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<document id="DAF2623F843387BC465BEAE9DF4F2E3D" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFE4073E578EFF3F7803FC36" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Rathbunaja bisarmata" docType="treatment" docVersion="3" lastPageNumber="212" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="211" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="174EED04B7E9B488CB7D4448A5A13E9C">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="FC799FB000C3823AF9F9719C12FC69A2">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="932FE55E2E1C75BF658FC7F8AC44C635">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="30B7847EF8DA49BAD10429EA76CD746F">Raffles Bulletin of Zoology</mods:title>
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<paragraph id="8BBFD45DFFE4073D578EFF3F7BC7FF76" blockId="101.[141,769,188,245]" box="[233,677,188,213]" pageId="101" pageNumber="211">
<heading id="D0F76331FFE4073D578EFF3F7BC7FF76" bold="true" box="[233,677,188,213]" centered="true" fontSize="10" level="2" pageId="101" pageNumber="211" reason="2">
<taxonomicName id="4C00AFDEFFE4073D578EFF3F7BC7FF76" ID-CoL="4RLXD" authority="(Rathbun, 1916)" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[233,677,188,213]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE4073D578EFF3F7BC7FF76" bold="true" box="[233,677,188,213]" pageId="101" pageNumber="211">
<emphasis id="B974084FFFE4073D578EFF3F7880FF77" bold="true" box="[233,482,188,212]" italics="true" pageId="101" pageNumber="211">Rathbunaja bisarmata</emphasis>
(
<bibRefCitation id="EF91A9ACFFE4073D5695FF3E7BFCFF76" author="Rathbun MJ" box="[498,670,189,213]" pageId="101" pageNumber="211" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C31A87D6FFE4073D57EAFF5E7A63FF56" box="[141,769,221,245]" pageId="101" pageNumber="211" type="description">
<paragraph id="8BBFD45DFFE4073D57EAFF5E7A63FF56" blockId="101.[141,769,188,245]" box="[141,769,221,245]" pageId="101" pageNumber="211">
(
<figureCitation id="133BC8D8FFE4073D57F2FF5E7847FF57" box="[149,293,221,245]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Figs. 59A, B</figureCitation>
,
<figureCitation id="133BC8D8FFE4073D5655FF5E78E1FF56" box="[306,387,221,245]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="101" pageNumber="211">61AD</figureCitation>
,
<figureCitation id="133BC8D8FFE4073D56F7FF5E7883FF57" box="[400,481,221,245]" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="101" pageNumber="211">63A, B</figureCitation>
,
<figureCitation id="133BC8D8FFE4073D5689FF5E7B5DFF57" box="[494,575,221,245]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="101" pageNumber="211">64A, B</figureCitation>
,
<figureCitation id="133BC8D8FFE4073D552BFF5E7BFCFF56" box="[588,670,221,245]" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="101" pageNumber="211">65AD</figureCitation>
,
<figureCitation id="133BC8D8FFE4073D55CDFF5E7B98FF56" box="[682,762,221,245]" captionStart="Fig" captionStartId="104.[113,148,1891,1913]" captionTargetBox="[114,1451,333,1876]" captionTargetId="figure-91@104.[95,1462,325,1884]" captionTargetPageId="104" captionText="Fig. 66. Left G1s, Rathbunaja species. AD, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; EG, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; HJ, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; KM, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan. Scales: A, H, K = 1.0 mm; E = 5.0 mm; BD, F, G, I, J, L, M= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384738" httpUri="https://zenodo.org/record/5384738/files/figure.png" pageId="101" pageNumber="211">66AG</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFE4073D57EFFE9F7B83FEED" pageId="101" pageNumber="211" type="reference_group">
<paragraph id="8BBFD45DFFE4073D57EFFE9F7B83FEED" blockId="101.[136,775,283,334]" pageId="101" pageNumber="211">
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<treatmentCitation id="0AA1F24CFFE4073D57EFFE9F7885FE92" author="Rathbun MJ" box="[136,487,283,305]" page="553" pageId="101" pageNumber="211" year="1916">
<taxonomicName id="4C00AFDEFFE4073D57EFFE9F7885FE92" ID-CoL="3XJS2" authority="Rathbun, 1916: 553" authorityName="Rathbun" authorityPageNumber="553" authorityYear="1916" box="[136,487,283,305]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE4073D57EFFE9F7843FE93" box="[136,289,283,305]" italics="true" pageId="101" pageNumber="211">Maja bisarmata</emphasis>
<bibRefCitation id="EF91A9ACFFE4073D5640FE987885FE92" author="Rathbun MJ" box="[295,487,283,305]" pageId="101" pageNumber="211" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916: 553</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="0AA1F24CFFE4073D5566FE9F7BC7FE92" author="Serene R" box="[513,677,283,306]" page="57" pageId="101" pageNumber="211" year="1968">
<bibRefCitation id="EF91A9ACFFE4073D5566FE9F7BC7FE92" author="Serene R" box="[513,677,283,306]" pageId="101" pageNumber="211" pagination="33 - 120" refId="ref88537" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1: 33 - 120." type="book chapter" year="1968">Serène, 1968: 57</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AA1F24CFFE4073D55D8FE987870FEEE" author="Griffin DJG" page="199" pageId="101" pageNumber="211" year="1976">
<bibRefCitation id="EF91A9ACFFE4073D55D8FE987870FEEE" author="Griffin DJG" pageId="101" pageNumber="211" pagination="179 - 222" refId="ref84048" refString="Griffin DJG (1976) Spider crabs of the family Majidae (Crustacea: Brachyura) from the Philippine Islands. Journal of Natural History, 10 (2): 179 - 222, figs. 1 - 11, tables 1, 2." type="journal article" year="1976">Griffin, 1976: 199</bibRefCitation>
</treatmentCitation>
,
<figureCitation id="133BC8D8FFE4073D5679FEBB7805FEED" box="[286,359,312,334]" captionStart="Fig" captionStartId="20.[113,148,1947,1969]" captionTargetBox="[178,1386,189,1931]" captionTargetId="figure-15@20.[152,1415,174,1934]" captionTargetPageId="20" captionText="Fig. 7. Left G1s. AC, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia [gonopods laterally inverted as left G1 missing, right G1 drawn]; DF, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; GI, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; JL, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; M, N, Maja cornuta (as Mamaia queketti, after Barnard, 1950: fig. 13a); OQ, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; RT, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo. Scale bars = 5.0 mm [A, D, G, J, O, R]; 1.0 mm [B, C, E, F, H, I, K, L, P, Q, S, T]." figureDoi="http://doi.org/10.5281/zenodo.5384604" httpUri="https://zenodo.org/record/5384604/files/figure.png" pageId="101" pageNumber="211">Fig. 7b</figureCitation>
.
<treatmentCitation id="0AA1F24CFFE4073D56E2FEBB7BC2FEEE" author="Griffin DJG &amp; Tranter HA" box="[389,672,312,334]" page="211" pageId="101" pageNumber="211" year="1986">
<bibRefCitation id="EF91A9ACFFE4073D56E2FEBB7BC2FEEE" author="Griffin DJG &amp; Tranter HA" box="[389,672,312,334]" pageId="101" pageNumber="211" pagination="1 - 335" refId="ref84094" refString="Griffin DJG &amp; Tranter HA (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie, 39 C 4: 1 - 335, 22 pls." type="journal article" year="1986">Griffin &amp; Tranter, 1986: 211</bibRefCitation>
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFE4073D57EFFEF57806FDEC" pageId="101" pageNumber="211" type="materials_examined">
<paragraph id="8BBFD45DFFE4073D57EFFEF57806FDEC" blockId="101.[136,775,374,591]" pageId="101" pageNumber="211">
<emphasis id="B974084FFFE4073D57EFFEF57811FE2D" bold="true" box="[136,371,374,398]" pageId="101" pageNumber="211">Material examined.</emphasis>
<materialsCitation id="3B68DE00FFE4073D5619FEF57806FDEC" collectingDate="1909-08-09" collectingDateMax="2004-03-02" collectingDateMin="1909-08-09" collectionCode="RV" collectorName="Philippines" country="Philippines" county="Point Tagolo Light" elevation="250" latitude="80.78333" location="Albatross" longLatPrecision="925" longitude="123.520836" municipality="northern Mindanao" pageId="101" pageNumber="211" specimenCode="USNM 48220, ZRC 2013.1274" specimenCount="1" specimenCount-male="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54BB6AFFFFE4073D5619FEF57888FE2D" box="[382,490,374,398]" pageId="101" pageNumber="211" type="holotype">Holotype</typeStatus>
: male (20.3 ×
<quantity id="4CF879B8FFE4073D55FFFEF57B9DFE2C" box="[664,767,374,399]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="101" pageNumber="211" unit="mm" value="15.6">15.6 mm</quantity>
) (
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), station 5519,
<collectingCounty id="62DEACD1FFE4073D5694FE157BB4FE0D" box="[499,726,406,430]" pageId="101" pageNumber="211">Point Tagolo Light</collectingCounty>
, off
<collectingMunicipality id="6BDB4E27FFE4073D57EFFE357804FE6D" box="[136,358,438,462]" pageId="101" pageNumber="211">northern Mindanao</collectingMunicipality>
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<collectorName id="26F5B18BFFE4073D55E6FE357A60FE6D" box="[641,770,438,462]" pageId="101" pageNumber="211">Philippines</collectorName>
, coll.
<collectionCode id="ED114C98FFE4073D57D9FE547981FE4D" box="[190,227,471,494]" country="Italy" name="Collection of Leptospira Strains" pageId="101" pageNumber="211">RV</collectionCode>
<emphasis id="B974084FFFE4073D578DFE557832FE4D" box="[234,336,470,494]" italics="true" pageId="101" pageNumber="211">
<location id="8EDF8286FFE4073D578DFE557832FE4D" LSID="urn:lsid:plazi:treatment:03A9654BFFE4073E578EFF3F7803FC36:8EDF8286FFE4073D578DFE557832FE4D" box="[234,336,470,494]" country="Philippines" county="Point Tagolo Light" latitude="80.78333" longLatPrecision="925" longitude="123.520836" municipality="northern Mindanao" name="Albatross" pageId="101" pageNumber="211" stateProvince="Bohol">Albatross</location>
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,
<date id="FFBEF29DFFE4073D5639FE54789DFE4C" box="[350,511,471,495]" pageId="101" pageNumber="211" value="1909-08-09">
<collectingDate id="EFFA0B75FFE4073D5639FE54789DFE4C" box="[350,511,471,495]" pageId="101" pageNumber="211" value="1909-08-09">9 August 1909</collectingDate>
</date>
.
<location id="8EDF8286FFE4073D556EFE557B36FE4D" LSID="urn:lsid:plazi:treatment:03A9654BFFE4073E578EFF3F7803FC36:8EDF8286FFE4073D556EFE557B36FE4D" box="[521,596,470,494]" country="Philippines" county="Point Tagolo Light" latitude="80.78333" longLatPrecision="925" longitude="123.520836" municipality="northern Mindanao" name="Others" pageId="101" pageNumber="211" stateProvince="Bohol">Others</location>
:
<emphasis id="B974084FFFE4073D5538FE557BBCFE4D" bold="true" box="[607,734,470,494]" pageId="101" pageNumber="211">
<collectingCountry id="F31794CDFFE4073D5538FE557BBCFE4D" box="[607,734,470,494]" name="Philippines" pageId="101" pageNumber="211">Philippines</collectingCountry>
</emphasis>
<specimenCount id="9D061FD4FFE4073D559EFE5479DEFDAD" pageId="101" pageNumber="211" type="male">1 male</specimenCount>
(37.1 ×
<quantity id="4CF879B8FFE4073D5670FE757815FDAC" box="[279,375,502,527]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="101" pageNumber="211" unit="mm" value="29.6">29.6 mm</quantity>
) (
<specimenCode id="DBA67C26FFE4073D56EAFE747B5DFDAC" box="[397,575,503,527]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="101" pageNumber="211">ZRC 2013.1274</specimenCode>
),
<location id="8EDF8286FFE4073D5537FE757A61FDAD" LSID="urn:lsid:plazi:treatment:03A9654BFFE4073E578EFF3F7803FC36:8EDF8286FFE4073D5537FE757A61FDAD" box="[592,771,502,526]" country="Philippines" county="Point Tagolo Light" latitude="80.78333" longLatPrecision="925" longitude="123.520836" municipality="northern Mindanao" name="Balicasag Island" pageId="101" pageNumber="211" stateProvince="Bohol">Balicasag Island</location>
,
<location id="8EDF8286FFE4073D57EFFD957981FD8D" LSID="urn:lsid:plazi:treatment:03A9654BFFE4073E578EFF3F7803FC36:8EDF8286FFE4073D57EFFD957981FD8D" box="[136,227,534,558]" country="Philippines" county="Point Tagolo Light" latitude="80.78333" longLatPrecision="925" longitude="123.520836" municipality="northern Mindanao" name="Panglao" pageId="101" pageNumber="211" stateProvince="Bohol">Panglao</location>
,
<collectingRegion id="49C41ABFFFE4073D5795FD95785AFD8D" box="[242,312,534,558]" country="Philippines" name="Bohol" pageId="101" pageNumber="211">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFE4073D5620FD9478AAFD8D" box="[327,456,535,559]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="101" pageNumber="211" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFE4073D5620FD9478AAFD8D" box="[327,456,535,559]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="101" pageNumber="211" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. fishermen with tangle nets,
<date id="FFBEF29DFFE4073D57A4FDB47802FDEC" box="[195,352,566,591]" pageId="101" pageNumber="211" value="2004-03-02">
<collectingDate id="EFFA0B75FFE4073D57A4FDB47802FDEC" box="[195,352,566,591]" pageId="101" pageNumber="211" value="2004-03-02">2 March 2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFE4073D57EFFDF57CA7FF56" pageId="101" pageNumber="211" type="diagnosis">
<paragraph id="8BBFD45DFFE4073D57EFFDF57A65FCED" blockId="101.[134,776,630,847]" pageId="101" pageNumber="211">
<emphasis id="B974084FFFE4073D57EFFDF57867FD2D" bold="true" box="[136,261,630,654]" pageId="101" pageNumber="211">Diagnosis.</emphasis>
Carapace relatively narrow (
<figureCitation id="133BC8D8FFE4073D5501FDF57B9BFD2C" box="[614,761,630,655]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Fig. 59A, B</figureCitation>
). Pseudorostral spines long, diverging (
<figureCitation id="133BC8D8FFE4073D5579FD157BC2FD0C" box="[542,672,662,687]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Fig. 59A, B</figureCitation>
). Median row with 2 median gastric spines; 2 cardiac spines; 2 intestinal spines (
<figureCitation id="133BC8D8FFE4073D57B0FD55783AFD4C" box="[215,344,726,751]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Fig. 59A, B</figureCitation>
). Basal antennal article with 2 relatively short distal spines not strongly diverging (
<figureCitation id="133BC8D8FFE4073D5508FD757B99FCAD" box="[623,763,758,782]" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="101" pageNumber="211">Fig. 61AD</figureCitation>
). Ischium of third maxilliped proportionately shorter (
<figureCitation id="133BC8D8FFE4073D55BDFC9579B5FCEC" captionStart="Fig" captionStartId="101.[135,170,1949,1971]" captionTargetBox="[134,1474,932,1933]" captionTargetId="figure-8@101.[134,1474,932,1934]" captionTargetPageId="101" captionText="Fig. 63. Third maxillipeds, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. kiiensis, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H, R. ursus n. sp., paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="101" pageNumber="211">Fig. 63A, B</figureCitation>
). Ambulatory merus relatively shorter, stouter (
<figureCitation id="133BC8D8FFE4073D55B6FCB57A65FCED" box="[721,775,822,846]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Figs.</figureCitation>
</paragraph>
<paragraph id="8BBFD45DFFE4073D5424FF3E7CA7FF56" blockId="101.[835,1477,189,245]" pageId="101" pageNumber="211">
59A, B,
<figureCitation id="133BC8D8FFE4073D54C5FF3E7A96FF76" box="[930,1012,189,213]" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="101" pageNumber="211">65B, D</figureCitation>
). Male telson subtriangular (
<figureCitation id="133BC8D8FFE4073D5248FF3E7CD6FF76" box="[1327,1460,189,213]" captionStart="Fig" captionStartId="102.[113,148,1977,1999]" captionTargetBox="[113,1451,1069,1962]" captionTargetId="figure-108@102.[113,1451,1068,1964]" captionTargetPageId="102" captionText="Fig. 64. Male anterior thoracic sternums and abdomens, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea; E, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384734" httpUri="https://zenodo.org/record/5384734/files/figure.png" pageId="101" pageNumber="211">Fig. 64A, B</figureCitation>
). G1 gently curved, distal part relatively shorter (
<figureCitation id="133BC8D8FFE4073D5251FF5E7CDBFF56" box="[1334,1465,221,245]" captionStart="Fig" captionStartId="104.[113,148,1891,1913]" captionTargetBox="[114,1451,333,1876]" captionTargetId="figure-91@104.[95,1462,325,1884]" captionTargetPageId="104" captionText="Fig. 66. Left G1s, Rathbunaja species. AD, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; EG, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; HJ, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; KM, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan. Scales: A, H, K = 1.0 mm; E = 5.0 mm; BD, F, G, I, J, L, M= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384738" httpUri="https://zenodo.org/record/5384738/files/figure.png" pageId="101" pageNumber="211">Fig. 66AG</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFE4073E5424FE9E7803FC36" lastPageId="102" lastPageNumber="212" pageId="101" pageNumber="211" type="discussion">
<paragraph id="8BBFD45DFFE4073D5424FE9E7D77FD35" blockId="101.[835,1477,285,662]" pageId="101" pageNumber="211">
<emphasis id="B974084FFFE4073D5424FE9E7AD0FE96" bold="true" box="[835,946,285,309]" pageId="101" pageNumber="211">Remarks.</emphasis>
This is a very poorly known species and although it has been treated as a valid
<taxonomicName id="4C00AFDEFFE4073D5310FEBD7C61FEF6" box="[1143,1283,317,341]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B974084FFFE4073D5310FEBD7DCDFEF6" box="[1143,1199,318,341]" italics="true" pageId="101" pageNumber="211">Maja</emphasis>
species
</taxonomicName>
by
<bibRefCitation id="EF91A9ACFFE4073D524BFEBE7CA0FEF6" author="Griffin DJG" box="[1324,1474,317,341]" pageId="101" pageNumber="211" pagination="179 - 222" refId="ref84048" refString="Griffin DJG (1976) Spider crabs of the family Majidae (Crustacea: Brachyura) from the Philippine Islands. Journal of Natural History, 10 (2): 179 - 222, figs. 1 - 11, tables 1, 2." type="journal article" year="1976">Griffin (1976)</bibRefCitation>
(who examined the type) and
<bibRefCitation id="EF91A9ACFFE4073D53F4FEDE7CCAFED6" author="Griffin DJG &amp; Tranter HA" box="[1171,1448,349,373]" pageId="101" pageNumber="211" pagination="1 - 335" refId="ref84094" refString="Griffin DJG &amp; Tranter HA (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie, 39 C 4: 1 - 335, 22 pls." type="journal article" year="1986">Griffin &amp; Tranter (1986)</bibRefCitation>
, a re-examination of the
<typeStatus id="54BB6AFFFFE4073D5352FEFE7DF0FE36" box="[1077,1170,381,405]" pageId="101" pageNumber="211" type="holotype">holotype</typeStatus>
shows otherwise. Although
<bibRefCitation id="EF91A9ACFFE4073D5422FE1E7A96FE16" author="Rathbun MJ" box="[837,1012,413,437]" pageId="101" pageNumber="211" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun (1916)</bibRefCitation>
placed this species in
<taxonomicName id="4C00AFDEFFE4073D539FFE1D7C53FE16" authorityName="Lamarck" authorityYear="1801" box="[1272,1329,414,437]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFE4073D539FFE1D7C53FE16" box="[1272,1329,414,437]" italics="true" pageId="101" pageNumber="211">Maja</emphasis>
</taxonomicName>
, its antennal flagellum is actually just outside the orbit. In fact, the species is very close to
<taxonomicName id="4C00AFDEFFE4073D548EFE5E7DADFE56" authorityName="T. Sakai" authorityYear="1969" box="[1001,1231,477,501]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE4073D548EFE5E7DADFE56" box="[1001,1231,477,501]" italics="true" pageId="101" pageNumber="211">Leptomithrax kiiensis</emphasis>
</taxonomicName>
, described by T.
<bibRefCitation id="EF91A9ACFFE4073D52E0FE5E7AE8FDB6" author="Sakai T" pageId="101" pageNumber="211" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai (1969)</bibRefCitation>
on the basis of a large male 55.0 by
<quantity id="4CF879B8FFE4073D5242FE7E7CE7FDB6" box="[1317,1413,509,533]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="101" pageNumber="211" unit="mm" value="44.5">44.5 mm</quantity>
from Kii Minable in
<collectingCountry id="F31794CDFFE4073D5489FD9D7D4CFD96" box="[1006,1070,542,565]" name="Japan" pageId="101" pageNumber="211">Japan</collectingCountry>
. Tune
<bibRefCitation id="EF91A9ACFFE4073D531FFD9E7D9FFD96" author="Sakai T" box="[1144,1277,541,565]" pageId="101" pageNumber="211" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai (1969</bibRefCitation>
,
<bibRefCitation id="EF91A9ACFFE4073D526EFD9E7C22FD96" author="Sakai T" box="[1289,1344,541,565]" pageId="101" pageNumber="211" pagination="1 - 16" refId="ref88450" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251." type="book chapter" year="1976">1976</bibRefCitation>
) placed this species in
<taxonomicName id="4C00AFDEFFE4073D54DCFDBE7D2EFDF6" authorityName="Miers" authorityYear="1876" box="[955,1100,573,597]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFE4073D54DCFDBE7D2EFDF6" box="[955,1100,573,597]" italics="true" pageId="101" pageNumber="211">Leptomithrax</emphasis>
</taxonomicName>
because its antennal flagellum is just outside the orbit, even if it is not to the same degree as other congeners.
</paragraph>
<paragraph id="8BBFD45DFFE4073E5422FD3E78DCFED6" blockId="101.[834,1476,701,853]" lastBlockId="102.[113,752,189,373]" lastPageId="102" lastPageNumber="212" pageId="101" pageNumber="211">
The
<typeStatus id="54BB6AFFFFE4073D541EFD3E7ABAFD76" box="[889,984,701,725]" pageId="101" pageNumber="211" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C00AFDEFFE4073D5327FD3D7D93FD76" authorityName="Rathbun" authorityYear="1916" box="[1088,1265,701,725]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE4073D5327FD3D7D93FD76" box="[1088,1265,701,725]" italics="true" pageId="101" pageNumber="211">Maja bisarmata</emphasis>
</taxonomicName>
is relatively small (
<figureCitation id="133BC8D8FFE4073D542CFD5E7ACCFD56" box="[843,942,733,757]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="101" pageNumber="211">Fig. 59A</figureCitation>
) and although its G1 is not fully developed (
<figureCitation id="133BC8D8FFE4073D52F0FD5E7AF5FCB6" captionStart="Fig" captionStartId="99.[135,170,2035,2057]" captionTargetBox="[134,1474,475,2019]" captionTargetId="figure-8@99.[134,1474,475,2020]" captionTargetPageId="99" captionText="Fig. 61. Antennae, antennules, orbits and epistomes, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; G, H, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384726" httpUri="https://zenodo.org/record/5384726/files/figure.png" pageId="101" pageNumber="211">Fig. 61AD</figureCitation>
), it shows enough features. A recent larger specimen from the
<collectingCountry id="F31794CDFFE4073D54CDFC9E7D40FC96" box="[938,1058,797,821]" name="Philippines" pageId="101" pageNumber="211">Philippines</collectingCountry>
agrees well with the type male in its carapace form, antennal basal article, epistome, structure of the third maxilliped, relatively short ambulatory legs, thoracic sternum, male abdomen and structure of the G1, and we are confident they are conspecific. These features make it unlikely that
<taxonomicName id="4C00AFDEFFE7073E5638FE9D7B6FFE96" authorityName="Rathbun" authorityYear="1916" box="[351,525,285,309]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE7073E5638FE9D7B6FFE96" box="[351,525,285,309]" italics="true" pageId="102" pageNumber="212">Maja bisarmata</emphasis>
</taxonomicName>
is the juvenile form of the more common new species described here from the
<collectingCountry id="F31794CDFFE7073E5716FEDE798EFED6" box="[113,236,349,373]" name="Philippines" pageId="102" pageNumber="212">Philippines</collectingCountry>
,
<taxonomicName id="4C00AFDEFFE7073E579FFEDE78D5FED6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[248,439,349,373]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B974084FFFE7073E579FFEDE78D5FED6" box="[248,439,349,373]" italics="true" pageId="102" pageNumber="212">Rathbunaja ursus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF7F84D5FFE4073D57E0F81E7B7EF7E0" ID-DOI="http://doi.org/10.5281/zenodo.5384730" ID-Zenodo-Dep="5384730" httpUri="https://zenodo.org/record/5384730/files/figure.png" pageId="101" pageNumber="211" startId="101.[135,170,1949,1971]" targetBox="[134,1474,932,1933]" targetPageId="101">
<paragraph id="8BBFD45DFFE4073D57E0F81E7B7EF7E0" blockId="101.[135,1474,1948,2115]" pageId="101" pageNumber="211">
Fig. 63. Third maxillipeds,
<taxonomicName id="4C00AFDEFFE4073D56FCF81E7B1EF811" box="[411,636,1949,1971]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B974084FFFE4073D56FCF81E7B69F811" box="[411,523,1949,1970]" italics="true" pageId="101" pageNumber="211">Rathbunaja</emphasis>
species. A
</taxonomicName>
,
<taxonomicName id="4C00AFDEFFE4073D55EEF81E7A6BF811" authorityName="Rathbun" authorityYear="1916" box="[649,777,1949,1970]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE4073D55EEF81E7A6BF811" box="[649,777,1949,1970]" italics="true" pageId="101" pageNumber="211">R. bisarmata</emphasis>
</taxonomicName>
, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B,
<taxonomicName id="4C00AFDEFFE4073D52CAF81E798AF86D" authorityName="Rathbun" authorityYear="1916" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE4073D52CAF81E798AF86D" italics="true" pageId="101" pageNumber="211">R. bisarmata</emphasis>
</taxonomicName>
, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C,
<taxonomicName id="4C00AFDEFFE4073D5425F8397AC5F86D" authorityName="T. Sakai" authorityYear="1969" box="[834,935,1977,1999]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE4073D5425F8397AC5F86D" box="[834,935,1977,1999]" italics="true" pageId="101" pageNumber="211">R. kiiensis</emphasis>
</taxonomicName>
, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; D,
<taxonomicName id="4C00AFDEFFE4073D57E0F854798EF848" authorityName="T. Sakai" authorityYear="1969" box="[135,236,2006,2028]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE4073D57E0F854798EF848" box="[135,236,2006,2028]" italics="true" pageId="101" pageNumber="211">R. kiiensis</emphasis>
</taxonomicName>
, ovigerous female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E,
<taxonomicName id="4C00AFDEFFE4073D5372F8547D04F84F" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1045,1126,2007,2028]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFE4073D5372F8547D04F84F" box="[1045,1126,2007,2028]" italics="true" pageId="101" pageNumber="211">R. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFE4073D5308F8547DC5F84F" box="[1135,1191,2007,2028]" pageId="101" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
, holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; F,
<taxonomicName id="4C00AFDEFFE4073D55E1F8777BBAF7AA" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[646,728,2036,2057]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFE4073D55E1F8777BBAF7AA" box="[646,728,2036,2057]" italics="true" pageId="101" pageNumber="211">R. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFE4073D5586F8777A7BF7AA" box="[737,793,2036,2057]" pageId="101" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
, paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; G,
<taxonomicName id="4C00AFDEFFE4073D57E0F79279BAF785" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[135,216,2065,2086]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFE4073D57E0F79279BAF785" box="[135,216,2065,2086]" italics="true" pageId="101" pageNumber="211">R. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFE4073D5787F7937875F786" box="[224,279,2064,2085]" pageId="101" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
, paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines; H,
<taxonomicName id="4C00AFDEFFE4073D54B4F7927D46F785" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[979,1060,2065,2086]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="101" pageNumber="211" phylum="Arthropoda" rank="species" species="ursus" status="sp. nov.">
<emphasis id="B974084FFFE4073D54B4F7927D46F785" box="[979,1060,2065,2086]" italics="true" pageId="101" pageNumber="211">R. ursus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFE4073D534BF7937D01F786" box="[1068,1123,2064,2085]" pageId="101" pageNumber="211" rank="species">n. sp.</taxonomicNameLabel>
, paratype ovigerous female (46.8 × 36.6 mm) (ZRC 2001.0594), Philippines.
</paragraph>
</caption>
<paragraph id="8BBFD45DFFE7073E5716FE1D7803FC36" blockId="102.[111,754,413,917]" pageId="102" pageNumber="212">
<taxonomicName id="4C00AFDEFFE7073E5716FE1D78DBFE16" authority="Rathbun, 1916" authorityName="Rathbun" authorityYear="1916" box="[113,441,413,437]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE7073E5716FE1D787AFE16" box="[113,280,413,437]" italics="true" pageId="102" pageNumber="212">Maja bisarmata</emphasis>
<bibRefCitation id="EF91A9ACFFE7073E567AFE1E78DBFE16" author="Rathbun MJ" box="[285,441,413,437]" pageId="102" pageNumber="212" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
</taxonomicName>
, is very close to
<taxonomicName id="4C00AFDEFFE7073E5507FE1E780EFE76" authority="T. Sakai, 1969" authorityName="T. Sakai" authorityYear="1969" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE7073E5507FE1E79A3FE76" italics="true" pageId="102" pageNumber="212">Leptomithrax kiiensis</emphasis>
T.
<bibRefCitation id="EF91A9ACFFE7073E578EFE3E780EFE76" author="Sakai T" box="[233,364,445,469]" pageId="102" pageNumber="212" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai, 1969</bibRefCitation>
</taxonomicName>
. Comparisons of Rathbuns species with material of
<taxonomicName id="4C00AFDEFFE7073E565DFE5D78D0FE56" authorityName="T. Sakai" authorityYear="1969" box="[314,434,477,501]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE7073E565DFE5D78D0FE56" box="[314,434,477,501]" italics="true" pageId="102" pageNumber="212">L. kiiensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F31794CDFFE7073E5567FE5D7B22FE56" box="[512,576,478,501]" name="Japan" pageId="102" pageNumber="212">Japan</collectingCountry>
as well as the descriptions and figures of the species by T.
<bibRefCitation id="EF91A9ACFFE7073E5501FE7E7B8EFDB6" author="Sakai T" box="[614,748,509,533]" pageId="102" pageNumber="212" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai (1969</bibRefCitation>
,
<bibRefCitation id="EF91A9ACFFE7073E5716FD9E79CFFD96" author="Sakai T" box="[113,173,541,565]" pageId="102" pageNumber="212" pagination="1 - 16" refId="ref88450" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. Kodansha Ltd., Tokyo. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251." type="book chapter" year="1976">1976</bibRefCitation>
),
<bibRefCitation id="EF91A9ACFFE7073E57A3FD9E780DFD96" author="Miyake S" box="[196,367,541,565]" pageId="102" pageNumber="212" refId="ref86514" refString="Miyake S (1983) Japanese Crustacean Decapods and Stomatopods in color. Vol. II, Brachyura (Crabs). Hoikusha Publishing Co Ltd., 277 pp." type="book" year="1983">Miyake (1983)</bibRefCitation>
and
<bibRefCitation id="EF91A9ACFFE7073E56C9FD9E7B20FD96" author="Ikeda H" box="[430,578,541,565]" pageId="102" pageNumber="212" refId="ref85235" refString="Ikeda H (1998) The Deep-Sea Crabs of Sagami Bay. Hayama Shiosai Museum, Kanagawa, Japan, 180 pp." type="book" year="1998">Ikeda (1998)</bibRefCitation>
show minimal differences. The main difference appears to be the relative lengths of the merus of the ambulatory leg, which in
<taxonomicName id="4C00AFDEFFE7073E55BFFDDD79A3FD36" authorityName="T. Sakai" authorityYear="1969" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE7073E55BFFDDD79A3FD36" italics="true" pageId="102" pageNumber="212">R. kiiensis</emphasis>
</taxonomicName>
, is proportionately longer and more slender (
<figureCitation id="133BC8D8FFE7073E55DDFDFE79DCFD16" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="102" pageNumber="212">Figs. 59C, E</figureCitation>
,
<figureCitation id="133BC8D8FFE7073E57ADFD1E7997FD16" box="[202,245,669,693]" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="102" pageNumber="212">65F</figureCitation>
) than that of
<taxonomicName id="4C00AFDEFFE7073E56EEFD1D7B76FD16" authorityName="Rathbun" authorityYear="1916" box="[393,532,669,693]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE7073E56EEFD1D7B76FD16" box="[393,532,669,693]" italics="true" pageId="102" pageNumber="212">R. bisarmata</emphasis>
</taxonomicName>
(
<figureCitation id="133BC8D8FFE7073E5544FD1E7BD2FD16" box="[547,688,669,693]" captionStart="Fig" captionStartId="97.[135,171,1927,1949]" captionTargetBox="[134,1474,268,1912]" captionTargetId="figure-8@97.[134,1474,266,1912]" captionTargetPageId="97" captionText="Fig. 59. General habitus, Rathbunaja species. A, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; B, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; C, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; D, R. kiiensis, female (43.8 × 37.3 mm) (NSMT-Cr 6419), Kyushu-Palau Ridge; E, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan; F, R. brevipes n. sp., holotype male (24.0 × 18.4 mm) (MNHN-IU-2011-2408), Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.5384720" httpUri="https://zenodo.org/record/5384720/files/figure.png" pageId="102" pageNumber="212">Figs. 59A, B</figureCitation>
,
<figureCitation id="133BC8D8FFE7073E55DAFD1E79EAFD76" captionStart="Fig" captionStartId="103.[135,171,1977,1999]" captionTargetBox="[185,1423,190,1962]" captionTargetId="figure-8@103.[185,1423,190,1963]" captionTargetPageId="103" captionText="Fig. 65. Chelipeds and ambulatory legs, Rathbunaja species. A, B, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; C, D, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; E, F, R. kiiensis, male (49.9 × 39.1 mm) (NSMT- Cr 5094), Japan; G, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; H, I, R. ursus n. sp., holotype male (50.1 × 41.5 mm) (NMCR, ex ZRC 2013.1275), Philippines; J, R. ursus n. sp., paratype male (41.6 × 32.8 mm) (ZRC 2013.1276), Philippines; K, R. ursus n. sp., paratype female (45.0 × 36.0 mm) (ZRC 2001.0594), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384736" httpUri="https://zenodo.org/record/5384736/files/figure.png" pageId="102" pageNumber="212">65B, D</figureCitation>
). The distal part of the G1 of
<taxonomicName id="4C00AFDEFFE7073E56B0FD3D7B24FD76" authorityName="T. Sakai" authorityYear="1969" box="[471,582,701,725]" class="Malacostraca" family="Majidae" genus="Leptomithrax" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B974084FFFE7073E56B0FD3D7B24FD76" box="[471,582,701,725]" italics="true" pageId="102" pageNumber="212">R. kiiensis</emphasis>
</taxonomicName>
is more curved and slightly longer than that of
<taxonomicName id="4C00AFDEFFE7073E56A9FD5D7B38FD56" authorityName="Rathbun" authorityYear="1916" box="[462,602,733,757]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="species" species="bisarmata">
<emphasis id="B974084FFFE7073E56A9FD5D7B38FD56" box="[462,602,733,757]" italics="true" pageId="102" pageNumber="212">R. bisarmata</emphasis>
</taxonomicName>
(
<figureCitation id="133BC8D8FFE7073E550EFD5E7B92FD56" box="[617,752,733,757]" captionStart="Fig" captionStartId="104.[113,148,1891,1913]" captionTargetBox="[114,1451,333,1876]" captionTargetId="figure-91@104.[95,1462,325,1884]" captionTargetPageId="104" captionText="Fig. 66. Left G1s, Rathbunaja species. AD, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; EG, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; HJ, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; KM, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan. Scales: A, H, K = 1.0 mm; E = 5.0 mm; BD, F, G, I, J, L, M= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384738" httpUri="https://zenodo.org/record/5384738/files/figure.png" pageId="102" pageNumber="212">Fig. 66HM</figureCitation>
versus
<figureCitation id="133BC8D8FFE7073E57DBFD7E785CFCB6" box="[188,318,765,789]" captionStart="Fig" captionStartId="104.[113,148,1891,1913]" captionTargetBox="[114,1451,333,1876]" captionTargetId="figure-91@104.[95,1462,325,1884]" captionTargetPageId="104" captionText="Fig. 66. Left G1s, Rathbunaja species. AD, R. bisarmata, holotype male (20.3 × 15.6 mm) (USNM 48220), Philippines; EG, R. bisarmata, male (37.1 × 29.6 mm) (ZRC 2013.1274), Philippines; HJ, R. kiiensis, holotype male (53.7 × 45.0 mm) (USNM 125885), Japan; KM, R. kiiensis, male (49.9 × 39.1 mm) (NSMT-Cr 5094), Japan. Scales: A, H, K = 1.0 mm; E = 5.0 mm; BD, F, G, I, J, L, M= 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5384738" httpUri="https://zenodo.org/record/5384738/files/figure.png" pageId="102" pageNumber="212">Fig. 66AG</figureCitation>
). As such, both species are recognised for the time being. As discussed under the genus, the distinctive features of these two species (as well as two other new ones here described) warrant the establishment of a new genus,
<taxonomicName id="4C00AFDEFFE7073E5716FCFE798FFC36" authorityName="Ng &amp; Forges" authorityYear="2015" box="[113,237,893,917]" class="Malacostraca" family="Majidae" genus="Rathbunaja" kingdom="Animalia" order="Decapoda" pageId="102" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFE7073E5716FCFE798FFC36" box="[113,237,893,917]" italics="true" pageId="102" pageNumber="212">Rathbunaja</emphasis>
</taxonomicName>
, for them.
</paragraph>
</subSubSection>
</treatment>
</document>