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<document id="6785C1C320357C7C3E1CDD1466B4476E" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFAD07745647FC5E7C70FB36" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Alcomaja desmondi Ng &amp; Forges 2015, n. sp." docType="treatment" docUuid="9465E42F-3FEC-4FE0-B6F0-C151940751AF" docUuidSource="ZooBank" docVersion="4" lastPageNumber="154" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="154" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="62A8873254E2BBA94496BD9ACBF86AB6">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="F6DE2D2B2A216B24CFD65C8C4C24F2A4">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="95475C9A69B7E0EE6F560728DE3B77DB">Forges, Bertrand Richer De</mods:namePart>
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<mods:title id="8D6156457DB8D71B296B2016E05200B4">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="25339A8F0AD2BBB369DDB8AB860FD56F">2015</mods:date>
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<treatment id="03A9654BFFAD07745647FC5E7C70FB36" LSID="urn:lsid:zoobank.org:act:9465E42F-3FEC-4FE0-B6F0-C151940751AF" httpUri="http://treatment.plazi.org/id/03A9654BFFAD07745647FC5E7C70FB36" lastPageNumber="154" pageId="44" pageNumber="154">
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<paragraph id="8BBFD45DFFAD07745647FC5E7B20FC56" blockId="44.[114,750,989,1045]" box="[288,578,989,1013]" pageId="44" pageNumber="154">
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<taxonomicName id="4C00AFDEFFAD07745647FC5E7899FC56" ID-CoL="BKVT" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[288,507,989,1013]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi" status="sp. nov.">
<emphasis id="B974084FFFAD07745647FC5E7899FC56" bold="true" box="[288,507,989,1013]" italics="true" pageId="44" pageNumber="154">Alcomaja desmondi</emphasis>
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n.
<emphasis id="B974084FFFAD07745545FC5E7B20FC56" bold="true" box="[546,578,989,1013]" pageId="44" pageNumber="154">sp.</emphasis>
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<subSubSection id="C31A87D6FFAD07745715FC7E7B8CFBB6" box="[114,750,1021,1045]" pageId="44" pageNumber="154" type="description">
<paragraph id="8BBFD45DFFAD07745715FC7E7B8CFBB6" blockId="44.[114,750,989,1045]" box="[114,750,1021,1045]" pageId="44" pageNumber="154">
(
<figureCitation id="133BC8D8FFAD0774571DFC7E7986FBB6" box="[122,228,1021,1045]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="44" pageNumber="154">Figs. 17F</figureCitation>
,
<figureCitation id="133BC8D8FFAD07745796FC7E785CFBB6" box="[241,318,1021,1045]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="44" pageNumber="154">20RT</figureCitation>
,
<figureCitation id="133BC8D8FFAD0774562DFC7E781AFBB6" box="[330,376,1021,1045]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="44" pageNumber="154">36L</figureCitation>
,
<figureCitation id="133BC8D8FFAD077456E2FC7E78D7FBB6" box="[389,437,1021,1045]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="44" pageNumber="154">39N</figureCitation>
,
<figureCitation id="133BC8D8FFAD077456A5FC7E7890FBB6" box="[450,498,1021,1045]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="44" pageNumber="154">45D</figureCitation>
,
<figureCitation id="133BC8D8FFAD07745698FC7E7B4DFBB6" box="[511,559,1021,1045]" captionStart="Fig" captionStartId="85.[135,170,1745,1767]" captionTargetBox="[136,1474,392,1730]" captionTargetId="figure-8@85.[136,1474,391,1732]" captionTargetPageId="85" captionText="Fig. 49. Male anterior thoracic sternums and abdomens, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A.gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; E, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; F, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; G, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; H, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384700" httpUri="https://zenodo.org/record/5384700/files/figure.png" pageId="44" pageNumber="154">49H</figureCitation>
,
<figureCitation id="133BC8D8FFAD0774555BFC7E7B0BFBB6" box="[572,617,1021,1045]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="44" pageNumber="154">52F</figureCitation>
,
<figureCitation id="133BC8D8FFAD07745512FC7E7BC7FBB6" box="[629,677,1021,1045]" captionStart="Fig" captionStartId="89.[135,170,1806,1828]" captionTargetBox="[136,1474,216,1791]" captionTargetId="figure-8@89.[136,1474,215,1791]" captionTargetPageId="89" captionText="Fig. 53. Chelae. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm), (ZRC 2008.0179), Spain; D, Maja cornuta, male (88.8 × 86.8 mm) (NHM 1928.12.1.177) (lectotype of Mamaia queketti Stebbing, 1908), South Africa; E, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; F, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; G, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; H, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; I, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; J, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; L, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; M, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; N, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; O, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; P, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384708" httpUri="https://zenodo.org/record/5384708/files/figure.png" pageId="44" pageNumber="154">53O</figureCitation>
,
<figureCitation id="133BC8D8FFAD077455D4FC7E7B84FBB6" box="[691,742,1021,1045]" captionStart="Fig" captionStartId="91.[135,171,1862,1884]" captionTargetBox="[170,1440,189,1847]" captionTargetId="figure-8@91.[170,1440,189,1847]" captionTargetPageId="91" captionText="Fig. 55. First ambulatory dactyli and propodi. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; G, Paramaja kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; H, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; I, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; J, Paramaja turgida n. sp., paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; K, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; L, Alcomaja irrorata, male (43.0 × 37.2 mm) (ZRC 2013.1223), Philippines; M, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; N, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384712" httpUri="https://zenodo.org/record/5384712/files/figure.png" pageId="44" pageNumber="154">55M</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFAD07745716FBBE7874FA16" pageId="44" pageNumber="154" type="materials_examined">
<paragraph id="8BBFD45DFFAD07745716FBBE7874FA16" blockId="44.[111,752,1085,1461]" pageId="44" pageNumber="154">
<emphasis id="B974084FFFAD07745716FBBE783DFBF6" bold="true" box="[113,351,1085,1109]" pageId="44" pageNumber="154">Material examined.</emphasis>
<materialsCitation id="3B68DE00FFAD0774560DFBBE79C8FB16" collectingDate="2002-06" collectionCode="NMCR" county="Balicasag Island" elevation="250" location="Philippines" municipality="Panglao" pageId="44" pageNumber="154" specimenCount="1" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54BB6AFFFFAD0774560DFBBE78B5FBF6" box="[362,471,1085,1109]" pageId="44" pageNumber="154" type="holotype">Holotype</typeStatus>
: male (35.4 ×
<quantity id="4CF879B8FFAD077455EEFBBE7B8CFBF6" box="[649,750,1085,1109]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="44" pageNumber="154" unit="mm" value="28.3">28.3 mm</quantity>
(
<collectionCode id="ED114C98FFAD0774571FFBDE79ACFBD6" box="[120,206,1117,1141]" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="44" pageNumber="154" type="Herbarium">NMCR</collectionCode>
),
<collectingCounty id="62DEACD1FFAD07745783FBDE78C2FBD6" box="[228,416,1117,1141]" pageId="44" pageNumber="154">Balicasag Island</collectingCounty>
,
<collectingMunicipality id="6BDB4E27FFAD077456C8FBDE7B68FBD6" box="[431,522,1117,1141]" pageId="44" pageNumber="154">Panglao</collectingMunicipality>
,
<collectingRegion id="49C41ABFFFAD0774557FFBDE7B3CFBD6" box="[536,606,1117,1141]" country="Philippines" name="Bohol" pageId="44" pageNumber="154">Bohol</collectingRegion>
,
<location id="8EDF8286FFAD0774550AFBDE7B89FBD6" LSID="urn:lsid:plazi:treatment:03A9654BFFAD07745647FC5E7C70FB36:8EDF8286FFAD0774550AFBDE7B89FBD6" box="[621,747,1117,1141]" county="Balicasag Island" municipality="Panglao" name="Philippines" pageId="44" pageNumber="154" stateProvince="Bohol">Philippines</location>
,
<quantity id="4CF879B8FFAD07745716FBFE7991FB36" box="[113,243,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="44" pageNumber="154" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation id="002D336EFFAD07745716FBFE7991FB36" box="[113,243,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="44" pageNumber="154" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, coll. local fishermen with tangle nets,
<date id="FFBEF29DFFAD077455DAFBFD79C8FB16" pageId="44" pageNumber="154" value="2002-06">
<collectingDate id="EFFA0B75FFAD077455DAFBFD79C8FB16" pageId="44" pageNumber="154" value="2002-06">June 2002</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B68DE00FFAD077457D1FB1D7B8EFB76" collectingDate="2002-06" collectionCode="ZRC" country="Philippines" county="Balicasag Island" elevation="250" location="Philippines" municipality="Panglao" pageId="44" pageNumber="154" specimenCode="ZRC 2013.1273" specimenCount="2" specimenCount-male="2" stateProvince="Bohol" typeStatus="paratype">
<typeStatus id="54BB6AFFFFAD077457D1FB1D7840FB15" box="[182,290,1182,1206]" pageId="44" pageNumber="154" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B974084FFFAD07745648FB1E78CCFB16" bold="true" box="[303,430,1181,1205]" pageId="44" pageNumber="154">
<collectingCountry id="F31794CDFFAD07745648FB1E78CCFB16" box="[303,430,1181,1205]" name="Philippines" pageId="44" pageNumber="154">Philippines</collectingCountry>
</emphasis>
<specimenCount id="9D061FD4FFAD077456A8FB1E7B41FB16" box="[463,547,1181,1205]" pageId="44" pageNumber="154" type="male">2 males</specimenCount>
(36.6 ×
<quantity id="4CF879B8FFAD077455E0FB1E7B89FB16" box="[647,747,1181,1205]" metricMagnitude="-2" metricUnit="m" metricValue="3.17" pageId="44" pageNumber="154" unit="mm" value="31.7">31.7 mm</quantity>
, 27.1 ×
<quantity id="4CF879B8FFAD077457D9FB3E787DFB76" box="[190,287,1213,1237]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="44" pageNumber="154" unit="mm" value="23.5">23.5 mm</quantity>
) (
<specimenCode id="DBA67C26FFAD07745652FB3E788AFB76" box="[309,488,1213,1237]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="44" pageNumber="154">ZRC 2013.1273</specimenCode>
), same data as holotype
</materialsCitation>
. —
<materialsCitation id="3B68DE00FFAD077457F4FB5E7870FA16" collectingDate="2001-11-28" collectingDateMax="2003-02" collectingDateMin="2001-11-28" collectionCode="ZRC" collectorName="Panglao &amp; M. Takeda &amp; H. Komatsu" country="Philippines" county="Balicasag Island" elevation="275" location="Balicasag Island" municipality="Panglao" pageId="44" pageNumber="154" specimenCode="ZRC 2001.0588" specimenCount="2" specimenCount-male="2" stateProvince="Bohol" typeStatus="paratype">
<specimenCount id="9D061FD4FFAD077457F4FB5E7984FB56" box="[147,230,1245,1269]" pageId="44" pageNumber="154" type="male">2 males</specimenCount>
(27.7 ×
<quantity id="4CF879B8FFAD07745625FB5E78C6FB56" box="[322,420,1245,1269]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="44" pageNumber="154" unit="mm" value="21.3">21.3 mm</quantity>
, 36.6 ×
<quantity id="4CF879B8FFAD0774569CFB5E7B3CFB56" box="[507,606,1245,1269]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="44" pageNumber="154" unit="mm" value="29.1">29.1 mm</quantity>
), 1 ovigerous female (38.0 ×
<quantity id="4CF879B8FFAD0774566AFB7E7809FAB6" box="[269,363,1277,1301]" metricMagnitude="-2" metricUnit="m" metricValue="3.26" pageId="44" pageNumber="154" unit="mm" value="32.6">32.6 mm</quantity>
) (
<specimenCode id="DBA67C26FFAD07745618FB7E7B4FFAB6" box="[383,557,1277,1301]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="44" pageNumber="154">ZRC 2001.0588</specimenCode>
),
<collectingCounty id="62DEACD1FFAD0774555AFB7E7B89FAB6" box="[573,747,1277,1301]" pageId="44" pageNumber="154">Balicasag Island</collectingCounty>
,
<collectingMunicipality id="6BDB4E27FFAD07745708FA9E79AAFA96" box="[111,200,1309,1333]" pageId="44" pageNumber="154">Panglao</collectingMunicipality>
,
<collectingRegion id="49C41ABFFFAD077457B5FA9E7874FA96" box="[210,278,1309,1333]" country="Philippines" name="Bohol" pageId="44" pageNumber="154">Bohol</collectingRegion>
,
<quantity id="4CF879B8FFAD07745647FA9E78EFFA96" box="[288,397,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" pageId="44" pageNumber="154" unit="m" value="275.0" valueMax="500.0" valueMin="50.0">
<elevation id="002D336EFFAD07745647FA9E78EFFA96" box="[288,397,1309,1333]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" pageId="44" pageNumber="154" unit="m" value="275.0" valueMax="500.0" valueMin="50.0">50500 m</elevation>
</quantity>
, coll. local fishermen with tangle nets,
<date id="FFBEF29DFFAD077457C9FABE78E8FAF6" box="[174,394,1341,1365]" pageId="44" pageNumber="154" value="2001-11-28">
<collectingDate id="EFFA0B75FFAD077457C9FABE78E8FAF6" box="[174,394,1341,1365]" pageId="44" pageNumber="154" value="2001-11-28">28 November 2001</collectingDate>
</date>
. — 1 ovigerous female (37.6 × 32.0 mm) (NSMT-Cr 22328),
<location id="8EDF8286FFAD077456AEFADE7BE0FAD6" LSID="urn:lsid:plazi:treatment:03A9654BFFAD07745647FC5E7C70FB36:8EDF8286FFAD077456AEFADE7BE0FAD6" box="[457,642,1373,1397]" country="Philippines" county="Balicasag Island" municipality="Panglao" name="Balicasag Island" pageId="44" pageNumber="154" stateProvince="Bohol">Balicasag Island</location>
,
<collectorName id="26F5B18BFFAD077455F6FADE7B89FAD6" box="[657,747,1373,1397]" pageId="44" pageNumber="154">Panglao</collectorName>
,
<collectingRegion id="49C41ABFFFAD07745716FAFE79D7FA36" box="[113,181,1405,1429]" country="Philippines" name="Bohol" pageId="44" pageNumber="154">Bohol</collectingRegion>
, coll. local fisherman via
<collectorName id="26F5B18BFFAD077456AAFAFD7B23FA36" box="[461,577,1405,1429]" pageId="44" pageNumber="154">M. Takeda</collectorName>
&amp;
<collectorName id="26F5B18BFFAD07745501FAFD7B89FA36" box="[614,747,1406,1429]" pageId="44" pageNumber="154">H. Komatsu</collectorName>
,
<date id="FFBEF29DFFAD07745716FA1E7870FA16" box="[113,274,1437,1461]" pageId="44" pageNumber="154" value="2003-02">
<collectingDate id="EFFA0B75FFAD07745716FA1E7870FA16" box="[113,274,1437,1461]" pageId="44" pageNumber="154" value="2003-02">February 2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFAD07745716FA5E7B7EF876" pageId="44" pageNumber="154" type="diagnosis">
<paragraph id="8BBFD45DFFAD07745716FA5E7B7EF876" blockId="44.[111,752,1501,2005]" pageId="44" pageNumber="154">
<emphasis id="B974084FFFAD07745716FA5E7984FA56" bold="true" box="[113,230,1501,1525]" pageId="44" pageNumber="154">Diagnosis.</emphasis>
Carapace pyriform; dorsal surface of carapace densely covered with long, stiff setae; with small rounded granules (
<figureCitation id="133BC8D8FFAD077457B0F99E7851F996" box="[215,307,1565,1589]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="44" pageNumber="154">Fig. 17F</figureCitation>
). Pseudorostral spines with rounded crosssection (
<figureCitation id="133BC8D8FFAD077457ADF9BE7849F9F6" box="[202,299,1597,1621]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="44" pageNumber="154">Fig. 36L</figureCitation>
). Antorbital spine strong, curved upwards, appears straight from dorsal view; intercalated spine acutely triangular; postorbital spine very long (
<figureCitation id="133BC8D8FFAD0774554FF9FE7BEFF936" box="[552,653,1661,1685]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="44" pageNumber="154">Fig. 36L</figureCitation>
). Lateral and branchial spines strong in juveniles and adults; 3 spines on median row: 2 gastric, 1 cardiac; 2 distinct spines on posterior carapace margin (
<figureCitation id="133BC8D8FFAD077456E9F95E7888F956" box="[398,490,1757,1781]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="44" pageNumber="154">Fig. 17F</figureCitation>
). Subhepatic region with numerous very small rounded granules on surface (
<figureCitation id="133BC8D8FFAD077455E5F97E7B81F8B6" box="[642,739,1789,1813]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="44" pageNumber="154">Fig. 39N</figureCitation>
). Basal antennal article with numerous granules (
<figureCitation id="133BC8D8FFAD0774551AF89E7B83F896" box="[637,737,1821,1845]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="44" pageNumber="154">Fig. 39N</figureCitation>
). Ischium of third maxilliped almost smooth to slightly rugose (
<figureCitation id="133BC8D8FFAD07745710F8DE79BEF8D6" box="[119,220,1885,1909]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="44" pageNumber="154">Fig. 45D</figureCitation>
). Ambulatory legs relatively short, first leg never twice length of carapace; surfaces strongly setose, covered with long coarse setae (
<figureCitation id="133BC8D8FFAD0774560AF81E78B7F816" box="[365,469,1949,1973]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="44" pageNumber="154">Figs. 17F</figureCitation>
,
<figureCitation id="133BC8D8FFAD07745686F81E7B76F816" box="[481,532,1949,1973]" captionStart="Fig" captionStartId="91.[135,171,1862,1884]" captionTargetBox="[170,1440,189,1847]" captionTargetId="figure-8@91.[170,1440,189,1847]" captionTargetPageId="91" captionText="Fig. 55. First ambulatory dactyli and propodi. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; G, Paramaja kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; H, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; I, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; J, Paramaja turgida n. sp., paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; K, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; L, Alcomaja irrorata, male (43.0 × 37.2 mm) (ZRC 2013.1223), Philippines; M, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; N, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384712" httpUri="https://zenodo.org/record/5384712/files/figure.png" pageId="44" pageNumber="154">55M</figureCitation>
). G1 almost straight with distinct ventral fold (
<figureCitation id="133BC8D8FFAD077456E9F83E7B6EF876" box="[398,524,1981,2005]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="44" pageNumber="154">Fig. 20RT</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFAD0774544AFF3E7C7EFEB6" pageId="44" pageNumber="154" type="etymology">
<paragraph id="8BBFD45DFFAD0774544AFF3E7C7EFEB6" blockId="44.[813,1452,189,278]" pageId="44" pageNumber="154">
<emphasis id="B974084FFFAD0774544AFF3E7ACDFF76" bold="true" box="[813,943,189,213]" pageId="44" pageNumber="154">Etymology.</emphasis>
The name is formed to honor Desmond Griffin, a carcinologist who devoted a large part of his life to the taxonomy of Indo-West Pacific majoid crabs.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFAD0774544AFEBE7C70FB36" pageId="44" pageNumber="154" type="discussion">
<paragraph id="8BBFD45DFFAD0774544AFEBE7D25FD76" blockId="44.[811,1452,317,725]" pageId="44" pageNumber="154">
<emphasis id="B974084FFFAD0774544AFEBE7AF9FEF6" bold="true" box="[813,923,317,341]" pageId="44" pageNumber="154">Remarks.</emphasis>
<taxonomicName id="4C00AFDEFFAD077454C6FEBE7D11FEF6" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[929,1139,317,341]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi" status="sp. nov.">
<emphasis id="B974084FFFAD077454C6FEBE7D11FEF6" box="[929,1139,317,341]" italics="true" pageId="44" pageNumber="154">Alcomaja desmondi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFAD0774531EFEBD7DD0FEF6" box="[1145,1202,318,341]" pageId="44" pageNumber="154" rank="species">n. sp.</taxonomicNameLabel>
superficially resembles
<taxonomicName id="4C00AFDEFFAD0774544AFEDE7DB7FED6" authority="(Rathbun, 1916)" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[813,1237,349,373]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="suluensis">
<emphasis id="B974084FFFAD0774544AFEDE7D69FED6" box="[813,1035,349,373]" italics="true" pageId="44" pageNumber="154">Holthuija suluensis</emphasis>
(
<bibRefCitation id="EF91A9ACFFAD07745379FEDE7DACFED6" author="Rathbun MJ" box="[1054,1230,349,373]" pageId="44" pageNumber="154" pagination="527 - 559" refId="ref87575" refString="Rathbun MJ (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135): 527 - 559." type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
)
</taxonomicName>
, in general shape, especially since both species have strong lateral and branchial spines as well as two gastric spines on the carapace. Excluding the generic level differences,
<taxonomicName id="4C00AFDEFFAD07745393FE3D7C1CFE76" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1268,1406,445,469]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD07745393FE3D7C1CFE76" box="[1268,1406,445,469]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
has a more rounded carapace, especially on the branchial regions; the pseudorostral spines are more diverging in
<emphasis id="B974084FFFAD0774544AFD9E7D68FD96" box="[813,1034,541,565]" italics="true" pageId="44" pageNumber="154">
<taxonomicName id="4C00AFDEFFAD0774544AFD9E7D61FD96" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[813,1027,541,565]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="suluensis">Holthuija suluensis</taxonomicName>
;
</emphasis>
the first gastric spine is placed more anteriorly in
<taxonomicName id="4C00AFDEFFAD077454A7FDBD7D28FDF6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[960,1098,573,597]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD077454A7FDBD7D28FDF6" box="[960,1098,573,597]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
; the antorbital spine is straight and directed outwards in
<taxonomicName id="4C00AFDEFFAD0774532EFDDD7DB3FDD6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1097,1233,605,629]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD0774532EFDDD7DB3FDD6" box="[1097,1233,605,629]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
(versus curved and directed anteriorly in
<taxonomicName id="4C00AFDEFFAD0774537DFDFD7DFDFD36" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[1050,1183,637,661]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="suluensis">
<emphasis id="B974084FFFAD0774537DFDFD7DFDFD36" box="[1050,1183,637,661]" italics="true" pageId="44" pageNumber="154">H. suluensis</emphasis>
</taxonomicName>
), and the setae covering the ambulatory legs in
<taxonomicName id="4C00AFDEFFAD07745357FD1D7DDAFD16" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1072,1208,669,693]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD07745357FD1D7DDAFD16" box="[1072,1208,669,693]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
are longer and stiffer compared to
<taxonomicName id="4C00AFDEFFAD077454D9FD3D7D22FD76" baseAuthorityName="Rathbun" baseAuthorityYear="1916" box="[958,1088,701,725]" class="Malacostraca" family="Majidae" genus="Holthuija" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="suluensis">
<emphasis id="B974084FFFAD077454D9FD3D7D22FD76" box="[958,1088,701,725]" italics="true" pageId="44" pageNumber="154">H. suluensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBFD45DFFAD0774544AFD7E7CEDFC76" blockId="44.[811,1452,765,982]" pageId="44" pageNumber="154">
Within
<taxonomicName id="4C00AFDEFFAD0774541BFD7E7A80FCB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[892,994,765,789]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFAD0774541BFD7E7A80FCB6" box="[892,994,765,789]" italics="true" pageId="44" pageNumber="154">Alcomaja</emphasis>
</taxonomicName>
,
<taxonomicName id="4C00AFDEFFAD07745489FD7D7D12FCB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1006,1136,765,789]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD07745489FD7D7D12FCB6" box="[1006,1136,765,789]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
stands out in several features. It is unique among members of the genus in possessing long stiff setae covering its carapace and legs; and the G1 structure is atypical, being almost straight, much stouter with an unusually broad ventral flap (
<figureCitation id="133BC8D8FFAD077453E1FCFE7C65FC36" box="[1158,1287,893,917]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; EG, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; HJ, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; KN, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; OQ, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; RT, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, KR]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="44" pageNumber="154">Fig. 20RT</figureCitation>
). Since the rest of its characters agree with
<taxonomicName id="4C00AFDEFFAD07745307FC1E7DA4FC16" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1120,1222,925,949]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFAD07745307FC1E7DA4FC16" box="[1120,1222,925,949]" italics="true" pageId="44" pageNumber="154">Alcomaja</emphasis>
</taxonomicName>
as presently defined, it seems best to leave it in this genus for the time being
</paragraph>
<paragraph id="8BBFD45DFFAD0774544AFC7E7C70FB36" blockId="44.[811,1452,1021,1173]" pageId="44" pageNumber="154">
The dense stiff setae on the specimens of
<taxonomicName id="4C00AFDEFFAD0774538AFC7D7C13FBB6" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1261,1393,1021,1045]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="44" pageNumber="154" phylum="Arthropoda" rank="species" species="desmondi">
<emphasis id="B974084FFFAD0774538AFC7D7C13FBB6" box="[1261,1393,1021,1045]" italics="true" pageId="44" pageNumber="154">A. desmondi</emphasis>
</taxonomicName>
from Balicasag trap a thick layer of silicate sponge spicules, which makes their handling very difficult. It appears to be a species living in the deep sea on the deep steep rocky slopes of Balicasag Island at depths exceeding
<specimenCount id="9D061FD4FFAD077453A1FBFE7C70FB36" box="[1222,1298,1149,1173]" pageId="44" pageNumber="154" type="generic">
<quantity id="4CF879B8FFAD077453A1FBFE7C6CFB36" box="[1222,1294,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="44" pageNumber="154" unit="m" value="300.0">300 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>