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<document id="29F28453D10ED66F9E8E99B9C1B662D3" ID-DOI="10.5281/zenodo.10134425" ID-ISSN="1942-1354" ID-Zenodo-Dep="10134425" ID-ZooBank="8F673BD6-7061-4B9E-9445-F8A5C21D5892" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699899090914" checkinUser="felipe" docAuthor="Hielkema, Meindert A." docDate="2023" docId="03904A124611FF93BAC2FED9FEBAFA10" docLanguage="en" docName="InsectaMundi.2023.1006.1-8.pdf" docOrigin="Insecta Mundi 2023 (6)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Cyclocephala kuijteni Hielkema 2023, new species" docType="treatment" docVersion="3" lastPageNumber="2" masterDocId="FFA9326A4612FF90BA02FFFDFFB8FFCA" masterDocTitle="Cyclocephala kuijteni (Scarabaeidae: Dynastinae: Cyclocephalini) a new species from Suriname" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1700278942298" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="28C85C92F954D1955C7CD1E8FCE37E29" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5501408B2F287D2CD36747FBDF05F97">
<mods:title id="DEFD4DC8B94121CA28AFBB23B8CCF66A">Cyclocephala kuijteni (Scarabaeidae: Dynastinae: Cyclocephalini) a new species from Suriname</mods:title>
</mods:titleInfo>
<mods:name id="FD4E96D39EC94569378AC71E841AE902" type="personal">
<mods:role id="AC2837DC29DA91F19AACC4A57A66FCF6">
<mods:roleTerm id="9A1CA5B9AA2E2F8C4E7FB33CD7FED0D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46268614CF4B8BE5AC755744173FC0A4">Hielkema, Meindert A.</mods:namePart>
<mods:affiliation id="468AA51FFDAF36B32D99743C278BFBED">Veenenburg 6, 2804 WX Gouda, The Netherlands</mods:affiliation>
</mods:name>
<mods:typeOfResource id="76BE48611F009245F9453243BD3C6885">text</mods:typeOfResource>
<mods:relatedItem id="76A001D46FCAB574DB4C7CDCA33CEB8E" type="host">
<mods:titleInfo id="395A85EEDB896D81952C3C0FA91F16AA">
<mods:title id="FEB50E580E02C7D79E1B845382EA04E1">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="F5C1825A7F2BECB04DF83CF4B1DF67FE">
<mods:date id="F40DF55065255D001F8247D657F86454">2023</mods:date>
<mods:detail id="20D90AA6A60FD7492E58E8C0A9EDDE31" type="pubDate">
<mods:number id="2FFC763099AC4023559D059D8DEB4969">2023-09-29</mods:number>
</mods:detail>
<mods:detail id="CC691EDDA91EECA05C4EDC5BD75CE0BF" type="volume">
<mods:number id="1E8A6119A14695D86C9CF6517A646823">2023</mods:number>
</mods:detail>
<mods:detail id="EEDF8D67DA750E45ABDD745603ED41FF" type="issue">
<mods:number id="C19C06EE2E828ABD28ED49F79D78AD07">6</mods:number>
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<mods:start id="11A412981FA1306A7B3D6D4AF37840B0">1</mods:start>
<mods:end id="2855E3048CD3F9B4CBD61FFAE34AE365">6</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="24D1807E8187C5C13F299C8B7910D7F2">journal article</mods:classification>
<mods:identifier id="ADBC80BF1B680A72623AB7606ED00972" type="DOI">10.5281/zenodo.10134425</mods:identifier>
<mods:identifier id="C5C5A93A77D7E7865DC308C242D59D9E" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="2E8288C067CD85E4C9656669823950B9" type="Zenodo-Dep">10134425</mods:identifier>
<mods:identifier id="62CD94759B2B665ACCD3147AC92D42CB" type="ZooBank">8F673BD6-7061-4B9E-9445-F8A5C21D5892</mods:identifier>
</mods:mods>
<treatment id="03904A124611FF93BAC2FED9FEBAFA10" LSID="urn:lsid:plazi:treatment:03904A124611FF93BAC2FED9FEBAFA10" httpUri="http://treatment.plazi.org/id/03904A124611FF93BAC2FED9FEBAFA10" lastPageNumber="2" pageId="3" pageNumber="2">
<subSubSection id="C323A88F4611FF93BAC2FED9FD4CFE8B" box="[192,756,292,321]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8B86FB044611FF93BAC2FED9FD4CFE8B" blockId="3.[192,756,292,355]" box="[192,756,292,321]" pageId="3" pageNumber="2">
<heading id="D0CE4C684611FF93BAC2FED9FD4CFE8B" box="[192,756,292,321]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
<emphasis id="B94D27164611FF93BAC2FED9FD4CFE8B" bold="true" box="[192,756,292,321]" pageId="3" pageNumber="2">
<taxonomicName id="4C3980874611FF93BAC2FED9FDEAFE8B" authority="Hielkema" authorityName="Hielkema" authorityYear="2023" box="[192,594,292,321]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="kuijteni" status="sp. nov.">
<emphasis id="B94D27164611FF93BAC2FED9FE76FE8B" bold="true" box="[192,462,292,321]" italics="true" pageId="3" pageNumber="2">Cyclocephala kuijteni</emphasis>
Hielkema
</taxonomicName>
,
<taxonomicNameLabel id="A27E9A6D4611FF93B85DFED8FD4CFE8B" box="[607,756,292,321]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C323A88F4611FF93BAC2FEB4FED3FEA9" box="[192,363,329,355]" pageId="3" pageNumber="2" type="description">
<paragraph id="8B86FB044611FF93BAC2FEB4FED3FEA9" blockId="3.[192,756,292,355]" box="[192,363,329,355]" pageId="3" pageNumber="2">
(
<figureCitation id="1302E7814611FF93BACAFEB4FEADFEA9" box="[200,277,329,355]" captionStart="Figure 1" captionStartId="4.[192,268,1866,1892]" captionTargetBox="[277,1355,217,1836]" captionTargetId="figure-42@4.[277,1355,213,1836]" captionTargetPageId="4" captionText="Figure 1. Cyclocephala spp., dorsal habitus. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, paratype." figureDoi="http://doi.org/10.5281/zenodo.10134427" httpUri="https://zenodo.org/record/10134427/files/figure.png" pageId="3" pageNumber="2">Fig. 1a</figureCitation>
,
<figureCitation id="1302E7814611FF93BB22FEB4FE84FEA9" box="[288,316,329,355]" captionStart="Figure 2" captionStartId="5.[192,268,925,951]" captionTargetBox="[318,1328,225,883]" captionTargetId="figure-43@5.[295,1337,213,895]" captionTargetPageId="5" captionText="Figure 2. Cyclocephala spp., lateral views of the aedeagi. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134429" httpUri="https://zenodo.org/record/10134429/files/figure.png" pageId="3" pageNumber="2">2a</figureCitation>
,
<figureCitation id="1302E7814611FF93BB45FEB4FEDAFEA9" box="[327,354,329,355]" captionStart="Figure 3" captionStartId="6.[192,268,1583,1609]" captionTargetBox="[365,1258,224,1562]" captionTargetId="figure-156@6.[316,1288,210,1583]" captionTargetPageId="6" captionText="Figure 3. Cyclocephala spp., frontal views of the parameres. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134431" httpUri="https://zenodo.org/record/10134431/files/figure.png" pageId="3" pageNumber="2">3a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323A88F4611FF93BAC2FE89FDC0FE3C" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8B86FB044611FF93BAC2FE89FDC0FE3C" blockId="3.[192,1441,371,502]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BAC2FE89FED1FE47" bold="true" box="[192,361,371,398]" pageId="3" pageNumber="2">Type material.</emphasis>
<materialsCitation id="3B51F1594611FF93BB72FE89FDC0FE3C" collectingDate="2010-01-09" collectionCode="UV, RMNH" country="Suriname" location="On" municipality="Label" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
<typeStatus id="548245A64611FF93BB72FE89FE60FE44" box="[368,472,372,398]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
male.
<collectingMunicipality id="6BE2617E4611FF93B821FE89FDD8FE44" box="[547,608,372,398]" pageId="3" pageNumber="2">Label</collectingMunicipality>
data, with a single slash (/) indicating a break between lines, and a double slash (//) indicating a different label: “
<collectingCountry id="F32EBB944611FF93B861FE6BFD69FE7A" box="[611,721,406,432]" name="Suriname" pageId="3" pageNumber="2">Suriname</collectingCountry>
,
<collectingRegion id="49FD35E64611FF93B8DEFE6AFCB6FE7B" box="[732,782,407,433]" country="Suriname" name="Para" pageId="3" pageNumber="2">Para</collectingRegion>
/
<locationDeviation id="9A34E2264611FF93B921FE6AFBB8FE7A" box="[803,1024,406,433]" location="Zanderij" pageId="3" pageNumber="2">
<quantity id="4CC156E14611FF93B921FE6AFCE4FE7A" box="[803,860,406,433]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="2" unit="km" value="1.0">1 km</quantity>
W of Zanderij
</locationDeviation>
/
<date id="FF87DDC44611FF93BE14FE6BFBCCFE7A" box="[1046,1140,406,432]" pageId="3" pageNumber="2" value="2010-01-09">
<collectingDate id="EFC3242C4611FF93BE14FE6BFBCCFE7A" box="[1046,1140,406,432]" pageId="3" pageNumber="2" value="2010-01-09">9-i-2010</collectingDate>
</date>
//
<location id="8EE6ADDF4611FF93BE91FE6AFB0FFE7B" LSID="urn:lsid:plazi:treatment:03904A124611FF93BAC2FED9FEBAFA10:8EE6ADDF4611FF93BE91FE6AFB0FFE7B" box="[1171,1207,407,433]" country="Suriname" municipality="Label" name="On" pageId="3" pageNumber="2" stateProvince="Para">On</location>
<collectionCode id="ED2863C14611FF93BEBCFE6AFB5EFE7B" box="[1214,1254,407,433]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
/HPL in savanna / forest // [red paper]
<typeStatus id="548245A64611FF93BBB0FE44FDFBFE19" box="[434,579,441,467]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D3F308D4611FF93B84BFE40FDE6FE19" box="[585,606,445,467]" count="1" pageId="3" pageNumber="2" type="male">♂</specimenCount>
/
<taxonomicName id="4C3980874611FF93B876FE44FCE4FE19" authorityName="Hielkema" authorityYear="2023" box="[628,860,441,467]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="kuijteni">
<emphasis id="B94D27164611FF93B876FE44FCE4FE19" box="[628,860,441,467]" italics="true" pageId="3" pageNumber="2">Cyclocephala kuijteni</emphasis>
</taxonomicName>
/ Hielkema 2023.” Aedeagus extracted and glued on separate card. Deposited at the
<collectionCode id="ED2863C14611FF93B81FFE21FDCDFE3C" box="[541,629,476,502]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="2" type="Museum">RMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C323A88F4611FF93BAC2FDFBFE6BFAF5" pageId="3" pageNumber="2" type="description">
<paragraph id="8B86FB044611FF93BAC2FDFBFE9CFDAC" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BAC2FDFBFEEAFDEA" bold="true" box="[192,338,518,544]" pageId="3" pageNumber="2">Description.</emphasis>
Male
<typeStatus id="548245A64611FF93BB95FDFBFE44FDEA" box="[407,508,518,544]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
: Length
<quantity id="4CC156E14611FF93B85EFDFAFD15FDEB" box="[604,685,519,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="mm" value="18.0">18 mm</quantity>
, width across humeri
<quantity id="4CC156E14611FF93B9A6FDFAFC51FDEB" box="[932,1001,519,545]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="2" unit="mm" value="8.0">8 mm</quantity>
. Color reddish brown. Basal part of the frons and elytra, except for the outer edges and sutural margin, darker brown. Outer edges of protibia darkened (
<figureCitation id="1302E7814611FF93BACAFDB1FEAEFDAC" box="[200,278,588,614]" captionStart="Figure 1" captionStartId="4.[192,268,1866,1892]" captionTargetBox="[277,1355,217,1836]" captionTargetId="figure-42@4.[277,1355,213,1836]" captionTargetPageId="4" captionText="Figure 1. Cyclocephala spp., dorsal habitus. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, paratype." figureDoi="http://doi.org/10.5281/zenodo.10134427" httpUri="https://zenodo.org/record/10134427/files/figure.png" pageId="3" pageNumber="2">Fig. 1a</figureCitation>
).
</paragraph>
<paragraph id="8B86FB044611FF93BB02FD92FF49FCDE" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FD92FEFAFD43" box="[256,322,623,649]" italics="true" pageId="3" pageNumber="2">Head.</emphasis>
Clypeus parabolic, apex weakly reflexed upwards. Apical and lateral edges rugulose, disc with moderately dense punctation, punctures unequal in size, becoming sparser centrally. Clypeofrontal suture biarcuate, distinct. Frons with a shallow furrow from clypeofrontal suture to halfway base, punctate, punctures unequal in size and sparser towards the center. Antennae with 10 antennomeres, club subequal in length to antennomeres 2–7.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FCE0FC93FCB1" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FCE0FECEFCFC" box="[256,374,797,822]" italics="true" pageId="3" pageNumber="2">Pronotum.</emphasis>
Shiny, sides evenly rounded, widest at one-third from base. Anterior angles acute, basal angles broadly rounded. Surface with minute punctures, set closer only along the sides, with some slightly larger punctures near the anterior angles. Base with marginal bead.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FC79FC0BFC54" blockId="3.[192,1441,518,1343]" box="[256,947,900,927]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FC79FECBFC54" box="[256,371,900,926]" italics="true" pageId="3" pageNumber="2">
<taxonomicName id="4C3980874611FF93BB02FC79FED7FC54" box="[256,367,900,926]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
.
</emphasis>
Base and center with moderately dense punctation.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FC5AFB8CFC2E" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FC5AFEF2FC0B" box="[256,330,935,961]" italics="true" pageId="3" pageNumber="2">Elytra.</emphasis>
With large annulate punctures arranged in vague rows, mixed with fine punctation, the larger punctures becoming smaller towards margins and apex, punctures on apex obsolete.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FC11FE13FBE0" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FC11FEC9FBCC" box="[256,369,1004,1030]" italics="true" pageId="3" pageNumber="2">
<taxonomicName id="4C3980874611FF93BB02FC11FED5FBCC" box="[256,365,1004,1030]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="2" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
.
</emphasis>
Regularly rounded with minute, sparse punctures; basal angles with moderately dense, larger, setiferous punctures.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FBCFFC64FB86" blockId="3.[192,1441,518,1343]" box="[256,988,1074,1100]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FBCFFE72FB86" box="[256,458,1074,1100]" italics="true" pageId="3" pageNumber="2">Prosternal process.</emphasis>
High, distally with long setae on posterior half.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FBA8FA19FB5B" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FBA8FE8FFBA4" box="[256,311,1109,1134]" italics="true" pageId="3" pageNumber="2">Legs.</emphasis>
Protibia with 2 apical teeth close together, basal tooth slightly removed from others, at about twothirds of tibial length. Protarsus enlarged, inside of tarsomere 4 with apical tooth, median claw narrowly incised.
</paragraph>
<paragraph id="8B86FB044611FF93BB02FB67FAF5FAD6" blockId="3.[192,1441,518,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FB67FED7FB7E" box="[256,367,1178,1204]" italics="true" pageId="3" pageNumber="2">Aedeagus.</emphasis>
Laterally (
<figureCitation id="1302E7814611FF93BBEBFB67FD81FB7E" box="[489,569,1178,1204]" captionStart="Figure 2" captionStartId="5.[192,268,925,951]" captionTargetBox="[318,1328,225,883]" captionTargetId="figure-43@5.[295,1337,213,895]" captionTargetPageId="5" captionText="Figure 2. Cyclocephala spp., lateral views of the aedeagi. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134429" httpUri="https://zenodo.org/record/10134429/files/figure.png" pageId="3" pageNumber="2">Fig. 2a</figureCitation>
) resembling that of
<taxonomicName id="4C3980874611FF93B920FB67FC32FB7E" authorityName="Endrodi" authorityYear="1975" box="[802,906,1178,1204]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hardyi">
<emphasis id="B94D27164611FF93B920FB67FC32FB7E" box="[802,906,1178,1204]" italics="true" pageId="3" pageNumber="2">C. hardyi</emphasis>
</taxonomicName>
(
<figureCitation id="1302E7814611FF93B999FB67FC55FB7E" box="[923,1005,1178,1204]" captionStart="Figure 2" captionStartId="5.[192,268,925,951]" captionTargetBox="[318,1328,225,883]" captionTargetId="figure-43@5.[295,1337,213,895]" captionTargetPageId="5" captionText="Figure 2. Cyclocephala spp., lateral views of the aedeagi. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134429" httpUri="https://zenodo.org/record/10134429/files/figure.png" pageId="3" pageNumber="2">Fig. 2b</figureCitation>
) and with a similar but much smaller forward-pointing dorsoapical process of the phallobase. The distal ends of the parameres (
<figureCitation id="1302E7814611FF93BEC7FB41FAACFB1D" box="[1221,1300,1212,1239]" captionStart="Figure 3" captionStartId="6.[192,268,1583,1609]" captionTargetBox="[365,1258,224,1562]" captionTargetId="figure-156@6.[316,1288,210,1583]" captionTargetPageId="6" captionText="Figure 3. Cyclocephala spp., frontal views of the parameres. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134431" httpUri="https://zenodo.org/record/10134431/files/figure.png" pageId="3" pageNumber="2">Fig. 3a</figureCitation>
) are shorter than those of
<taxonomicName id="4C3980874611FF93BB5AFB1DFE05FB33" authorityName="Endrodi" authorityYear="1975" box="[344,445,1247,1274]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hardyi">
<emphasis id="B94D27164611FF93BB5AFB1DFE05FB33" box="[344,445,1247,1274]" italics="true" pageId="3" pageNumber="2">C. hardyi</emphasis>
</taxonomicName>
(
<figureCitation id="1302E7814611FF93BBCEFB22FDA3FB33" box="[460,539,1247,1273]" captionStart="Figure 3" captionStartId="6.[192,268,1583,1609]" captionTargetBox="[365,1258,224,1562]" captionTargetId="figure-156@6.[316,1288,210,1583]" captionTargetPageId="6" captionText="Figure 3. Cyclocephala spp., frontal views of the parameres. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134431" httpUri="https://zenodo.org/record/10134431/files/figure.png" pageId="3" pageNumber="2">Fig. 3b</figureCitation>
), with the tips not touching. The aedeagi of
<taxonomicName id="4C3980874611FF93BE01FB1DFB38FB33" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1027,1152,1248,1274]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="castanea">
<emphasis id="B94D27164611FF93BE01FB1DFB38FB33" box="[1027,1152,1248,1274]" italics="true" pageId="3" pageNumber="2">C. castanea</emphasis>
</taxonomicName>
(
<figureCitation id="1302E7814611FF93BE8CFB22FB63FB33" box="[1166,1243,1247,1273]" captionStart="Figure 2" captionStartId="5.[192,268,925,951]" captionTargetBox="[318,1328,225,883]" captionTargetId="figure-43@5.[295,1337,213,895]" captionTargetPageId="5" captionText="Figure 2. Cyclocephala spp., lateral views of the aedeagi. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134429" httpUri="https://zenodo.org/record/10134429/files/figure.png" pageId="3" pageNumber="2">Fig. 2c</figureCitation>
) and
<taxonomicName id="4C3980874611FF93BF1BFB1DFA18FB33" authorityName="Joly" authorityYear="2000" box="[1305,1440,1247,1274]" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="B94D27164611FF93BF1BFB1DFA18FB33" box="[1305,1440,1247,1274]" italics="true" pageId="3" pageNumber="2">C. pygidialis</emphasis>
</taxonomicName>
(
<figureCitation id="1302E7814611FF93BACAFAFFFEAFFAD6" box="[200,279,1282,1308]" captionStart="Figure 2" captionStartId="5.[192,268,925,951]" captionTargetBox="[318,1328,225,883]" captionTargetId="figure-43@5.[295,1337,213,895]" captionTargetPageId="5" captionText="Figure 2. Cyclocephala spp., lateral views of the aedeagi. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134429" httpUri="https://zenodo.org/record/10134429/files/figure.png" pageId="3" pageNumber="2">Fig. 2d</figureCitation>
) do not have a dorsoapical process, and their parameres are shaped much differently (
<figureCitation id="1302E7814611FF93BED6FAFFFAF9FAD6" box="[1236,1345,1282,1308]" captionStart="Figure 3" captionStartId="6.[192,268,1583,1609]" captionTargetBox="[365,1258,224,1562]" captionTargetId="figure-156@6.[316,1288,210,1583]" captionTargetPageId="6" captionText="Figure 3. Cyclocephala spp., frontal views of the parameres. a) C. kuijteni n. sp., holotype. b) C. hardyi Endrödi, paratype. c) C. castanea (Olivier). d) C. pygidialis Joly, from Joly (2000)." figureDoi="http://doi.org/10.5281/zenodo.10134431" httpUri="https://zenodo.org/record/10134431/files/figure.png" pageId="3" pageNumber="2">Fig. 3c–d</figureCitation>
).
</paragraph>
<paragraph id="8B86FB044611FF93BB02FAD8FE6BFAF5" blockId="3.[192,1441,518,1343]" box="[256,467,1317,1343]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BB02FAD8FEF6FAF5" box="[256,334,1317,1343]" italics="true" pageId="3" pageNumber="2">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C323A88F4611FF93BAC2FAB2FE73FA47" pageId="3" pageNumber="2" type="distribution">
<paragraph id="8B86FB044611FF93BAC2FAB2FE73FA47" blockId="3.[192,1441,1359,1421]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BAC2FAB2FEE3FAA3" bold="true" box="[192,347,1359,1385]" pageId="3" pageNumber="2">Distribution.</emphasis>
Only known from near the village of Zanderij in the district of
<collectingRegion id="49FD35E64611FF93BE53FAADFB3DFAA0" box="[1105,1157,1360,1386]" country="Suriname" name="Para" pageId="3" pageNumber="2">Para</collectingRegion>
,
<collectingCountry id="F32EBB944611FF93BE96FAB2FABBFAA3" box="[1172,1283,1359,1385]" name="Suriname" pageId="3" pageNumber="2">Suriname</collectingCountry>
. Coordinates:
<geoCoordinate id="EE0D9DC34611FF93BAC2FA8FFE8EFA47" box="[192,310,1394,1421]" degrees="5" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="2" precision="15" seconds="37" value="5.4602776">5°27′37″N</geoCoordinate>
,
<geoCoordinate id="EE0D9DC34611FF93BB42FA8FFE7EFA47" box="[320,454,1394,1421]" degrees="55" direction="west" minutes="13" orientation="longitude" pageId="3" pageNumber="2" precision="15" seconds="21" value="-55.2225">55°13′21″W</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C323A88F4611FF93BAC2FA60FEBAFA10" pageId="3" pageNumber="2" type="etymology">
<paragraph id="8B86FB044611FF93BAC2FA60FEBAFA10" blockId="3.[192,1439,1437,1498]" pageId="3" pageNumber="2">
<emphasis id="B94D27164611FF93BAC2FA60FEFCFA7D" bold="true" box="[192,324,1437,1463]" pageId="3" pageNumber="2">Etymology.</emphasis>
This species is named in honor of the late Piet Kuijten, my mentor in scarabs and friend for many years.
</paragraph>
</subSubSection>
</treatment>
</document>
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