treatments-xml/data/03/92/87/039287A1954BFFACCFF7596BFDE92643.xml
2024-06-21 12:22:17 +02:00

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<mods:title id="6B0F2727CEC9C2CFA012703D9158F733">An annotated checklist of the chondrichthyans of South Africa</mods:title>
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<taxonomicName id="4C3B4D34954BFFACCFF7596BFD9E23EF" ID-CoL="3LDLV" authority="Fowler, 1934" authorityName=": Fowler" authorityYear="1934" box="[151,615,870,897]" class="Elasmobranchii" family="Dalatiidae" genus="Heteroscymnoides" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="marleyi">
<emphasis id="B94FEAA5954BFFACCFF7596BFD9E23EF" bold="true" box="[151,615,870,897]" pageId="34" pageNumber="35">
<emphasis id="B94FEAA5954BFFACCFF7596BFE3023EE" bold="true" box="[151,457,870,897]" italics="true" pageId="34" pageNumber="35">Heteroscymnoides marleyi</emphasis>
<bibRefCitation id="EFAA4B46954BFFACCEB0596BFD9E23EF" author="Fowler, H. W." box="[464,615,870,896]" pageId="34" pageNumber="35" pagination="233 - 367" refId="ref101068" refString="Fowler, H. W. (1934 a) Descriptions of new fishes obtained 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Proceedings of the Academy of Natural Sciences of Philadelphia, 85 (for 1933), 233 - 367." type="journal article" year="1934">Fowler, 1934</bibRefCitation>
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<paragraph id="8B8436B7954BFFACCFF75986FE5D23CB" blockId="34.[151,1437,870,1581]" box="[151,420,906,933]" pageId="34" pageNumber="35">Longnose Pygmy Shark</paragraph>
</subSubSection>
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<paragraph id="8B8436B7954BFFACCFF759A3FCE72383" blockId="34.[151,1437,870,1581]" pageId="34" pageNumber="35">
<taxonomicName id="4C3B4D34954BFFACCFF759A3FD5923A7" ID-CoL="3LDLV" authority="Fowler, 1934: 240" authorityName=": Fowler" authorityPageNumber="240" authorityYear="1934" box="[151,672,942,968]" class="Elasmobranchii" family="Dalatiidae" genus="Heteroscymnoides" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="marleyi">
<emphasis id="B94FEAA5954BFFACCFF759A3FE3D23A7" box="[151,452,942,968]" italics="true" pageId="34" pageNumber="35">Heteroscymnoides marleyi</emphasis>
<materialsCitation id="3B533CEA954BFFACCEAC59A3FD5923A7" ID-GBIF-Occurrence="3072574801" box="[460,672,942,968]" collectionCode="ANSP" location="Point Ocean Beach" municipality="southwestern Indian Ocean." pageId="34" pageNumber="35" specimenCode="ANSP 53046" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFAA4B46954BFFACCEAC59A3FD5923A7" author="Fowler, H. W." box="[460,672,942,968]" pageId="34" pageNumber="35" pagination="233 - 367" refId="ref101068" refString="Fowler, H. W. (1934 a) Descriptions of new fishes obtained 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Proceedings of the Academy of Natural Sciences of Philadelphia, 85 (for 1933), 233 - 367." type="journal article" year="1934">Fowler, 1934: 240</bibRefCitation>
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, fig. 4.
<typeStatus id="54808815954BFFACCD9A59A3FC9E23A7" box="[762,871,942,968]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB9D9ECC954BFFACCC1159A3FBF323A6" box="[881,1034,942,969]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="34" pageNumber="35">ANSP 53046</specimenCode>
. Type locality:
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, Durban, KwaZulu-Natal,
<collectingMunicipality id="6BE0ACCD954BFFACCE8659DFFCE72383" box="[486,798,978,1004]" pageId="34" pageNumber="35">southwestern Indian Ocean.</collectingMunicipality>
</paragraph>
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<emphasis id="B94FEAA5954BFFACCFF759FBFEA5247F" bold="true" box="[151,348,1014,1040]" pageId="34" pageNumber="35">Local synonymy</emphasis>
:
<taxonomicName id="4C3B4D34954BFFACCE0D59FAFD94247F" authority="Tanaka, 1912" box="[365,621,1014,1040]" class="Elasmobranchii" family="Dalatiidae" genus="Heteroscymnus" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="longus">
<emphasis id="B94FEAA5954BFFACCE0D59FAFD94247F" box="[365,621,1014,1040]" italics="true" pageId="34" pageNumber="35">Heteroscymnus longus</emphasis>
</taxonomicName>
:
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<bibRefCitation id="EFAA4B46954BFFACCD1D59FBFCA6247F" author="Fowler" box="[637,863,1014,1041]" firstAuthor="Fowler" page="191" pageId="34" pageNumber="35" pagination="187 - 268" refId="ref100995" refString="Fowler, H. W. (1925 b) Fishes from Natal, Zululand and Portuguese East Africa. Proceedings of the Academy of Natural Sciences, Philadelphia, 1925, 77, 187 - 268." type="journal article" year="1925" yearSuffix="b">Fowler, 1925b: 191</bibRefCitation>
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;
<treatmentCitation id="0A9A10A6954BFFACCC0D59FBFBA2247F" box="[877,1115,1014,1040]" pageId="34" pageNumber="35">
<bibRefCitation id="EFAA4B46954BFFACCC0D59FBFBA2247F" author="Barnard" box="[877,1115,1014,1040]" firstAuthor="Barnard" page="1013" pageId="34" pageNumber="35" pagination="419 - 1065" refId="ref92485" refString="Barnard, K. H. (1927) A monograph of the marine fishes of South Africa. Part II (Teleostei - Discocephali to end. Appendix). Annals of the South African Museum 21, 419 - 1065." type="journal article" year="1927">Barnard, 1927: 1013</bibRefCitation>
</treatmentCitation>
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.
<taxonomicName id="4C3B4D34954BFFACCB0859FBFE6E245B" ID-CoL="3LDLV" authority=": Fowler, 1934: 240" authorityName=": Fowler" authorityPageNumber="240" authorityYear="1934" class="Elasmobranchii" family="Dalatiidae" genus="Heteroscymnoides" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="marleyi">
<emphasis id="B94FEAA5954BFFACCB0859FBFA6C247F" box="[1128,1429,1014,1040]" italics="true" pageId="34" pageNumber="35">Heteroscymnoides marleyi</emphasis>
:
<bibRefCitation id="EFAA4B46954BFFACCFA65E17FE6E245B" author="Fowler, H. W." box="[198,407,1050,1076]" pageId="34" pageNumber="35" pagination="233 - 367" refId="ref101068" refString="Fowler, H. W. (1934 a) Descriptions of new fishes obtained 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Proceedings of the Academy of Natural Sciences of Philadelphia, 85 (for 1933), 233 - 367." type="journal article" year="1934">Fowler, 1934: 240</bibRefCitation>
</taxonomicName>
, fig. 4.;
<bibRefCitation id="EFAA4B46954BFFACCE945E17FD4E245A" author="Smith, J. L. B." box="[500,695,1050,1077]" pageId="34" pageNumber="35" refId="ref114538" refString="Smith, J. L. B. (1949 a) The Sea Fishes of Southern Africa. South Africa Central News Agency Ltd., 550 pp." type="book" year="1949">Smith, 1949a: 56</bibRefCitation>
, fig. 45; Bigelow &amp; Schroder, 1957: 132, figs. 15f, g, 16e;
<bibRefCitation id="EFAA4B46954BFFACCA325E17FED42436" author="Smith, J. L. B." pageId="34" pageNumber="35" refId="ref115700" refString="Smith, J. L. B. (1965) The Sea Fishes of Southern Africa. 5 th Edition. Central News Agency Ltd., 580 pp." type="book" year="1965">Smith, 1965: 56</bibRefCitation>
, fig. 45;
<bibRefCitation id="EFAA4B46954BFFACCEF85E32FD712437" author="Bass" box="[408,648,1086,1113]" etAl="et al." firstAuthor="Bass" page="49" pageId="34" pageNumber="35" pagination="1 - 103" refId="ref93155" refString="Bass, A. J., D'Aubrey, J. D. &amp; Kistnasamy, N. (1976) Sharks of the east coast of southern Africa. VI. The families Oxynotidae, Squalidae, Dalatiidae and Echinorhinidae. Investigational Report. Oceanographic Research Institute, Durban, 45, 1 - 103." type="journal article" year="1976">
Bass
<emphasis id="B94FEAA5954BFFACCEB65E32FDF02437" box="[470,521,1086,1112]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 1976: 49
</bibRefCitation>
, fig. 35;
<bibRefCitation id="EFAA4B46954BFFACCD925E33FC0B2437" author="Compagno" box="[754,1010,1086,1112]" firstAuthor="Compagno" page="92" pageId="34" pageNumber="35" pagination="1 - 250" refId="ref95434" refString="Compagno, L. J. V. (1984 a) FAO Species Catalogue. Sharks of the World. An Annotated and Illustrated Catalogue of Shark Species Known to Date. FAO Fisheries Synopsis. Vol. 4. No. 125. Part 1. Hexanchiformes to Lamniformes. FAO, Rome, pp. 1 - 250." type="book chapter" year="1984" yearSuffix="a">Compagno, 1984a: 92</bibRefCitation>
, fig.;
<bibRefCitation id="EFAA4B46954BFFACCB565E32FADF2436" author="Bass, A. J. &amp; Compagno, L. J. V." box="[1078,1318,1086,1113]" editor="Smith, M. M. &amp; Heemstra, P. C." journalOrPublisher="Macmillan, Johannesburg" pageId="34" pageNumber="35" pagination="104 - 105" refId="ref93292" refString="Bass, A. J. &amp; Compagno, L. J. V. (1986) Families Echinorhinidae, Proscyllidae, Odontaspidiidae, Mitsukurinidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smith's Sea Fishes. Macmillan, Johannesburg, pp. 63 + 103 + 104 - 105." title="Families Echinorhinidae, Proscyllidae, Odontaspidiidae, Mitsukurinidae" type="book chapter" volumeTitle="Smith's Sea Fishes" year="1986">
Bass
<emphasis id="B94FEAA5954BFFACCB145E32FB512437" box="[1140,1192,1086,1112]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 1986: 58
</bibRefCitation>
, fig. 5.19;
<bibRefCitation id="EFAA4B46954BFFACCFA65E6FFE012413" author="Compagno" box="[198,504,1122,1149]" etAl="et al." firstAuthor="Compagno" page="34" pageId="34" pageNumber="35" refId="ref96138" refString="Compagno, L. J. V., Ebert, D. A. &amp; Smale, M. J. (1989) Guide to the Sharks and Rays of Southern Africa. Struik Publishers, Cape Town, 158 pp." type="book" year="1989">
Compagno
<emphasis id="B94FEAA5954BFFACCE295E6EFE852413" box="[329,380,1122,1148]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 1989: 34
</bibRefCitation>
, pl.;
<bibRefCitation id="EFAA4B46954BFFACCD525E6FFCC92412" author="Compagno" box="[562,816,1122,1149]" firstAuthor="Compagno" page="115" pageId="34" pageNumber="35" pagination="75 - 120" refId="ref95720" refString="Compagno, L. J. V. (1999) An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa, 54, 75 - 120. https: // doi. org / 10.1080 / 00359199909520406" type="journal article" year="1999">Compagno, 1999: 115</bibRefCitation>
;
<bibRefCitation id="EFAA4B46954BFFACCC5D5E6FFB842413" author="Compagno" box="[829,1149,1122,1149]" etAl="et al." firstAuthor="Compagno" page="126" pageId="34" pageNumber="35" refId="ref95897" refString="Compagno, L., Dando, M. &amp; Fowler, S. (2005) Field Guide to the Sharks of the World. Harper Collins Publishers Ltd, London, 368 pp." type="book" year="2005">
<treatmentCitation id="0A9A10A6954BFFACCC5D5E6FFBBC2412" box="[829,1093,1122,1149]" pageId="34" pageNumber="35">
Compagno
<emphasis id="B94FEAA5954BFFACCCA05E6EFC0A2413" box="[960,1011,1122,1148]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2005
</treatmentCitation>
: 126
</bibRefCitation>
, fig., pl. 14;
<bibRefCitation id="EFAA4B46954BFFACCA715E6FFF0824CF" author="Ebert D. A." pageId="34" pageNumber="35" refId="ref98340" refString="Ebert D. A. (2013) Deep-sea cartilaginous fishes of the Indian Ocean. Vol. 1. Sharks. FAO Species Catalogue for Fishery Purposes No. 8. Vol. 1. FAO, Rome, 256 pp." type="book" year="2013">Ebert, 2013: 113</bibRefCitation>
, fig. 152;
<bibRefCitation id="EFAA4B46954BFFACCE045E8BFDD224CF" author="Ebert, D. A. &amp; Fowler, S. &amp; Compagno, L. J. V." box="[356,555,1158,1185]" pageId="34" pageNumber="35" refId="ref99433" refString="Ebert, D. A., Fowler, S. &amp; Compagno, L. J. V. (2013) Sharks of the World: A Fully Illustrated Guide to the Sharks of the World. Wild Nature Press, Plymouth, 528 pp." type="book" year="2013">
Ebert
<emphasis id="B94FEAA5954BFFACCEC85E8AFE2324CF" box="[424,474,1158,1184]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2013
</bibRefCitation>
a: 169, fig., pl. 17; Ebert &amp; Mostada, 2013: 48, fig.;
<bibRefCitation id="EFAA4B46954BFFACCB175E8BFA6E24CF" author="Ebert &amp; Dando" box="[1143,1431,1158,1184]" firstAuthor="Ebert" page="83" pageId="34" pageNumber="35" refId="ref99385" refString="Ebert, D. A. &amp; Dando, M. (2014) On Board Guide for the Identification of Pelagic Sharks and Rays of the Western Indian Ocean. SmartFish Programme, FAO, Rome &amp; Indian Ocean Commission, Port Louis, 109 pp." type="book" year="2014">Ebert &amp; Dando, 2014: 83</bibRefCitation>
, fig.;
<bibRefCitation id="EFAA4B46954BFFACCF9D5EA7FE4624AB" author="Ebert, D. A." box="[253,447,1194,1221]" pageId="34" pageNumber="35" refId="ref98428" refString="Ebert, D. A. (2015) Deep-sea cartilaginous fishes of the Southeastern Atlantic Ocean. FAO Species Catalogue for Fishery Purposes No. 9. FAO, Rome, 251 pp." type="book" year="2015">Ebert, 2015: 108</bibRefCitation>
, fig. 126; Ebert &amp; Mostada, 2015: 41, fig.;
<bibRefCitation id="EFAA4B46954BFFACCCDA5EA7FAF624AB" author="Ebert &amp; van Hees" box="[954,1295,1194,1221]" firstAuthor="Ebert" page="144" pageId="34" pageNumber="35" pagination="141 - 156" refId="ref99877" refString="Ebert, D. A. &amp; van Hees, K. E. (2015) Beyond jaws: rediscovering the &quot; Lost Sharks &quot; of southern Africa. African Journal of Marine Science, 37, 141 - 156. https: // doi. org / 10.2989 / 1814232 X. 2015.1048730" type="journal article" year="2015">Ebert &amp; van Hees, 2015: 144</bibRefCitation>
;
<bibRefCitation id="EFAA4B46954BFFACCA7D5EA7FEC32487" author="Weigmann, S." pageId="34" pageNumber="35" pagination="837 - 1037" refId="ref118762" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88, 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann, 2016: 892</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321653C954BFFACCFF75EFFFD532562" box="[151,682,1266,1293]" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8B8436B7954BFFACCFF75EFFFD532562" blockId="34.[151,1437,870,1581]" box="[151,682,1266,1293]" pageId="34" pageNumber="35">
<emphasis id="B94FEAA5954BFFACCFF75EFFFDFB2563" bold="true" box="[151,514,1266,1292]" pageId="34" pageNumber="35">
<collectingCountry id="F32C7627954BFFACCFF75EFFFED62563" box="[151,303,1266,1292]" name="South Africa" pageId="34" pageNumber="35">South Africa</collectingCountry>
voucher material
</emphasis>
:
<collectionCode id="ED2AAE72954BFFACCD6F5EFFFDAF2563" box="[527,598,1266,1292]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="34" pageNumber="35">ANSP</collectionCode>
53046.
</paragraph>
</subSubSection>
<subSubSection id="C321653C954BFFACCFF75F1BFAB6255F" box="[151,1359,1302,1329]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8B8436B7954BFFACCFF75F1BFAB6255F" blockId="34.[151,1437,870,1581]" box="[151,1359,1302,1329]" pageId="34" pageNumber="35">
<emphasis id="B94FEAA5954BFFACCFF75F1BFE3D255F" bold="true" box="[151,452,1302,1328]" pageId="34" pageNumber="35">
<collectingCountry id="F32C7627954BFFACCFF75F1BFED6255F" box="[151,303,1302,1328]" name="South Africa" pageId="34" pageNumber="35">South Africa</collectingCountry>
distribution
</emphasis>
: Known from a single specimen picked up on a beach at Durban (
<bibRefCitation id="EFAA4B46954BFFACCBC85F1BFABB255F" author="Fowler, H. W." box="[1192,1346,1302,1328]" pageId="34" pageNumber="35" pagination="233 - 367" refId="ref101068" refString="Fowler, H. W. (1934 a) Descriptions of new fishes obtained 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Proceedings of the Academy of Natural Sciences of Philadelphia, 85 (for 1933), 233 - 367." type="journal article" year="1934">Fowler, 1934</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321653C954BFFACCFF75F37FDE92643" pageId="34" pageNumber="35" type="discussion">
<subSubSection id="C321653C954BFFACCFF75F37FA812666" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8B8436B7954BFFACCFF75F37FA812666" blockId="34.[151,1437,870,1581]" pageId="34" pageNumber="35">
<emphasis id="B94FEAA5954BFFACCFF75F37FEFD253B" bold="true" box="[151,260,1338,1364]" pageId="34" pageNumber="35">Remarks</emphasis>
: The
<typeStatus id="54808815954BFFACCE265F37FE51253B" box="[326,424,1338,1364]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
was picked up on a beach near Vetches Pier, Durban, on
<date id="FF851077954BFFACCB4D5F37FB25253B" box="[1069,1244,1338,1365]" pageId="34" pageNumber="35" value="1923-01-04">4 January 1923</date>
. The
<quantity id="4CC39B52954BFFACCA7A5F37FA8D253A" box="[1306,1396,1338,1365]" metricMagnitude="-1" metricUnit="m" metricValue="1.26" pageId="34" pageNumber="35" unit="cm" value="12.6">12.6 cm</quantity>
TL female specimen was collected by H.W. Bell-Marley who sent it to H.W. Fowler who originally mistook it for
<taxonomicName id="4C3B4D34954BFFACCFA65F8EFD9E25F3" authority="Tanaka, 1912" authorityName="Tanaka" authorityYear="1912" box="[198,615,1410,1436]" class="Elasmobranchii" family="Dalatiidae" genus="Heteroscymnus" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="longus">
<emphasis id="B94FEAA5954BFFACCFA65F8EFE3C25F3" box="[198,453,1410,1436]" italics="true" pageId="34" pageNumber="35">Heteroscymnus longus</emphasis>
<bibRefCitation id="EFAA4B46954BFFACCEAC5F8FFD9E25F3" author="Tanaka, S." box="[460,615,1410,1436]" pageId="34" pageNumber="35" pagination="145 - 164" refId="ref116851" refString="Tanaka, S. (1912) Figures and descriptions of the fishes of Japan, including the Riukiu Islands, Bonin Islands, Formosa, Kurile Islands, Korea, and southern Sakhalin. Figures and Descriptions of the Fishes of Japan, 9, 145 - 164. https: // doi. org / 10.5962 / bhl. title. 13715" type="journal article" year="1912">Tanaka, 1912</bibRefCitation>
</taxonomicName>
; a species referable to the genus
<taxonomicName id="4C3B4D34954BFFACCC805F8EFBA125F2" authorityName="Lesueur" authorityYear="1818" box="[992,1112,1411,1437]" class="Elasmobranchii" family="Somniosidae" genus="Somniosus" kingdom="Animalia" order="Squaliformes" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="B94FEAA5954BFFACCC805F8EFBA125F2" box="[992,1112,1411,1437]" italics="true" pageId="34" pageNumber="35">Somniosus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFAA4B46954BFFACCB075F8FFAF025F3" author="Fowler, H. W." box="[1127,1289,1410,1436]" pageId="34" pageNumber="35" pagination="233 - 367" refId="ref101068" refString="Fowler, H. W. (1934 a) Descriptions of new fishes obtained 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Proceedings of the Academy of Natural Sciences of Philadelphia, 85 (for 1933), 233 - 367." type="journal article" year="1934">Fowler (1934)</bibRefCitation>
later revised his original identification and described it as a new genus and species. The species appears to be wide-ranging in the Southern Hemisphere with the only six known specimens occurring in three widely dispersed locations in the southwestern Indian Ocean, southeastern Atlantic Ocean, and southeastern Pacific Ocean (
<bibRefCitation id="EFAA4B46954BFFACCB835FE3FA942666" author="Ebert, D. A." box="[1251,1389,1518,1545]" pageId="34" pageNumber="35" refId="ref98428" refString="Ebert, D. A. (2015) Deep-sea cartilaginous fishes of the Southeastern Atlantic Ocean. FAO Species Catalogue for Fishery Purposes No. 9. FAO, Rome, 251 pp." type="book" year="2015">Ebert, 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321653C954BFFACCFF75C1FFDE92643" box="[151,528,1554,1581]" pageId="34" pageNumber="35" type="multiple">
<paragraph id="8B8436B7954BFFACCFF75C1FFDE92643" blockId="34.[151,1437,870,1581]" box="[151,528,1554,1581]" pageId="34" pageNumber="35">
<emphasis id="B94FEAA5954BFFACCFF75C1FFE7D2643" bold="true" box="[151,388,1554,1580]" pageId="34" pageNumber="35">Conservation status</emphasis>
: LC (2019).
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>