treatments-xml/data/03/92/7F/03927F0EFFE46059FCC9FB9D6F8FD74C.xml

<document id="5C9641BCB2AF6AAA775506C107FB0955" ID-CLB-Dataset="5419" ID-DOI="10.1046/j.1096-3642.2002.00039.x" ID-GBIF-Dataset="8acc9095-eaff-47d7-b3da-91b6c2fb636e" ID-ISSN="0024-4082" ID-Zenodo-Dep="4634200" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1616618193905" checkinUser="carolina" docAuthor="Valdés, Ángel" docDate="2002" docId="03927F0EFFE46059FCC9FB9D6F8FD74C" docLanguage="en" docName="ZoolJLinnSoc.136.535-636.pdf" docOrigin="Zoological Journal of the Linnean Society 136 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00039.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Doris pseudoargus RAPP 1827" docType="treatment" docVersion="11" lastPageNumber="551" masterDocId="FFAB0776FFEF6049FF87FFCE6F73D33D" masterDocTitle="A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)" masterLastPageNumber="636" masterPageNumber="535" pageNumber="546" updateTime="1692298629198" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="5EA9BCBCC0B1A2E3EFF83184B1BAF69E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="55FD0FCFFAE54CD44CF3DF19AC2BB0D3">
<mods:title id="4598649E03EB7C5FF94E50214BB873A7">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
</mods:titleInfo>
<mods:name id="85F6F4B3B72DEFC86DAF79A4985F351E" type="personal">
<mods:role id="A1F9E6DF3CB2C61735E88EF997ED7B5A">
<mods:roleTerm id="8B78BA6737E2AAE35C0C98D7122CF989">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E8E81EEAC4721903E567B9C0E5A1571">Valdés, Ángel</mods:namePart>
</mods:name>
<mods:typeOfResource id="BF96FCAA5F484479305A838A068C3F3E">text</mods:typeOfResource>
<mods:relatedItem id="B2ED0ED7A309A8A50F5FE41406D1E549" type="host">
<mods:titleInfo id="C8FC947803F8BB1960DC75540F38760B">
<mods:title id="1C200BD947BFD8D2A965615919075273">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="E9671F67898D96FD69A360EC10C8CB3C">
<mods:date id="0040A8CE04F92D0795FE9DE1EFFF4418">2002</mods:date>
<mods:detail id="E098616C4DF2B67D6D2A4F31E2A85FB7" type="pubDate">
<mods:number id="61FA168C497D0EFC48C4A95A0DCDA80D">2002-12-31</mods:number>
</mods:detail>
<mods:detail id="A9B3E20638385E866298CBD31DE17E59" type="volume">
<mods:number id="490C9A1C88D2090E9056EA853CA43EC0">136</mods:number>
</mods:detail>
<mods:detail id="981A65CAF60B9816A6BE10A06FB86BA1" type="issue">
<mods:number id="C4BA24D7A8D661998C908037AA4064D6">4</mods:number>
</mods:detail>
<mods:extent id="98B72555B4B423BB04CAE0790D1BA793" unit="page">
<mods:start id="AD30726E496E88DCDEF7227DEA9FD39A">535</mods:start>
<mods:end id="9CBAE2A062B18C198BA742E523E7BAE8">636</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C8DBEDCC9345025C0A4FD4E25F11912A">
<mods:url id="72CF3DB8C6DD206FC519914D6703E87D">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00039.x</mods:url>
</mods:location>
<mods:classification id="3ABBDDE57AA7B2D03679FAE0BE9A896A">journal article</mods:classification>
<mods:identifier id="93C3E9DEFA36E024238886D7124B5C08" type="CLB-Dataset">5419</mods:identifier>
<mods:identifier id="5E9E9B63A363BB8ECD709F69BCA988BD" type="DOI">10.1046/j.1096-3642.2002.00039.x</mods:identifier>
<mods:identifier id="A7A0A407112EA053BB7D43ED0350AF10" type="GBIF-Dataset">8acc9095-eaff-47d7-b3da-91b6c2fb636e</mods:identifier>
<mods:identifier id="7859D03952D24C7BA68800A20B249179" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="C83C0BBF0192F4669DDD7661C3EFDD2A" type="Zenodo-Dep">4634200</mods:identifier>
</mods:mods>
<treatment id="03927F0EFFE46059FCC9FB9D6F8FD74C" ID-DOI="http://doi.org/10.5281/zenodo.5110262" ID-GBIF-Taxon="212430591" ID-Zenodo-Dep="5110262" LSID="urn:lsid:plazi:treatment:03927F0EFFE46059FCC9FB9D6F8FD74C" httpUri="http://treatment.plazi.org/id/03927F0EFFE46059FCC9FB9D6F8FD74C" lastPageId="16" lastPageNumber="551" pageId="11" pageNumber="546">
<subSubSection id="C3219D93FFE46042FCC9FB9D6AF3D757" box="[846,1408,1106,1130]" pageId="11" pageNumber="546" type="nomenclature">
<paragraph id="8B84CE18FFE46042FCC9FB9D6AF3D757" blockId="11.[846,1408,1106,1130]" box="[846,1408,1106,1130]" pageId="11" pageNumber="546">
<heading id="D0CC7974FFE46042FCC9FB9D6AF3D757" box="[846,1408,1106,1130]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="546" reason="5">
<taxonomicName id="4C3BB59BFFE46042FCC9FB9D6BBDD757" ID-CoL="6DGPS" ID-ENA="1677693" authority="RAPP, 1827" authorityYear="1827" box="[846,1230,1106,1130]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE46042FCC9FB9D6B30D755" box="[846,1091,1107,1130]" italics="true" pageId="11" pageNumber="546">DORIS PSEUDOARGUS</emphasis>
<bibRefCitation id="EFAAB3E9FFE46042FBCCFB9C6BBDD757" author="RAPP" box="[1099,1230,1106,1130]" firstAuthor="RAPP" pageId="11" pageNumber="546" pagination="515 - 522" refId="ref58646" refString="Rapp WL. 1827. Uber das Molluskengeschlecht Doris und Beschreibung einiger neuen Arten desselben. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 13: 515 - 522, plates 26 - 27." type="journal article" year="1827">RAPP, 1827</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1300D29DFFE46042FB58FB9C6A4CD757" box="[1247,1343,1106,1130]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.4634212" httpUri="https://zenodo.org/record/4634212/files/figure.png" pageId="11" pageNumber="546">FIGS 4A</figureCitation>
, 
<figureCitation id="1300D29DFFE46042FACBFB9C6A2FD757" box="[1356,1372,1106,1130]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="11" pageNumber="546">5</figureCitation>
, 
<figureCitation id="1300D29DFFE46042FAEDFB9C6A0BD757" box="[1386,1400,1106,1130]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="11" pageNumber="546">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFE46042FCB4FBB56BABD7D1" pageId="11" pageNumber="546" type="reference_group">
<paragraph id="8B84CE18FFE46042FCB4FBB56BB9D7AD" blockId="11.[819,1408,1146,1260]" box="[819,1226,1146,1168]" pageId="11" pageNumber="546">
<treatmentCitationGroup id="AB2BE936FFE46042FCB4FBB56BB9D7AD" box="[819,1226,1146,1168]" pageId="11" pageNumber="546">
<treatmentCitation id="0A9AE809FFE46042FCB4FBB56BB5D7AD" author="Rapp WL" box="[819,1222,1146,1168]" page="519" pageId="11" pageNumber="546" year="1827">
<taxonomicName id="4C3BB59BFFE46042FCB4FBB56BB5D7AD" ID-CoL="6DGPS" authority="Rapp, 1827: 519" authorityName="Rapp" authorityPageNumber="519" authorityYear="1827" box="[819,1222,1146,1168]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE46042FCB4FBB56B71D7B2" box="[819,1026,1146,1168]" italics="true" pageId="11" pageNumber="546">Doris pseudoargus</emphasis>
<bibRefCitation id="EFAAB3E9FFE46042FB8DFBB56BB5D7AD" author="Rapp" box="[1034,1222,1146,1168]" firstAuthor="Rapp" page="519" pageId="11" pageNumber="546" pagination="515 - 522" refId="ref58646" refString="Rapp WL. 1827. Uber das Molluskengeschlecht Doris und Beschreibung einiger neuen Arten desselben. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 13: 515 - 522, plates 26 - 27." type="journal article" year="1827">Rapp, 1827: 519</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFE46042FCB4FB546BC2D793" blockId="11.[819,1408,1146,1260]" box="[819,1201,1177,1199]" pageId="11" pageNumber="546">
<treatmentCitationGroup id="AB2BE936FFE46042FCB4FB546BC2D793" box="[819,1201,1177,1199]" pageId="11" pageNumber="546">
<treatmentCitation id="0A9AE809FFE46042FCB4FB546BDFD793" author="Leuckart FS" box="[819,1196,1177,1199]" page="14" pageId="11" pageNumber="546" year="1828">
<taxonomicName id="4C3BB59BFFE46042FCB4FB546BDFD793" ID-CoL="37DF4" authority="Leuckart, 1828: 14" authorityName="Leuckart" authorityPageNumber="14" authorityYear="1828" box="[819,1196,1177,1199]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="flavipes">
<emphasis id="B94F120AFFE46042FCB4FB546CB8D793" box="[819,971,1177,1199]" italics="true" pageId="11" pageNumber="546">Doris flavipes</emphasis>
<bibRefCitation id="EFAAB3E9FFE46042FC54FB576BDFD793" author="Leuckart" box="[979,1196,1177,1199]" firstAuthor="Leuckart" page="14" pageId="11" pageNumber="546" refId="ref56877" refString="Leuckart FS. 1828. Breves Animalium quorundam maxima ex parte marinorum descriptiones. Heidelbergae: A. Osswaldi." type="book" year="1828">Leuckart, 1828: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFE46042FCB4FB766AF3D7F0" blockId="11.[819,1408,1146,1260]" box="[819,1408,1208,1230]" pageId="11" pageNumber="546">
<treatmentCitationGroup id="AB2BE936FFE46042FCB4FB766AF3D7F0" box="[819,1408,1208,1230]" pageId="11" pageNumber="546">
<treatmentCitation id="0A9AE809FFE46042FCB4FB766B9BD7F0" author="Delle Chiaje S" box="[819,1256,1208,1230]" page="19" pageId="11" pageNumber="546" year="1841">
<taxonomicName id="4C3BB59BFFE46042FCB4FB766B9BD7F0" ID-CoL="37DGL" authority="Delle Chiaje, 1841: 19" authorityName="Delle Chiaje" authorityPageNumber="19" authorityYear="1841" box="[819,1256,1208,1230]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="leuckartii">
<emphasis id="B94F120AFFE46042FCB4FB766C92D7F0" box="[819,993,1208,1229]" italics="true" pageId="11" pageNumber="546">Doris leuckartii</emphasis>
<bibRefCitation id="EFAAB3E9FFE46042FC6EFB766B9BD7F0" author="Delle Chiaje" box="[1001,1256,1208,1230]" firstAuthor="Delle Chiaje" page="19" pageId="11" pageNumber="546" refId="ref55179" refString="Delle Chiaje S. 1841. Animale senza vetebre del Regno di Napoli. Naples: Batelli." type="book" year="1841">Delle Chiaje, 1841: 19</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 40, fig. 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B84CE18FFE46042FCB4FB196BABD7D1" blockId="11.[819,1408,1146,1260]" box="[819,1240,1238,1260]" pageId="11" pageNumber="546">
<treatmentCitationGroup id="AB2BE936FFE46042FCB4FB196BABD7D1" box="[819,1240,1238,1260]" pageId="11" pageNumber="546">
<treatmentCitation id="0A9AE809FFE46042FCB4FB196BA7D7D1" author="Verany DB" box="[819,1236,1238,1260]" page="22" pageId="11" pageNumber="546" year="1846">
<taxonomicName id="4C3BB59BFFE46042FCB4FB196BA7D7D1" ID-CoL="37DLD" authority="Verany, 1846: 21 - 22" authorityName="Verany" authorityPageNumber="21 - 22" authorityYear="1846" box="[819,1236,1238,1260]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="schembrii">
<emphasis id="B94F120AFFE46042FCB4FB196C90D7D6" box="[819,995,1238,1260]" italics="true" pageId="11" pageNumber="546">Doris schembrii</emphasis>
<bibRefCitation id="EFAAB3E9FFE46042FC6CFB196BA7D7D1" author="Verany" box="[1003,1236,1238,1260]" firstAuthor="Verany" page="21 - 22" pageId="11" pageNumber="546" pagination="89 - 109" refId="ref60000" refString="Verany DB. 1846. Catalogo degli animali invertebrati marini del Golfo di Genova e Nizza. Guida di Genova 1: 89 - 109, plates 2 - 4." type="journal article" year="1846">Verany, 1846: 21–22</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFE46045FCB4FAFD6D52D698" lastPageId="12" lastPageNumber="547" pageId="11" pageNumber="546" type="materials_examined">
<paragraph id="8B84CE18FFE46042FCB4FAFD6CBDD67A" blockId="11.[819,1435,1330,1720]" box="[819,974,1330,1352]" pageId="11" pageNumber="546">
<emphasis id="B94F120AFFE46042FCB4FAFD6CBDD67A" box="[819,974,1330,1352]" italics="true" pageId="11" pageNumber="546">Type material</emphasis>
</paragraph>
<paragraph id="8B84CE18FFE46042FCB4FA9C6B6DD585" blockId="11.[819,1435,1330,1720]" pageId="11" pageNumber="546">
<taxonomicName id="4C3BB59BFFE46042FCB4FA9C6B38D65A" authority="Rapp" authorityName="Rapp" authorityYear="1827" box="[819,1099,1361,1383]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE46042FCB4FA9C6B77D65B" box="[819,1028,1361,1383]" italics="true" pageId="11" pageNumber="546">Doris pseudoargus</emphasis>
Rapp
</taxonomicName>
, the type material, collected from Le Havre, 
<collectingCountry id="F32C8E88FFE46042FC7AFABE6B3DD6B8" box="[1021,1102,1392,1413]" name="France" pageId="11" pageNumber="546">France</collectingCountry>
, is untraceable. 
<typeStatus id="548070BAFFE46042FA9AFABE6AE8D6B8" box="[1309,1435,1392,1413]" pageId="11" pageNumber="546">NEOTYPE</typeStatus>
(here designated): Locmariaquer, 
<collectingCountry id="F32C8E88FFE46042FB48FA416A53D699" box="[1231,1312,1423,1444]" name="France" pageId="11" pageNumber="546">France</collectingCountry>
, 
<date id="FF85E8D8FFE46042FAB2FA416C1FD6FF" pageId="11" pageNumber="546" value="1972-04-13">13 April 1972</date>
, one specimen, 
<quantity id="4CC363FDFFE46042FBAFFA636B05D6FE" box="[1064,1142,1453,1475]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="546" unit="mm" value="22.0">22 mm</quantity>
preserved length, leg. P. Bouchet (
<collectionCode id="ED2A56DDFFE46042FC23FA026C8ED6DF" box="[932,1021,1484,1506]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="546">MNHN</collectionCode>
). 
<taxonomicName id="4C3BB59BFFE46042FB93FA026A54D6DF" authority="Leuckart" authorityName="Leuckart" authorityYear="1828" box="[1044,1319,1484,1506]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="flavipes">
<emphasis id="B94F120AFFE46042FB93FA026BC2D6DC" box="[1044,1201,1484,1505]" italics="true" pageId="11" pageNumber="546">Doris flavipes</emphasis>
Leuckart
</taxonomicName>
, the type material collected from the Mediterranean Sea is untraceable. 
<taxonomicName id="4C3BB59BFFE46042FC49F9C46A57D522" authority="Delle Chiaje" authorityName="Delle Chiaje" authorityYear="1841" box="[974,1316,1545,1567]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="leuckartii">
<emphasis id="B94F120AFFE46042FC49F9C46BF2D523" box="[974,1153,1545,1567]" italics="true" pageId="11" pageNumber="546">Doris leuckartii</emphasis>
Delle Chiaje
</taxonomicName>
, the type material collected from Nice, 
<collectingCountry id="F32C8E88FFE46042FB17F9E66B92D500" box="[1168,1249,1576,1597]" name="France" pageId="11" pageNumber="546">France</collectingCountry>
, is untraceable. 
<taxonomicName id="4C3BB59BFFE46042FCB4F9896B4DD561" authority="Verany" authorityName="Verany" authorityYear="1846" box="[819,1086,1606,1628]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="schembrii">
<emphasis id="B94F120AFFE46042FCB4F9896C97D566" box="[819,996,1606,1628]" italics="true" pageId="11" pageNumber="546">Doris schembrii</emphasis>
Verany
</taxonomicName>
, 
<typeStatus id="548070BAFFE46042FBCCF9896BA4D561" box="[1099,1239,1607,1628]" pageId="11" pageNumber="546">SYNTYPES</typeStatus>
: Gulf of Geneva, 
<collectingCountry id="F32C8E88FFE46042FCB4F9AB6C1BD546" box="[819,872,1637,1659]" name="Italy" pageId="11" pageNumber="546">Italy</collectingCountry>
, two specimens (
<collectionCode id="ED2A56DDFFE46042FBB2F9AB6BFDD546" box="[1077,1166,1637,1659]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="546">MNHN</collectionCode>
). The type material of 
<taxonomicName id="4C3BB59BFFE46042FCB4F94A6B44D5A7" authority="Leach" authorityName="Leach" box="[819,1079,1668,1690]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="11" pageNumber="546" phylum="Mollusca" rank="species" species="britannica">
<emphasis id="B94F120AFFE46042FCB4F94A6C99D5A4" box="[819,1002,1668,1689]" italics="true" pageId="11" pageNumber="546">Doris britannica</emphasis>
Leach
</taxonomicName>
could not be located at 
<collectionCode id="ED2A56DDFFE46042FACFF94A6AE9D5A4" box="[1352,1434,1668,1689]" pageId="11" pageNumber="546">BMNH</collectionCode>
and is probably lost.
</paragraph>
<paragraph id="8B84CE18FFE46042FCB4F9306B62D42E" blockId="11.[819,1434,1790,1904]" box="[819,1041,1790,1811]" pageId="11" pageNumber="546">
<emphasis id="B94F120AFFE46042FCB4F9306B62D42E" box="[819,1041,1790,1811]" italics="true" pageId="11" pageNumber="546">Additional material</emphasis>
</paragraph>
<paragraph id="8B84CE18FFE46045FCB4F8D06D52D698" blockId="11.[819,1434,1790,1904]" lastBlockId="12.[142,759,1392,1445]" lastPageId="12" lastPageNumber="547" pageId="11" pageNumber="546">
<materialsCitation id="3B53C445FFE46042FCB4F8D06B8AD44D" ID-GBIF-Occurrence="3323414302" collectingDate="1987-08-16" collectionCode="CASIZ" collectorName="A. Valdes" country="Spain" lastPageId="11" lastPageNumber="547" location="Las Llanas Beach" municipality="Muros de Nalon" pageId="11" pageNumber="546" specimenCode="121105" specimenCount="1" stateProvince="Asturias">
<location id="8EE498C3FFE46042FCB4F8D06B75D40E" LSID="urn:lsid:plazi:treatment:03927F0EFFE46059FCC9FB9D6F8FD74C:8EE498C3FFE46042FCB4F8D06B75D40E" box="[819,1030,1821,1843]" country="Spain" municipality="Muros de Nalon" name="Las Llanas Beach" pageId="11" pageNumber="546" stateProvince="Asturias">Las Llanas Beach</location>
, 
<collectingMunicipality id="6BE05462FFE46042FB92F8D06BA0D40E" box="[1045,1235,1821,1843]" pageId="11" pageNumber="546">Muros de Nalón</collectingMunicipality>
, 
<collectingRegion id="49FF00FAFFE46042FB67F8D36A30D40E" box="[1248,1347,1821,1843]" country="Spain" name="Asturias" pageId="11" pageNumber="546">Asturias</collectingRegion>
, 
<collectingCountry id="F32C8E88FFE46042FAD5F8D36AE4D40E" box="[1362,1431,1821,1843]" name="Spain" pageId="11" pageNumber="546">Spain</collectingCountry>
, 
<collectingDate id="EFC11130FFE46042FCB4F8F26C88D46C" box="[819,1019,1852,1874]" pageId="11" pageNumber="546" value="1987-08-16">
<date id="FF85E8D8FFE46042FCB4F8F26C88D46C" box="[819,1019,1852,1874]" pageId="11" pageNumber="546" value="1987-08-16">16 August 1987</date>
</collectingDate>
, 
<specimenCount id="9D3D0591FFE46042FB96F8F26BC6D46F" box="[1041,1205,1852,1874]" pageId="11" pageNumber="546">one specimen</specimenCount>
, 
<quantity id="4CC363FDFFE46042FB4CF8F26A6AD46C" box="[1227,1305,1852,1873]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="546" unit="mm" value="17.0">17 mm</quantity>
preserved length, leg. 
<collectorName id="26CEABCEFFE46042FC38F8956B5FD44D" box="[959,1068,1882,1904]" pageId="11" pageNumber="546">A. Valdés</collectorName>
(
<collectionCode id="ED2A56DDFFE46042FBB8F8956BFFD44D" box="[1087,1164,1883,1904]" pageId="11" pageNumber="546">CASIZ</collectionCode>
<specimenCode id="DB9D6663FFE46042FB10F8946B9ED44D" box="[1175,1261,1882,1904]" pageId="11" pageNumber="546">121105</specimenCode>
).
</materialsCitation>
<materialsCitation id="3B53C445FFE46045FA83F8946D52D698" ID-GBIF-Occurrence="3323414310" collectingDate="1987-08-16" collectionCode="CASIZ" collectorName="F. M. MacFarland" country="Italy" lastPageId="12" lastPageNumber="547" location="Naples" pageId="11" pageNumber="546" specimenCode="081871" specimenCount="1">
<location id="8EE498C3FFE46042FA83F8946A26D44D" LSID="urn:lsid:plazi:treatment:03927F0EFFE46059FCC9FB9D6F8FD74C:8EE498C3FFE46042FA83F8946A26D44D" box="[1284,1365,1882,1904]" country="Italy" name="Naples" pageId="11" pageNumber="546">Naples</location>
, 
<collectingCountry id="F32C8E88FFE46042FAE3F8946AE9D44D" box="[1380,1434,1882,1904]" name="Italy" pageId="11" pageNumber="546">Italy</collectingCountry>
1902–03, 
<specimenCount id="9D3D0591FFE36045FF7AFABE6EE9D6BB" box="[253,410,1392,1414]" pageId="12" pageNumber="547">one specimen</specimenCount>
, 
<quantity id="4CC363FDFFE36045FE20FABE6E86D6B8" box="[423,501,1392,1413]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="547" unit="mm" value="33.0">33 mm</quantity>
preserved length, leg. 
<collectorName id="26CEABCEFFE36045FF09FA416E2AD699" box="[142,345,1422,1444]" pageId="12" pageNumber="547">F. M. MacFarland</collectorName>
(
<collectionCode id="ED2A56DDFFE36045FEEEFA416EC5D699" box="[361,438,1423,1444]" pageId="12" pageNumber="547">CASIZ</collectionCode>
<specimenCode id="DB9D6663FFE36045FE38FA416D66D698" box="[447,533,1423,1445]" pageId="12" pageNumber="547">081871</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF449E90FFE36045FF09FBB56BFDD63E" ID-DOI="http://doi.org/10.5281/zenodo.4634212" ID-Zenodo-Dep="4634212" httpUri="https://zenodo.org/record/4634212/files/figure.png" pageId="12" pageNumber="547" startId="12.[142,220,1147,1166]" targetBox="[142,1422,195,1115]" targetPageId="12">
<paragraph id="8B84CE18FFE36045FF09FBB56BFDD63E" blockId="12.[142,1423,1147,1284]" pageId="12" pageNumber="547">
<emphasis id="B94F120AFFE36045FF09FBB56F85D7B3" bold="true" box="[142,246,1147,1166]" pageId="12" pageNumber="547">Figure 4.</emphasis>
Living animals. A, 
<taxonomicName id="4C3BB59BFFE36045FE53FBB26DE9D7B3" authorityName="Rapp" authorityYear="1827" box="[468,666,1147,1166]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE36045FE53FBB26DE9D7B3" box="[468,666,1147,1166]" italics="true" pageId="12" pageNumber="547">Doris pseudoargus</emphasis>
</taxonomicName>
(CASIZ 121105). B, 
<taxonomicName id="4C3BB59BFFE36045FC06FBB26B5BD7B3" authorityName="Risbec" authorityYear="1928" box="[897,1064,1147,1166]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="immonda">
<emphasis id="B94F120AFFE36045FC06FBB26B5BD7B3" box="[897,1064,1147,1166]" italics="true" pageId="12" pageNumber="547">Doris immonda</emphasis>
</taxonomicName>
(CASIZ 089023), photo by T. M. Gosliner. C, 
<taxonomicName id="4C3BB59BFFE36045FE88FB576EC4D796" baseAuthorityName="Pease" baseAuthorityYear="1860" box="[271,439,1176,1195]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="granulosa">
<emphasis id="B94F120AFFE36045FE88FB576EC4D796" box="[271,439,1176,1195]" italics="true" pageId="12" pageNumber="547">Doris granulosa</emphasis>
</taxonomicName>
(CASIZ 073536), photo by T. M. Gosliner. D, 
<taxonomicName id="4C3BB59BFFE36045FC17FB566B0BD796" authorityName="Bergh" authorityYear="1877" box="[912,1144,1176,1195]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="boholiensis">
<emphasis id="B94F120AFFE36045FC17FB566B0BD796" box="[912,1144,1176,1195]" italics="true" pageId="12" pageNumber="547">Discodoris boholiensis</emphasis>
</taxonomicName>
(CASIZ 083654), photo by T. M. Gosliner. E, 
<taxonomicName id="4C3BB59BFFE36045FEC8FB786DF8D7F4" authority=", San Pedrillo" authorityName="San Pedrillo" box="[335,651,1206,1225]" class="Gastropoda" family="Discodorididae" genus="Discodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="ketos">
<emphasis id="B94F120AFFE36045FEC8FB786E8AD7F4" box="[335,505,1206,1225]" italics="true" pageId="12" pageNumber="547">Discodoris ketos</emphasis>
, San Pedrillo
</taxonomicName>
, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, 
<taxonomicName id="4C3BB59BFFE36045FB45FB786AFDD7F4" authorityName="Behrens &amp; Henderson" authorityYear="1981" box="[1218,1422,1206,1225]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="rubescens">
<emphasis id="B94F120AFFE36045FB45FB786AFDD7F4" box="[1218,1422,1206,1225]" italics="true" pageId="12" pageNumber="547">Thordisa rubescens</emphasis>
</taxonomicName>
(CASIZ 015860), photo by T. M. Gosliner. G, 
<taxonomicName id="4C3BB59BFFE36045FDD9FB1D6C3DD7DB" authorityName="Bergh" authorityYear="1879" box="[606,846,1235,1254]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="antillensis">
<emphasis id="B94F120AFFE36045FDD9FB1D6C3DD7DB" box="[606,846,1235,1254]" italics="true" pageId="12" pageNumber="547">Aphelodoris antillensis</emphasis>
</taxonomicName>
(CASIZ 077289), photo by T. M. Gosliner. H, 
<taxonomicName id="4C3BB59BFFE36045FAA1FB1D6E68D63E" authorityName="Bergh" authorityYear="1880" class="Gastropoda" family="Discodorididae" genus="Peltodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="atromaculata">
<emphasis id="B94F120AFFE36045FAA1FB1D6E68D63E" italics="true" pageId="12" pageNumber="547">Peltodoris atromaculata</emphasis>
</taxonomicName>
(CASIZ 119474). I, 
<taxonomicName id="4C3BB59BFFE36045FE68FB3E6C65D639" authority=", Monterey" authorityName="Monterey" box="[495,790,1264,1284]" class="Gastropoda" family="Discodorididae" genus="Peltodoris" kingdom="Animalia" order="Nudibranchia" pageId="12" pageNumber="547" phylum="Mollusca" rank="species" species="nobilis">
<emphasis id="B94F120AFFE36045FE68FB3E6DD7D63E" box="[495,676,1264,1283]" italics="true" pageId="12" pageNumber="547">Peltodoris nobilis</emphasis>
, Monterey
</taxonomicName>
Bay, California, photo by A. Smith.
</paragraph>
</caption>
<subSubSection id="C3219D93FFE36046FF09FA246EDDD709" lastPageId="15" lastPageNumber="550" pageId="12" pageNumber="547" type="description">
<paragraph id="8B84CE18FFE36045FF09FA246E04D6C2" blockId="12.[142,758,1514,1904]" box="[142,375,1514,1535]" pageId="12" pageNumber="547">
<emphasis id="B94F120AFFE36045FF09FA246E04D6C2" box="[142,375,1514,1535]" italics="true" pageId="12" pageNumber="547">External morphology</emphasis>
</paragraph>
<paragraph id="8B84CE18FFE36045FF09F9C76A6BD6DF" blockId="12.[142,758,1514,1904]" lastBlockId="12.[806,1423,1392,1628]" pageId="12" pageNumber="547">
The general colour of the living animals varies from yellowish to pale brown, with pale purple, whitish, green, dark brown or reddish irregular patches on the dorsum (
<figureCitation id="1300D29DFFE36045FF7FF9AB6E3FD546" box="[248,332,1637,1659]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.4634212" httpUri="https://zenodo.org/record/4634212/files/figure.png" pageId="12" pageNumber="547">Fig. 4A</figureCitation>
). In some specimens there are only dark brown patches. The rhinophores and the gill are yellowish to pale brown. The whole dorsum is covered with rounded and simple tubercles, all of them similar in size (
<figureCitation id="1300D29DFFE36045FF6EF92E6E4CD5CB" box="[233,319,1760,1782]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="12" pageNumber="547">Fig. 5D</figureCitation>
). The largest tubercles are those situated in the central region of the body. The rhinophoral and branchial sheaths have several tubercles which are slightly stalked but otherwise similar to the rest of the dorsal tubercles. There are 8–9 tripinnate branchial leaves, forming a circle. The anal papilla is prominent, situated in the centre of the branchial circle of leaves. The rhinophores are elongate, having 14 lamellae in a 17-mm preserved length specimen.
</paragraph>
<paragraph id="8B84CE18FFE36045FCB8FA246CC6D561" blockId="12.[806,1423,1392,1628]" pageId="12" pageNumber="547">
Ventrally, there are no oral tentacles, but two blunt prolongations on each side of the mouth opening (
<figureCitation id="1300D29DFFE36045FCA9F9E66CF2D503" box="[814,897,1576,1598]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="12" pageNumber="547">Fig. 6E</figureCitation>
). The anterior border of the foot is grooved but not notched.
</paragraph>
<paragraph id="8B84CE18FFE36045FCA1F96C6CFAD58A" blockId="12.[806,1422,1698,1904]" box="[806,905,1698,1719]" pageId="12" pageNumber="547">
<emphasis id="B94F120AFFE36045FCA1F96C6CFAD58A" box="[806,905,1698,1719]" italics="true" pageId="12" pageNumber="547">Anatomy</emphasis>
</paragraph>
<paragraph id="8B84CE18FFE36047FCA1F90F6CFCD40E" blockId="12.[806,1422,1698,1904]" lastBlockId="14.[806,1422,1729,1904]" lastPageId="14" lastPageNumber="549" pageId="12" pageNumber="547">
The posterior end of the glandular portion of the oral tube has six strong retractor muscles (
<figureCitation id="1300D29DFFE36045FB60F92E6A49D5CB" box="[1255,1338,1760,1782]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="12" pageNumber="547">Fig. 6D</figureCitation>
) which attach to the body wall. Two long salivary glands connect with the buccal bulb at each side of the oesophageal junction. The buccal bulb is several times longer than the glandular portion of the oral tube. The labial cuticle is smooth. The radular formula is 41 ¥ 73.0. 
<quantity id="4CC363FDFFE16047FD5CF90F6FD6D5CB" metricMagnitude="0" metricUnit="m" metricValue="1.8542" pageId="14" pageNumber="549" unit="in" value="73.0">73 in</quantity>
a 33-mm long specimen. Rachidian teeth are absent. The lateral teeth are narrow and elongate, having a single cusp and lacking denticles (
<figureCitation id="1300D29DFFE16047FD11F8D36D98D40E" box="[662,747,1821,1843]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="14" pageNumber="549">Fig. 5A</figureCitation>
). The teeth from the middle portion of the half-row are larger than those closer to the medial portion of the radula (
<figureCitation id="1300D29DFFE16047FC06F90F6CA6D5EA" box="[897,981,1729,1751]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="14" pageNumber="549">Fig. 5B</figureCitation>
). The outermost teeth are smaller and have a number of thin denticles (
<figureCitation id="1300D29DFFE16047FB18F92E6B80D5CB" box="[1183,1267,1760,1782]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="14" pageNumber="549">Fig. 5C</figureCitation>
). The oesophagus is short and connects directly to the stomach (
<figureCitation id="1300D29DFFE16047FCA9F8D36CF0D40E" box="[814,899,1821,1843]" captionStart="Figure 5" captionStartId="13.[155,233,1731,1750]" captionTargetBox="[155,1435,194,1698]" captionTargetId="figure-75@13.[155,1435,194,1698]" captionTargetPageId="13" captionText="Figure 5. Doris pseudoargus (CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="14" pageNumber="549">Fig. 5A</figureCitation>
).
</paragraph>
<caption id="DF449E90FFE26044FF1CF90D6E5FD42C" ID-DOI="http://doi.org/10.5281/zenodo.4634216" ID-Zenodo-Dep="4634216" httpUri="https://zenodo.org/record/4634216/files/figure.png" pageId="13" pageNumber="548" startId="13.[155,233,1731,1750]" targetBox="[155,1435,194,1698]" targetPageId="13">
<paragraph id="8B84CE18FFE26044FF1CF90D6E5FD42C" blockId="13.[155,1435,1731,1809]" pageId="13" pageNumber="548">
<emphasis id="B94F120AFFE26044FF1CF90D6E70D5EB" bold="true" box="[155,259,1731,1750]" pageId="13" pageNumber="548">Figure 5.</emphasis>
<taxonomicName id="4C3BB59BFFE26044FE88F90A6EA3D5EB" authorityName="Rapp" authorityYear="1827" box="[271,464,1731,1750]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="548" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE26044FE88F90A6EA3D5EB" box="[271,464,1731,1750]" italics="true" pageId="13" pageNumber="548">Doris pseudoargus</emphasis>
</taxonomicName>
(CASIZ 081871), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 75 Mm. C, outer lateral teeth; scale bar = 43 Mm. D, dorsal tubercles; scale bar = 750 Mm.
</paragraph>
</caption>
<caption id="DF449E90FFE16047FF09F9D36A51D551" ID-DOI="http://doi.org/10.5281/zenodo.4634218" ID-Zenodo-Dep="4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="14" pageNumber="549" startId="14.[142,220,1565,1584]" targetBox="[330,1251,199,1529]" targetPageId="14">
<paragraph id="8B84CE18FFE16047FF09F9D36A51D551" blockId="14.[142,1423,1565,1644]" pageId="14" pageNumber="549">
<emphasis id="B94F120AFFE16047FF09F9D36F85D50C" bold="true" box="[142,246,1565,1585]" pageId="14" pageNumber="549">Figure 6.</emphasis>
<taxonomicName id="4C3BB59BFFE16047FE8EF9D06EB8D50C" authorityName="Rapp" authorityYear="1827" box="[265,459,1566,1585]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="549" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE16047FE8EF9D06EB8D50C" box="[265,459,1566,1585]" italics="true" pageId="14" pageNumber="549">Doris pseudoargus</emphasis>
</taxonomicName>
(CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8B84CE18FFE16046FCB8F8F26D91D131" blockId="14.[806,1422,1729,1904]" lastBlockId="15.[155,771,196,1076]" lastPageId="15" lastPageNumber="550" pageId="14" pageNumber="549">
The ampulla is convoluted and branches into a short oviduct and the prostate (
<figureCitation id="1300D29DFFE16047FBD7F8946BD7D44D" box="[1104,1188,1882,1904]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="14" pageNumber="549">Fig. 6C</figureCitation>
). The oviduct enters the female gland mass near to its centre. The prostate is tubular, very long, folded and granular (
<figureCitation id="1300D29DFFE06046FD00FF2C6DA8D3C5" box="[647,731,226,248]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="15" pageNumber="550">Fig. 6B</figureCitation>
). It connects with a long duct that narrows and expands again into the huge ejaculatory portion of the deferent duct. The muscular deferent duct opens into a short common atrium with the vagina. The vagina is long and wide. Near to its proximal end it joins the duct connecting the bursa copulatrix and the seminal receptacle. The uterine duct also leads from this duct. The bursa copulatrix is irregular in shape, about 10 times larger than the seminal receptacle (
<figureCitation id="1300D29DFFE06046FD06FE386DA6D131" box="[641,725,502,524]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="15" pageNumber="550">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="8B84CE18FFE06046FF33FDD86E2CD0E5" blockId="15.[155,771,196,1076]" pageId="15" pageNumber="550">
In the central nervous system (
<figureCitation id="1300D29DFFE06046FD9EFDDB6D19D116" box="[537,618,533,555]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="15" pageNumber="550">Fig. 6F</figureCitation>
) the cerebral and pleural ganglia are fused and distinct from the pedal ganglia. There are four cerebral nerves leading from each cerebral ganglion, and three pleural nerves leading from the left pleural ganglion and two from the right one. There is no separate abdominal ganglion on the right side of the visceral loop. The buccal ganglia are near to the rest of the central nervous system, joined to the cerebral ganglia by two relatively short nerves. Gastro-oesophageal, rhinophoral and optical ganglia are present. The pedal ganglia are clearly separated, having five nerves leading from the left ganglion and four from the right one. The pedal and parapedal commissures are enveloped together with the visceral loop.
</paragraph>
<paragraph id="8B84CE18FFE06046FF33FC2F6EDDD709" blockId="15.[155,771,196,1076]" pageId="15" pageNumber="550">
The circulatory system (
<figureCitation id="1300D29DFFE06046FE49FC2F6D52D0CA" box="[462,545,993,1015]" captionStart="Figure 6" captionStartId="14.[142,220,1565,1584]" captionTargetBox="[330,1251,199,1529]" captionTargetId="figure-198@14.[302,1262,179,1533]" captionTargetPageId="14" captionText="Figure 6. Doris pseudoargus (CASIZ 081871). A, general view of the anatomy; scale bar = 2 mm. B, reproductive system; scale bar = 2 mm. C, detail of several reproductive organs; scale bar = 2 mm. D, lateral view of the buccal bulb; scale bar = 2 mm. E, ventral view of the mouth area; scale bar = 5 mm. E, central nervous system; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634218" httpUri="https://zenodo.org/record/4634218/files/figure.png" pageId="15" pageNumber="550">Fig. 6A</figureCitation>
) consists of a large heart and a single large blood gland situated over the central nervous system.
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFE06059FF1CFBB46F8FD74C" lastPageId="16" lastPageNumber="551" pageId="15" pageNumber="550" type="discussion">
<paragraph id="8B84CE18FFE06046FF1CFBB46F8ED7B2" blockId="15.[155,771,1146,1904]" box="[155,253,1146,1167]" pageId="15" pageNumber="550">
<emphasis id="B94F120AFFE06046FF1CFBB46F8ED7B2" box="[155,253,1146,1167]" italics="true" pageId="15" pageNumber="550">Remarks</emphasis>
</paragraph>
<paragraph id="8B84CE18FFE06046FF1CFB546D4BD699" blockId="15.[155,771,1146,1904]" pageId="15" pageNumber="550">
<taxonomicName id="4C3BB59BFFE06046FF1CFB546E8DD793" authority="Muller, 1778" authorityName="Muller" authorityYear="1778" box="[155,510,1177,1199]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FF1CFB546E13D793" box="[155,352,1177,1199]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
<bibRefCitation id="EFAAB3E9FFE06046FEECFB576E8DD793" author="Muller" box="[363,510,1177,1199]" firstAuthor="Muller" pageId="15" pageNumber="550" pagination="48 - 54" refId="ref57558" refString="Muller OF. 1778. Molluscorum marinorum Norvagiae, Decas 1. Nova Acta Physico-Medica Academiae Cesariae Leopoldino-Carolinae 6: 48 - 54." type="journal article" year="1778">Müller, 1778</bibRefCitation>
</taxonomicName>
was described on the basis of an undetermined number of specimens collected from 
<collectingCountry id="F32C8E88FFE06046FEA1FB196E0CD7D1" box="[294,383,1239,1260]" name="Norway" pageId="15" pageNumber="550">Norway</collectingCountry>
. 
<bibRefCitation id="EFAAB3E9FFE06046FE0CFB186D5AD7D0" author="Muller" box="[395,553,1238,1261]" firstAuthor="Muller" pageId="15" pageNumber="550" pagination="48 - 54" refId="ref57558" refString="Muller OF. 1778. Molluscorum marinorum Norvagiae, Decas 1. Nova Acta Physico-Medica Academiae Cesariae Leopoldino-Carolinae 6: 48 - 54." type="journal article" year="1778">Müller (1778)</bibRefCitation>
described the animals as golden, patelliform, with the dorsum covered with numerous hair-like yellowish tubercles. The description of the animals clearly represents a species of phanerobranch dorid, probably a member of the genera 
<taxonomicName id="4C3BB59BFFE06046FF76FABE6D4DD6BB" authority="J. E. Gray, 1850" authorityName="J. E. Gray" authorityYear="1850" box="[241,574,1392,1414]" class="Gastropoda" family="Dorididae" genus="Acanthodoris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FF76FABE6EF7D6B8" box="[241,388,1392,1413]" italics="true" pageId="15" pageNumber="550">Acanthodoris</emphasis>
J.E. 
<bibRefCitation id="EFAAB3E9FFE06046FE47FABE6D4DD6BB" author="Gray ME" bookContentInfo="3 plates" box="[448,574,1392,1414]" journalOrPublisher="London: Longman, Brown, Green and Longmans" pageId="15" pageNumber="550" pagination="1 - 124" part="4" refId="ref56447" refString="Gray ME. 1842 - 1850. Figures of Molluscous Animals, Selected from Various Authors. Etched for the Use of Students. London: Longman, Brown, Green and Longmans, Vol. 1, plates 1 - 78 [1842], Vol. 2. plates 79 - 199 [1850], Vol. 3 plates 200 - 312 [1850], Vol. 4, pp. 1 - 124 [1850]." title="Figures of Molluscous Animals, Selected from Various Authors. Etched for the Use of Students" type="book chapter" year="1850">Gray, 1850</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3BB59BFFE06046FDCBFABE6FA1D699" authority="Bergh, 1878" authorityName="Bergh" authorityYear="1878" class="Gastropoda" family="Dorididae" genus="Adalaria" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FDCBFABE6DC3D6B8" box="[588,688,1392,1413]" italics="true" pageId="15" pageNumber="550">Adalaria</emphasis>
<bibRefCitation id="EFAAB3E9FFE06046FD31FABE6FA1D699" author="Bergh R" pageId="15" pageNumber="550" pagination="62 - 65" refId="ref53874" refString="Bergh R. 1878 a. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 2, heft 13. Wiesbaden: Kreidel, 547 - 602, plates 62 - 65." type="book chapter" year="1878">Bergh, 1878</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="4C3BB59BFFE06046FF7CFA406D40D699" authority="Blainville, 1816" authorityName="Blainville" authorityYear="1816" box="[251,563,1422,1444]" class="Gastropoda" family="Dorididae" genus="Onchidoris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FF7CFA406E06D69E" box="[251,373,1422,1443]" italics="true" pageId="15" pageNumber="550">Onchidoris</emphasis>
Blainville, 1816
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFE06046FF33FA606E54D429" blockId="15.[155,771,1146,1904]" pageId="15" pageNumber="550">
Years later, 
<bibRefCitation id="EFAAB3E9FFE06046FEBEFA636EA6D6FE" author="Cuvier" box="[313,469,1453,1475]" firstAuthor="Cuvier" pageId="15" pageNumber="550" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier (1804)</bibRefCitation>
reported 
<taxonomicName id="4C3BB59BFFE06046FDC6FA606E5FD6DC" authority="Muller, 1778" authorityName="Muller" authorityYear="1778" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FDC6FA606C71D6FF" box="[577,770,1453,1475]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
<bibRefCitation id="EFAAB3E9FFE06046FF1CFA026E5FD6DC" author="Muller" box="[155,300,1484,1506]" firstAuthor="Muller" pageId="15" pageNumber="550" pagination="48 - 54" refId="ref57558" refString="Muller OF. 1778. Molluscorum marinorum Norvagiae, Decas 1. Nova Acta Physico-Medica Academiae Cesariae Leopoldino-Carolinae 6: 48 - 54." type="journal article" year="1778">Müller, 1778</bibRefCitation>
</taxonomicName>
from the Atlantic coast of 
<collectingCountry id="F32C8E88FFE06046FDEFFA026DCBD6DC" box="[616,696,1484,1505]" name="France" pageId="15" pageNumber="550">France</collectingCountry>
based on two newly collected specimens, but indicating that his material was clearly different from 
<bibRefCitation id="EFAAB3E9FFE06046FD21F9C76F96D503" author="Muller's" firstAuthor="Muller's" pageId="15" pageNumber="550" pagination="48 - 54" refId="ref57558" refString="Muller OF. 1778. Molluscorum marinorum Norvagiae, Decas 1. Nova Acta Physico-Medica Academiae Cesariae Leopoldino-Carolinae 6: 48 - 54." type="journal article" year="1778">Müller’s (1778)</bibRefCitation>
. The animals described by Cuvier are large cryptobranch dorids with the dorsum covered with rounded tubercles. 
<bibRefCitation id="EFAAB3E9FFE06046FE13F9A86D5DD546" author="Rapp" box="[404,558,1637,1659]" firstAuthor="Rapp" pageId="15" pageNumber="550" pagination="515 - 522" refId="ref58646" refString="Rapp WL. 1827. Uber das Molluskengeschlecht Doris und Beschreibung einiger neuen Arten desselben. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 13: 515 - 522, plates 26 - 27." type="journal article" year="1827">Rapp (1827)</bibRefCitation>
described 
<taxonomicName id="4C3BB59BFFE06046FD40F9A86E55D5A4" authorityName="Rapp" authorityYear="1827" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE06046FD40F9A86E55D5A4" italics="true" pageId="15" pageNumber="550">Doris pseudoargus</emphasis>
</taxonomicName>
from Le Havre, 
<collectingCountry id="F32C8E88FFE06046FE72F94A6D35D5A4" box="[501,582,1668,1689]" name="France" pageId="15" pageNumber="550">France</collectingCountry>
, with the same characteristics of the specimens studied by 
<bibRefCitation id="EFAAB3E9FFE06046FD32F96C6F96D5EA" author="Cuvier" firstAuthor="Cuvier" pageId="15" pageNumber="550" pagination="447 - 473" refId="ref55151" refString="Cuvier GL. 1804. Memoire sur le genre Doris. Annales du Museum National d'Histoire Naturelle 4: 447 - 473, plates 73 - 74." type="journal article" year="1804">Cuvier (1804)</bibRefCitation>
: ‘ash colour with dull reddish spots’, and therefore this is the first valid introduction of a name for this species.
</paragraph>
<paragraph id="8B84CE18FFE06046FF33F8D36A3BD3C4" blockId="15.[155,771,1146,1904]" lastBlockId="15.[819,1436,196,1904]" pageId="15" pageNumber="550">
<bibRefCitation id="EFAAB3E9FFE06046FF33F8D36E18D40E" author="Johnston" box="[180,363,1821,1843]" firstAuthor="Johnston" pageId="15" pageNumber="550" pagination="44 - 56" refId="ref56705" refString="Johnston G. 1838. Miscellanea zoologica. Annals of Natural History; or, Magazine of Zoology, Botany, and Geology 1: 44 - 56." type="journal article" year="1838">Johnston (1838)</bibRefCitation>
introduced the names 
<taxonomicName id="4C3BB59BFFE06046FDF6F8D06C70D40F" box="[625,771,1821,1843]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="britannica">
<emphasis id="B94F120AFFE06046FDF6F8D06C70D40F" box="[625,771,1821,1843]" italics="true" pageId="15" pageNumber="550">D. britannica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3BB59BFFE06046FF4BF8F36E27D46C" box="[204,340,1852,1874]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="montagui">
<emphasis id="B94F120AFFE06046FF4BF8F36E27D46C" box="[204,340,1852,1874]" italics="true" pageId="15" pageNumber="550">D. montagui</emphasis>
</taxonomicName>
, without a description and in the synonymy of 
<taxonomicName id="4C3BB59BFFE06046FE96F8956ECBD452" authorityName="Muller" authorityYear="1778" box="[273,440,1882,1904]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FE96F8956ECBD452" box="[273,440,1882,1904]" italics="true" pageId="15" pageNumber="550">D. Tuberculata</emphasis>
</taxonomicName>
. Therefore they are 
<emphasis id="B94F120AFFE06046FD29F8956C1ED3E4" box="[686,877,196,1904]" italics="true" pageId="15" pageNumber="550">nomina nuda</emphasis>
and if they have not been used as valid before 1961 they are also not available (
<bibRefCitation id="EFAAB3E9FFE06046FB33FF2D6A4ED3C4" author="ICZN" box="[1204,1341,227,249]" firstAuthor="ICZN" pageId="15" pageNumber="550" refId="ref56591" refString="ICZN. 1999. International Code of Zoological Nomenclature. London: International Trust for Zoological Nomenclature." type="book" year="1999">ICZN, 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B84CE18FFE06046FCCBFECC6CF9D131" blockId="15.[819,1436,196,1904]" pageId="15" pageNumber="550">
In the following years most authors referred to this species as 
<taxonomicName id="4C3BB59BFFE06046FC64FEEE6BD6D208" authorityName="Muller" authorityYear="1778" box="[995,1189,288,309]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FC64FEEE6BD6D208" box="[995,1189,288,309]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
, but with authorship of Cuvier. Examples include 
<bibRefCitation id="EFAAB3E9FFE06046FB24FEF06AE5D268" author="Delle Chiaje" box="[1187,1430,318,341]" firstAuthor="Delle Chiaje" pageId="15" pageNumber="550" refId="ref55179" refString="Delle Chiaje S. 1841. Animale senza vetebre del Regno di Napoli. Naples: Batelli." type="book" year="1841">Delle Chiaje (1841)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FCB4FE936C9FD24E" author="Bergh" box="[819,1004,349,371]" firstAuthor="Bergh" pageId="15" pageNumber="550" pagination="66 - 68" refId="ref53915" refString="Bergh R. 1878 b. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 2, heft 14. Wiesbaden: Kreidel, 603 - 645, plates 66 - 68." type="book chapter" year="1878" yearSuffix="b">Bergh (1878b)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB8BFE936BD9D24E" author="Eliot" box="[1036,1194,349,371]" firstAuthor="Eliot" pageId="15" pageNumber="550" pagination="411 - 438" refId="ref55677" refString="Eliot CN. 1910. Nudibranchs collected by Mr. Stanley Gardiner from the Indian Ocean in H. M. S. Sealark in Percy Sladen trust expedition to the Indian Ocean in 1905, under the leadership of Mr. J Stanley Gardiner 2. Transactions of the Linnaean Society of London, Zoology 13: 411 - 438." type="journal article" year="1910">Eliot (1910)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB4DFE936AE8D24E" author="Vayssiere" box="[1226,1435,349,371]" firstAuthor="Vayssiere" pageId="15" pageNumber="550" pagination="11" refId="ref59908" refString="Vayssiere A. 1913. Mollusques de la France et des regions voisines, t. 1. Amphineures, Gasteropodes opisthobranches, Heteropodes, Marseniades et Oncidiides. In: Richard J, ed. Encyclopedie Scientifique; Bibliotheque d'Oceanographie Physique, 11. Paris: O. Doin et fils." type="book chapter" year="1913">Vayssière (1913)</bibRefCitation>
<bibRefCitation id="EFAAB3E9FFE06046FCB4FEB26B7DD2AF" author="O'Donoghue" box="[819,1038,380,402]" firstAuthor="O'Donoghue" pageId="15" pageNumber="550" pagination="713 - 841" refId="ref57773" refString="O'Donoghue CH. 1929. Report on the Opisthobranchiata. Zoological results of the Cambridge Expedition to the Suez Canal, 1924 - 38. Transactions of the Zoological Society of London 22: 713 - 841." type="journal article" year="1929">O’Donoghue (1929)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB9DFEB26B90D2AF" author="Pruvot-Fol" box="[1050,1251,380,402]" firstAuthor="Pruvot-Fol" pageId="15" pageNumber="550" pagination="209 - 261" refId="ref58562" refString="Pruvot-Fol A. 1935. Les doridiens de Cuvier publies dans les Annales du Museum en 1804, etude critique et historique. Journal de Conchyliologie 78: 209 - 261." type="journal article" year="1935">Pruvot-Fol (1935)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB68FEB26AE4D2AF" author="Odhner" box="[1263,1431,380,402]" firstAuthor="Odhner" pageId="15" pageNumber="550" pagination="1 - 92" refId="ref57662" refString="Odhner N. 1939. Opisthobranchiate Mollusca from the western and northern coasts of Norway. Det Kgl Norske Videnskabers Selskabs Skrifter 1939 (1): 1 - 92." type="journal article" year="1939">Odhner (1939)</bibRefCitation>
. The scientific influence of Cuvier’s papers probably explains why subsequently many authors applied the name 
<taxonomicName id="4C3BB59BFFE06046FC20FE166B1AD2D0" authorityName="Muller" authorityYear="1778" box="[935,1129,472,493]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FC20FE166B1AD2D0" box="[935,1129,472,493]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
to this cryptobranch dorid species.
</paragraph>
<paragraph id="8B84CE18FFE06046FCCBFDDB6B98D0E5" blockId="15.[819,1436,196,1904]" pageId="15" pageNumber="550">
The usage of the name 
<taxonomicName id="4C3BB59BFFE06046FBC9FDD86A7CD117" authorityName="Muller" authorityYear="1778" box="[1102,1295,533,555]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FBC9FDD86A7CD117" box="[1102,1295,533,555]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
for this species was challenged by the British School. Early on, 
<bibRefCitation id="EFAAB3E9FFE06046FCB4FD9C6BE3D154" author="Iredale &amp; O'Donoghue" box="[819,1168,594,617]" firstAuthor="Iredale" pageId="15" pageNumber="550" pagination="195 - 233" refId="ref56678" refString="Iredale T, O'Donoghue CH. 1923. List of British nudibranchiate Mollusca. Proceedings of the Malacological Society of London 15: 195 - 233." type="journal article" year="1923">Iredale &amp; O’Donoghue (1923)</bibRefCitation>
for some unexplained reason decided that the animals named 
<taxonomicName id="4C3BB59BFFE06046FB7AFDBC6C2CD198" authorityName="Muller" authorityYear="1778" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FB7AFDBC6C2CD198" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
by Cuvier are a different species from specimens identified as such by Alder &amp; Hancock and Eliot; they used the unavailable name 
<taxonomicName id="4C3BB59BFFE06046FBECFD006A51D1DF" box="[1131,1314,717,739]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="britannica">
<emphasis id="B94F120AFFE06046FBECFD006A51D1DF" box="[1131,1314,717,739]" italics="true" pageId="15" pageNumber="550">Doris britannica</emphasis>
</taxonomicName>
, combined with the genus name 
<taxonomicName id="4C3BB59BFFE06046FBABFD226BD3D03C" authorityName="Bergh" authorityYear="1878" box="[1068,1184,748,769]" class="Gastropoda" family="Discodorididae" genus="Archidoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FBABFD226BD3D03C" box="[1068,1184,748,769]" italics="true" pageId="15" pageNumber="550">Archidoris</emphasis>
</taxonomicName>
, for the latter. On the other hand, 
<bibRefCitation id="EFAAB3E9FFE06046FC41FCC46BE0D01C" author="Pruvot-Fol" box="[966,1171,778,801]" firstAuthor="Pruvot-Fol" pageId="15" pageNumber="550" pagination="746 - 755" refId="ref58491" refString="Pruvot-Fol A. 1931. Notes de systematique sur les opisthobranches. Bulletin du Museum National d'Histoire Naturelle 2 (8): 746 - 755." type="journal article" year="1931">Pruvot-Fol (1931)</bibRefCitation>
argued that all these animals belonged to the same species - 
<taxonomicName id="4C3BB59BFFE06046FB7AFCE46C2CD060" authorityName="Muller" authorityYear="1778" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FB7AFCE46C2CD060" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
with authorship of Cuvier the valid name. The name 
<taxonomicName id="4C3BB59BFFE06046FC06FCA96B33D046" box="[897,1088,870,892]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="britannica">
<emphasis id="B94F120AFFE06046FC06FCA96B33D046" box="[897,1088,870,892]" italics="true" pageId="15" pageNumber="550">Doris britannica</emphasis>
</taxonomicName>
very rarely appears in the literature. 
<bibRefCitation id="EFAAB3E9FFE06046FC34FC4B6B0DD0A6" author="Thompson" box="[947,1150,901,923]" firstAuthor="Thompson" pageId="15" pageNumber="550" pagination="83 - 84" refId="ref59564" refString="Thompson TE. 1966. Development and life history of Archidoris pseudoargus. Malacologia 5: 83 - 84." type="journal article" year="1966">Thompson (1966)</bibRefCitation>
reintroduced the usage of the name 
<taxonomicName id="4C3BB59BFFE06046FC57FC6A6BD0D084" authorityName="Rapp" authorityYear="1827" box="[976,1187,932,953]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE06046FC57FC6A6BD0D084" box="[976,1187,932,953]" italics="true" pageId="15" pageNumber="550">Doris pseudoargus</emphasis>
</taxonomicName>
, also combined with 
<taxonomicName id="4C3BB59BFFE06046FCB4FC0C6CD4D0EA" authorityName="Bergh" authorityYear="1878" box="[819,935,962,983]" class="Gastropoda" family="Discodorididae" genus="Archidoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FCB4FC0C6CD4D0EA" box="[819,935,962,983]" italics="true" pageId="15" pageNumber="550">Archidoris</emphasis>
</taxonomicName>
, but without a justification.
</paragraph>
<paragraph id="8B84CE18FFE06046FCCBFC2F6CCAD500" blockId="15.[819,1436,196,1904]" pageId="15" pageNumber="550">
Both 
<taxonomicName id="4C3BB59BFFE06046FC0CFC2C6B28D0CB" authorityName="Rapp" authorityYear="1827" box="[907,1115,993,1015]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE06046FC0CFC2C6B28D0CB" box="[907,1115,993,1015]" italics="true" pageId="15" pageNumber="550">Doris pseudoargus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3BB59BFFE06046FB1DFC2C6A2DD0CB" authorityName="Muller" authorityYear="1778" box="[1178,1374,993,1015]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FB1DFC2C6A2DD0CB" box="[1178,1374,993,1015]" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
have been equally used in modern literature, usually combined with the genus name 
<taxonomicName id="4C3BB59BFFE06046FB02FBD06B8AD70E" authorityName="Bergh" authorityYear="1878" box="[1157,1273,1054,1075]" class="Gastropoda" family="Discodorididae" genus="Archidoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFE06046FB02FBD06B8AD70E" box="[1157,1273,1054,1075]" italics="true" pageId="15" pageNumber="550">Archidoris</emphasis>
</taxonomicName>
. Examples of the former in taxonomic papers include 
<bibRefCitation id="EFAAB3E9FFE06046FAABFBF36B5BD74F" author="Schmekel &amp; Portmann" firstAuthor="Schmekel" pageId="15" pageNumber="550" refId="ref59291" refString="Schmekel L, Portmann A. 1982. Opisthobranchia des Mittelmeeres. Nudibranchia und Sacoglossa. Berlin: Springer-Verlag." type="book" year="1982">Schmekel &amp; Portmann (1982)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FBB9FB926AE4D74F" author="Thompson &amp; Brown" box="[1086,1431,1116,1138]" firstAuthor="Thompson" pageId="15" pageNumber="550" refId="ref59658" refString="Thompson TE, Brown GH. 1984. Biology of Opisthobranch Molluscs, 2. London: The Ray Society." type="book" year="1984">Thompson &amp; Brown (1984)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FCB4FBB46B07D7AC" author="Cattaneo-Vietti" box="[819,1140,1146,1169]" etAl="et al." firstAuthor="Cattaneo-Vietti" pageId="15" pageNumber="550" refId="ref54881" refString="Cattaneo-Vietti R, Chemello R, Giannuzzi-Savelli R. 1990. Atlas of Mediterranean nudibranchs. Rome: La Conchiglia, 14 plates." type="book" year="1990">
Cattaneo-Vietti 
<emphasis id="B94F120AFFE06046FC6BFBB56B6FD7B2" box="[1004,1052,1146,1168]" italics="true" pageId="15" pageNumber="550">et al</emphasis>
. (1990)
</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB07FBB46AE5D7AC" author="Picton &amp; Morrow" box="[1152,1430,1146,1169]" firstAuthor="Picton" pageId="15" pageNumber="550" refId="ref58435" refString="Picton BE, Morrow CC. 1994. A Field Guide to the Nudibranchs of the British Isles. London: Immel." type="book" year="1994">Picton &amp; Morrow (1994)</bibRefCitation>
; examples of the latter include 
<bibRefCitation id="EFAAB3E9FFE06046FB2CFB546A59D792" author="Ros" box="[1195,1322,1177,1199]" firstAuthor="Ros" pageId="15" pageNumber="550" pagination="269 - 371" refId="ref58779" refString="Ros J. 1975. Opistobranquios (Gastropoda. Euthyneura) del litoral Iberico. Investigaciones Pesqueras 39: 269 - 371." type="journal article" year="1975">Ros (1975)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FABCFB576C0FD7F3" author="Barletta" firstAuthor="Barletta" pageId="15" pageNumber="550" pagination="1 - 124" refId="ref53717" refString="Barletta G. 1981. Gasteropodi nudi (Pleurobranchomorpha, Sacoglossa, Aplysiomorpha e Nudibranchia). Quaderni della Civica Stazione Idrobiologica di Milano 9: 1 - 124." type="journal article" year="1981">Barletta (1981)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FC00FB766BC5D7F3" author="Swennen &amp; Dekker" box="[903,1206,1208,1230]" firstAuthor="Swennen" pageId="15" pageNumber="550" pagination="1 - 52" refId="ref59368" refString="Swennen C, Dekker R. 1987. De Nederlandse Zeenaaktslakken (Gastropoda Opisthobranchia: Sacoglossa en Nudibranchia). Wetenschappelijke Mededelingen Koninklijke Nederlandse Natuurhistorische Vereniging 183: 1 - 52." type="journal article" year="1987">Swennen &amp; Dekker (1987)</bibRefCitation>
, 
<bibRefCitation id="EFAAB3E9FFE06046FB46FB766B3ED7D0" author="Sabelli B &amp; Giannuzzi-Savelli R &amp; Bedulli D" pageId="15" pageNumber="550" refId="ref59088" refString="Sabelli B, Giannuzzi-Savelli R, Bedulli D. 1990. Catalogo annotato dei molluschi marini del Mediterraneo. Annotated Check-List of Mediterranean Marine Mollusks, 1. Bologna: Societa Italiana di Malacologia and Edizioni Libreria Nauralistica Bolognese." type="journal volume" year="1990">Sabelli, Giannuzzi-Savelli &amp; Bedulli (1990)</bibRefCitation>
. In addition, most papers on physiology, ecology or histology of this species have used the former (
<bibRefCitation id="EFAAB3E9FFE06046FC78FADA6BB3D617" author="Thompson" box="[1023,1216,1300,1322]" firstAuthor="Thompson" pageId="15" pageNumber="550" pagination="83 - 84" refId="ref59564" refString="Thompson TE. 1966. Development and life history of Archidoris pseudoargus. Malacologia 5: 83 - 84." type="journal article" year="1966">Thompson, 1966</bibRefCitation>
; 
<bibRefCitation id="EFAAB3E9FFE06046FB49FADA6A3FD614" author="Rose" box="[1230,1356,1300,1321]" firstAuthor="Rose" pageId="15" pageNumber="550" pagination="185 - 204" refId="ref58802" refString="Rose RM. 1971. Patterned activity of the buccal ganglion of the nudibranch mollusc Archidoris pseudoargus. Journal of Experimental Biology 55: 185 - 204." type="journal article" year="1971">Rose, 1971</bibRefCitation>
; 
<bibRefCitation id="EFAAB3E9FFE06046FADDFADA6C1FD675" author="Potts" firstAuthor="Potts" pageId="15" pageNumber="550" pagination="399 - 407" refId="ref58458" refString="Potts GW. 1983. The respiration of Onchidoris bilamellata and Archidoris pseudoargus (Doridacea). Journal of the Marine Biological Association of the United Kingdom 63: 399 - 407." type="journal article" year="1983">Potts, 1983</bibRefCitation>
; 
<bibRefCitation id="EFAAB3E9FFE06046FCF0FAFD6C8DD675" author="Jonas" box="[887,1022,1330,1352]" firstAuthor="Jonas" pageId="15" pageNumber="550" pagination="207 - 216" refId="ref56734" refString="Jonas M. 1986. Ultrastructure of the gill epithelia of the dorid nudibranchs Archidoris pseudoargus (von Rapp, 1827) and Peltodoris atromaculata Bergh, 1880 (Gastropoda: Opisthobranchia). The Veliger 29: 207 - 216." type="journal article" year="1986">Jonas, 1986</bibRefCitation>
), whereas biochemistry papers have used the latter (
<bibRefCitation id="EFAAB3E9FFE06046FC72FA9F6BA3D65A" author="Cimino" box="[1013,1232,1361,1383]" etAl="et al." firstAuthor="Cimino" pageId="15" pageNumber="550" pagination="1642 - 1646" refId="ref55028" refString="Cimino G, Crispino A, Gavagnin M, Trivellone E, Zubia E, Martinez E, Ortca J. 1993. Archidorin - a new ichthyotoxic diacylglycerol from the Atlantic dorid nudibranch Archidoris tuberculata. Journal of Natural Products 56: 1642 - 1646." type="journal article" year="1993">
Cimino 
<emphasis id="B94F120AFFE06046FBD3FA9C6BF7D65B" box="[1108,1156,1361,1383]" italics="true" pageId="15" pageNumber="550">et al</emphasis>
. 1993
</bibRefCitation>
). In no cases did authors specify their reasons for using one or the other name, which increased the general confusion. Because both names are currently in use, the maintenance of the usage of the valid name for this species, 
<taxonomicName id="4C3BB59BFFE06046FAD8FA026CCDD6C2" authorityName="Rapp" authorityYear="1827" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE06046FAD8FA026CCDD6C2" italics="true" pageId="15" pageNumber="550">Doris pseudoargus</emphasis>
</taxonomicName>
, would certainly not cause a larger disruption than the validation of the name 
<taxonomicName id="4C3BB59BFFE06046FAD8F9C46CC1D500" authorityName="Muller" authorityYear="1778" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FAD8F9C46CC1D500" italics="true" pageId="15" pageNumber="550">Doris tuberculata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFE06059FCCBF9896EEDD3C5" blockId="15.[819,1436,196,1904]" lastBlockId="16.[142,759,196,1137]" lastPageId="16" lastPageNumber="551" pageId="15" pageNumber="550">
<taxonomicName id="4C3BB59BFFE06046FCCBF9896B50D566" authorityName="Rapp" authorityYear="1827" box="[844,1059,1606,1628]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFE06046FCCBF9896B50D566" box="[844,1059,1606,1628]" italics="true" pageId="15" pageNumber="550">Doris pseudoargus</emphasis>
</taxonomicName>
is a well-known species that ranges from Nordkapp (
<collectingCountry id="F32C8E88FFE06046FBD9F9AB6BC8D546" box="[1118,1211,1637,1659]" name="Norway" pageId="15" pageNumber="550">Norway</collectingCountry>
), 
<collectingCountry id="F32C8E88FFE06046FB50F9AB6A58D546" box="[1239,1323,1637,1659]" name="Iceland" pageId="15" pageNumber="550">Iceland</collectingCountry>
and the Faroes to the Mediterranean Sea (
<bibRefCitation id="EFAAB3E9FFE06046FB7DF94A6CB5D584" author="Thompson &amp; Brown" firstAuthor="Thompson" pageId="15" pageNumber="550" refId="ref59658" refString="Thompson TE, Brown GH. 1984. Biology of Opisthobranch Molluscs, 2. London: The Ray Society." type="book" year="1984">Thompson &amp; Brown, 1984</bibRefCitation>
). The name 
<taxonomicName id="4C3BB59BFFE06046FBD3F96D6B82D58A" authorityName="Muller" authorityYear="1778" box="[1108,1265,1698,1720]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="550" phylum="Mollusca" rank="species" species="tuberculata">
<emphasis id="B94F120AFFE06046FBD3F96D6B82D58A" box="[1108,1265,1698,1720]" italics="true" pageId="15" pageNumber="550">D. tuberculata</emphasis>
</taxonomicName>
has been used for specimens that occur beyond the geographical range of this species. 
<bibRefCitation id="EFAAB3E9FFE06046FBA4F92E6BBED5CB" author="Savigny" box="[1059,1229,1760,1782]" firstAuthor="Savigny" pageId="15" pageNumber="550" refId="ref59128" refString="Savigny JC. 1817. Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise, publiee par ordre du gouvernement. Histoire Naturelle Planches, Vol. 2. Paris, Imprimerie royale." type="book" year="1817">Savigny (1817)</bibRefCitation>
reported this species from the Red Sea, 
<bibRefCitation id="EFAAB3E9FFE06046FBC0F9306BADD428" author="Bergh" box="[1095,1246,1790,1813]" firstAuthor="Bergh" pageId="15" pageNumber="550" pagination="125 - 233" refId="ref54333" refString="Bergh R. 1894. XIII. Die Opisthobranchen. Reports of the dredging operations off the West coast of Central America to the Galapagos, to the West coast of Mexico, and in the Gulf of California, in charge of Alexander Agassiz, carried on by the U. S. Fish Commission Steamer ' Albatross', during 1891, Lieut. Commander Z. L. Tanner, U. S. N., commanding. Bulletin of the Museum of Comparative Zoology 25: 125 - 233, plates 1 - 12." type="journal article" year="1894">Bergh (1894)</bibRefCitation>
from the North Pacific and 
<bibRefCitation id="EFAAB3E9FFE06046FC38F8D36B18D40E" author="Lemche" box="[959,1131,1821,1843]" firstAuthor="Lemche" pageId="15" pageNumber="550" pagination="1" refId="ref56838" refString="Lemche HM. 1929. LIII. Gastropoda Opisthobranchiata. In: Sparck R, Tuxen SL, eds. The Zoology of the Faroes, Vol. 3 (1). Copenhagen: Host &amp; Son." type="book chapter" year="1929">Lemche (1929)</bibRefCitation>
from the Gulf of Mexico. These three records are probably misidentifications (see 
<bibRefCitation id="EFAAB3E9FFE06046FCEFF8946B5BD44D" author="Pruvot-Fol" box="[872,1064,1882,1904]" firstAuthor="Pruvot-Fol" pageId="15" pageNumber="550" pagination="209 - 261" refId="ref58562" refString="Pruvot-Fol A. 1935. Les doridiens de Cuvier publies dans les Annales du Museum en 1804, etude critique et historique. Journal de Conchyliologie 78: 209 - 261." type="journal article" year="1935">Pruvot-Fol, 1935</bibRefCitation>
and 
<bibRefCitation id="EFAAB3E9FFE06046FBE2F8946AE5D44D" author="Thompson &amp; Brown" box="[1125,1430,1882,1904]" firstAuthor="Thompson" pageId="15" pageNumber="550" refId="ref59658" refString="Thompson TE, Brown GH. 1984. Biology of Opisthobranch Molluscs, 2. London: The Ray Society." type="book" year="1984">Thompson &amp; Brown, 1984</bibRefCitation>
, who have also listed several other synonyms for this species discussed here).
</paragraph>
<paragraph id="8B84CE18FFFF6059FF20FECC6DC1D2D0" blockId="16.[142,759,196,1137]" pageId="16" pageNumber="551">
<taxonomicName id="4C3BB59BFFFF6059FF20FECC6D7AD22A" authority="Verany, 1846" authorityName="Verany" authorityYear="1846" box="[167,521,257,279]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="schembrii">
<emphasis id="B94F120AFFFF6059FF20FECC6E2CD22B" box="[167,351,257,279]" italics="true" pageId="16" pageNumber="551">Doris schembrii</emphasis>
<bibRefCitation id="EFAAB3E9FFFF6059FEEAFECC6D7AD22A" author="Verany" box="[365,521,257,279]" firstAuthor="Verany" pageId="16" pageNumber="551" pagination="89 - 109" refId="ref60000" refString="Verany DB. 1846. Catalogo degli animali invertebrati marini del Golfo di Genova e Nizza. Guida di Genova 1: 89 - 109, plates 2 - 4." type="journal article" year="1846">Verany, 1846</bibRefCitation>
</taxonomicName>
was originally described with the same external features of 
<taxonomicName id="4C3BB59BFFFF6059FD1DFEEE6F94D26E" authorityName="Rapp" authorityYear="1827" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFFF6059FD1DFEEE6F94D26E" italics="true" pageId="16" pageNumber="551">A. pseudoargus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFAAB3E9FFFF6059FEAAFEF16EB5D269" author="Verany" box="[301,454,318,340]" firstAuthor="Verany" pageId="16" pageNumber="551" pagination="89 - 109" refId="ref60000" refString="Verany DB. 1846. Catalogo degli animali invertebrati marini del Golfo di Genova e Nizza. Guida di Genova 1: 89 - 109, plates 2 - 4." type="journal article" year="1846">Verany, 1846</bibRefCitation>
), and the re-examination of its 
<typeStatus id="548070BAFFFF6059FF52FE906E75D24E" box="[213,262,350,371]" pageId="16" pageNumber="551">type</typeStatus>
material confirmed that these names are synonyms. Also, the original descriptions of 
<taxonomicName id="4C3BB59BFFFF6059FD3DFEB26F91D292" authorityName="Leuckart" authorityYear="1828" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="flavipes">
<emphasis id="B94F120AFFFF6059FD3DFEB26F91D292" italics="true" pageId="16" pageNumber="551">Doris flavipes</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFAAB3E9FFFF6059FEA7FE546EBCD28C" author="Leuckart" box="[288,463,410,433]" firstAuthor="Leuckart" pageId="16" pageNumber="551" refId="ref56877" refString="Leuckart FS. 1828. Breves Animalium quorundam maxima ex parte marinorum descriptiones. Heidelbergae: A. Osswaldi." type="book" year="1828">Leuckart, 1828</bibRefCitation>
) and 
<taxonomicName id="4C3BB59BFFFF6059FD92FE556DB0D292" authorityName="Delle Chiaje" authorityYear="1841" box="[533,707,410,432]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="leuckartii">
<emphasis id="B94F120AFFFF6059FD92FE556DB0D292" box="[533,707,410,432]" italics="true" pageId="16" pageNumber="551">Doris leuckartii</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFAAB3E9FFFF6059FF09FE776E17D2F2" author="Delle Chiaje" box="[142,356,441,463]" firstAuthor="Delle Chiaje" pageId="16" pageNumber="551" refId="ref55179" refString="Delle Chiaje S. 1841. Animale senza vetebre del Regno di Napoli. Naples: Batelli." type="book" year="1841">Delle Chiaje, 1841</bibRefCitation>
) clearly show that they should be regarded as junior synonyms of 
<taxonomicName id="4C3BB59BFFFF6059FD83FE166DD8D2D0" authorityName="Rapp" authorityYear="1827" box="[516,683,472,493]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFFF6059FD83FE166DD8D2D0" box="[516,683,472,493]" italics="true" pageId="16" pageNumber="551">A. pseudoargus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFFF6059FF20FE396F8FD74C" blockId="16.[142,759,196,1137]" pageId="16" pageNumber="551">
<taxonomicName id="4C3BB59BFFFF6059FF20FE396D06D131" authority="Alder &amp; Hancock, 1844" authorityName="Alder &amp; Hancock" authorityYear="1844" box="[167,629,502,524]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="flammea">
<emphasis id="B94F120AFFFF6059FF20FE396E23D136" box="[167,336,502,524]" italics="true" pageId="16" pageNumber="551">Doris flammea</emphasis>
Alder &amp; Hancock, 1844
</taxonomicName>
and 
<taxonomicName id="4C3BB59BFFFF6059FD3DFE396E91D117" authority="Alder &amp; Hancock, 1844" authorityName="Alder &amp; Hancock" authorityYear="1844" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="mera">
<emphasis id="B94F120AFFFF6059FD3DFE396FB5D116" italics="true" pageId="16" pageNumber="551">Doris mera</emphasis>
Alder &amp; Hancock, 1844
</taxonomicName>
have been regarded as synonyms of 
<taxonomicName id="4C3BB59BFFFF6059FEA2FDFB6EBDD174" authorityName="Rapp" authorityYear="1827" box="[293,462,564,586]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFFF6059FEA2FDFB6EBDD174" box="[293,462,564,586]" italics="true" pageId="16" pageNumber="551">D. pseudoargus</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFAAB3E9FFFF6059FD8EFDFA6FBBD154" author="Thompson &amp; Brown" firstAuthor="Thompson" pageId="16" pageNumber="551" refId="ref59658" refString="Thompson TE, Brown GH. 1984. Biology of Opisthobranch Molluscs, 2. London: The Ray Society." type="book" year="1984">Thompson &amp; Brown, 1984</bibRefCitation>
). However, the original description of these species (
<bibRefCitation id="EFAAB3E9FFFF6059FF4EFDBF6EA5D1BA" author="Alder &amp; Hancock" box="[201,470,625,647]" firstAuthor="Alder" pageId="16" pageNumber="551" refId="ref52940" refString="Alder J, Hancock A. 1845 - 1855. A monograph of the British Nudibranchiate Mollusca: with figures of all the species. Ray Society, London. Dates of Publication: part 1, fam. 1 (plates 4 - 5, 26), fam. 3 (plates 3, 21, 24, 26, 34 - 36) [1845]; part 2, fam. 1 (plates 10, 13, 18, 23), fam. 3 (plates 1 - 2, 4, 6, 12, 15, 23, 30, 42) [1845]; part 3, fam. 1 (plates 6, 8, 19, 25), fam. 2 (pl. 3), fam 3 (plates 1 a, 7, 19, 28, 31, 33) [1846]; part 4, fam. 1 (plates 7, 14, 20, 21, 24), fam. 2 (pl. 5), fam. 3 (plates 11, 13 - 14, 20, 25, 40) [1848]; part 5, fam. 1 (plates 1 - 2, 12, 15 - 16, 22), fam. 2 (pl. 4), fam. 3 (plates 5, 16 - 17, 27, 37, 39, 43) [1851]; part 6, fam. 1 (plates 3, 9, 11, 17), fam. 3 (plates 9 - 10, 18, 22, 29, 32, 41, 44) [1854]; part 7, fam. 1 (plates 21 a, 27), fam. 2 (plates 1 - 2), fam. 3 (plates 38 a, 45 - 48), appendix. 1 - 54, i - xl [1855]." type="journal volume" year="1845">Alder &amp; Hancock, 1845</bibRefCitation>
-55) shows that they are externally very different from 
<taxonomicName id="4C3BB59BFFFF6059FE78FD5F6DDBD198" authorityName="Rapp" authorityYear="1827" box="[511,680,656,678]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFFF6059FE78FD5F6DDBD198" box="[511,680,656,678]" italics="true" pageId="16" pageNumber="551">D. pseudoargus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C3BB59BFFFF6059FD3EFD5E6F9DD1FE" authorityName="Alder &amp; Hancock" authorityYear="1844" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="flammea">
<emphasis id="B94F120AFFFF6059FD3EFD5E6F9DD1FE" italics="true" pageId="16" pageNumber="551">Doris flammea</emphasis>
</taxonomicName>
is a bright orange-scarlet species, occasionally blotched with purple. The dorsum is covered with short, obtuse, spiculose tubercles. The rhinophores are large, tapering, orange with 10 or 11 scarlet lamellae. There are nine scarlet branchial leaves. This description resembles 
<taxonomicName id="4C3BB59BFFFF6059FEFBFC876DB3D060" authority="Risso, 1818" authorityName="Risso" authorityYear="1818" box="[380,704,840,862]" class="Gastropoda" family="Discodorididae" genus="Rostanga" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="rubra">
<emphasis id="B94F120AFFFF6059FEFBFC876D5DD060" box="[380,558,840,862]" italics="true" pageId="16" pageNumber="551">Rostanga rubra</emphasis>
<bibRefCitation id="EFAAB3E9FFFF6059FDBDFC866DB3D060" author="Risso" box="[570,704,840,862]" firstAuthor="Risso" pageId="16" pageNumber="551" pagination="368 - 377" refId="ref58708" refString="Risso A. 1818. Memoire sur quelques Gasteropodes nouveaux, Nudibranches et Tectibranches observes dans la mer de Nice. Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts 87: 368 - 377." type="journal article" year="1818">Risso, 1818</bibRefCitation>
</taxonomicName>
, but whether these two names are synonyms requires further investigation. 
<taxonomicName id="4C3BB59BFFFF6059FEFFFC486E88D0A6" authorityName="Alder &amp; Hancock" authorityYear="1844" box="[376,507,902,923]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="mera">
<emphasis id="B94F120AFFFF6059FEFFFC486E88D0A6" box="[376,507,902,923]" italics="true" pageId="16" pageNumber="551">Doris mera</emphasis>
</taxonomicName>
was described as a white species, ‘rather broad and elevated on the back’. This is very different from 
<taxonomicName id="4C3BB59BFFFF6059FE4CFC0D6D01D0EA" authorityName="Rapp" authorityYear="1827" box="[459,626,962,984]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="pseudoargus">
<emphasis id="B94F120AFFFF6059FE4CFC0D6D01D0EA" box="[459,626,962,984]" italics="true" pageId="16" pageNumber="551">D. pseudoargus</emphasis>
</taxonomicName>
, which is a brownish species. Also, the dorsal tubercles of 
<taxonomicName id="4C3BB59BFFFF6059FD27FC2C6D85D0CA" authorityName="Alder &amp; Hancock" authorityYear="1844" box="[672,758,994,1015]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="mera">
<emphasis id="B94F120AFFFF6059FD27FC2C6D85D0CA" box="[672,758,994,1015]" italics="true" pageId="16" pageNumber="551">D. mera</emphasis>
</taxonomicName>
were described as being moderately sized, unequal and round. This is very similar to 
<taxonomicName id="4C3BB59BFFFF6059FDB5FBD06EDAD76E" authority="(Alder &amp; Hancock, 1854)" baseAuthorityName="Alder &amp; Hancock" baseAuthorityYear="1854" class="Gastropoda" family="Cadlinidae" genus="Aldisa" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="zetlandica">
<emphasis id="B94F120AFFFF6059FDB5FBD06D84D70E" box="[562,759,1054,1075]" italics="true" pageId="16" pageNumber="551">Aldisa zetlandica</emphasis>
(
<bibRefCitation id="EFAAB3E9FFFF6059FF11FBF36ED2D76E" author="Alder &amp; Hancock" box="[150,417,1085,1107]" firstAuthor="Alder" pageId="16" pageNumber="551" pagination="102 - 105" refId="ref53330" refString="Alder J, Hancock A. 1854. Notice of some new species of British Nudibranchiata. Annals and Magazine of Natural History 2 (14): 102 - 105." type="journal article" year="1854">Alder &amp; Hancock, 1854</bibRefCitation>
)
</taxonomicName>
, for which 
<taxonomicName id="4C3BB59BFFFF6059FDA1FBF06D0FD76E" authorityName="Alder &amp; Hancock" authorityYear="1844" box="[550,636,1086,1107]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="551" phylum="Mollusca" rank="species" species="mera">
<emphasis id="B94F120AFFFF6059FDA1FBF06D0FD76E" box="[550,636,1086,1107]" italics="true" pageId="16" pageNumber="551">D. mera</emphasis>
</taxonomicName>
could be a synonym.
</paragraph>
</subSubSection>
</treatment>
</document>