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<mods:title id="3E6AA9D69F535B76E7B16847F6D2E827">A review of the deepwater cardinalfish genus Epigonus (Perciformes: Epigonidae) of the Western Indian Ocean, with description of two new species</mods:title>
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<emphasis id="B97FD23C6E78407CFF185E7615449D94" bold="true" box="[151,585,620,646]" pageId="21" pageNumber="282">
<emphasis id="B97FD23C6E78407CFF185E7616779D94" bold="true" box="[151,378,620,646]" italics="true" pageId="21" pageNumber="282">Epigonus robustus</emphasis>
(
<bibRefCitation id="EF9A73DF6E78407CFE065E76154C9D94" box="[393,577,620,646]" pageId="21" pageNumber="282" refId="ref17595" refString="Barnard, K. H. (1927) Diagnoses of new genera and species of South African marine fishes. Annals and Magazine of Natural History, Series 9, 20 (115), 66 - 79. https: // doi. org / 10.1080 / 00222932708655566" type="journal article">Barnard, 1927</bibRefCitation>
)
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(English name: Robust Cardinalfish; Robust Deepsea Cardinal) (
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,
<figureCitation id="133012AB6E78407CFF755EA916149DD9" box="[250,281,691,715]" captionStart="FIGURE17" captionStartId="22.[151,250,1718,1740]" captionTargetBox="[188,1398,558,1667]" captionTargetId="figure@22.[188,1398,529,1667]" captionTargetPageId="22" captionText="FIGURE17.Epigonus robustus, SAIAB 38259, 1of 2 specimens,155.0 mm SL.A:lateral view;B: ventral view of symphysis of lower jaw (arrows show the nub-like structures); C: SAM 13080, holotype, standard length unknown." httpUri="https://zenodo.org/record/1182203/files/figure.png" pageId="21" pageNumber="282">17</figureCitation>
; Tables 1, 2)
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<emphasis id="B97FD23C6E78407CFF185EE016879C02" box="[151,394,762,784]" italics="true" pageId="21" pageNumber="282">Parahynnodus robustus</emphasis>
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<bibRefCitation id="EF9A73DF6E78407CFE1B5EE015219C02" box="[404,556,762,784]" pageId="21" pageNumber="282" refId="ref17595" refString="Barnard, K. H. (1927) Diagnoses of new genera and species of South African marine fishes. Annals and Magazine of Natural History, Series 9, 20 (115), 66 - 79. https: // doi. org / 10.1080 / 00222932708655566" type="journal article">Barnard, 1927</bibRefCitation>
: 70
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(original description;
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locality: off
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Point,
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,
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depth;
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:
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13080).
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<taxonomicName id="4C0B75AD6E78407CFF185F2315159C5C" ID-CoL="6FTL8" authority=": Mayer, 1974: 189" authorityName=": Mayer" authorityPageNumber="189" authorityYear="1974" box="[151,536,824,846]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B97FD23C6E78407CFF185F23165E9C5C" box="[151,339,824,846]" italics="true" pageId="21" pageNumber="282">Epigonus robustus</emphasis>
:
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: 189
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, fig. 20 (description,
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,
<collectingCountry id="F31C4EBE6E78407CFCD35F2214EE9C5C" box="[860,995,824,846]" name="South Africa" pageId="21" pageNumber="282">
<collectingRegion id="49CFC0CC6E78407CFCD35F22149A9C5C" box="[860,919,824,846]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Africa
</collectingCountry>
);
<treatmentCitation id="0AAA283F6E78407CFC7A5F2313909C5C" author="" box="[1013,1181,824,846]" page="16" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFC7A5F2313749C5C" box="[1013,1145,824,846]" pageId="21" pageNumber="282" refId="ref20112" refString="Mayer, G. F. (1975) Results of the research cruises of FRV &quot; Walther Herwig &quot; to South America. XXXIX. The Epigonine fauna of the South Atlantic, with a key to the genera and a redescription of Rosenblattia robusta Mead and De Falla. Archiv fur Fischereiwissenschaft, 26, 13 - 28." type="journal article">Mayer, 1975</bibRefCitation>
: 16
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(brief description,
<collectingRegion id="49CFC0CC6E78407CFAEE5F2212919C5C" box="[1377,1436,824,846]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Atlantic);
<treatmentCitation id="0AAA283F6E78407CFEBE5F4C16DB9C7E" author="" box="[305,470,854,876]" page="559" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFEBE5F4C16AE9C7E" box="[305,419,854,876]" pageId="21" pageNumber="282" refId="ref18641" refString="Gon, O. (1986) Family No. 175: Apogonidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smiths' Sea Fishes. Macmillan South Africa Ltd, Johannesburg, pp. 546 - 561." type="book chapter">Gon, 1986</bibRefCitation>
: 559
</treatmentCitation>
(
<collectingCountry id="F31C4EBE6E78407CFE695F4C157C9C7E" box="[486,625,854,876]" name="South Africa" pageId="21" pageNumber="282">
<collectingRegion id="49CFC0CC6E78407CFE695F4C152E9C7E" box="[486,547,854,876]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Africa
</collectingCountry>
);
<treatmentCitation id="0AAA283F6E78407CFD095F4C14659C7E" author="" box="[646,872,854,876]" page="200" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFD095F4C14389C7E" box="[646,821,854,876]" pageId="21" pageNumber="282" refId="ref20875" refString="Nakamura, I. (1986) Apogonidae. In: Nakamura, I. (Ed.), Important Fishes Trawled off Patagonia. Japan Marine Fisheries Resources Research Center, Tokyo, pp. 200 - 203." type="book chapter">Nakamura, 1986</bibRefCitation>
: 200
</treatmentCitation>
(description and photograph, from southern
<collectingCountry id="F31C4EBE6E78407CFACE5F4C12729C7E" box="[1345,1407,854,876]" name="Brazil" pageId="21" pageNumber="282">Brazil</collectingCountry>
to
<collectingCountry id="F31C4EBE6E78407CFF485F6F16219C99" box="[199,300,885,907]" name="Argentina" pageId="21" pageNumber="282">Argentine</collectingCountry>
Patagonia); Parin &amp; Abramov, 1986b: 189 (description, southern
<collectingCountry id="F31C4EBE6E78407CFC795F6F13589C99" box="[1014,1109,885,907]" name="Australia" pageId="21" pageNumber="282">Australia</collectingCountry>
,
<collectingRegion id="49CFC0CC6E78407CFBEB5F6C13BB9C99" box="[1124,1206,886,907]" country="Papua New Guinea" name="Western" pageId="21" pageNumber="282">Western</collectingRegion>
Indian Ocean,
<collectingCountry id="F31C4EBE6E78407CFAEE5F6F16009CB8" name="South Africa" pageId="21" pageNumber="282">
<collectingRegion id="49CFC0CC6E78407CFAEE5F6F12919C99" box="[1377,1436,885,907]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Africa
</collectingCountry>
);
<treatmentCitation id="0AAA283F6E78407CFE905F8E15209CB8" author="" box="[287,557,916,938]" page="553" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFE905F8E16F79CB8" box="[287,506,916,938]" pageId="21" pageNumber="282" refId="ref17359" refString="Allen, G. R. &amp; Cross, N. (1989) Apogonidae. In: Paxton, J. R., Hoese, D. F., Allen, G. R. &amp; Hanley, J. E. (Eds.), Zoological Catalogue of Australia. 7. Pisces. Petromyzontidae to Carangidae. Australian Government Publishing Service, Canberra, pp. 544 - 549." type="book chapter">Allen &amp; Cross, 1989</bibRefCitation>
: 553
</treatmentCitation>
(list,
<collectingCountry id="F31C4EBE6E78407CFDE45F8E15C19CB8" box="[619,716,916,938]" name="Australia" pageId="21" pageNumber="282">Australia</collectingCountry>
); Pequeño, 1989: 60 (list,
<collectingCountry id="F31C4EBE6E78407CFC6F5F8E13169CB8" box="[992,1051,916,938]" name="Chile" pageId="21" pageNumber="282">Chile</collectingCountry>
); Paulin,
<emphasis id="B97FD23C6E78407CFBF15F8F13A09CB8" box="[1150,1197,916,938]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 1989: 180 (key,
<collectingCountry id="F31C4EBE6E78407CFAE35F8F16109CDA" name="New Zealand" pageId="21" pageNumber="282">New Zealand</collectingCountry>
);
<treatmentCitation id="0AAA283F6E78407CFEA15FA816F29CDA" author="" box="[302,511,946,968]" page="100" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFEA15FA816DD9CDA" box="[302,464,946,968]" pageId="21" pageNumber="282" refId="ref17192" refString="Abramov, A. A. (1992) Species composition and distribution of Epigonus (Epigonidae) in the world ocean. Journal of Ichthyology, 32, 94 - 108." type="journal article">Abramov, 1992</bibRefCitation>
: 100
</treatmentCitation>
(key);
<treatmentCitation id="0AAA283F6E78407CFDC75FA8144D9CDA" author="" box="[584,832,946,968]" page="563" pageId="21" pageNumber="282">
<bibRefCitation id="EF9A73DF6E78407CFDC75FA8141D9CDA" box="[584,784,946,968]" pageId="21" pageNumber="282" refId="ref18595" refString="Gomon, M. F., Glover, J. C. M. &amp; Kuiter, R. H. (1994) The Fishes of Australia's South Coast. Flora and Fauna of South Australia Handbooks Committee. Adelaide, 992 pp." type="book">
Gomon
<emphasis id="B97FD23C6E78407CFD155FA915CA9CDA" box="[666,711,946,968]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 1994
</bibRefCitation>
: 563
</treatmentCitation>
(key and brief description, southern
<collectingCountry id="F31C4EBE6E78407CFB3A5FA8121B9CDA" box="[1205,1302,946,968]" name="Australia" pageId="21" pageNumber="282">Australia</collectingCountry>
); Williams
<emphasis id="B97FD23C6E78407CFA055FA917D69CF5" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 1996: 153 (list,
<collectingRegion id="49CFC0CC6E78407CFE165FC8155B9CF5" box="[409,598,977,999]" country="Australia" name="Western Australia" pageId="21" pageNumber="282">
<collectingRegion id="49CFC0CC6E78407CFE165FC816E69CF5" box="[409,491,978,999]" country="Papua New Guinea" name="Western" pageId="21" pageNumber="282">Western</collectingRegion>
<collectingCountry id="F31C4EBE6E78407CFE7A5FCB155B9CF5" box="[501,598,977,999]" name="Australia" pageId="21" pageNumber="282">Australia</collectingCountry>
</collectingRegion>
);
<bibRefCitation id="EF9A73DF6E78407CFDE55FCB14309CF5" box="[618,829,977,999]" pageId="21" pageNumber="282" refId="ref17719" refString="Bianchi, G., Carpenter, K. E., Roux, J. - P., Molloy, F. J., Boyer, D. &amp; Boyer, H. J. (Eds.) (1999) FAO Species Identification Field Guide for Fishery Purposes. The Living Marine Resources of Namibia. FAO, Rome, 256 pp." type="book">
Bianchi
<emphasis id="B97FD23C6E78407CFD4D5FC815FC9CF5" box="[706,753,977,999]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 1999
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: (list,
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); Trunov, 1999: 466 (list, Atlantic Ocean);
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: 32
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(list, southern
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);
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Hoese
<emphasis id="B97FD23C6E78407CFD605FEB14129B14" box="[751,799,1008,1030]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 2006
</bibRefCitation>
: 1114
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(list,
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);
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Gomon
<emphasis id="B97FD23C6E78407CFB3A5FEB13E89B14" box="[1205,1253,1008,1030]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
., 2008
</bibRefCitation>
: 563
</treatmentCitation>
(list,
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);
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<bibRefCitation id="EF9A73DF6E78407CFEB3581415339B36" box="[316,574,1038,1060]" pageId="21" pageNumber="282" refId="ref20944" refString="Okamoto, M. (2011) A new species of deepwater cardinalfish, Epigonus mayeri, from the eastern Central Atlantic, and redescription of Epigonus heracleus Parin and Abramov 1986 (Perciformes: Epigonidae). Ichthyological Research, 58, 101 - 108. https: // doi. org / 10.1007 / s 10228 - 010 - 0195 - x" type="journal article">
Okamoto &amp;
<collectingRegion id="49CFC0CC6E78407CFE30581416F09B36" box="[447,509,1038,1060]" country="Japan" name="Fukui" pageId="21" pageNumber="282">Fukui</collectingRegion>
, 2011
</bibRefCitation>
: 391
</treatmentCitation>
(key); Okamoto, 2012: 252, fig. 10-m (key and photograph); Stewart &amp; Gon, 2015: 1225: (description and photograph,
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).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E78407CFF18587513009A0A" pageId="21" pageNumber="282" type="materials_examined">
<paragraph id="8BB40E2E6E78407CFF18587513009A0A" blockId="21.[151,1436,1135,2025]" pageId="21" pageNumber="282">
<materialsCitation id="3B6304736E78407CFF18587512959B9A" ID-GBIF-Occurrence="1820370033" box="[151,1432,1135,1160]" collectionCode="SAM" country="South Africa" elevation="841" location="Cape" pageId="21" pageNumber="282" specimenCount="1" stateProvince="Cape" typeStatus="holotype">
<emphasis id="B97FD23C6E78407CFF185875168D9B9A" bold="true" box="[151,384,1135,1160]" pageId="21" pageNumber="282">Material examined.</emphasis>
<collectionCode id="ED1A96EB6E78407CFE09586A16CE9B9A" box="[390,451,1136,1160]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="21" pageNumber="282">SAM</collectionCode>
13080,
<typeStatus id="54B0B08C6E78407CFD925875158F9B9A" box="[541,642,1135,1160]" pageId="21" pageNumber="282" type="holotype">holotype</typeStatus>
,
<quantity id="4CF3A3CB6E78407CFD02586A15E49B95" box="[653,745,1136,1160]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="282" unit="mm" value="170.0">170 mm</quantity>
SL, Off
<collectingRegion id="49CFC0CC6E78407CFCC4586A14899B9A" box="[843,900,1136,1160]" country="South Africa" name="Western Cape" pageId="21" pageNumber="282">Cape</collectingRegion>
Point,
<collectingCountry id="F31C4EBE6E78407CFC5C587513689B9A" box="[979,1125,1135,1160]" name="South Africa" pageId="21" pageNumber="282">South Africa</collectingCountry>
,
<elevation id="0026E91D6E78407CFBFF586A13BA9B95" box="[1136,1207,1136,1160]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="21" pageNumber="282" unit="m" value="841.0">
<quantity id="4CF3A3CB6E78407CFBFF586A13BA9B95" box="[1136,1207,1136,1160]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="21" pageNumber="282" unit="m" value="841.0">841 m</quantity>
</elevation>
depth,
<elevation id="0026E91D6E78407CFA86586A125D9B95" box="[1289,1360,1136,1160]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="21" pageNumber="282" unit="m" value="841.0">
<quantity id="4CF3A3CB6E78407CFA86586A125D9B95" box="[1289,1360,1136,1160]" metricMagnitude="2" metricUnit="m" metricValue="8.41" pageId="21" pageNumber="282" unit="m" value="841.0">841 m</quantity>
</elevation>
depth
</materialsCitation>
;
<materialsCitation id="3B6304736E78407CFF18588E165D9BC2" ID-GBIF-Occurrence="1820370032" collectingDate="1999-11-12" collectionCode="CSIRO" country="South Africa" elevation="897" location="western South Indian Ocean" pageId="21" pageNumber="282" specimenCount="1" stateProvince="South" typeStatus="holotype">
<collectionCode id="ED1A96EB6E78407CFF18588E17E79BBE" box="[151,234,1172,1196]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="21" pageNumber="282">CSIRO</collectionCode>
H 5375-02,
<quantity id="4CF3A3CB6E78407CFE06588E150C9BBE" box="[393,513,1172,1196]" metricMagnitude="-1" metricUnit="m" metricValue="1.847" pageId="21" pageNumber="282" unit="mm" value="184.7">184.7 mm</quantity>
SL, 39°01 S, 46°32 E,
<location id="8ED458F56E78407CFCA4588F13709BBE" LSID="urn:lsid:plazi:treatment:03A2BF386E78407FFF185E76158A9EC2:8ED458F56E78407CFCA4588F13709BBE" box="[811,1149,1172,1197]" country="South Africa" name="western South Indian Ocean" pageId="21" pageNumber="282" stateProvince="South">
western
<collectingRegion id="49CFC0CC6E78407CFC00588E14DD9BBF" box="[911,976,1172,1197]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Indian Ocean
</location>
,
<elevation id="0026E91D6E78407CFB01588E122C9BBE" box="[1166,1313,1172,1197]" metricMagnitude="2" metricUnit="m" metricValue="8.965" metricValueMax="10.17" metricValueMin="7.76" pageId="21" pageNumber="282" unit="m" value="896.5" valueMax="1017.0" valueMin="776.0">
<quantity id="4CF3A3CB6E78407CFB01588E122C9BBE" box="[1166,1313,1172,1197]" metricMagnitude="2" metricUnit="m" metricValue="8.965" metricValueMax="10.17" metricValueMin="7.76" pageId="21" pageNumber="282" unit="m" value="896.5" valueMax="1017.0" valueMin="776.0">7761017 m</quantity>
</elevation>
depth,
<collectingDate id="EFF1D1066E78407CFA0F588E165D9BC2" pageId="21" pageNumber="282" value="1999-11-12">
<date id="FFB528EE6E78407CFA0F588E165D9BC2" pageId="21" pageNumber="282" value="1999-11-12">12 November 1999</date>
</collectingDate>
</materialsCitation>
; CSIRO H 5398-01,
<quantity id="4CF3A3CB6E78407CFDB158AD15BC9BDD" box="[574,689,1207,1232]" metricMagnitude="-1" metricUnit="m" metricValue="1.767" pageId="21" pageNumber="282" unit="mm" value="176.7">176.7 mm</quantity>
SL,
<materialsCitation id="3B6304736E78407CFD6858A2134B9BE6" ID-GBIF-Occurrence="1820370041" collectingDate="1999-11-13" collectionCode="CSIRO" country="South Africa" elevation="889" location="western South Indian Ocean" pageId="21" pageNumber="282" specimenCount="3" stateProvince="South" typeStatus="holotype">
<collectionCode id="ED1A96EB6E78407CFD6858A214379BC2" box="[743,826,1208,1232]" country="Australia" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="21" pageNumber="282">CSIRO</collectionCode>
H 5398-02,
<specimenCount id="9D0DC5A76E78407CFC4658A2135A9BC2" box="[969,1111,1207,1232]" pageId="21" pageNumber="282" type="generic">3 specimens</specimenCount>
,
<quantity id="4CF3A3CB6E78407CFBEC58A2122F9BDD" box="[1123,1314,1208,1232]" metricMagnitude="-1" metricUnit="m" metricValue="1.938" metricValueMax="1.972" metricValueMin="1.904" pageId="21" pageNumber="282" unit="mm" value="193.8" valueMax="197.2" valueMin="190.4">190.4197.2 mm</quantity>
SL, 39°01 S, 46°32 E,
<location id="8ED458F56E78407CFEA958C715639BE6" LSID="urn:lsid:plazi:treatment:03A2BF386E78407FFF185E76158A9EC2:8ED458F56E78407CFEA958C715639BE6" box="[294,622,1244,1269]" country="South Africa" name="western South Indian Ocean" pageId="21" pageNumber="282" stateProvince="South">
western
<collectingRegion id="49CFC0CC6E78407CFE0858C616C59BE7" box="[391,456,1244,1269]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Indian Ocean
</location>
,
<elevation id="0026E91D6E78407CFDF358C614019BE6" box="[636,780,1244,1269]" metricMagnitude="2" metricUnit="m" metricValue="8.895" metricValueMax="10.67" metricValueMin="7.12" pageId="21" pageNumber="282" unit="m" value="889.5" valueMax="1067.0" valueMin="712.0">
<quantity id="4CF3A3CB6E78407CFDF358C614019BE6" box="[636,780,1244,1269]" metricMagnitude="2" metricUnit="m" metricValue="8.895" metricValueMax="10.67" metricValueMin="7.12" pageId="21" pageNumber="282" unit="m" value="889.5" valueMax="1067.0" valueMin="712.0">7121067 m</quantity>
</elevation>
depth,
<collectingDate id="EFF1D1066E78407CFCEA58C6134B9BE6" box="[869,1094,1244,1269]" pageId="21" pageNumber="282" value="1999-11-13">
<date id="FFB528EE6E78407CFCEA58C6134B9BE6" box="[869,1094,1244,1269]" pageId="21" pageNumber="282" value="1999-11-13">13 November 1999</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B6304736E78407CFBDB58C613059A0A" ID-GBIF-Occurrence="1820370043" collectionCode="SAIAB" country="South Africa" latitude="-32.400833" location="West Coast Slope" longLatPrecision="12" longitude="16.233334" pageId="21" pageNumber="282" specimenCode="SAIAB 38259" specimenCount="2" stateProvince="West" typeStatus="holotype">
<specimenCode id="DBADA6556E78407CFBDB58C613F69BE7" box="[1108,1275,1244,1269]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="21" pageNumber="282">SAIAB 38259</specimenCode>
,
<specimenCount id="9D0DC5A76E78407CFA8658C612959BE7" box="[1289,1432,1244,1269]" pageId="21" pageNumber="282" type="generic">2 specimens</specimenCount>
, 150.7155.0 mm SL,
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,
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,
<location id="8ED458F56E78407CFD2D591A14649A0A" LSID="urn:lsid:plazi:treatment:03A2BF386E78407FFF185E76158A9EC2:8ED458F56E78407CFD2D591A14649A0A" box="[674,873,1279,1304]" country="South Africa" latitude="-32.400833" longLatPrecision="12" longitude="16.233334" name="West Coast Slope" pageId="21" pageNumber="282" stateProvince="West">
<collectingRegion id="49CFC0CC6E78407CFD2D591A15D49A05" box="[674,729,1280,1303]" country="Tanzania" name="Mjini Magharibi" pageId="21" pageNumber="282">West</collectingRegion>
<collectingRegion id="49CFC0CC6E78407CFD6F591A142D9A0A" box="[736,800,1280,1304]" country="Tanzania" name="Pwani" pageId="21" pageNumber="282">Coast</collectingRegion>
Slope
</location>
,
<collectingCountry id="F31C4EBE6E78407CFCFA58E513059A0A" box="[885,1032,1279,1304]" name="South Africa" pageId="21" pageNumber="282">South Africa</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E78407CFF48593E16A59ABA" pageId="21" pageNumber="282" type="diagnosis">
<paragraph id="8BB40E2E6E78407CFF48593E16A59ABA" blockId="21.[151,1436,1135,2025]" pageId="21" pageNumber="282">
<emphasis id="B97FD23C6E78407CFF48593E164C9A2F" bold="true" box="[199,321,1316,1341]" pageId="21" pageNumber="282">Diagnosis.</emphasis>
Dorsal-fin rays VII-I, 9; pectoral-fin rays 1618; total gill rakers 2933; vertebrae 11 + 14; pyloric caeca 67; pored lateral-line scales 4650 + 35; pungent opercular spine present; blunt maxillary mustache-like processes present; ribs on last abdominal vertebra present; tongue toothless; two nub-like structures present on symphysis of lower jaw.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E78407CFF4859AE14289802" pageId="21" pageNumber="282" type="description">
<paragraph id="8BB40E2E6E78407CFF4859AE14289802" blockId="21.[151,1436,1135,2025]" pageId="21" pageNumber="282">
<emphasis id="B97FD23C6E78407CFF4859AE16D19ADF" bold="true" box="[199,476,1460,1485]" pageId="21" pageNumber="282">Measurements (% SL)</emphasis>
; counts are given in Table 1. Head length 31.534.2; head width 15.017.1; head height 14.916.4; body depth 18.822.5; body width 13.616.2; caudal-peduncle depth 9.210.8; caudal-peduncle length 24.330.6; orbital diameter 12.214.3; interorbital width 7.38.4; postorbital length 12.515.3; upper-jaw length 12.614.4; lower-jaw length 14.015.3; snout length 6.37.7; pre-first dorsal-fin length 36.839.4; presecond dorsal-fin length 56.260.3; pre-pectoral-fin length 31.233.5; pre-pelvic-fin length 34.137.0; pre-anus length 55.461.4; pre-anal-fin length 62.471.8; first spine length on first dorsal fin 1.23.1; second spine length on first dorsal fin 10.716.0; third spine length on first dorsal fin 13.719.3; second dorsal-fin spine length 10.0 15.7; first anal-fin spine length 2.54.4; second anal-fin spine length 10.716.3; pelvic-fin spine length 11.316.8; first dorsal-fin base length 10.313.6; second dorsal-fin base length 9.112.0; anal-fin base length 8.810.4; pectoral-fin length 18.821.4; pelvic-fin length 17.123.0.
</paragraph>
</subSubSection>
<subSubSection id="C3115DA56E78407FFF485B06158A9EC2" lastPageId="22" lastPageNumber="283" pageId="21" pageNumber="282" type="distribution">
<paragraph id="8BB40E2E6E78407CFF485B061312986F" blockId="21.[151,1436,1135,2025]" pageId="21" pageNumber="282">
<emphasis id="B97FD23C6E78407CFF485B06166D9827" bold="true" box="[199,352,1820,1845]" pageId="21" pageNumber="282">Distribution.</emphasis>
<collectingRegion id="49CFC0CC6E78407CFEE25B0616A39827" box="[365,430,1820,1845]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Indian Ocean (
<bibRefCitation id="EF9A73DF6E78407CFDE15B06140C9826" box="[622,769,1820,1844]" pageId="21" pageNumber="282" refId="ref20071" refString="Mayer, G. F. (1974) A revision of the cardinalfish genus Epigonus (Perciformes, Apogonidae), with descriptions of two new species. Bulletin Museum of Comparative Zoology, 146, 147 - 203." type="journal article">Mayer 1974</bibRefCitation>
;
<bibRefCitation id="EF9A73DF6E78407CFC9C5B0614819827" box="[787,908,1820,1845]" pageId="21" pageNumber="282" refId="ref18641" refString="Gon, O. (1986) Family No. 175: Apogonidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smiths' Sea Fishes. Macmillan South Africa Ltd, Johannesburg, pp. 546 - 561." type="book chapter">Gon 1986</bibRefCitation>
; Parin &amp; Abramov 1986b;
<bibRefCitation id="EF9A73DF6E78407CFB535B0612829826" box="[1244,1423,1820,1845]" pageId="21" pageNumber="282" refId="ref17192" refString="Abramov, A. A. (1992) Species composition and distribution of Epigonus (Epigonidae) in the world ocean. Journal of Ichthyology, 32, 94 - 108." type="journal article">Abramov 1992</bibRefCitation>
);
<collectingCountry id="F31C4EBE6E78407CFF185B2517F3984A" box="[151,254,1855,1880]" name="Australia" pageId="21" pageNumber="282">Australia</collectingCountry>
(
<bibRefCitation id="EF9A73DF6E78407CFE815B5A16D2984A" box="[270,479,1855,1880]" pageId="21" pageNumber="282" refId="ref19284" refString="Hoese, D. F., Bray, D. J., Allen, G. R. &amp; Cross, N. J. (2006) Epigonidae. Deepsea cardinalfishes, deepwater cardinalfishes. In: Hoese, D. F., Bray, D. J., Paxton, J. R. &amp; Allen, G. R. (Eds.), Zoological Catalogue of Australia. Vol. 35. Fishes. Parts 1 - 3. CSIRO Publishing and the Australian Biological Resources Study, Collingwood, pp. 1113 - 1115." type="book chapter">
Hoese
<emphasis id="B97FD23C6E78407CFED05B5B169F984A" box="[351,402,1856,1880]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="EF9A73DF6E78407CFE625B5A15C6984A" box="[493,715,1856,1880]" pageId="21" pageNumber="282" refId="ref18553" refString="Gomon, M. F., Bray, D. J. &amp; Kuiter, R. H. (Eds.) (2008) Fishes of Australian's Southern Coast. New Holland Publishers, Sydney, 928 pp." type="book">
Gomon
<emphasis id="B97FD23C6E78407CFDC55B5B1573984A" box="[586,638,1856,1880]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
. 2008
</bibRefCitation>
);
<collectingCountry id="F31C4EBE6E78407CFD6F5B5B1474984A" box="[736,889,1855,1880]" name="New Zealand" pageId="21" pageNumber="282">New Zealand</collectingCountry>
(Paulin
<emphasis id="B97FD23C6E78407CFC535B5B131D984A" box="[988,1040,1856,1880]" italics="true" pageId="21" pageNumber="282">et al</emphasis>
. 1989; Stewart &amp; Gon 2015); and western
<collectingRegion id="49CFC0CC6E78407CFF7A5B7E163B986F" box="[245,310,1892,1917]" country="Tanzania" name="Kusini Unguja" pageId="21" pageNumber="282">South</collectingRegion>
Atlantic (
<bibRefCitation id="EF9A73DF6E78407CFE295B7E1552986F" box="[422,607,1892,1917]" pageId="21" pageNumber="282" refId="ref20875" refString="Nakamura, I. (1986) Apogonidae. In: Nakamura, I. (Ed.), Important Fishes Trawled off Patagonia. Japan Marine Fisheries Resources Research Center, Tokyo, pp. 200 - 203." type="book chapter">Nakamura 1986</bibRefCitation>
;
<bibRefCitation id="EF9A73DF6E78407CFDE55B7E141A986E" box="[618,791,1892,1917]" pageId="21" pageNumber="282" refId="ref17192" refString="Abramov, A. A. (1992) Species composition and distribution of Epigonus (Epigonidae) in the world ocean. Journal of Ichthyology, 32, 94 - 108." type="journal article">Abramov 1992</bibRefCitation>
), at
<quantity id="4CF3A3CB6E78407CFCC95B7E14DE986E" box="[838,979,1892,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.7" metricValueMin="0.5" pageId="21" pageNumber="282" unit="m" value="1100.0" valueMax="1700.0" valueMin="500.0">5001700 m</quantity>
depth.
</paragraph>
<paragraph id="8BB40E2E6E78407FFF485B9D158A9EC2" blockId="21.[151,1436,1135,2025]" lastBlockId="22.[151,1437,151,464]" lastPageId="22" lastPageNumber="283" pageId="21" pageNumber="282">
<emphasis id="B97FD23C6E78407CFF485B9D151998B2" bold="true" box="[199,532,1927,1952]" pageId="21" pageNumber="282">Comparisons and Remarks.</emphasis>
<taxonomicName id="4C0B75AD6E78407CFD955B9215E798B2" authorityName=": Mayer" authorityYear="1974" box="[538,746,1928,1952]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B97FD23C6E78407CFD955B9215E798B2" box="[538,746,1928,1952]" italics="true" pageId="21" pageNumber="282">Epigonus robustus</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C0B75AD6E78407CFC495B931366988D" baseAuthorityName="Giglioli" baseAuthorityYear="1880" box="[966,1131,1928,1952]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="constanciae">
<emphasis id="B97FD23C6E78407CFC495B931366988D" box="[966,1131,1928,1952]" italics="true" pageId="21" pageNumber="282">E. constanciae</emphasis>
</taxonomicName>
group by having a pungent opercular spine (
<figureCitation id="133012AB6E78407CFEDE5BB616B498D6" box="[337,441,1964,1989]" captionStart="FIGURE17" captionStartId="22.[151,250,1718,1740]" captionTargetBox="[188,1398,558,1667]" captionTargetId="figure@22.[188,1398,529,1667]" captionTargetPageId="22" captionText="FIGURE17.Epigonus robustus, SAIAB 38259, 1of 2 specimens,155.0 mm SL.A:lateral view;B: ventral view of symphysis of lower jaw (arrows show the nub-like structures); C: SAM 13080, holotype, standard length unknown." httpUri="https://zenodo.org/record/1182203/files/figure.png" pageId="21" pageNumber="282">Fig. 17A</figureCitation>
). The species is similar to
<taxonomicName id="4C0B75AD6E78407CFD6E5BB7145598D6" authorityName="Scott" authorityYear="1962" box="[737,856,1964,1988]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="lenimen">
<emphasis id="B97FD23C6E78407CFD6E5BB7145598D6" box="[737,856,1964,1988]" italics="true" pageId="21" pageNumber="282">E. lenimen</emphasis>
</taxonomicName>
in having 11 + 14 vertebra; however, it differs from
<taxonomicName id="4C0B75AD6E78407CFF185BCB161F98FA" authorityName="Scott" authorityYear="1962" box="[151,274,2000,2024]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="lenimen">
<emphasis id="B97FD23C6E78407CFF185BCB161F98FA" box="[151,274,2000,2024]" italics="true" pageId="21" pageNumber="282">E. lenimen</emphasis>
</taxonomicName>
by having two nub-like structures on the symphysis of lower jaw (vs. absent in
<taxonomicName id="4C0B75AD6E78407CFB355BCB123898FA" authorityName="Scott" authorityYear="1962" box="[1210,1333,2000,2024]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="21" pageNumber="282" phylum="Chordata" rank="species" species="lenimen">
<emphasis id="B97FD23C6E78407CFB355BCB123898FA" box="[1210,1333,2000,2024]" italics="true" pageId="21" pageNumber="282">E. lenimen</emphasis>
</taxonomicName>
) and the orbital diameter shorter than postorbital length (vs. orbital diameter longer than postorbital length in
<taxonomicName id="4C0B75AD6E7B407FFA9D5C8312819FA2" authorityName="Scott" authorityYear="1962" box="[1298,1420,152,176]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="lenimen">
<emphasis id="B97FD23C6E7B407FFA9D5C8312819FA2" box="[1298,1420,152,176]" italics="true" pageId="22" pageNumber="283">E. lenimen</emphasis>
</taxonomicName>
). The other five species of the group in the
<collectingRegion id="49CFC0CC6E7B407FFD085CA715EF9FC6" box="[647,738,189,212]" country="Papua New Guinea" name="Western" pageId="22" pageNumber="283">Western</collectingRegion>
Indian Ocean,
<taxonomicName id="4C0B75AD6E7B407FFC185CA7133D9FC6" authorityName="Okamoto &amp; Gon" authorityYear="2018" box="[919,1072,188,212]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="bispinosus">
<emphasis id="B97FD23C6E7B407FFC185CA7133D9FC6" box="[919,1072,188,212]" italics="true" pageId="22" pageNumber="283">E. bispinosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B75AD6E7B407FFBCE5CA7139D9FC6" authorityName="Okamoto &amp; Gon" authorityYear="2018" box="[1089,1168,188,212]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="idai">
<emphasis id="B97FD23C6E7B407FFBCE5CA7139D9FC6" box="[1089,1168,188,212]" italics="true" pageId="22" pageNumber="283">E. idai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B75AD6E7B407FFB2F5CA7127E9FC6" authorityName="Parin &amp; Abramov" authorityYear="1986" box="[1184,1395,188,212]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="marimonticolus">
<emphasis id="B97FD23C6E7B407FFB2F5CA7127E9FC6" box="[1184,1395,188,212]" italics="true" pageId="22" pageNumber="283">E. marimonticolus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B75AD6E7B407FFA0B5CA7160C9FEA" authorityName="Mayer" authorityYear="1974" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="pectinifer">
<emphasis id="B97FD23C6E7B407FFA0B5CA7160C9FEA" italics="true" pageId="22" pageNumber="283">E. pectinifer</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C0B75AD6E7B407FFEB15CFB16EA9FEA" authorityName="Parin &amp; Abramov" authorityYear="1986" box="[318,487,224,248]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="waltersensis">
<emphasis id="B97FD23C6E7B407FFEB15CFB16EA9FEA" box="[318,487,224,248]" italics="true" pageId="22" pageNumber="283">E. waltersensis</emphasis>
</taxonomicName>
, have 10 + 15 vertebrae (vs. 11 + 14 vertebrae in
<taxonomicName id="4C0B75AD6E7B407FFB9D5CFB139D9FEA" authorityName=": Mayer" authorityYear="1974" box="[1042,1168,224,248]" class="Actinopterygii" family="Epigonidae" genus="Epigonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="22" pageNumber="283" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B97FD23C6E7B407FFB9D5CFB139D9FEA" box="[1042,1168,224,248]" italics="true" pageId="22" pageNumber="283">E. robustus</emphasis>
</taxonomicName>
). Several years ago, the second author examined the
<typeStatus id="54B0B08C6E7B407FFE6E5D1E154B9E0F" box="[481,582,260,285]" pageId="22" pageNumber="283" type="holotype">holotype</typeStatus>
, or rather what of left of it (
<figureCitation id="133012AB6E7B407FFC1F5D1E14F49E0E" box="[912,1017,260,285]" captionStart="FIGURE17" captionStartId="22.[151,250,1718,1740]" captionTargetBox="[188,1398,558,1667]" captionTargetId="figure@22.[188,1398,529,1667]" captionTargetPageId="22" captionText="FIGURE17.Epigonus robustus, SAIAB 38259, 1of 2 specimens,155.0 mm SL.A:lateral view;B: ventral view of symphysis of lower jaw (arrows show the nub-like structures); C: SAM 13080, holotype, standard length unknown." httpUri="https://zenodo.org/record/1182203/files/figure.png" pageId="22" pageNumber="283">Fig. 17C</figureCitation>
). The specimen was collected on
<date id="FFB528EE6E7B407FFA0F5D1E162B9E52" pageId="22" pageNumber="283" value="1903-08-20">20 August 1903</date>
during marine surveys conducted in the period
<date id="FFB528EE6E7B407FFCB55D3214B59E52" box="[826,952,296,320]" pageId="22" pageNumber="283" value="1897" valueMax="1904">18971904</date>
by the Pieter Faure, the Government steam trawler, under the leadership of J.D.F. Gilchrist who was then Director of the Governments Marine Biological Survey as well as Honorary Keeper of the Department of Marine Invertebrates of the
<collectingRegion id="49CFC0CC6E7B407FFBC05D75139D9E9A" box="[1103,1168,367,392]" country="Tanzania" name="Kusini Unguja" pageId="22" pageNumber="283">South</collectingRegion>
African Museum,
<collectingRegion id="49CFC0CC6E7B407FFAED5D6A12919E9A" box="[1378,1436,368,392]" country="South Africa" name="Western Cape" pageId="22" pageNumber="283">Cape</collectingRegion>
Town (
<bibRefCitation id="EF9A73DF6E7B407FFF6A5D8E16DE9EBE" box="[229,467,404,429]" pageId="22" pageNumber="283" refId="ref18961" refString="Gon, O. &amp; Skelton, P. H. (1997) A history of the fish collections of South Africa. In: Pietsch, T. W. &amp; Anderson, W. D. Jr. (Eds.), Collection Building in Ichthyology and Herpetology. American Society of Ichthyologists and Herpetologists, Special Publication No. 3, pp. 133 - 168." type="book chapter">Gon &amp; Skelton 1997</bibRefCitation>
). In addition to the parts of the fish shown in
<figureCitation id="133012AB6E7B407FFC5E5D8E133B9EBE" box="[977,1078,404,429]" captionStart="FIGURE17" captionStartId="22.[151,250,1718,1740]" captionTargetBox="[188,1398,558,1667]" captionTargetId="figure@22.[188,1398,529,1667]" captionTargetPageId="22" captionText="FIGURE17.Epigonus robustus, SAIAB 38259, 1of 2 specimens,155.0 mm SL.A:lateral view;B: ventral view of symphysis of lower jaw (arrows show the nub-like structures); C: SAM 13080, holotype, standard length unknown." httpUri="https://zenodo.org/record/1182203/files/figure.png" pageId="22" pageNumber="283">Fig. 17C</figureCitation>
, the jar of SAM 13080 contains scattered bones and scales of the specimens.
</paragraph>
</subSubSection>
</treatment>
</document>