1163,445,469]" country="Philippines" name="Bohol" pageId="106" pageNumber="216">Bohol</collectingRegion>
, coll. fishermen with tangle nets, via
<collectorName id="26F5B18BFFEB073254F4FE5D7D75FE56" box="[915,1047,478,501]" pageId="106" pageNumber="216">H. Komatsu</collectorName>
&amp;
<collectorName id="26F5B18BFFEB07325327FE5D7DD8FE56" box="[1088,1210,477,501]" pageId="106" pageNumber="216">M. Takeda</collectorName>
,
<date id="FFBEF29DFFEB073253AFFE5E7C0EFE56" box="[1224,1388,477,501]" pageId="106" pageNumber="216" value="2003-02">
<collectingDate id="EFFA0B75FFEB073253AFFE5E7C0EFE56" box="[1224,1388,477,501]" pageId="106" pageNumber="216" value="2003-02">February 2003</collectingDate>
</date>
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFEB073252F9FE5E7CC8FD96" collectingDate="2004-05-30" collectionCode="ZRC" collectorName="J. Arbasto" country="Philippines" location="Panglao" municipality="Maribohoc Bay" pageId="106" pageNumber="216" specimenCode="ZRC 2013.1277" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFEB073252F9FE5E7A03FDB6" pageId="106" pageNumber="216" type="male">1 male</specimenCount>
(47.5 ×
<quantity id="4CF879B8FFEB073254D8FE7E7D42FDB6" box="[959,1056,509,533]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" pageId="106" pageNumber="216" unit="mm" value="39.5">39.5 mm</quantity>
) (
<specimenCode id="DBA67C26FFEB07325350FE7E7D88FDB6" box="[1079,1258,509,533]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="106" pageNumber="216">ZRC 2013.1277</specimenCode>
),
<collectingMunicipality id="6BDB4E27FFEB0732539AFE7E7CC5FDB5" box="[1277,1447,509,534]" pageId="106" pageNumber="216">Maribohoc Bay</collectingMunicipality>
,
<location id="8EDF8286FFEB07325449FD9E7AE5FD96" LSID="urn:lsid:plazi:treatment:03A9654BFFEB0732564EF8FE7CC9F936:8EDF8286FFEB07325449FD9E7AE5FD96" box="[814,903,541,565]" country="Philippines" municipality="Maribohoc Bay" name="Panglao" pageId="106" pageNumber="216" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFEB073254F5FD9E7AB4FD96" box="[914,982,541,565]" country="Philippines" name="Bohol" pageId="106" pageNumber="216">Bohol</collectingRegion>
, coll.
<collectorName id="26F5B18BFFEB0732537FFD9D7DEAFD96" box="[1048,1160,541,565]" pageId="106" pageNumber="216">J. Arbasto</collectorName>
, tangle nets,
<date id="FFBEF29DFFEB07325271FD9E7CC8FD96" box="[1302,1450,541,566]" pageId="106" pageNumber="216" value="2004-05-30">
<collectingDate id="EFFA0B75FFEB07325271FD9E7CC8FD96" box="[1302,1450,541,566]" pageId="106" pageNumber="216" value="2004-05-30">30 May 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFEB0732544AFDDE7C56FCD6" pageId="106" pageNumber="216" type="diagnosis">
<paragraph id="8BBFD45DFFEB0732544AFDDE7C56FCD6" blockId="106.[811,1452,605,885]" pageId="106" pageNumber="216">
<emphasis id="B974084FFFEB0732544AFDDE7AC2FDD6" bold="true" box="[813,928,605,629]" pageId="106" pageNumber="216">Diagnosis.</emphasis>
Carapace relatively broad (
<figureCitation id="133BC8D8FFEB073253D9FDDE7C69FDD6" box="[1214,1291,605,629]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Fig. 60</figureCitation>
). Pseudorostral spines long, diverging (
<figureCitation id="133BC8D8FFEB07325341FDFE7D14FD36" box="[1062,1142,637,661]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Fig. 60</figureCitation>
). Median row with 2 median gastric spines; 2 cardiac spines; 2 intestinal spines (
<figureCitation id="133BC8D8FFEB07325237FD1E7CFDFD16" box="[1360,1439,669,693]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Fig. 60</figureCitation>
). Basal antennal article with 2 relatively long distal spines, strongly diverging (
<figureCitation id="133BC8D8FFEB07325363FD5E7D09FD56" box="[1028,1131,733,757]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="106" pageNumber="216">Fig. 61G</figureCitation>
). Ischium of third maxilliped long (
<figureCitation id="133BC8D8FFEB07325417FD7E7A9AFCB6" box="[880,1016,765,789]" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="106" pageNumber="216">Fig. 63E–H</figureCitation>
). Ambulatory legs (notably merus and propodus) relatively long (
<figureCitation id="133BC8D8FFEB0732533DFC9E7DD8FC96" box="[1114,1210,797,821]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Figs. 60</figureCitation>
,
<figureCitation id="133BC8D8FFEB073253AFFC9E7C70FC96" box="[1224,1298,797,821]" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="106" pageNumber="216">65I–K</figureCitation>
). Male telson appears semicircular (
<figureCitation id="133BC8D8FFEB0732534FFCBE7DF2FCF6" box="[1064,1168,829,853]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64E</figureCitation>
). G1 very long, strongly curved, slender, tip bent inwards (
<figureCitation id="133BC8D8FFEB073253C6FCDE7C4AFCD6" box="[1185,1320,861,885]" captionStart="Fig" captionStartId="105.[135,170,1959,1981]" captionTargetBox="[137,1474,266,1944]" captionTargetId="figure-8@105.[127,1495,262,1947]" captionTargetPageId="105" captionText="Fig. 67. Left G1s, Rathbunaja species. A–D, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; E–G, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea. A, E = 5.0 mm; B-G = 1.0 mm; F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384740" httpUri="https://zenodo.org/record/5384740/files/figure.png" pageId="106" pageNumber="216">Fig. 67A–D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFEB0732544AFC1E7DE7FC56" pageId="106" pageNumber="216" type="etymology">
<paragraph id="8BBFD45DFFEB0732544AFC1E7DE7FC56" blockId="106.[811,1452,925,1013]" pageId="106" pageNumber="216">
<emphasis id="B974084FFFEB0732544AFC1E7ACCFC16" bold="true" box="[813,942,925,949]" pageId="106" pageNumber="216">Etymology.</emphasis>
The name “ursus” alludes to the thick tomentum covering the animal like the fur of a bear, «ursus». The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFEB0732544AFB9E7CC9F936" pageId="106" pageNumber="216" type="discussion">
<paragraph id="8BBFD45DFFEB0732544AFB9E7CC9F936" blockId="106.[811,1454,1053,1685]" pageId="106" pageNumber="216">
<emphasis id="B974084FFFEB0732544AFB9E7AF9FB96" bold="true" box="[813,923,1053,1077]" pageId="106" pageNumber="216">Remarks.</emphasis>
The discovery of
<taxonomicName id="4C00AFDEFFEB0732530EFB9E7C4BFB96" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1129,1321,1053,1077]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFEB0732530EFB9E7C4BFB96" box="[1129,1321,1053,1077]" italics="true" pageId="106" pageNumber="216">Rathbunaja ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFEB07325255FB9D7C0CFB96" box="[1330,1390,1054,1077]" pageId="106" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
from Balicasag,
<collectingCountry id="F31794CDFFEB073254C3FBBE7D7DFBF6" box="[932,1055,1085,1109]" name="Philippines" pageId="106" pageNumber="216">Philippines</collectingCountry>
, is rather surprising especially since there was also a specimen of
<taxonomicName id="4C00AFDEFFEB0732530CFBDD7CCEFBD6" authority="(Rathbun, 1916)" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[1131,1452,1117,1141]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFEB0732530CFBDD7D96FBD6" box="[1131,1268,1117,1141]" italics="true" pageId="106" pageNumber="216">R. bisarmata</emphasis>
(
<bibRefCitation id="EF91A9ACFFEB07325264FBDE7CC6FBD6" author="Rathbun MJ" box="[1283,1444,1117,1141]" pageId="106" pageNumber="216" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
)
</taxonomicName>
collected from the same area. Most specimens of
<taxonomicName id="4C00AFDEFFEB07325249FBFE7CC8FB36" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1326,1450,1149,1173]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFEB07325249FBFE7CC8FB36" box="[1326,1450,1149,1173]" italics="true" pageId="106" pageNumber="216">Rathbunaja</emphasis>
</taxonomicName>
from the
<collectingCountry id="F31794CDFFEB073254F0FB1E7D73FB16" box="[919,1041,1181,1205]" name="Philippines" pageId="106" pageNumber="216">Philippines</collectingCountry>
actually belong to this new species. The two species are similar and are not easy to separate; In
<taxonomicName id="4C00AFDEFFEB07325429FB5D7ACEFB56" authorityName="Ng &amp; Forges" authorityYear="2015" box="[846,940,1246,1269]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B974084FFFEB07325429FB5D7ACEFB56" box="[846,940,1246,1269]" italics="true" pageId="106" pageNumber="216">R. ursus</emphasis>
</taxonomicName>
, however, the carapace is relatively broader (
<figureCitation id="133BC8D8FFEB07325452FB7E7AE7FAB6" box="[821,901,1277,1301]" captionStart="Fig" captionStartId="98.[113,148,2006,2028]" captionTargetBox="[113,1451,706,1986]" captionTargetId="figure-350@98.[113,1451,703,1987]" captionTargetPageId="98" captionText="Fig. 60. General habitus. A, Rathbunaja ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; B, Rathbunaja ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; C, Rathbunaja ursus n. sp., paratype female (49.4 × 38.4 mm) (NSMT-Cr 15386), Philippines; D, Rathbunaja ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines" figureDoi="http://doi.org/10.5281/zenodo.5384724" httpUri="https://zenodo.org/record/5384724/files/figure.png" pageId="106" pageNumber="216">Fig. 60</figureCitation>
versus
<figureCitation id="133BC8D8FFEB073254BBFB7E7D01FAB6" box="[988,1123,1277,1301]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="106" pageNumber="216">Fig. 59A, B</figureCitation>
), the two distal spines on the basal antennal article are more strongly diverging (
<figureCitation id="133BC8D8FFEB0732522BFA9E7CC9FA96" box="[1356,1451,1309,1333]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="106" pageNumber="216">Fig. 61G</figureCitation>
versus
<figureCitation id="133BC8D8FFEB0732541DFABE7A9CFAF6" box="[890,1022,1341,1365]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="106" pageNumber="216">Fig. 61A, C</figureCitation>
), the ischium of the third maxilliped is proportionately longer (
<figureCitation id="133BC8D8FFEB0732535EFADE7DDEFAD6" box="[1081,1212,1373,1397]" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="106" pageNumber="216">Fig. 63E–H</figureCitation>
versus
<figureCitation id="133BC8D8FFEB07325271FADE7CFDFAD6" box="[1302,1439,1373,1397]" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="106" pageNumber="216">Fig. 63A, B</figureCitation>
), the anterior thoracic sternum (sternites 1–4) is relatively longer (
<figureCitation id="133BC8D8FFEB073254E6FA1E7A9CFA16" box="[897,1022,1437,1461]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64E, F</figureCitation>
versus
<figureCitation id="133BC8D8FFEB07325334FA1E7DB4FA16" box="[1107,1238,1437,1461]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64A, B</figureCitation>
), the male telson is proportionately broader (
<figureCitation id="133BC8D8FFEB07325326FA3E7DA3FA76" box="[1089,1217,1469,1493]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64E, F</figureCitation>
versus
<figureCitation id="133BC8D8FFEB0732527EFA3E7CC2FA76" box="[1305,1440,1469,1493]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64A, B</figureCitation>
), and the lateral margins of male abdominal somites 5 and 6 are more concave (
<figureCitation id="133BC8D8FFEB07325365FA7E7DE3F9B6" box="[1026,1153,1533,1557]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64E, F</figureCitation>
versus
<figureCitation id="133BC8D8FFEB073253BEFA7E7C3DF9B6" box="[1241,1375,1533,1557]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="106" pageNumber="216">Fig. 64A, B</figureCitation>
). Most significantly, their G1 structures are completely different, with that of
<taxonomicName id="4C00AFDEFFEB073254D6F9BD7D6BF9F6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[945,1033,1598,1621]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="106" pageNumber="216" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B974084FFFEB073254D6F9BD7D6BF9F6" box="[945,1033,1598,1621]" italics="true" pageId="106" pageNumber="216">R. ursus</emphasis>
</taxonomicName>
much longer, with the distal part more strongly curved and slender (
<figureCitation id="133BC8D8FFEB07325330F9DE7DB6F9D6" box="[1111,1236,1629,1653]" captionStart="Fig" captionStartId="105.[135,170,1959,1981]" captionTargetBox="[137,1474,266,1944]" captionTargetId="figure-8@105.[127,1495,262,1947]" captionTargetPageId="105" captionText="Fig. 67. Left G1s, Rathbunaja species. A–D, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; E–G, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea. A, E = 5.0 mm; B-G = 1.0 mm; F, G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384740" httpUri="https://zenodo.org/record/5384740/files/figure.png" pageId="106" pageNumber="216">Fig. 67A–D</figureCitation>
versus
<figureCitation id="133BC8D8FFEB07325246F9DE7CC3F9D6" box="[1313,1441,1629,1653]" captionStart="Fig" captionStartId="104.[113,148,1891,1913]" captionTargetBox="[114,1451,333,1876]" captionTargetId="figure-91@104.[95,1462,325,1884]" captionTargetPageId="104" captionText="Fig. 66. Left G1s, Rathbunaja species. A–D, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; E–G, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; H–J, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; K–M, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan. Scales: A, H, K = 1.0 mm; E = 5.0 mm; B–D, F, G, I, J, L, M= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384738" httpUri="https://zenodo.org/record/5384738/files/figure.png" pageId="106" pageNumber="216">Fig. 66A–G</figureCitation>
). These differences cannot be explained by size or maturity.
</paragraph>
</subSubSection>
</treatment>
</document>
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