510 lines
66 KiB
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510 lines
66 KiB
XML
<document id="0F1F6BCDB39FFB02362CD869D1C222C5" ID="10.11646/zootaxa.4277.4.1SLASH11426" ID-CLB-Dataset="32677" ID-DOI="10.11646/zootaxa.4277.4.1" ID-GBIF-Dataset="bc0690ff-ca74-4ba8-bc70-7376f04cf203" ID-ISSN="1175-5326" ID-Zenodo-Dep="829833" ID-ZooBank="1A18DA8-2AFF-4998-AE87-00D713EC531D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497944538512" checkinUser="plazi" docAuthor="Garg, Sonali & Biju, S. D." docDate="2017" docId="039287B73132FF81FF5DB255D1E8FD07" docLanguage="en" docName="zootaxa.4277.4.1SLASH11426.pdf" docOrigin="Zootaxa 4277 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fejervarya kadar Garg & Biju 2017, sp. nov." docType="treatment" docUuid="A9F6F16C-839B-4B13-95CF-A5DCA9165D68" docUuidSource="ZooBank" docVersion="8" lastPageNumber="478" masterDocId="FFABFFCF312AFF9AFFCAB132D536FFAF" masterDocTitle="Description of four new species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on morphological affinities of Fejervarya species in the Western Ghats" masterLastPageNumber="490" masterPageNumber="451" pageNumber="475" updateTime="1698447367985" updateUser="ExternalLinkService">
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<mods:title id="44A9989BC9DAFE96059E39861C91B77D">Description of four new species of Burrowing Frogs in the Fejervarya rufescens complex (Dicroglossidae) with notes on morphological affinities of Fejervarya species in the Western Ghats</mods:title>
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<mods:namePart id="841CF96937CE79AB92394ABDEE4C9343">Garg, Sonali</mods:namePart>
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<mods:namePart id="838D75DB673B94859D74D4AB8AE7259F">Biju, S. D.</mods:namePart>
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<mods:title id="40A22F66D630C614098B0C6106DADD95">Zootaxa</mods:title>
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<mods:date id="816FAD9FE89794C2D3576F0ABCCFD778">2017</mods:date>
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<treatment id="039287B73132FF81FF5DB255D1E8FD07" ID-DOI="http://doi.org/10.5281/zenodo.6052416" ID-GBIF-Taxon="132549775" ID-Zenodo-Dep="6052416" LSID="urn:lsid:zoobank.org:act:A9F6F16C-839B-4B13-95CF-A5DCA9165D68" httpUri="http://treatment.plazi.org/id/039287B73132FF81FF5DB255D1E8FD07" lastPageId="27" lastPageNumber="478" pageId="24" pageNumber="475">
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<subSubSection id="C321652A3132FF82FF5DB255D4A1FC68" pageId="24" pageNumber="475" type="nomenclature">
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<paragraph id="8B8436A13132FF82FF5DB255D4FBFC2D" blockId="24.[151,1189,871,1003]" box="[151,461,871,898]" pageId="24" pageNumber="475">
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<heading id="D0CC81CD3132FF82FF5DB255D4FBFC2D" bold="true" box="[151,461,871,898]" fontSize="11" level="1" pageId="24" pageNumber="475" reason="1">
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<emphasis id="B94FEAB33132FF82FF5DB255D4FBFC2D" bold="true" box="[151,461,871,898]" pageId="24" pageNumber="475">
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<taxonomicName id="4C3B4D223132FF82FF5DB255D45EFC2E" ID-CoL="6HR3H" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[151,360,871,897]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
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<emphasis id="B94FEAB33132FF82FF5DB255D45EFC2E" bold="true" box="[151,360,871,897]" italics="true" pageId="24" pageNumber="475">Fejervarya kadar</emphasis>
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<taxonomicNameLabel id="A27C57C83132FF82FEA5B25AD4FBFC2D" box="[367,461,872,898]" pageId="24" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8436A13132FF82FF5DB2B8D4A1FC68" blockId="24.[151,1189,871,1003]" pageId="24" pageNumber="475">
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http://zoobank.org/urn:lsid:zoobank.org:act:A9F6F16C-839B-4B13-95CF-A5DCA9165D68
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<taxonomicName id="4C3B4D223132FF82FF5DB29CD5EDFC68" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[151,219,942,967]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="485" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">Kadar</taxonomicName>
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Burrowing Frog
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</paragraph>
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</subSubSection>
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<subSubSection id="C321652A3132FF82FF5DB2E0D4F3FC44" box="[151,453,978,1003]" pageId="24" pageNumber="475" type="description">
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<paragraph id="8B8436A13132FF82FF5DB2E0D4F3FC44" blockId="24.[151,1189,871,1003]" box="[151,453,978,1003]" pageId="24" pageNumber="475">
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(
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<tableCitation id="C6B9031A3132FF82FF54B2E0D42BFC44" box="[158,285,978,1003]" captionStart-0="TABLE 1" captionStart-1="TABLE 4" captionStart-2="TABLE 5" captionStart-3="TABLE 6" captionStart-4="TABLE 7" captionStartId-0="11.[125,217,515,539]" captionStartId-1="14.[151,239,940,963]" captionStartId-2="15.[151,239,581,604]" captionStartId-3="15.[151,239,1081,1104]" captionStartId-4="18.[119,204,562,584]" captionText-0="TABLE 1. List of DNA sequences used in the study. Taxa are arranged in the order of their phylogenetic position in Figure 1 (Top to bottom)." captionText-1="TABLE 4. Factor loadings and percent variance explained by Principal Component (PC) 1 – 5 for 17 size-corrected morphometric variables from males of five species in the Fejervarya rufescens group. Values in bold indicate variables with the highest loadings." captionText-2="TABLE 5. Scores for discriminant function analysis of the principal components (PC 1 – 5). Values in bold indicate variables with the highest loadings for discriminant function roots of eigenvalue> 1.0." captionText-3="TABLE 6. Classification matrix for adult male specimens of the Fejervarya rufescens group from the discriminant function analysis." captionText-4="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="24" pageNumber="475">Tables 1–7</tableCitation>
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;
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<figureCitation id="13002A243132FF82FEE3B2E0D4A5FC44" box="[297,403,978,1003]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionStartId-2="6.[151,250,1765,1787]" captionStartId-3="8.[151,250,1820,1842]" captionStartId-4="9.[151,250,1518,1540]" captionStartId-5="10.[151,250,1726,1748]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetBox-2="[151,1436,197,1739]" captionTargetBox-3="[151,1436,180,1813]" captionTargetBox-4="[151,1436,193,1495]" captionTargetBox-5="[158,1434,882,1697]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetId-2="figure@6.[151,1436,193,1744]" captionTargetId-3="figure@8.[151,1436,180,1818]" captionTargetId-4="figure@9.[151,1436,193,1497]" captionTargetId-5="figure@10.[151,1437,879,1704]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." captionText-2="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." captionText-3="FIGURE 4. Four morphologically recognized Fejervarya groups in the Western Ghats. A – B. Fejervarya nilagirica group: A. Fejervarya nilagirica in life (male, SDBDU 2014.2772); B. Fejervarya keralensis in life (male, SDBDU 2015.3139); C. Fejervarya rufescens group, Fejervarya rufescens in life (male, SDBDU 2015.3070); D. Fejervarya sahyadris group, Fejervarya sahyadris in life (male, SDBDU 2015.3046); E – F. Fejervarya syhadrensis group: E. Fejervarya syhadrensis in life (male, SDBDU 2015.2934); F. Fejervarya granosa in life (male, SDBDU 2014.2541). Numbers on the panels represents: 1. dorsal view, 2. ventral view of foot showing the inner metatarsal tubercle, 3. schematic illustration of foot webbing, 4. thighs showing the marking and colour pattern, 5. lateral view showing the groin markings." captionText-4="FIGURE 5. Morphological comparison between members of the genus Sphaerotheca and Fejervarya rufescens group of the Western Ghats. A – H. Genus Sphaerotheca: A – E. Sphaerotheca cf. breviceps, in life (SDBDU 2014.2822): A. dorsolateral view, B. ventral view, C. ventral skin texture, D. lateral view showing the groin markings, E. thighs showing the marking and colour pattern; F – H. Inner metatarsal tubercle: F. Sphaerotheca breviceps (syntype ZMB 3351), G. S. dobsonii (Holotype BMNH 1947.2.28.45), H. S. rolandae (Holotype BMNH 1973.3024). I – R. Fejervarya rufescens group: I – K. F. rufescens in life (SDBDU 2015.3070): I. dorsolateral view, J. ventral view, K. ventral skin texture; L – M. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940): L. lateral view showing the groin markings, M. thighs showing the marking and colour pattern; N – R. Inner metatarsal tubercle: N. F. cepfi sp. nov. (Holotype ZSI / WGRC / V / A / 937), O. F. kadar sp. nov. (Holotype ZSI / WGRC / V / A / 940), P. F. manoharani sp. nov. (Holotype ZSI / WGRC / V / A / 945), Q. F. neilcoxi sp. nov. (Holotype ZSI / WGRC / V / A / 951), R. F. rufescens (SDBDU 2015.3070)." captionText-5="FIGURE 6. PCA plot for the first two principal component axes showing morphometric differentiation among adult males of five species in the Fejervarya rufescens groups of the Western Ghats. Factor loadings for each component are provided in Table 4." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" httpUri-2="https://zenodo.org/record/829839/files/figure.png" httpUri-3="https://zenodo.org/record/829841/files/figure.png" httpUri-4="https://zenodo.org/record/829843/files/figure.png" httpUri-5="https://zenodo.org/record/829845/files/figure.png" pageId="24" pageNumber="475">Figs. 1–6</figureCitation>
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,
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<figureCitation id="13002A243132FF82FE55B2E1D48AFC44" box="[415,444,979,1003]" captionStart="FIGURE 10" captionStartId="25.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1924]" captionTargetId="figure@25.[151,1437,193,1924]" captionTargetPageId="25" captionText="FIGURE 10. Holotype of Fejervarya kadar sp. nov. (ZSI / WGRC / V / A / 940): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829853/files/figure.png" pageId="24" pageNumber="475">10</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C321652A3132FF82FF5DB528D159FBD5" pageId="24" pageNumber="475" type="etymology">
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<paragraph id="8B8436A13132FF82FF5DB528D159FBD5" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
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<emphasis id="B94FEAB33132FF82FF5DB528D428FB9C" bold="true" box="[151,286,1050,1075]" pageId="24" pageNumber="475">Etymology.</emphasis>
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The species is named after the
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<taxonomicName id="4C3B4D223132FF82FD49B528D7F1FB9C" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[643,711,1050,1075]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="485" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">Kadar</taxonomicName>
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tribe of
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<collectingRegion id="49FFF8433132FF82FCE3B528D640FB9C" box="[809,886,1050,1075]" country="India" name="Kerala" pageId="24" pageNumber="475">Kerala</collectingRegion>
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, who live in the Vazhachal forest where the
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<typeStatus id="548088033132FF82FAA6B529D0AAFB9C" box="[1388,1436,1051,1075]" pageId="24" pageNumber="475">type</typeStatus>
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series was collected. We enjoyed their support and hospitality during amphibian field studies in the region. The specific epithet
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<taxonomicName id="4C3B4D223132FF82FE82B550D4BFFBD4" authorityName="Garg & Biju" authorityYear="2017" box="[328,393,1122,1147]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar">
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<emphasis id="B94FEAB33132FF82FE82B550D4BFFBD4" box="[328,393,1122,1147]" italics="true" pageId="24" pageNumber="475">kadar</emphasis>
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</taxonomicName>
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is treated as an invariable noun in apposition to the generic name.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321652A3132FF82FF0DB5B4D722FA80" pageId="24" pageNumber="475" type="materials_examined">
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<paragraph id="8B8436A13132FF82FF0DB5B4D5E0FB48" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
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<materialsCitation id="3B533CFC3132FF82FF0DB5B4D5E0FB48" ID-GBIF-Occurrence="1572402623" collectingDate="2015-09-12" collectionCode="WGRC" country="India" county="Thrissur district" elevation="577" latitude="10.283806" location="Thavalakuzhipara" longLatPrecision="1" longitude="76.68511" municipality="Vazhachal forest" pageId="24" pageNumber="475" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
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<typeStatus id="548088033132FF82FF0DB5B4D400FB30" box="[199,310,1158,1183]" pageId="24" pageNumber="475" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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ZSI/
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<collectionCode id="ED2AAE643132FF82FEBCB5B5D4FAFB30" box="[374,460,1159,1183]" pageId="24" pageNumber="475">WGRC</collectionCode>
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/V/A/940, an adult male, from
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, (
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,
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asl
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</elevation>
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),
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<collectingMunicipality id="6BE0ACDB3132FF82FF21B598D490FB6C" box="[235,422,1194,1219]" pageId="24" pageNumber="475">Vazhachal forest</collectingMunicipality>
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division,
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<collectingCounty id="62E54E2D3132FF82FDDCB598D7FAFB6C" box="[534,716,1194,1219]" pageId="24" pageNumber="475">Thrissur district</collectingCounty>
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,
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<collectingRegion id="49FFF8433132FF82FD1DB598D614FB6C" box="[727,802,1194,1219]" country="India" name="Kerala" pageId="24" pageNumber="475">Kerala</collectingRegion>
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state,
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<collectingCountry id="F32C76313132FF82FCA1B598D691FB6C" box="[875,935,1194,1219]" name="India" pageId="24" pageNumber="475">India</collectingCountry>
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, collected by SDB and SG on
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<collectingDate id="EFC1E9893132FF82FACBB599D5E5FB48" pageId="24" pageNumber="475" value="2015-09-12">
|
||
<date id="FF8510613132FF82FACBB599D5E5FB48" pageId="24" pageNumber="475" value="2015-09-12">12 September 2015</date>
|
||
</collectingDate>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B8436A13132FF82FF0DB5C0D722FA80" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB5C0D470FAA4" bold="true" box="[199,326,1266,1291]" pageId="24" pageNumber="475">Paratypes.</emphasis>
|
||
ZSI/WGRC/V/A/941–942, two adult males and ZSI/WGRC/V/A/943–944, two adult females, collected along with the holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A3132FF82FF0DB408D1F9F9A8" pageId="24" pageNumber="475" type="description">
|
||
<paragraph id="8B8436A13132FF82FF0DB408D1F9F9A8" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB408D489FAFC" bold="true" box="[199,447,1338,1363]" pageId="24" pageNumber="475">Genetic relationship.</emphasis>
|
||
Phylogenetically,
|
||
<taxonomicName id="4C3B4D223132FF82FD44B40ED664FAFC" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[654,850,1338,1363]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FD44B40ED664FAFC" box="[654,850,1338,1363]" italics="true" pageId="24" pageNumber="475">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FC93B408D685FAFC" box="[857,947,1338,1363]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FC93B408D685FAFC" bold="true" box="[857,947,1338,1363]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is nested in the
|
||
<taxonomicName id="4C3B4D223132FF82FBADB40ED062FAFC" authorityName="Jerdon" authorityYear="1853" box="[1127,1364,1339,1363]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB33132FF82FBADB40ED062FAFC" box="[1127,1364,1339,1363]" italics="true" pageId="24" pageNumber="475">Fejervarya rufescens</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<figureCitation id="13002A243132FF82FF55B46CD43EFAD8" box="[159,264,1374,1399]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1885,1907]" captionStartId-1="5.[151,250,1618,1640]" captionTargetBox-0="[161,1435,204,1854]" captionTargetBox-1="[155,1424,193,1582]" captionTargetId-0="figure@4.[151,1436,193,1864]" captionTargetId-1="figure@5.[151,1436,193,1597]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Maximum Likelihood phylogram based on the 16 S mitochondrial DNA dataset of 545 basepairs. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. The uncorrected pairwise distances between species in the Fejervarya rufescens group are shown near the clade labels. Voucher number and locality details of the samples are referenced in Table 1." captionText-1="FIGURE 2. Phylogenetic analyses. A. Maximum Likelihood phylogram based on mitochondrial (16 S, COI and Cytb) and nuclear (Rag 1) DNA of 2314 basepairs, showing phylogenetic relationships among members of the Fejervarya rufescens group. Bayesian Posterior Probabilities (BPP) and RaxML bootstrap values of> 50 % are indicated above and below the branches, respectively. Fejervarya granosa (2) was used as the outgroup taxon. Voucher number and locality details of the samples are referenced in Table 1; B. Median Joining haplotype network inferred from 554 basepair fragment of the nuclear Rag 1 gene. Circle sizes are proportional to the number of haplotypes, branch lengths are proportional to the number of mutational steps, and black bars represent mutational steps. Number of individuals used for each species: Fejervarya cepfi (3), Fejervarya kadar (1), Fejervarya manoharani (3), Fejervarya neilcoxi (2), and Fejervarya rufescens (5)." httpUri-0="https://zenodo.org/record/829835/files/figure.png" httpUri-1="https://zenodo.org/record/829837/files/figure.png" pageId="24" pageNumber="475">Figs. 1–2</figureCitation>
|
||
) of the
|
||
<collectingRegion id="49FFF8433132FF82FE95B46DD48CFAD9" box="[351,442,1375,1398]" country="Sri Lanka" name="Western" pageId="24" pageNumber="475">Western</collectingRegion>
|
||
Ghats. The average uncorrected pairwise genetic divergences with other members of the group are: 3.2% (range 2.8–3.9%,
|
||
<emphasis id="B94FEAB33132FF82FDFAB4B6D775FA34" box="[560,579,1412,1435]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 7) for 16S, 9.5% (range 9.3–9.7%,
|
||
<emphasis id="B94FEAB33132FF82FBC4B4B6D117FA34" box="[1038,1057,1412,1435]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 5) for COI, and 8.2% (range 8.0–8.4%,
|
||
<emphasis id="B94FEAB33132FF82FEDBB495D412FA11" box="[273,292,1447,1470]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 5) for Cytb with
|
||
<taxonomicName id="4C3B4D223132FF82FDCAB495D7B0FA10" authorityName="Jerdon" authorityYear="1853" box="[512,646,1447,1471]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB33132FF82FDCAB495D7B0FA10" box="[512,646,1447,1471]" italics="true" pageId="24" pageNumber="475">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
; and 6.2% (6.2–6.3%,
|
||
<emphasis id="B94FEAB33132FF82FC41B495D6A8FA11" box="[907,926,1447,1470]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 4) for 16S, 12.1% (range 12.0–12.3%,
|
||
<emphasis id="B94FEAB33132FF82FAB8B495D0B3FA11" box="[1394,1413,1447,1470]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3) for COI, and 13.7% (range 13.6–13.7%,
|
||
<emphasis id="B94FEAB33132FF82FDB7B4FED7A6FA4C" box="[637,656,1484,1507]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3) for Cytb with
|
||
<taxonomicName id="4C3B4D223132FF82FCA0B4FED68AFA4C" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[874,956,1483,1507]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FCA0B4FED68AFA4C" box="[874,956,1483,1507]" italics="true" pageId="24" pageNumber="475">F. cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FC09B4F8D12BFA4C" box="[963,1053,1482,1507]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FC09B4F8D12BFA4C" bold="true" box="[963,1053,1482,1507]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Table 2). For comparison with
|
||
<taxonomicName id="4C3B4D223132FF82FA4DB4FED416F9A8" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FA4DB4FED416F9A8" italics="true" pageId="24" pageNumber="475">F. manoharani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FEEDB4DCD4B6F9A8" box="[295,384,1518,1543]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FEEDB4DCD4B6F9A8" bold="true" box="[295,384,1518,1543]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3B4D223132FF82FE7CB4DDD711F9A8" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[438,551,1518,1543]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FE7CB4DDD711F9A8" box="[438,551,1518,1543]" italics="true" pageId="24" pageNumber="475">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FDF8B4DCD7BDF9A8" box="[562,651,1518,1543]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FDF8B4DCD7BDF9A8" bold="true" box="[562,651,1518,1543]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, see ‘Genetic relationship’ section of those species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A3132FF80FF0DB720D0ABFC02" lastPageId="26" lastPageNumber="477" pageId="24" pageNumber="475" type="diagnosis">
|
||
<paragraph id="8B8436A13132FF82FF0DB720D0A4F888" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB720D477F984" bold="true" box="[199,321,1554,1579]" pageId="24" pageNumber="475">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C3B4D223132FF82FE9BB726D729F984" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[337,543,1554,1579]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FE9BB726D729F984" box="[337,543,1554,1579]" italics="true" pageId="24" pageNumber="475">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FDFAB720D7A5F984" box="[560,659,1554,1579]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FDFAB720D7A5F984" bold="true" box="[560,659,1554,1579]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from known congeners by the following combination of morphological characters: (1) medium male adult size (SVL 31.0–
|
||
<quantity id="4CC39B443132FF82FBFBB704D1A0F9E1" box="[1073,1174,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" pageId="24" pageNumber="475" unit="mm" value="33.5">33.5 mm</quantity>
|
||
,
|
||
<emphasis id="B94FEAB33132FF82FB6BB705D182F9E1" box="[1185,1204,1591,1614]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3); (2) stout body; (3) snout rounded in dorsal view and obtuse in lateral view; (4) presence of rictal gland at labial commissure of the mouth; (5) eye length shorter than snout length (male EL/SL ratio 88.9–93.3%,
|
||
<emphasis id="B94FEAB33132FF82FBC5B74DD114F939" box="[1039,1058,1663,1686]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3); (6) tympanum diameter less than half of eye length, (male TYD/EL ratio 37.5–40.5%,
|
||
<emphasis id="B94FEAB33132FF82FCEEB796D601F914" box="[804,823,1700,1723]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3); (7) inter upper eyelid width narrower than upper eyelid width (male IUE/UEW ratio 53.1–60.0%,
|
||
<emphasis id="B94FEAB33132FF82FD76B7F5D7F9F971" box="[700,719,1735,1758]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3) and internarial distance (male IUE/IN ratio 63.0–69.2%,
|
||
<emphasis id="B94FEAB33132FF82FA43B7F5D0AAF971" box="[1417,1436,1735,1758]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3); (8) thigh length nearly equal to shank length (male TL/SHL ratio 99.3–99.4%,
|
||
<emphasis id="B94FEAB33132FF82FB85B7DED154F8AC" box="[1103,1122,1772,1795]" italics="true" pageId="24" pageNumber="475">N</emphasis>
|
||
= 3); (9) prominent shovelshaped inner metatarsal tubercle prominent and small outer metatarsal tubercle; (10) webbing between toes small.
|
||
</paragraph>
|
||
<paragraph id="8B8436A13132FF82FF0DB600D7BDF83C" blockId="24.[151,1437,1050,2011]" pageId="24" pageNumber="475">
|
||
<emphasis id="B94FEAB33132FF82FF0DB600D724F8E4" bold="true" box="[199,530,1842,1867]" pageId="24" pageNumber="475">Morphological comparison.</emphasis>
|
||
Based on the overall morphology and comparable body size,
|
||
<taxonomicName id="4C3B4D223132FF82FB1CB606D0AAF8E4" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[1238,1436,1842,1867]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FB1CB606D0AAF8E4" box="[1238,1436,1842,1867]" italics="true" pageId="24" pageNumber="475">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FF5DB664D5C0F8C0" box="[151,246,1878,1903]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FF5DB664D5C0F8C0" bold="true" box="[151,246,1878,1903]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
could be confused with the known species
|
||
<taxonomicName id="4C3B4D223132FF82FCC2B665D6A5F8C0" authorityName="Jerdon" authorityYear="1853" box="[776,915,1879,1903]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB33132FF82FCC2B665D6A5F8C0" box="[776,915,1879,1903]" italics="true" pageId="24" pageNumber="475">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
and three new species
|
||
<taxonomicName id="4C3B4D223132FF82FB7BB665D033F8C0" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[1201,1285,1879,1903]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="cepfi" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FB7BB665D033F8C0" box="[1201,1285,1879,1903]" italics="true" pageId="24" pageNumber="475">F. cepfi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FADFB664D042F8C0" box="[1301,1396,1878,1903]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FADFB664D042F8C0" bold="true" box="[1301,1396,1878,1903]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3B4D223132FF82FA4DB665D416F83C" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="manoharani" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FA4DB665D416F83C" italics="true" pageId="24" pageNumber="475">F. manoharani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FEEDB648D4B6F83C" box="[295,384,1914,1939]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FEEDB648D4B6F83C" bold="true" box="[295,384,1914,1939]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3B4D223132FF82FE7CB64ED71DF83C" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[438,555,1914,1939]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="neilcoxi" status="sp. nov.">
|
||
<emphasis id="B94FEAB33132FF82FE7CB64ED71DF83C" box="[438,555,1914,1939]" italics="true" pageId="24" pageNumber="475">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83132FF82FDF8B648D7BDF83C" box="[562,651,1914,1939]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="B94FEAB33132FF82FDF8B648D7BDF83C" bold="true" box="[562,651,1914,1939]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B8436A13132FF80FF0DB6ADD42DFE1C" blockId="24.[151,1437,1050,2011]" lastBlockId="26.[151,1437,151,2023]" lastPageId="26" lastPageNumber="477" pageId="24" pageNumber="475">
|
||
However,
|
||
<taxonomicName id="4C3B4D223132FF82FEF4B6ADD730F818" authorityName="Garg & Biju" authorityYear="2017" box="[318,518,1950,1975]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="kadar">
|
||
<emphasis id="B94FEAB33132FF82FEF4B6ADD730F818" box="[318,518,1950,1975]" italics="true" pageId="24" pageNumber="475">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B4D223132FF82FD6FB6ADD618F818" authorityName="Jerdon" authorityYear="1853" box="[677,814,1951,1975]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rufescens">
|
||
<emphasis id="B94FEAB33132FF82FD6FB6ADD618F818" box="[677,814,1951,1975]" italics="true" pageId="24" pageNumber="475">F. rufescens</emphasis>
|
||
</taxonomicName>
|
||
by its dorsal skin with scattered glandular warts (vs. shagreened to granular); snout obtuse in lateral view (vs. rounded) and rounded in dorsal view (vs. subovoid); tympanum to eye distance relatively longer or nearly equal to tympanum diameter, male TYE
|
||
<quantity id="4CC39B443130FF80FB01B1A5D060FF00" box="[1227,1366,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="26" pageNumber="477" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
|
||
, TYD
|
||
<quantity id="4CC39B443130FF80FF5DB189D416FF7C" box="[151,288,187,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="26" pageNumber="477" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
|
||
, TYE/TYD ratio 80.0–93.3%,
|
||
<emphasis id="B94FEAB33130FF80FDB1B18FD7B8FF7B" box="[635,654,189,212]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. relatively shorter, male TYE
|
||
<quantity id="4CC39B443130FF80FBF2B18ED1F7FF7C" box="[1080,1217,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="26" pageNumber="477" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.1–1.3 mm</quantity>
|
||
, TYD
|
||
<quantity id="4CC39B443130FF80FAC4B189D0AEFF7C" box="[1294,1432,187,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="26" pageNumber="477" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.6–2.2 mm</quantity>
|
||
, TYE/TYD ratio 50.0–76.5%,
|
||
<emphasis id="B94FEAB33130FF80FE2EB1D2D4C1FF58" box="[484,503,224,247]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 6); thigh length shorter than foot length, male TL
|
||
<quantity id="4CC39B443130FF80FB8AB1EDD1D3FF58" box="[1088,1253,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.54" metricValueMin="1.45" pageId="26" pageNumber="477" unit="mm" value="14.95" valueMax="15.4" valueMin="14.5">14.5–15.4 mm</quantity>
|
||
, FOL
|
||
<quantity id="4CC39B443130FF80FAE6B1EDD5F3FEB4" metricMagnitude="-2" metricUnit="m" metricValue="1.59" metricValueMax="1.61" metricValueMin="1.5699999999999998" pageId="26" pageNumber="477" unit="mm" value="15.9" valueMax="16.1" valueMin="15.7">15.7–16.1 mm</quantity>
|
||
, TL/FOL ratio 92.4–95.7%,
|
||
<emphasis id="B94FEAB33130FF80FDC5B036D714FEB4" box="[527,546,260,283]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. nearly equal, male TL
|
||
<quantity id="4CC39B443130FF80FC5FB031D10AFEB4" box="[917,1084,259,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.58" metricValueMin="1.47" pageId="26" pageNumber="477" unit="mm" value="15.25" valueMax="15.8" valueMin="14.7">14.7–15.8 mm</quantity>
|
||
, FOL
|
||
<quantity id="4CC39B443130FF80FB42B031D019FEB4" box="[1160,1327,259,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.535" metricValueMax="1.59" metricValueMin="1.48" pageId="26" pageNumber="477" unit="mm" value="15.35" valueMax="15.9" valueMin="14.8">14.8–15.9 mm</quantity>
|
||
, TL/FOL ratio 98.0–99.4%,
|
||
<emphasis id="B94FEAB33130FF80FEBFB019D4BEFEED" box="[373,392,299,322]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 6); and relatively more webbing between toes, male
|
||
<date id="FF8510613130FF80FBF0B018D167FEED" box="[1082,1105,298,322]" pageId="26" pageNumber="477">I2</date>
|
||
– –2–
|
||
<date id="FF8510613130FF80FBB6B018D1A8FEED" box="[1148,1182,298,322]" pageId="26" pageNumber="477">II2</date>
|
||
– –
|
||
<date id="FF8510613130FF80FB7EB018D1D3FEED" box="[1204,1253,294,322]" bridgedPair="-" pageId="26" pageNumber="477">3–III</date>
|
||
2–3 2/3
|
||
<date id="FF8510613130FF80FAEFB018D067FEED" box="[1317,1361,298,322]" pageId="26" pageNumber="477">IV3</date>
|
||
2/3–
|
||
<date id="FF8510613130FF80FABDB018D0AEFEED" box="[1399,1432,298,322]" pageId="26" pageNumber="477">2V</date>
|
||
, specifically the fourth toe webbing slightly above the third subarticular tubercle on either side (vs. less, male
|
||
<date id="FF8510613130FF80FF5DB045D598FE20" box="[151,174,375,399]" pageId="26" pageNumber="477">I2</date>
|
||
– –2
|
||
<date id="FF8510613130FF80FF18B045D5C4FE20" box="[210,242,375,399]" pageId="26" pageNumber="477">II2</date>
|
||
– –
|
||
<date id="FF8510613130FF80FEC3B045D40DFE20" box="[265,315,371,399]" bridgedPair="-" pageId="26" pageNumber="477">3–III</date>
|
||
2–
|
||
<date id="FF8510613130FF80FE9DB045D4B7FE20" box="[343,385,375,399]" pageId="26" pageNumber="477">3IV</date>
|
||
3–
|
||
<date id="FF8510613130FF80FE56B045D489FE20" box="[412,447,375,399]" pageId="26" pageNumber="477">2V</date>
|
||
, specifically the fourth toe webbing extending up to the second subarticular tubercle on either side).
|
||
</paragraph>
|
||
<caption id="DF4466293133FF83FF5DB6ABD6E6F843" httpUri="https://zenodo.org/record/829853/files/figure.png" pageId="25" pageNumber="476" targetBox="[151,1436,193,1924]" targetPageId="25">
|
||
<paragraph id="8B8436A13133FF83FF5DB6ABD6E6F843" blockId="25.[151,1436,1945,2028]" pageId="25" pageNumber="476">
|
||
<emphasis id="B94FEAB33133FF83FF5DB6ABD429F801" bold="true" box="[151,287,1945,1967]" pageId="25" pageNumber="476">FIGURE 10.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C3B4D223133FF83FE0AB6A8D744F800" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[448,626,1945,1967]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB33133FF83FE0AB6A8D744F800" box="[448,626,1945,1967]" italics="true" pageId="25" pageNumber="476">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83133FF83FDB2B6ABD7FEF801" box="[632,712,1945,1966]" pageId="25" pageNumber="476" rank="species">
|
||
<emphasis id="B94FEAB33133FF83FDB2B6ABD7FEF801" bold="true" box="[632,712,1945,1966]" pageId="25" pageNumber="476">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(ZSI/WGRC/V/A/940):
|
||
<emphasis id="B94FEAB33133FF83FC0DB6ABD6E9F801" bold="true" box="[967,991,1945,1966]" pageId="25" pageNumber="476">A.</emphasis>
|
||
dorsolateral view, in life;
|
||
<emphasis id="B94FEAB33133FF83FB20B6ABD037F801" bold="true" box="[1258,1281,1945,1966]" pageId="25" pageNumber="476">B.</emphasis>
|
||
dorsal view, in preservation;
|
||
<emphasis id="B94FEAB33133FF83FEEBB68AD40CF861" bold="true" box="[289,314,1976,1998]" pageId="25" pageNumber="476">C.</emphasis>
|
||
ventral view, in preservation;
|
||
<emphasis id="B94FEAB33133FF83FDA6B68AD7B3F862" bold="true" box="[620,645,1976,1997]" pageId="25" pageNumber="476">D.</emphasis>
|
||
lateral view of head, in preservation;
|
||
<emphasis id="B94FEAB33133FF83FBCEB68AD12DF862" bold="true" box="[1028,1051,1976,1997]" pageId="25" pageNumber="476">E.</emphasis>
|
||
ventral view of hand, in preservation;
|
||
<emphasis id="B94FEAB33133FF83FF5DB6E5D59EF843" bold="true" box="[151,168,2007,2028]" pageId="25" pageNumber="476">F.</emphasis>
|
||
ventral view of foot, in preservation;
|
||
<emphasis id="B94FEAB33133FF83FDE0B6E4D708F843" bold="true" box="[554,574,2006,2028]" pageId="25" pageNumber="476">G.</emphasis>
|
||
schematic illustration of foot webbing.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8436A13130FF80FF0DB08DD17EFC29" blockId="26.[151,1437,151,2023]" pageId="26" pageNumber="477">
|
||
<taxonomicName id="4C3B4D223130FF80FF0DB08DD4BBFE78" authorityName="Garg & Biju" authorityYear="2017" box="[199,397,446,471]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="26" pageNumber="477" phylum="Chordata" rank="species" species="kadar">
|
||
<emphasis id="B94FEAB33130FF80FF0DB08DD4BBFE78" box="[199,397,446,471]" italics="true" pageId="26" pageNumber="477">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B4D223130FF80FDECB08DD74CFE79" authorityName="Garg & Biju" authorityYear="2017" box="[550,634,446,470]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="26" pageNumber="477" phylum="Chordata" rank="species" species="cepfi">
|
||
<emphasis id="B94FEAB33130FF80FDECB08DD74CFE79" box="[550,634,446,470]" italics="true" pageId="26" pageNumber="477">F. cepfi</emphasis>
|
||
</taxonomicName>
|
||
by its dorsal skin with relatively more prominent glandular warts (vs. less prominent); snout rounded in dorsal view (vs. subovoid); inter upper eyelid width narrower than upper eyelid width, male IUE
|
||
<quantity id="4CC39B443130FF80FE90B334D4D2FDB2" box="[346,484,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="26" pageNumber="477" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
|
||
, UEW 3.0–
|
||
<quantity id="4CC39B443130FF80FDA0B334D7F4FDB2" box="[618,706,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="26" pageNumber="477" unit="mm" value="3.3">3.3 mm</quantity>
|
||
, IUE/UEW ratio 53.1–60.0%,
|
||
<emphasis id="B94FEAB33130FF80FBD7B334D106FDB2" box="[1053,1072,518,541]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. nearly equal, male IUE
|
||
<quantity id="4CC39B443130FF80FF5DB31BD412FDEE" box="[151,292,553,578]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="26" pageNumber="477" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
, UEW
|
||
<quantity id="4CC39B443130FF80FEB4B31BD73AFDEE" box="[382,524,553,578]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.6" metricValueMin="2.5" pageId="26" pageNumber="477" unit="mm" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 mm</quantity>
|
||
, IUE/UEW ratio 96.0–96.2%,
|
||
<emphasis id="B94FEAB33130FF80FCBEB318D6B1FDEE" box="[884,903,554,577]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 2); inter upper eyelid width narrower than internarial distance, male IUE
|
||
<quantity id="4CC39B443130FF80FE33B37CD7B3FDCA" box="[505,645,590,614]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="26" pageNumber="477" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
|
||
, IN
|
||
<quantity id="4CC39B443130FF80FD70B37FD671FDCA" box="[698,839,589,614]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="26" pageNumber="477" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 mm</quantity>
|
||
, IUE/IN ratio 63.0–69.2%,
|
||
<emphasis id="B94FEAB33130FF80FB4DB37CD1ACFDCA" box="[1159,1178,590,613]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. nearly equal, male IUE
|
||
<quantity id="4CC39B443130FF80FEC0B342D4A5FD27" box="[266,403,624,649]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="26" pageNumber="477" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
, IN
|
||
<quantity id="4CC39B443130FF80FE08B342D72CFD27" box="[450,538,624,649]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="26" pageNumber="477" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, IUE/IN ratio 96–100%,
|
||
<emphasis id="B94FEAB33130FF80FCFEB340D671FD26" box="[820,839,626,649]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 2); tympanum diameter less than half of horizontal diameter of eye, male TYD
|
||
<quantity id="4CC39B443130FF80FE1EB3A6D768FD03" box="[468,606,660,685]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="26" pageNumber="477" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
|
||
, EL 4.0–
|
||
<quantity id="4CC39B443130FF80FD0FB3A7D628FD03" box="[709,798,661,685]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="26" pageNumber="477" unit="mm" value="4.2">4.2 mm</quantity>
|
||
, TYD/EL ratio 37.5–40.5%,
|
||
<emphasis id="B94FEAB33130FF80FBAFB3A4D14EFD02" box="[1125,1144,662,685]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. nearly half, male TYD
|
||
<quantity id="4CC39B443130FF80FF10B38BD403FD7F" box="[218,309,697,721]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="26" pageNumber="477" unit="mm" value="1.8">1.8 mm</quantity>
|
||
, EL
|
||
<quantity id="4CC39B443130FF80FEBAB38AD4CAFD7F" box="[368,508,696,721]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="26" pageNumber="477" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
|
||
, TYD/EL ratio 51.4–52.9%,
|
||
<emphasis id="B94FEAB33130FF80FC84B38BD657FD7F" box="[846,865,697,720]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 2); horizontal diameter of eye relatively larger compared to snout length, male EL 4.0–
|
||
<quantity id="4CC39B443130FF80FDAEB3EED788FD5B" box="[612,702,732,756]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="26" pageNumber="477" unit="mm" value="4.2">4.2 mm</quantity>
|
||
, SL
|
||
<quantity id="4CC39B443130FF80FD3FB3EED6B6FD5B" box="[757,896,732,757]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="26" pageNumber="477" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
|
||
, EL/SL ratio 88.9–93.3%,
|
||
<emphasis id="B94FEAB33130FF80FB7EB3EFD1F1FD5B" box="[1204,1223,733,756]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. relatively smaller, male EL
|
||
<quantity id="4CC39B443130FF80FE96B3CDD4D2FCB8" box="[348,484,767,792]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="26" pageNumber="477" unit="mm" value="3.45" valueMax="3.5" valueMin="3.4">3.4–3.5 mm</quantity>
|
||
, SL
|
||
<quantity id="4CC39B443130FF80FDDDB3CDD796FCB8" box="[535,672,767,792]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="26" pageNumber="477" unit="mm" value="4.699999999999999" valueMax="4.8" valueMin="4.6">4.6–4.8 mm</quantity>
|
||
, EL/SL ratio 72.9–73.9%,
|
||
<emphasis id="B94FEAB33130FF80FC06B233D6E9FCB7" box="[972,991,769,792]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 2); thigh length nearly equal to shank length, male TL
|
||
<quantity id="4CC39B443130FF80FE94B211D731FC94" box="[350,519,803,828]" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.54" metricValueMin="1.45" pageId="26" pageNumber="477" unit="mm" value="14.95" valueMax="15.4" valueMin="14.5">14.5–15.4 mm</quantity>
|
||
, SHL
|
||
<quantity id="4CC39B443130FF80FD9DB211D637FC94" box="[599,769,803,828]" metricMagnitude="-2" metricUnit="m" metricValue="1.505" metricValueMax="1.55" metricValueMin="1.46" pageId="26" pageNumber="477" unit="mm" value="15.05" valueMax="15.5" valueMin="14.6">14.6–15.5 mm</quantity>
|
||
, TL/SHL ratio 99.3–99.4%,
|
||
<emphasis id="B94FEAB33130FF80FB99B217D150FC93" box="[1107,1126,805,828]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 3 (vs. shorter, male TL
|
||
<quantity id="4CC39B443130FF80FF5DB275D40AFCF0" box="[151,316,839,864]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" metricValueMax="1.54" metricValueMin="1.48" pageId="26" pageNumber="477" unit="mm" value="15.1" valueMax="15.4" valueMin="14.8">14.8–15.4 mm</quantity>
|
||
, SHL
|
||
<quantity id="4CC39B443130FF80FE4CB275D71DFCF0" box="[390,555,839,864]" metricMagnitude="-2" metricUnit="m" metricValue="1.6049999999999998" metricValueMax="1.64" metricValueMin="1.5699999999999998" pageId="26" pageNumber="477" unit="mm" value="16.049999999999997" valueMax="16.4" valueMin="15.7">15.7–16.4 mm</quantity>
|
||
, TL/SHL ratio 93.9–94.3%,
|
||
<emphasis id="B94FEAB33130FF80FCA5B27AD6B4FCF0" box="[879,898,840,863]" italics="true" pageId="26" pageNumber="477">N</emphasis>
|
||
= 2); and relatively less webbing between toes, male
|
||
<date id="FF8510613130FF80FF1EB25CD5DAFC29" box="[212,236,878,902]" pageId="26" pageNumber="477">I2</date>
|
||
– –2–
|
||
<date id="FF8510613130FF80FEDDB25CD40EFC29" box="[279,312,878,902]" pageId="26" pageNumber="477">II2</date>
|
||
– –
|
||
<date id="FF8510613130FF80FE85B25CD4B6FC29" box="[335,384,874,902]" bridgedPair="-" pageId="26" pageNumber="477">3–III</date>
|
||
2–3 2/3
|
||
<date id="FF8510613130FF80FE0AB25CD4DDFC29" box="[448,491,878,902]" pageId="26" pageNumber="477">IV3</date>
|
||
2/3–
|
||
<date id="FF8510613130FF80FDDBB25CD704FC29" box="[529,562,878,902]" pageId="26" pageNumber="477">2V</date>
|
||
(vs. more, male
|
||
<date id="FF8510613130FF80FD25B25CD630FC29" box="[751,774,878,902]" pageId="26" pageNumber="477">I1</date>
|
||
+–2–
|
||
<date id="FF8510613130FF80FCF9B25CD665FC29" box="[819,851,878,902]" pageId="26" pageNumber="477">II1</date>
|
||
+–
|
||
<date id="FF8510613130FF80FCA6B25CD6A8FC29" box="[876,926,874,902]" bridgedPair="-" pageId="26" pageNumber="477">3–III</date>
|
||
2–
|
||
<date id="FF8510613130FF80FC76B25CD6DEFC29" box="[956,1000,878,902]" pageId="26" pageNumber="477">3IV</date>
|
||
3–1 1/
|
||
<date id="FF8510613130FF80FBD6B24FD10AFC29" box="[1052,1084,878,907]" pageId="26" pageNumber="477">2V</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A13130FF80FF0DB2A7D0ABFC02" blockId="26.[151,1437,151,2023]" box="[199,1437,916,941]" pageId="26" pageNumber="477">
|
||
For differences with
|
||
<taxonomicName id="4C3B4D223130FF80FE7AB2A7D78AFC02" authorityName="Garg & Biju" authorityYear="2017" box="[432,700,916,941]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="26" pageNumber="477" phylum="Chordata" rank="species" species="manoharani">
|
||
<emphasis id="B94FEAB33130FF80FE7AB2A7D78AFC02" box="[432,700,916,941]" italics="true" pageId="26" pageNumber="477">Fejervarya manoharani</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D223130FF80FD38B2A7D650FC02" authorityName="Garg & Biju" authorityYear="2017" box="[754,870,916,941]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="26" pageNumber="477" phylum="Chordata" rank="species" species="neilcoxi">
|
||
<emphasis id="B94FEAB33130FF80FD38B2A7D650FC02" box="[754,870,916,941]" italics="true" pageId="26" pageNumber="477">F. neilcoxi</emphasis>
|
||
</taxonomicName>
|
||
, see ‘morphological comparison’ of those species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A3130FF81FF0DB28AD650FE03" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477" type="description">
|
||
<paragraph id="8B8436A13130FF80FF0DB28AD02FF9B9" blockId="26.[151,1437,151,2023]" pageId="26" pageNumber="477">
|
||
<emphasis id="B94FEAB33130FF80FF0DB28AD7CAFC7E" bold="true" box="[199,764,951,977]" pageId="26" pageNumber="477">
|
||
Description of
|
||
<typeStatus id="548088033130FF80FEB3B28AD4D6FC7E" box="[377,480,952,977]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
<emphasis id="B94FEAB33130FF80FE22B285D7C3FC7F" bold="true" box="[488,757,951,976]" italics="true" pageId="26" pageNumber="477">(measurements in mm)</emphasis>
|
||
.
|
||
</emphasis>
|
||
Adult male (SVL 33.5), rather stout; head wider than long (HW 12.7, HL 12.1); snout rounded in dorsal view, obtuse in lateral view, snout length (SL 4.5) longer than horizontal diameter of eye (EL 4.2); loreal region acute, rounded canthus rostralis; interorbital space flat, narrower than (IUE 1.8) upper eyelid width (UEW 3.3) and internarial distance (IN 2.8); nostril oval, nearly as close to the tip of snout (NS 2.0) as to the eye (EN 2.1); tympanum (TYD 1.7) 40.5% of eye diameter (EL 4.2); tympanum-eye distance (TYE 1.5) 88.2% of the tympanum diameter (TYD 1.7); supratympanic fold well developed, extends from posterior corner of eye to near the shoulder; vomerine ridge present, bearing small teeth, at an angle of 45° to the body axis, as close to choanae as to each other; tongue moderately large, emarginated, bearing no median lingual process; rictal gland present at labial commissure of the mouth. Arms short, forearm length (FAL 6.2) shorter than hand length (HAL 7.1); relative length of fingers IV<II<I<III (FL I 4.0, FL II 3.4, FL III 4.4, FL IV 2.5); finger tips rounded, slightly enlarged without discs, fingers without fringes, webbing between fingers absent; subarticular tubercles prominent, circular; one distinct palmar tubercle, oval, bifid; supernumerary tubercles absent. Hind limbs short, thigh (TL 15.4) subequal to shank (SHL 15.5) and shorter than foot (FOL 16.1), distance from the base of tarsus to the tip of toe IV (TFOL 21.7); toes long, relative length of toes I<II<V<III<IV; toe tips rounded, slightly enlarged without discs, toes without fringes, webbing between toes small:
|
||
<date id="FF8510613130FF80FC3EB481D13DFA64" box="[1012,1035,1459,1483]" pageId="26" pageNumber="477">I2</date>
|
||
– –2–
|
||
<date id="FF8510613130FF80FBFDB481D16EFA64" box="[1079,1112,1459,1483]" pageId="26" pageNumber="477">II2</date>
|
||
– –
|
||
<date id="FF8510613130FF80FBA5B481D196FA64" box="[1135,1184,1455,1483]" bridgedPair="-" pageId="26" pageNumber="477">3–III</date>
|
||
2–3 2/3
|
||
<date id="FF8510613130FF80FB2AB481D03DFA64" box="[1248,1291,1459,1483]" pageId="26" pageNumber="477">IV3</date>
|
||
2/3–
|
||
<date id="FF8510613130FF80FAF8B481D063FA64" box="[1330,1365,1459,1483]" pageId="26" pageNumber="477">2V</date>
|
||
; inner toe length (ITL 2.7); subarticular tubercles prominent, all present, circular; inner metatarsal tubercle prominent, shovel-shaped (IMT 2.1); outer metatarsal small (OMT 0.7), rounded; supernumerary tubercles absent.
|
||
</paragraph>
|
||
<paragraph id="8B8436A13130FF80FF0DB713D6A4F967" blockId="26.[151,1437,151,2023]" pageId="26" pageNumber="477">
|
||
Skin of snout shagreened with scattered glandular projections, upper eyelids prominently tuberculate, anterior and posterior parts of back, and upper and lower parts of flank shagreened with prominent granular projections and scattered glandular warts; interrupted inverse V-shaped ridge on the center of dorsum; dorsal surfaces of forelimb, thigh and shank shagreened with prominent granular projections. Ventral surface of throat, chest, belly and limbs smooth; fejervaryan line present on both sides of the belly (
|
||
<figureCitation id="13002A243130FF80FCFBB782D6B2F967" box="[817,900,1712,1737]" captionStart="FIGURE 10" captionStartId="25.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1924]" captionTargetId="figure@25.[151,1437,193,1924]" captionTargetPageId="25" captionText="FIGURE 10. Holotype of Fejervarya kadar sp. nov. (ZSI / WGRC / V / A / 940): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829853/files/figure.png" pageId="26" pageNumber="477">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A13130FF81FF0DB7E1D6A1FF57" blockId="26.[151,1437,151,2023]" lastBlockId="27.[151,1437,151,680]" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
|
||
<emphasis id="B94FEAB33130FF80FF0DB7E1D487F943" bold="true" box="[199,433,1747,1772]" pageId="26" pageNumber="477">
|
||
Colour of
|
||
<typeStatus id="548088033130FF80FE8EB7E1D49BF943" box="[324,429,1747,1772]" pageId="26" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B94FEAB33130FF80FE70B7E7D733F943" box="[442,517,1747,1772]" italics="true" pageId="26" pageNumber="477">In life.</emphasis>
|
||
Dorsum, upper eyelids, lateral side of snout brownish-grey with prominent dark brown mottling and blotches (
|
||
<figureCitation id="13002A243130FF80FDCDB7C5D757F8BF" box="[519,609,1783,1808]" captionStart="FIGURE 10" captionStartId="25.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1924]" captionTargetId="figure@25.[151,1437,193,1924]" captionTargetPageId="25" captionText="FIGURE 10. Holotype of Fejervarya kadar sp. nov. (ZSI / WGRC / V / A / 940): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829853/files/figure.png" pageId="26" pageNumber="477">Fig. 10</figureCitation>
|
||
A); upper and lower lip with prominent greyish-brown cross bands; tympanum light brown with a dark greyish-brown patch; anterior parts of flank light greyish-brown with dark grey reticulations, forelimbs and hind limbs (including toes) lighter brown compared to dorsum with greyish-brown transverse bands; groin off-white; webbing light grey; anterior part of thigh light yellowish with faint grey reticulation. Ventral surface of throat light flesh colour with two lateral black calling patches; belly white; forearm and foreleg light flesh red in colour with dark brown mottling on the margins.
|
||
<emphasis id="B94FEAB33130FF80FBD9B699D189F86D" box="[1043,1215,1963,1986]" italics="true" pageId="26" pageNumber="477">In preservation</emphasis>
|
||
. Dorsum dark grey with blackish-brown patches, forelimbs and hind limbs lighter brown in colour than dorsum with dark grey transverse bands, posterior parts of thigh light brown with faint dark brown reticulations. Ventral surface of throat light greyish-brown with two lateral black calling patches on either side of the throat, belly off-white, and margins of the limbs light brown with dark greyish-brown mottling (
|
||
<figureCitation id="13002A243131FF81FCFDB1EDD6BCFF57" box="[823,906,223,248]" captionStart="FIGURE 10" captionStartId="25.[151,250,1945,1967]" captionTargetBox="[151,1436,193,1924]" captionTargetId="figure@25.[151,1437,193,1924]" captionTargetPageId="25" captionText="FIGURE 10. Holotype of Fejervarya kadar sp. nov. (ZSI / WGRC / V / A / 940): A. dorsolateral view, in life; B. dorsal view, in preservation; C. ventral view, in preservation; D. lateral view of head, in preservation; E. ventral view of hand, in preservation; F. ventral view of foot, in preservation; G. schematic illustration of foot webbing." httpUri="https://zenodo.org/record/829853/files/figure.png" pageId="27" pageNumber="478">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8436A13131FF81FF0DB031D15EFECB" blockId="27.[151,1437,151,680]" pageId="27" pageNumber="478">
|
||
<emphasis id="B94FEAB33131FF81FF0DB031D47CFEB3" bold="true" box="[199,330,259,284]" pageId="27" pageNumber="478">Variations.</emphasis>
|
||
Morphometric data from three adult males and two adult females, including the holotype, is given in
|
||
<tableCitation id="C6B9031A3131FF81FF7EB015D43AFE90" box="[180,268,295,320]" captionStart="TABLE 7" captionStartId="18.[119,204,562,584]" captionText="TABLE 7. Morphometric measurements for five species in the Fejervarya rufescens group. All measurements are in millimeters (mm). The status of each specimen is given the museum number: Holotype (HT), Paratype (PT), Neotype (NT), Referred specimen (RS)." pageId="27" pageNumber="478">Table 7</tableCitation>
|
||
.
|
||
<emphasis id="B94FEAB33131FF81FEDDB01AD4F5FE90" box="[279,451,296,319]" italics="true" pageId="27" pageNumber="478">In preservation</emphasis>
|
||
. ZSI/WGRC/V/A/942: dorsum grey with scattered faint black spots; ZSI/WGRC/V/A/ 944: dorsum light grey with dark grey spots and more prominent glandular projections.
|
||
</paragraph>
|
||
<paragraph id="8B8436A13131FF81FF0DB05DD650FE03" blockId="27.[151,1437,151,680]" pageId="27" pageNumber="478">
|
||
<emphasis id="B94FEAB33131FF81FF0DB05DD712FE27" bold="true" box="[199,548,367,392]" pageId="27" pageNumber="478">Secondary sexual characters.</emphasis>
|
||
<emphasis id="B94FEAB33131FF81FDE6B05DD753FE27" box="[556,613,367,392]" italics="true" pageId="27" pageNumber="478">Male</emphasis>
|
||
: nuptial pad on finger I present; distinct calling patches on either side of the throat.
|
||
<emphasis id="B94FEAB33131FF81FEC5B0A1D455FE03" box="[271,355,403,428]" italics="true" pageId="27" pageNumber="478">Female</emphasis>
|
||
(ZSI/WGRC/V/A/944): mature ovum absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321652A3131FF81FF0DB085D1E8FD07" pageId="27" pageNumber="478" type="distribution">
|
||
<paragraph id="8B8436A13131FF81FF0DB085D1E8FD07" blockId="27.[151,1437,151,680]" pageId="27" pageNumber="478">
|
||
<emphasis id="B94FEAB33131FF81FF0DB085D762FE7F" bold="true" box="[199,596,439,464]" pageId="27" pageNumber="478">Distribution and natural history.</emphasis>
|
||
<taxonomicName id="4C3B4D223131FF81FD96B08AD615FE7F" authority="Garg & Biju, 2017" authorityName="Garg & Biju" authorityYear="2017" box="[604,803,439,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="27" pageNumber="478" phylum="Chordata" rank="species" species="kadar" status="sp. nov.">
|
||
<emphasis id="B94FEAB33131FF81FD96B08AD615FE7F" box="[604,803,439,464]" italics="true" pageId="27" pageNumber="478">Fejervarya kadar</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57C83131FF81FCE6B085D6BEFE7F" box="[812,904,439,464]" pageId="27" pageNumber="478" rank="species">
|
||
<emphasis id="B94FEAB33131FF81FCE6B085D6BEFE7F" bold="true" box="[812,904,439,464]" pageId="27" pageNumber="478">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is currently known only from its
|
||
<typeStatus id="548088033131FF81FAC4B08AD008FE7F" box="[1294,1342,440,464]" pageId="27" pageNumber="478">type</typeStatus>
|
||
locality Thavalakuzhipara (Vazhachal forest), south of Palghat gap in the
|
||
<collectingRegion id="49FFF8433131FF81FCB1B0EED6E0FE5C" box="[891,982,476,499]" country="Sri Lanka" name="Western" pageId="27" pageNumber="478">Western</collectingRegion>
|
||
Ghats state of
|
||
<collectingRegion id="49FFF8433131FF81FB48B0E9D1FBFE5B" box="[1154,1229,475,500]" country="India" name="Kerala" pageId="27" pageNumber="478">Kerala</collectingRegion>
|
||
(
|
||
<figureCitation id="13002A243131FF81FB17B0E9D014FE5C" box="[1245,1314,475,500]" captionStart="FIGURE 3" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[151,1436,197,1739]" captionTargetId="figure@6.[151,1436,193,1744]" captionTargetPageId="6" captionText="FIGURE 3. Geographical distribution of five species in Fejervarya rufescens group of the Western Ghats, Peninsular India. The Western Ghats biodiversity hotspot region is shown in orange colour. Locality details are referenced in Table 3." httpUri="https://zenodo.org/record/829839/files/figure.png" pageId="27" pageNumber="478">Fig. 3</figureCitation>
|
||
). The
|
||
<typeStatus id="548088033131FF81FAA6B0EED0AAFE5B" box="[1388,1436,476,500]" pageId="27" pageNumber="478">type</typeStatus>
|
||
series was collected from rocky surfaces of an open hill stream with small puddles or narrow water channels flowing down the slopes at the time of collection. The specific site is located inside an evergreen forest patch close to the Thavalakuzhipara tribal colony. Animals were sensitive to movement around them and tried to hide in nearby ground vegetation or grasses when approached. A few male individuals were found calling erratically but breeding activity was not observed. Collections were made in the late evening between 18:00–20:00 hours.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |