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<mods:title id="9726815C31DD48D9602A6B4C562ADCC3">Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea)</mods:title>
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<mods:namePart id="9568BEE5EAFFF05389CA257CA458887D">Park, Kyu-Tek</mods:namePart>
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<mods:namePart id="AC5C17788DE3F7F5CD09542A2FC0E889">Koo, Jun-Mo</mods:namePart>
<mods:affiliation id="E37003F8D926D2DDA617F6AC24C5F5D3">Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea.</mods:affiliation>
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<mods:namePart id="51915569F3F626BF4D9C5515A68CE065">Minet, Joël</mods:namePart>
<mods:affiliation id="98D113DEC5758EA1EEEF8FF730277947">Muséum National dHistoire Naturelle (ISYEB, UMR 7205), Entomologie (C. P. 50), 45, rue Buffon, F- 75005 Paris, France.</mods:affiliation>
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<subSubSection id="C3219BF7FFA9FFC25CE19FB5FBA6FEE8" pageId="30" pageNumber="181" type="nomenclature">
<paragraph id="8B84C87CFFA9FFC25CE19FB5FCC0FF0F" blockId="30.[151,1030,223,286]" box="[151,864,223,249]" pageId="30" pageNumber="181">
<heading id="D0CC7F10FFA9FFC25CE19FB5FCC0FF0F" bold="true" box="[151,864,223,249]" fontSize="11" level="1" pageId="30" pageNumber="181" reason="1">
<emphasis id="B94F146EFFA9FFC25CE19FB5FCC0FF0F" bold="true" box="[151,864,223,249]" pageId="30" pageNumber="181">
Subfamily
<taxonomicName id="4C3BB3FFFFA9FFC25D689FB5FD15FF0F" ID-CoL="9FPWM" authority="Minet &amp; Park" authorityName="Minet &amp; Park" box="[286,693,223,249]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="subFamily" subFamily="Adelomomphinae">Adelomomphinae Minet &amp; Park</taxonomicName>
, subfam. nov.
</emphasis>
</heading>
</paragraph>
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LSID:
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urn:lsid:zoobank.org:act:
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<subSubSection id="C3219BF7FFA9FFC25CE19E22FC39FE96" box="[151,921,328,352]" pageId="30" pageNumber="181" type="description">
<paragraph id="8B84C87CFFA9FFC25CE19E22FC39FE96" blockId="30.[151,921,328,352]" box="[151,921,328,352]" pageId="30" pageNumber="181">
Type-genus:
<taxonomicName id="4C3BB3FFFFA9FFC25D6C9E22FDE1FE96" authority="Minet &amp; Park" authorityName="Minet &amp; Park" authorityYear="2020" box="[282,577,328,352]" class="Insecta" family="Momphidae" genus="Adelomompha" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94F146EFFA9FFC25D6C9E22FE0CFE96" box="[282,428,328,352]" italics="true" pageId="30" pageNumber="181">Adelomompha</emphasis>
Minet &amp; Park
</taxonomicName>
,
<taxonomicNameLabel id="A27CA915FFA9FFC25E3D9E22FD08FEA9" box="[587,680,328,352]" pageId="30" pageNumber="181" rank="genus">gen. nov.</taxonomicNameLabel>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFA9FFC25CE19EE7FE68FC8B" pageId="30" pageNumber="181" type="diagnosis">
<paragraph id="8B84C87CFFA9FFC25CE19EE7FE68FC8B" blockId="30.[151,1437,397,1252]" pageId="30" pageNumber="181">
<emphasis id="B94F146EFFA9FFC25CE19EE7FEB2FE51" bold="true" box="[151,274,397,423]" pageId="30" pageNumber="181">Diagnosis.</emphasis>
This group share with the other
<taxonomicName id="4C3BB3FFFFA9FFC25EFA9EE4FCB3FE5E" box="[652,787,398,424]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="family">Momphidae</taxonomicName>
the following traits: (1) the compound eye is notched in the vicinity of the antennal foramen (as in Fig. II-18.4Q of
<bibRefCitation id="EFAAB58DFFA9FFC25F4C9ED8FBABFE3A" author="Komai, F. &amp; Yoshiyasu, Y. &amp; Nasu, Y. &amp; Saito, T." box="[826,1035,434,460]" pageId="30" pageNumber="181" refId="ref23117" refString="Komai, F., Yoshiyasu, Y., Nasu, Y. &amp; Saito, T. (Eds.) (2011) A guide to the Lepidoptera of Japan. Tokai University Press, Hadano, Kanagawa, xx + 1307 pp." type="book" year="2011">
Komai
<emphasis id="B94F146EFFA9FFC25FF99ED9FC61FE3A" box="[911,961,434,460]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2011
</bibRefCitation>
), (2) the head is smooth-scaled, (3) the ocelli are absent, (4) in the hind wing, the anterior cubital vein (CuA) is somewhat arched and has its distal section (proximad of CuA2) fairly close and parallel to the anal margin (as in, e.g.,
<figureCitation id="1300D4F9FFA9FFC258579E90FB25FDE2" box="[1057,1157,506,532]" captionStart="FIGURE 2" captionStartId="6.[151,250,1904,1928]" captionTargetBox="[185,1352,469,1863]" captionTargetId="figure-145@6.[151,1436,452,1880]" captionTargetPageId="6" captionText="FIGURE 2. Parkiana janineae (Viette, 1954), comb. nov.: A, holotype; B, labels; C, female genitalia, prep. J. Minet no. 1666; D, ditto, close-up of segment VIII (ventral view); E, ditto, corpus bursae; F, ditto, ductus bursae (anterior region); G, ditto, abdomen. Scale bar for the genitalia: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406464" httpUri="https://zenodo.org/record/4406464/files/figure.png" pageId="30" pageNumber="181">Figs. 2A</figureCitation>
,
<figureCitation id="1300D4F9FFA9FFC258E79E90FB12FDE3" box="[1169,1202,506,533]" captionStart="FIGURE 5" captionStartId="10.[151,250,1409,1433]" captionTargetBox="[166,1422,432,1343]" captionTargetId="figure-118@10.[151,1435,417,1386]" captionTargetPageId="10" captionText="FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406470" httpUri="https://zenodo.org/record/4406470/files/figure.png" pageId="30" pageNumber="181">5A</figureCitation>
and
<figureCitation id="1300D4F9FFA9FFC2589F9E90FAAAFDE2" box="[1257,1290,506,532]" captionStart="FIGURE 8" captionStartId="13.[151,250,1245,1269]" captionTargetBox="[170,1424,282,1202]" captionTargetId="figure-46@13.[151,1436,273,1222]" captionTargetPageId="13" captionText="FIGURE 8. Torodora hiarakella (Viette, 1988), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. P. Viette no. 5755; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4406476" httpUri="https://zenodo.org/record/4406476/files/figure.png" pageId="30" pageNumber="181">8A</figureCitation>
of
<bibRefCitation id="EFAAB58DFFA9FFC259479E90FF73FDCE" author="Luz, F. A. &amp; Goncalves, G. L. &amp; Moreira, G. R. P. &amp; Becker, V. O." pageId="30" pageNumber="181" pagination="97 - 127" refId="ref23423" refString="Luz, F. A., Goncalves, G. L., Moreira, G. R. P. &amp; Becker, V. O. (2014) Three new cecidogenous species of Palaeomystella Fletcher (Lepidoptera, Momphidae) from the Brazilian Atlantic Rain Forest. ZooKeys, 433, 97 - 127. https: // doi. org / 10.3897 / zookeys. 433.7379" type="journal article" year="2014">
Luz
<emphasis id="B94F146EFFA9FFC259129E91FA35FDE2" box="[1380,1429,506,532]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2014
</bibRefCitation>
), (5) the abdominal terga IIVII have large areas of close-set spiniform scales, (6) the apodemes of abdominal sternum II are practically absent in the male, (7) the aedeagus has largely desclerotized walls, (8) the vinculum has narrow arms, (9) when observed in lateral view, the uncus is long, slender, pointed apically, curved anteriorly, and separated from the tegumen by a well indicated furrow, (10) the gnathos is absent. Nevertheless the subfamily
<taxonomicName id="4C3BB3FFFFA9FFC25CCD9DC4FE24FD3E" box="[187,388,686,712]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="subFamily" subFamily="Adelomomphinae">Adelomomphinae</taxonomicName>
differs from the other
<taxonomicName id="4C3BB3FFFFA9FFC25EF19DC4FCAEFD3E" box="[647,782,686,712]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="family">Momphidae</taxonomicName>
(here placed in subfamily
<taxonomicName id="4C3BB3FFFFA9FFC258369DC4FB6AFD3E" authorityName="Herrich-Schäffer" authorityYear="1857" box="[1088,1226,686,712]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="subFamily" subFamily="Momphinae">Momphinae</taxonomicName>
, revised status) by the absence in male and female genitalia of, respectively, a modified transtilla (typically bearing, in
<taxonomicName id="4C3BB3FFFFA9FFC2588C9DB8FA24FD1A" authorityName="Herrich-Schäffer" authorityYear="1857" box="[1274,1412,722,748]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="subFamily" subFamily="Momphinae">Momphinae</taxonomicName>
, a central, scobinate plate, which is sometimes erroneously regarded as part of a gnathos:
<bibRefCitation id="EFAAB58DFFA9FFC258129D9CFAF3FCE6" author="Koster, J. C. &amp; Sinev S. Yu." box="[1124,1363,758,784]" pageId="30" pageNumber="181" refId="ref23205" refString="Koster, J. C. &amp; Sinev S. Yu. (2003) Momphidae, Batrachedridae, Stathmopodidae, Agonoxenidae, Cosmopterigidae, Chrysopeleiidae. In: Huemer, P., Karsholt, O. &amp; Lyneborg, L. (Eds), Microlepidoptera of Europe. Vol. 5. Apollo Books, Stenstrup, 387 pp." type="book" year="2003">Koster &amp; Sinev 2003</bibRefCitation>
) and a pair of basically sickle-shaped signa (secondarily lost in two species of
<taxonomicName id="4C3BB3FFFFA9FFC25FB59C70FABDFCC2" authority="Fletcher, 1940" authorityName="Fletcher" authorityYear="1940" box="[963,1309,794,820]" class="Insecta" family="Agonoxenidae" genus="Palaeomystella" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="B94F146EFFA9FFC25FB59C70FBD0FCC2" box="[963,1136,794,820]" italics="true" pageId="30" pageNumber="181">Palaeomystella</emphasis>
Fletcher, 1940
</taxonomicName>
: see
<bibRefCitation id="EFAAB58DFFA9FFC259239C70FF56FCAE" author="Luz, F. A. &amp; Goncalves, G. L. &amp; Moreira, G. R. P. &amp; Becker, V. O." pageId="30" pageNumber="181" pagination="97 - 127" refId="ref23423" refString="Luz, F. A., Goncalves, G. L., Moreira, G. R. P. &amp; Becker, V. O. (2014) Three new cecidogenous species of Palaeomystella Fletcher (Lepidoptera, Momphidae) from the Brazilian Atlantic Rain Forest. ZooKeys, 433, 97 - 127. https: // doi. org / 10.3897 / zookeys. 433.7379" type="journal article" year="2014">
Luz
<emphasis id="B94F146EFFA9FFC259FE9C71FF0DFCAE" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2014
</bibRefCitation>
, and
<bibRefCitation id="EFAAB58DFFA9FFC25D469C54FDA2FCAF" author="Basilio, D. S. &amp; Casagrande, M. M. &amp; Bordignon, S. A. L. &amp; Moreira, G. R. P." box="[304,514,830,857]" pageId="30" pageNumber="181" pagination="188 - 196" refId="ref22147" refString="Basilio, D. S., Casagrande, M. M., Bordignon, S. A. L. &amp; Moreira, G. R. P. (2015) Description and life history of a new cecidogenous species of Palaeomystella Fletcher (Lepidoptera, Momphidae) from Brazil. Revista Brasileira de Entomologia, 59, 188 - 196. https: // doi. org / 10.1016 / j. rbe. 2015.06.005" type="journal article" year="2015">
Basilio
<emphasis id="B94F146EFFA9FFC25DF09C55FE17FCAE" box="[390,439,830,856]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2015
</bibRefCitation>
). These last two genitalia traits had been regarded as momphid autapomorphies by
<bibRefCitation id="EFAAB58DFFA9FFC25CE19C08FE64FC8B" author="Heikkila, M. &amp; Mutanen, M. &amp; Kekkonen, M. &amp; Kaila, L." box="[151,452,866,893]" pageId="30" pageNumber="181" pagination="563 - 589" refId="ref22760" refString="Heikkila, M., Mutanen, M., Kekkonen, M. &amp; Kaila, L. (2014) Morphology reinforces proposed molecular phylogenetic affinities: a revised classification for Gelechioidea (Lepidoptera). Cladistics, 30 (6), 563 - 589. https: // doi. org / 10.1111 / cla. 12064" type="journal article" year="2014">
Heikkilä
<emphasis id="B94F146EFFA9FFC25C899C09FE90FC8A" box="[255,304,866,892]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. (2014: 582)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3219BF7FFA9FFC25CB19CEFFB53FB12" pageId="30" pageNumber="181" type="description">
<paragraph id="8B84C87CFFA9FFC25CB19CEFFB53FB12" blockId="30.[151,1437,397,1252]" pageId="30" pageNumber="181">
<emphasis id="B94F146EFFA9FFC25CB19CEFFEF9FC69" bold="true" box="[199,345,901,927]" pageId="30" pageNumber="181">Description.</emphasis>
Since the subfamily
<taxonomicName id="4C3BB3FFFFA9FFC25E3C9CECFCB3FC56" box="[586,787,902,928]" class="Insecta" family="Momphidae" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="181" phylum="Arthropoda" rank="subFamily" subFamily="Adelomomphinae">Adelomomphinae</taxonomicName>
is monotypic, its description does not differ from that of the type-genus (see hereafter). We just summarize here its main traits. Head smooth-scaled. Compound eye notched dorsally. Antenna longer than forewing in both sexes; scape without a pecten. Labial palpus sickle-shaped. Foreleg well developed, with an epiphysis; hindtibia of male dorsally fringed with long piliform scales; spur formula: 0-2-4. Forewing venation: R
<subScript id="17BFCA39FFA9FFC25DFA9B42FE34FBC0" attach="left" box="[396,404,1064,1078]" fontSize="6" pageId="30" pageNumber="181">4</subScript>
stalked with R
<subScript id="17BFCA39FFA9FFC25E4B9B42FDE5FBC0" attach="left" box="[573,581,1064,1078]" fontSize="6" pageId="30" pageNumber="181">5</subScript>
, the latter running to the costa; only two M veins. Hind wing venation complete; Rs to costa; M
<subScript id="17BFCA39FFA9FFC25DF49B26FE2AFBAC" attach="left" box="[386,394,1100,1114]" fontSize="6" pageId="30" pageNumber="181">1</subScript>
and M
<subScript id="17BFCA39FFA9FFC25DAD9B26FE43FBAC" attach="left" box="[475,483,1100,1114]" fontSize="6" pageId="30" pageNumber="181">2</subScript>
short-stalked. Abdominal terga II to VII with wide spinose areas. Antero-abdominal sternum with venulae and sexually dimorphic apodemes (practically absent in the male). Male genitalia: aedeagus without an anterior coecum and sclerotized only dorsally; arms of vinculum narrow; saccus, transtilla and gnathos absent; valve broad, undivided, provided with a large, rounded central structure; uncus elongate. Female genitalia: corpus bursae without sickle-shaped structures but with a small, plate-like, toothed/spinose signum.
</paragraph>
</subSubSection>
</treatment>
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