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<document id="B553CF0765C92F1108B0D5530655C0B1" ID-DOI="10.11646/zootaxa.3894.1.11" ID-GBIF-Dataset="6c37cabe-0df5-462b-9f82-b5acfc8c37cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="229191" ID-ZooBank="78A995DA-7D11-48CE-A547-C7653E505720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460997608442" checkinUser="plazi" docAuthor="Golovatch, Sergei I. &amp; Vandenspiegel, Didier" docDate="2014" docId="0389DE08734B2551FF35FA3EFDADC5A0" docLanguage="en" docName="zt03894p151.pdf" docOrigin="Zootaxa 3894 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Koponenius biramus Golovatch &amp; Vandenspiegel, 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="149" masterDocId="FFB0A670734C2559FFA2FD62FFEBC76C" masterDocTitle="Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)" masterLastPageNumber="151" masterPageNumber="141" pageNumber="148" updateTime="1698563275390" updateUser="plazi">
<mods:mods id="5C98CF81D0EB52F0FBDB4294F0EFD9F0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="98A9A7F1B09518CF40DC578E4434E974">Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="6F069F13729B24D72AF4637462D573FE" type="personal">
<mods:role id="B692E48161BED96EEBF60AC37ED30DD8">
<mods:roleTerm id="54844F4948BC9D75D6B64C2FAE3B8E0C">Author</mods:roleTerm>
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<mods:namePart id="3C3AEC3DD4BC54269C05B8B2DB2AC22D">Golovatch, Sergei I.</mods:namePart>
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<mods:name id="CF3E97DBFB80B9F8E3A87C72E0DC1951" type="personal">
<mods:role id="0D71BE47B8343807F4C3AF81F7EA7AFD">
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<mods:namePart id="4BED404066C3A601BB905B4B648CF3C5">Vandenspiegel, Didier</mods:namePart>
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<mods:typeOfResource id="97617DDDD30DC120E8E3C99C86C49C15">text</mods:typeOfResource>
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<mods:title id="86C997E21CE9A116E61C1FF7F6704AF3">Zootaxa</mods:title>
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<mods:part id="37BAB6A1DCA6EABD6F11DB25D8D81C1D">
<mods:date id="A3816BB6C3998E4C649FB5AB2B5DE8E1">2014</mods:date>
<mods:detail id="D9752C59EB3D74A7AF3BD5D313B2C348" type="volume">
<mods:number id="0FDF26E8966E08C790B518A09CC36D1C">3894</mods:number>
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<mods:classification id="F011D2D21F038723BA7C69BB8D9BB46B">journal article</mods:classification>
<mods:identifier id="CF4EA7CC558FAAE0F74F49D54D45D163" type="DOI">10.11646/zootaxa.3894.1.11</mods:identifier>
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<treatment id="0389DE08734B2551FF35FA3EFDADC5A0" ID-DOI="http://doi.org/10.5281/zenodo.6126356" ID-GBIF-Taxon="119597642" ID-Zenodo-Dep="6126356" LSID="urn:lsid:plazi:treatment:0389DE08734B2551FF35FA3EFDADC5A0" httpUri="http://treatment.plazi.org/id/0389DE08734B2551FF35FA3EFDADC5A0" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148">
<subSubSection id="C33A3C95734B255EFF35FA3EFF12C0F4" pageId="7" pageNumber="148" type="nomenclature">
<paragraph id="8B9F6F1E734B255EFF35FA3EFE1CC01A" blockId="7.[151,503,1884,1944]" box="[151,503,1884,1910]" pageId="7" pageNumber="148">
<heading id="D0D7D872734B255EFF35FA3EFE1CC01A" bold="true" box="[151,503,1884,1910]" fontSize="11" level="1" pageId="7" pageNumber="148" reason="1">
<emphasis id="B954B30C734B255EFF35FA3EFE1CC01A" bold="true" box="[151,503,1884,1910]" pageId="7" pageNumber="148">
<taxonomicName id="4C20149D734B255EFF35FA3EFE79C01A" ID-CoL="849LX" box="[151,402,1884,1910]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="148" phylum="Arthropoda" rank="species" species="biramus" status="sp. nov.">
<emphasis id="B954B30C734B255EFF35FA3EFE79C01A" bold="true" box="[151,402,1884,1910]" italics="true" pageId="7" pageNumber="148">Koponenius biramus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2670E77734B255EFE38FA3EFE1CC01A" box="[410,503,1884,1910]" pageId="7" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9F6F1E734B255EFF35FA1DFF12C0F4" blockId="7.[151,503,1884,1944]" box="[151,249,1919,1944]" pageId="7" pageNumber="148">
<figureCitation id="131B739B734B255EFF35FA1DFF12C0F4" box="[151,249,1919,1944]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 8" captionStartId-0="5.[151,249,1330,1352]" captionStartId-1="6.[151,249,1763,1785]" captionStartId-2="8.[151,249,1889,1911]" captionTargetBox-0="[179,1392,465,1290]" captionTargetBox-1="[151,1436,195,1737]" captionTargetBox-2="[154,1426,783,1812]" captionTargetId-0="figure@5.[151,1436,447,1305]" captionTargetId-1="figure@6.[151,1436,195,1738]" captionTargetId-2="figure@8.[151,1436,769,1868]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionText-0="FIGURE 5. Habitus of Koponenius biramus sp. nov., male paratype, lateral view. Picture by K. Makarov, not taken to scale." captionText-1="FIGURE 6. Koponenius biramus sp. nov., male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A G, I L, 0.2 mm; N, 0.05 mm; L, 0.02 mm." captionText-2="FIGURE 8. Koponenius biramus sp. nov., male paratype. A: leg 6, lateral view. B: left gonopod, submesal view. Scale bar: 0.2 mm. Designations explained in text." httpUri-0="https://zenodo.org/record/229196/files/figure.png" httpUri-1="https://zenodo.org/record/229197/files/figure.png" httpUri-2="https://zenodo.org/record/229198/files/figure.png" pageId="7" pageNumber="148">Figs 58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A3C95734B2551FF35FAA5FC47C794" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148" type="materials_examined">
<paragraph id="8B9F6F1E734B2551FF35FAA5FC1FC7B9" blockId="7.[151,1436,1991,2016]" lastBlockId="8.[151,1437,151,717]" lastPageId="8" lastPageNumber="149" pageId="7" pageNumber="148">
<emphasis id="B954B30C734B255EFF35FAA5FE68C08C" bold="true" box="[151,387,1991,2016]" pageId="7" pageNumber="148">Material examined:</emphasis>
<typeStatus id="549BD1BC734B255EFE28FAA5FE19C08C" box="[394,498,1991,2016]" pageId="7" pageNumber="148" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED31F7DB734B255EFDE2FAAAFD96C08C" LSID="urn:lsid:biocol.org:col:34838" box="[576,637,1992,2016]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="148">SMF</collectionCode>
),
<collectingCountry id="F3372F8E734B255EFD33FAA5FD3CC08C" box="[657,727,1991,2016]" name="Nepal" pageId="7" pageNumber="148">Nepal</collectingCountry>
, Lalitpur District, Godawari,
<geoCoordinate id="EE1409D9734B255EFB81FAA5FB6CC08C" box="[1059,1159,1991,2016]" direction="north" orientation="latitude" pageId="7" pageNumber="148" precision="925" value="27.6">27°36´N</geoCoordinate>
,
<geoCoordinate id="EE1409D9734B255EFB30FAA5FB19C08C" box="[1170,1266,1991,2016]" direction="east" orientation="longitude" pageId="7" pageNumber="148" precision="925" value="85.4">85°24´E</geoCoordinate>
, foot of Mount Phulchoki, ca
<quantity id="4CD8C2FB73442551FE97FDF5FE67C7C3" box="[309,396,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="8" pageNumber="149" unit="m" value="1750.0">1750 m</quantity>
, secondary broadleaved forest,
<date id="FF9E49DE73442551FD4EFDFAFC78C7DC" box="[748,915,152,176]" pageId="8" pageNumber="149" value="1970-08-09">9 August 1970</date>
, leg. J. Martens.
<typeStatus id="549BD1BC73442551FBEDFDFAFB2BC7DC" box="[1103,1216,152,176]" pageId="8" pageNumber="149" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D26A49773442551FB68FDFAFADAC7DC" box="[1226,1329,151,176]" pageId="8" pageNumber="149" type="male">12 males</specimenCount>
,
<specimenCount id="9D26A49773442551FA99FDFAFA77C7DC" box="[1339,1436,151,176]" pageId="8" pageNumber="149" type="female">1 female</specimenCount>
(
<collectionCode id="ED31F7DB73442551FF3DFDDEFF37C7B8" LSID="urn:lsid:biocol.org:col:34838" box="[159,220,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="149">SMF</collectionCode>
),
<specimenCount id="9D26A49773442551FF4DFDDEFEAEC7B9" box="[239,325,188,213]" pageId="8" pageNumber="149" type="male">2 males</specimenCount>
(
<collectionCode id="ED31F7DB73442551FEF7FDDEFE59C7B8" box="[341,434,188,212]" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="8" pageNumber="149">ZMUM</collectionCode>
),
<specimenCount id="9D26A49773442551FE66FDDEFDFBC7B9" box="[452,528,188,213]" pageId="8" pageNumber="149" type="male">1 male</specimenCount>
(SEM), same data, together with
<typeStatus id="549BD1BC73442551FC28FDDEFC1BC7B9" box="[906,1008,188,213]" pageId="8" pageNumber="149" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B9F6F1E73442551FF65FDBDFC47C794" blockId="8.[151,1437,151,717]" box="[199,940,223,248]" pageId="8" pageNumber="149">
<emphasis id="B954B30C73442551FF65FDBDFEFDC794" bold="true" box="[199,278,223,248]" pageId="8" pageNumber="149">Name:</emphasis>
To emphasize the biramous gonopod telopodite; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C33A3C9573442551FF65FC66FC2CC62C" pageId="8" pageNumber="149" type="diagnosis">
<paragraph id="8B9F6F1E73442551FF65FC66FC2CC62C" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="149">
<emphasis id="B954B30C73442551FF65FC66FEA8C671" bold="true" box="[199,323,260,285]" pageId="8" pageNumber="149">Diagnosis:</emphasis>
Differs from the sole congener, the
<typeStatus id="549BD1BC73442551FD5DFC67FCC4C671" box="[767,815,261,285]" pageId="8" pageNumber="149">type</typeStatus>
species
<taxonomicName id="4C20149D73442551FC3BFC67FBC2C670" box="[921,1065,261,284]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
<emphasis id="B954B30C73442551FC3BFC67FBC2C670" box="[921,1065,261,284]" italics="true" pageId="8" pageNumber="149">K. unicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2670E7773442551FB94FC67FB7EC670" box="[1078,1173,261,284]" pageId="8" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
, in the lack of a fore projection on the collum, as well as by the biramous gonopod telopodite.
</paragraph>
</subSubSection>
<subSubSection id="C33A3C9573442551FF65FC2EFDADC5A0" pageId="8" pageNumber="149" type="description">
<paragraph id="8B9F6F1E73442551FF65FC2EFCC5C6C1" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="149">
<emphasis id="B954B30C73442551FF65FC2EFEB0C609" bold="true" box="[199,347,332,357]" pageId="8" pageNumber="149">Description:</emphasis>
Length in both sexes ca
<quantity id="4CD8C2FB73442551FD29FC2EFCE2C608" box="[651,777,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="8" pageNumber="149" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
, width of pro- and metazonae 1.21.3 and
<quantity id="4CD8C2FB73442551FAA8FC2EFA73C608" box="[1290,1432,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="149" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.61.8 mm</quantity>
, respectively.
<typeStatus id="549BD1BC73442551FE8FFC0DFE7EC6E4" box="[301,405,367,392]" pageId="8" pageNumber="149" type="holotype">Holotype</typeStatus>
ca
<quantity id="4CD8C2FB73442551FE62FC12FDFBC6EB" box="[448,528,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="149" unit="mm" value="11.0">11 mm</quantity>
long, 1.2 and
<quantity id="4CD8C2FB73442551FD1BFC0DFCFAC6EB" box="[697,785,367,392]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="149" unit="mm" value="1.6">1.6 mm</quantity>
wide on pro- and metazonae, respectively. Coloration in alcohol rather uniformly grey-brown to dark bown (
<figureCitation id="131B739B73442551FD7EFCF6FCCBC6C1" box="[732,800,404,429]" captionStart="FIGURE 5" captionStartId="5.[151,249,1330,1352]" captionTargetBox="[179,1392,465,1290]" captionTargetId="figure@5.[151,1436,447,1305]" captionTargetPageId="5" captionText="FIGURE 5. Habitus of Koponenius biramus sp. nov., male paratype, lateral view. Picture by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229196/files/figure.png" pageId="8" pageNumber="149">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B9F6F1E73442551FF65FCD5FC8EC6BC" blockId="8.[151,1437,151,717]" box="[199,869,439,464]" pageId="8" pageNumber="149">
All characters as in
<taxonomicName id="4C20149D73442551FE07FCDBFDDBC6A3" box="[421,560,440,464]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="149" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
<emphasis id="B954B30C73442551FE07FCDBFDDBC6A3" box="[421,560,440,464]" italics="true" pageId="8" pageNumber="149">K. unicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2670E7773442551FD94FCDAFD64C6A3" box="[566,655,440,463]" pageId="8" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
, except as follows.
</paragraph>
<paragraph id="8B9F6F1E73442551FF65FCBEFCE7C50C" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="149">
Collums fore margin regularly rounded, faintly lobulated, but covering the head from above, totally devoid of a fore protuberance (
<figureCitation id="131B739B73442551FEDFFC9DFE28C574" box="[381,451,511,536]" captionStart="FIGURE 5" captionStartId="5.[151,249,1330,1352]" captionTargetBox="[179,1392,465,1290]" captionTargetId="figure@5.[151,1436,447,1305]" captionTargetPageId="5" captionText="FIGURE 5. Habitus of Koponenius biramus sp. nov., male paratype, lateral view. Picture by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229196/files/figure.png" pageId="8" pageNumber="149">Figs 5</figureCitation>
&amp;
<figureCitation id="131B739B73442551FE47FC9DFE18C574" box="[485,499,511,536]" captionStart="FIGURE 6" captionStartId="6.[151,249,1763,1785]" captionTargetBox="[151,1436,195,1737]" captionTargetId="figure@6.[151,1436,195,1738]" captionTargetPageId="6" captionText="FIGURE 6. Koponenius biramus sp. nov., male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A G, I L, 0.2 mm; N, 0.05 mm; L, 0.02 mm." httpUri="https://zenodo.org/record/229197/files/figure.png" pageId="8" pageNumber="149">6</figureCitation>
A, D, G, J). Paraterga more wing-shaped and their caudolateral corners usually more acute (
<figureCitation id="131B739B73442551FF46FF46FECCC551" box="[228,295,548,573]" captionStart="FIGURE 6" captionStartId="6.[151,249,1763,1785]" captionTargetBox="[151,1436,195,1737]" captionTargetId="figure@6.[151,1436,195,1738]" captionTargetPageId="6" captionText="FIGURE 6. Koponenius biramus sp. nov., male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A G, I L, 0.2 mm; N, 0.05 mm; L, 0.02 mm." httpUri="https://zenodo.org/record/229197/files/figure.png" pageId="8" pageNumber="149">Fig. 6</figureCitation>
AF, H, I). Metatergal bosses usually more strongly obliterated (
<figureCitation id="131B739B73442551FBB2FF46FBB2C551" box="[1040,1113,548,573]" captionStart="FIGURE 5" captionStartId="5.[151,249,1330,1352]" captionTargetBox="[179,1392,465,1290]" captionTargetId="figure@5.[151,1436,447,1305]" captionTargetPageId="5" captionText="FIGURE 5. Habitus of Koponenius biramus sp. nov., male paratype, lateral view. Picture by K. Makarov, not taken to scale." httpUri="https://zenodo.org/record/229196/files/figure.png" pageId="8" pageNumber="149">Figs 5</figureCitation>
&amp;
<figureCitation id="131B739B73442551FB20FF46FB64C551" box="[1154,1167,548,573]" captionStart="FIGURE 6" captionStartId="6.[151,249,1763,1785]" captionTargetBox="[151,1436,195,1737]" captionTargetId="figure@6.[151,1436,195,1738]" captionTargetPageId="6" captionText="FIGURE 6. Koponenius biramus sp. nov., male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A G, I L, 0.2 mm; N, 0.05 mm; L, 0.02 mm." httpUri="https://zenodo.org/record/229197/files/figure.png" pageId="8" pageNumber="149">6</figureCitation>
AF, J, K). Limbus not crenulate, but long, dense, sharp blades (
<figureCitation id="131B739B73442551FDFCFF25FD48C50C" box="[606,675,583,608]" captionStart="FIGURE 6" captionStartId="6.[151,249,1763,1785]" captionTargetBox="[151,1436,195,1737]" captionTargetId="figure@6.[151,1436,195,1738]" captionTargetPageId="6" captionText="FIGURE 6. Koponenius biramus sp. nov., male paratype. A, D, G, J: anterior body part, dorsal, lateral, ventral and frontoventral views, respectively. B, E, H: midbody segments, dorsal, lateral and ventral views, respectively. C, F, I: posterior body part, dorsal, lateral and ventral views, respectively. K: cross-section of a midbody segment. L: tergal texture, dorsal view. M: ozopore region, dorsolateral view. Scale bars: H, 0.5 mm; A G, I L, 0.2 mm; N, 0.05 mm; L, 0.02 mm." httpUri="https://zenodo.org/record/229197/files/figure.png" pageId="8" pageNumber="149">Figs 6</figureCitation>
L &amp; 7A).
</paragraph>
<paragraph id="8B9F6F1E73442551FF65FF0EFDADC5A0" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="149">
Gonopods (Figs 7CF &amp; 8B) with two conspicuous, oblique, mesal, prefemoral ribs basal to main sulcus (
<emphasis id="B954B30C73442551FA2AFF0EFA78C5E9" bold="true" box="[1416,1427,620,645]" pageId="8" pageNumber="149">s</emphasis>
) demarcating a slender, biramous, equally long acropodite consisting of a subflagelliform solenomere (
<emphasis id="B954B30C73442551FA94FFEDFAA2C5C4" bold="true" box="[1334,1353,655,680]" pageId="8" pageNumber="149">sl</emphasis>
) and a similarly slender and long process (
<emphasis id="B954B30C73442551FD84FFD6FDDEC5A1" bold="true" box="[550,565,692,717]" pageId="8" pageNumber="149">p</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>