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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.210.3042" ID-PMC="PMC3406452" ID-Pensoft-Pub="1313-2970-210-19" ID-Pensoft-UUID="BD1FF0348E7DFFF87D580B20FFF06438" ID-PubMed="22859895" ID-Zenodo-Dep="577401" ID-ZooBank="DF32BC38E8394DC48EA0FB6508EC7B2B" ModsDocID="1313-2970-210-19" checkinTime="1451248849322" checkinUser="pensoft" docAuthor="Valdes, Pavel" docDate="2012" docId="175459E19565B9924855B8D6BB883E30" docLanguage="en" docName="ZooKeys 210: 19-67" docOrigin="ZooKeys 210" docPubDate="2012-07-24" docSource="http://dx.doi.org/10.3897/zookeys.210.3042" docTitle="Semiardistomis Kult 1950" docType="treatment" docVersion="5" id="BD1FF0348E7DFFF87D580B20FFF06438" lastPageNumber="29" masterDocId="BD1FF0348E7DFFF87D580B20FFF06438" masterDocTitle="Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)" masterLastPageNumber="67" masterPageNumber="19" pageNumber="22" updateTime="1668154243518" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Notes about morphological features of the Western Hemisphere subtribe Ardistomina, and revision of genus Semiardistomis Kult (Coleoptera, Carabidae, Scaritinae, Clivinini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Valdes, Pavel</mods:namePart>
<mods:affiliation>Gertrudis 365 apto 5 e / D'Strampes y Goicuria, Cp 10500, Habana, Cuba</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-07-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>210</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>67</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.210.3042</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.210.3042</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-210-19</mods:identifier>
<mods:identifier type="ZooBank">DF32BC38E8394DC48EA0FB6508EC7B2B</mods:identifier>
<mods:identifier type="Pensoft-UUID">BD1FF0348E7DFFF87D580B20FFF06438</mods:identifier>
<mods:identifier type="Zenodo-Dep">577401</mods:identifier>
</mods:mods>
<subSection lastPageId="11" lastPageNumber="30" pageId="3" pageNumber="22" type="taxonomy">
<treatment ID-GBIF-Taxon="152036405" LSID="urn:lsid:plazi:treatment:175459E19565B9924855B8D6BB883E30" httpUri="http://treatment.plazi.org/id/175459E19565B9924855B8D6BB883E30" lastPageId="11" lastPageNumber="29" pageId="3" pageNumber="22">
<subSubSection pageId="3" pageNumber="22" type="nomenclature">
<paragraph pageId="3" pageNumber="22">
Genus
<taxonomicName LSID="175459E1-9565-B992-4855-B8D6BB883E30" authority="Kult, 1950" authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Semiardistomis" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">Semiardistomis Kult, 1950</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="22" type="reference_group">
<paragraph pageId="3" pageNumber="22">
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Semiardistomis">Ardistomis (Semiardistomis)</taxonomicName>
Kult, 1950: 301. Type species:
<taxonomicName authorityName="Chaudoir" authorityYear="1837" class="Hexapoda" family="Carabidae" genus="Clivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="3" pageNumber="22">Clivina labialis</emphasis>
</taxonomicName>
Chaudoir, 1837, designated by
<bibRefCitation author="Kult, K" journalOrPublisher="Arthropoda" pageId="34" pageNumber="53" pagination="299 - 325" refId="B20" refString="Kult, K, 1950. New Neotropical species of the group Ardistominina (Carabidae, Col.). Arthropoda 1: 299 - 325" title="New Neotropical species of the group Ardistominina (Carabidae, Col.)." volume="1" year="1950">Kult (1950</bibRefCitation>
: 301).
</paragraph>
<paragraph pageId="3" pageNumber="22">
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Ardistomiellus">Ardistomis (Ardistomiellus)</taxonomicName>
Kult, 1950: 303. Type Species:
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="viridis">
<emphasis italics="true" pageId="3" pageNumber="22">Ardistomis viridis</emphasis>
</taxonomicName>
Say, 1823, designated by
<bibRefCitation author="Kult, K" journalOrPublisher="Arthropoda" pageId="34" pageNumber="53" pagination="299 - 325" refId="B20" refString="Kult, K, 1950. New Neotropical species of the group Ardistominina (Carabidae, Col.). Arthropoda 1: 299 - 325" title="New Neotropical species of the group Ardistominina (Carabidae, Col.)." volume="1" year="1950">Kult (1950</bibRefCitation>
: 303). Synonymy established by
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Quaestiones entomologicae" pageId="34" pageNumber="53" pagination="131 - 348" refId="B37" refString="Whitehead, DR, 1972. Classification, phylogeny, and zoogeography of Schizogenius Putzeys (Coleoptera: Carabidae: Scaritini). Quaestiones entomologicae 8: 131 - 348" title="Classification, phylogeny, and zoogeography of Schizogenius Putzeys (Coleoptera: Carabidae: Scaritini)." volume="8" year="1972">Whitehead [in Reichardt] (1977</bibRefCitation>
: 392).
</paragraph>
<paragraph pageId="3" pageNumber="22">
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">Semiardistomis</taxonomicName>
Kult.
<bibRefCitation author="Nichols, SW" editor="Liebherr, JK" journalOrPublisher="Comstock Publishing Associates, Ithaca" pageId="34" pageNumber="53" pagination="71 - 120" refId="B30" refString="Nichols, SW, 1988b. Kaleidoscopic biogeography of West Indian Scaritinae (Coleoptera: Carabidae). In: Liebherr, JK, Ed., Zoogeography of Caribbean Insects. Comstock Publishing Associates, Ithaca: 71 - 120" title="Kaleidoscopic biogeography of West Indian Scaritinae (Coleoptera: Carabidae)." volumeTitle="Zoogeography of Caribbean Insects." year="1988 b">Nichols (1988b</bibRefCitation>
: 98);
<bibRefCitation author="Ball, GE" editor="Arnett RH, Jr" journalOrPublisher="Volume 1. CRC Press, Boca Raton" pageId="34" pageNumber="53" pagination="32 - 132" refId="B3" refString="Ball, GE, Bousquet, Y, 2001. Carabidae Latreille, 1810. In: Arnett RH, Jr, Thomas, MC, Eds., American beetles: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Volume 1. CRC Press, Boca Raton: 32 - 132" title="Carabidae Latreille, 1810." volumeTitle="American beetles: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia." year="2001">Ball and Bousquet (2001</bibRefCitation>
: 45, 74);
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Zootaxa" pageId="34" pageNumber="53" pagination="1 - 29" refId="B7" refString="Bousquet, Y, 2006. Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico. Zootaxa 1308: 1 - 29" title="Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico." volume="1308" year="2006">Bousquet (2006</bibRefCitation>
: 10);
<bibRefCitation author="Valdes, P" journalOrPublisher="Zootaxa" pageId="34" pageNumber="53" pagination="23 - 33" refId="B35" refString="Valdes, P, 2007. Ardistomis quixotei (Coleoptera: Carabidae: Clivinini), a new Middle American species: structural and habitat features of adults, larvae and pupae, comparisons with previously described species, and notes about classification and biogeography. Zootaxa 1497: 23 - 33" title="Ardistomis quixotei (Coleoptera: Carabidae: Clivinini), a new Middle American species: structural and habitat features of adults, larvae and pupae, comparisons with previously described species, and notes about classification and biogeography." volume="1497" year="2007">
<normalizedToken originalValue="Valdés">Valdes</normalizedToken>
(2007
</bibRefCitation>
: 24);
<bibRefCitation author="Erwin, TL" journalOrPublisher="Pensoft, Sofia-Moscow" pageId="34" pageNumber="53" refId="B15" refString="Erwin, TL, 2011. A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - loxomeriformes, melaeniformes). Pensoft, Sofia-Moscow, 412 pp." title="A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life Volume III (Carabidae - loxomeriformes, melaeniformes)." year="2011">Erwin (2011</bibRefCitation>
: 233).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="22" type="diagnostic combination">
<paragraph pageId="3" pageNumber="22">Diagnostic combination.</paragraph>
<paragraph pageId="3" pageNumber="22">
Features of adults used in recognition of this genus include: antennomere 2 shorter than antennomere 3 (
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="3" pageNumber="22">Fig. 1</figureCitation>
,
<emphasis bold="true" pageId="3" pageNumber="22">a2, a3</emphasis>
); glossal sclerite of ligula (
<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="3" pageNumber="22">Figs 13B, 14B</figureCitation>
) with a secondary pair of setae; mentum with median carina (
<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="3" pageNumber="22">Figs 13C, 14C</figureCitation>
) extended distal in form of an appendiform keel; protibia (
<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="3" pageNumber="22">Figs 23, 24</figureCitation>
) with basal half of ventral surface with a group of setae.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="27" pageId="3" pageNumber="22" type="description">
<paragraph pageId="3" pageNumber="22">Description.</paragraph>
<paragraph pageId="3" pageNumber="22">
<emphasis bold="true" pageId="3" pageNumber="22">Adult. Body shape, color and size</emphasis>
(
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="3" pageNumber="22">Figs 1, 2</figureCitation>
,
<figureCitation captionStart="Figures 37–38" captionStartId="F9" captionText="Figures 37 - 38. 37 Semiardistomis cordicollis (Putzeys) 38 Semiardistomis exspectatus sp. n. Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19667" pageId="3" pageNumber="22">37</figureCitation>
-
<figureCitation captionStart="Figures 39–40" captionStartId="F10" captionText="Figures 39 - 40. 39 Semiardistomis darlingtoni (Kult) (teneral) 40 Semiardistomis subglabra (van Emden). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19668" pageId="3" pageNumber="22">41</figureCitation>
,
<figureCitation captionStart="Figures 45–46" captionStartId="F12" captionText="Figures 45 - 46. 45 Semiardistomis jedlickai (Kult) 46 Semiardistomis maindroni (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19670" pageId="3" pageNumber="22">44</figureCitation>
-
<figureCitation captionStart="Figures 51–52" captionStartId="F15" captionText="Figures 51 - 52. 51 Semiardistomis cyaneolimbatus (Chevrolat) 52 Semiardistomis pallipes (Dejean). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19673" pageId="3" pageNumber="22">52</figureCitation>
): Body pedunculate. Body size small, ranging from 4.0 to 7.5 mm. Body monochromous, dark with metallic greenish and/ or brassy luster, shiny in species without microsculpture; appendages reddish brown. One species,
<taxonomicName baseAuthorityName="Fleutiaux &amp; Salle" baseAuthorityYear="1889" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="laevistriatus">
<emphasis italics="true" pageId="3" pageNumber="22">Semiardistomis laevistriatus</emphasis>
</taxonomicName>
(Fleutiaux &amp;
<normalizedToken originalValue="Sallé">Salle</normalizedToken>
), is dark brown a color also found in monticolous brachypterous species of the genus
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="22">Ardistomis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22">
<emphasis bold="true" pageId="3" pageNumber="22">Microsculpture:</emphasis>
Frontoclypeus mostly smooth; supraantennal lobes smooth; vertex with mesh pattern isodiametric; gena with mesh pattern isodiametric; gula with mesh pattern transverse. Mandibles smooth; submentum and mentum with mesh pattern isodiametric. Pronotal disc generally smooth or with mesh pattern
<pageBreakToken pageId="4" pageNumber="23" start="start">isodiametric</pageBreakToken>
, microlines very shallow, marginally and smooth at the center; proepisternum smooth or with isodiametric mesh pattern, submarginal band of microsculpture absent; prosternum smooth or with mesh pattern transverse. Metasternum smooth or with mesh pattern transverse. Abdominal sterna smooth or with mesh pattern transverse. Elytra smooth or with isodiametric mesh pattern covering entirely or only part of the disc.
</paragraph>
<paragraph pageId="4" pageNumber="23">
<emphasis bold="true" pageId="4" pageNumber="23">Chaetotaxy:</emphasis>
Two pair of supraorbital setae (except for
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="4" pageNumber="23">Semiardistomis puncticollis</emphasis>
</taxonomicName>
); pronotal disc with two pair of marginal setae (except for
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="4" pageNumber="23">Semiardistomis puncticollis</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="viridis">
<emphasis italics="true" pageId="4" pageNumber="23">Semiardistomis viridis</emphasis>
</taxonomicName>
); except for hirsute species, setae on elytral disc located in interval 3 and
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis italics="true" pageId="4" pageNumber="23">Semiardistomis darlingtoni</emphasis>
</taxonomicName>
(Kult) also in interval 5; abdominal sterna IV-VII with accessory setae.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
<emphasis bold="true" pageId="4" pageNumber="23">Head capsule:</emphasis>
Clypeus with anterior margin concave medially; lateral lobes distinct, projected at the same level or below anterior margin. Antennal lobes prominent, sometimes partially sulcate medially in basal half. Frontoclypeus smooth except in
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis italics="true" pageId="4" pageNumber="23">Semiardistomis rugosus</emphasis>
</taxonomicName>
(Putzeys). Frontal impressions deep and wide from base of antennal lobes to anterior margin of clypeus. Gena with sides subparallel.
<emphasis bold="true" pageId="4" pageNumber="23">Eyes:</emphasis>
Hemispherical, prominent.
<emphasis bold="true" pageId="4" pageNumber="23">Antennae</emphasis>
(
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="4" pageNumber="23">Figs 1, 2</figureCitation>
): Filiform to submoniliform. Antennomere 1 (
<emphasis bold="true" pageId="4" pageNumber="23">a1</emphasis>
) with
<pageBreakToken pageId="5" pageNumber="24" start="start">single</pageBreakToken>
preapical seta. Antennomere 2 (
<emphasis bold="true" pageId="5" pageNumber="24">a2</emphasis>
) with few setae, shorter than antennomere 3 (
<emphasis bold="true" pageId="5" pageNumber="24">a3</emphasis>
). Antennomeres 4-11 densely setose, setae short.
</paragraph>
<paragraph lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24">
<emphasis bold="true" pageId="5" pageNumber="24">Mouthparts:</emphasis>
<emphasis bold="true" pageId="5" pageNumber="24">Labrum</emphasis>
(
<figureCitation captionStart="Figures 3–5" captionStartId="F2" captionText="Figures 3 - 5. 3 - 4 Labrum (dorsal aspect) 3 Semiardistomis glabratus (Putzeys) 4 Semiardistomis labialis (Chaudoir) 5 RightMaxilla (dorsal aspect) of Semiardistomis labialis (Chaudoir). Legend: Labrum: ds dorsal seta es epipharyngeal seta lp labral dentiform projection. Maxilla: c cardo g 1 galeomere 1 g 2 galeomere 2 lac lacinia mp 2 mp 3 mp 4 maxillary palpomeres 2 - 4 respectively pf palpifer s stipes. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19660" pageId="5" pageNumber="24">Figs 3, 4</figureCitation>
): anterior margin angulate, dentiform medially (
<emphasis bold="true" pageId="5" pageNumber="24">dp</emphasis>
); dorsal setae (
<emphasis bold="true" pageId="5" pageNumber="24">ds</emphasis>
) 7, as for most
<taxonomicName authorityName="Rafinesque" authorityYear="1815" lsidName="" pageId="5" pageNumber="24" rank="tribe" tribe="Clivinini">Clivinini</taxonomicName>
; apical portion of epipharyngeal setae (
<emphasis bold="true" pageId="5" pageNumber="24">es</emphasis>
) serrulate anteriolaterally.
<emphasis bold="true" pageId="5" pageNumber="24">Maxillae</emphasis>
(
<figureCitation captionStart="Figures 6–12" captionStartId="F3" captionText="Figures 6 - 12. Mandibles ofArdistomina, Clivinina and Dyschiriina with length / width ratios: A left, dorsal aspect B right, dorsal aspect C left, ventral aspect and D right, ventral aspect) 6 Semiardistomis labialis (Chaudoir) 7 Ardistomis fasciolatus Putzeys 8 Aspidoglossa mexicana (Chaudoir) 9 Oxydrepanus rufus (Putzeys) 10 Clivina dentipes Dejean 11 Schizogenius arimao Darlington 12 Dyschiriodes larochellei Bousquet. Legend: Occlusal teeth: m molar pm premolar rt retinacular tt terebral. Ventral surface: vg ventral groove mt microtrichia. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19661" pageId="5" pageNumber="24">Fig. 5</figureCitation>
) with lacinia (
<emphasis bold="true" pageId="5" pageNumber="24">lac</emphasis>
) apical tooth sharp and curved; galeomere 1 (
<emphasis bold="true" pageId="5" pageNumber="24">g1</emphasis>
) slender, a little longer than galeomere 2 (
<emphasis bold="true" pageId="5" pageNumber="24">g2</emphasis>
), the latter flat, broad and sinuate in outline; palpomere 2 (
<emphasis bold="true" pageId="5" pageNumber="24">mp2</emphasis>
) thick and triangular in outline; palpomere 3 (
<emphasis bold="true" pageId="5" pageNumber="24">mp3</emphasis>
) subequal in length to palpomere 4 (
<emphasis bold="true" pageId="5" pageNumber="24">mp4</emphasis>
); palpomere 4 apically acuminate. Stipes (
<emphasis bold="true" pageId="5" pageNumber="24">st</emphasis>
) with 4 setae, palpifer (
<emphasis bold="true" pageId="5" pageNumber="24">pf</emphasis>
) with one seta. (This structure shows little difference from other clivinines being most similar to that of
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="24">Ardistomis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="5" pageNumber="24">Mandibles</emphasis>
(
<figureCitation captionStart="Figures 6–12" captionStartId="F3" captionText="Figures 6 - 12. Mandibles ofArdistomina, Clivinina and Dyschiriina with length / width ratios: A left, dorsal aspect B right, dorsal aspect C left, ventral aspect and D right, ventral aspect) 6 Semiardistomis labialis (Chaudoir) 7 Ardistomis fasciolatus Putzeys 8 Aspidoglossa mexicana (Chaudoir) 9 Oxydrepanus rufus (Putzeys) 10 Clivina dentipes Dejean 11 Schizogenius arimao Darlington 12 Dyschiriodes larochellei Bousquet. Legend: Occlusal teeth: m molar pm premolar rt retinacular tt terebral. Ventral surface: vg ventral groove mt microtrichia. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19661" pageId="5" pageNumber="24">Figs 6A-6D</figureCitation>
): elongate, about 1.9 times longer than wide; ventral groove (
<emphasis bold="true" pageId="5" pageNumber="24">vg</emphasis>
) moderate in length, with short microtrichia (
<emphasis bold="true" pageId="5" pageNumber="24">mt</emphasis>
); left mandible with terebral ridge curved, the terebral tooth (
<emphasis bold="true" pageId="5" pageNumber="24">tt</emphasis>
) blunt and moderate in size, visible from ventral view, retinaculum (
<emphasis bold="true" pageId="5" pageNumber="24">rt</emphasis>
) broad, premolar tooth (
<emphasis bold="true" pageId="5" pageNumber="24">pm</emphasis>
) absent, the molar tooth (
<emphasis bold="true" pageId="5" pageNumber="24">m</emphasis>
) broad and rounded; right mandible with terebral ridge more or less straight, the terebral tooth small, not visible from ventral view, the retinaculum with acute anterior tooth, premolar tooth absent, and the molar tooth broad and rounded.
<emphasis bold="true" pageId="5" pageNumber="24">Labium</emphasis>
(
<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="5" pageNumber="24">Figs 13A-C, 14A-C</figureCitation>
) with mental-submental suture complete; submentum with paramedian projections distinct, two pair of setae; mentum (
<emphasis bold="true" pageId="5" pageNumber="24">m</emphasis>
) trapezoidal, about 1.5 times wider than long; two pair of setae; lateral lobes with anterior margins angulate; mental tooth (
<emphasis bold="true" pageId="5" pageNumber="24">mt</emphasis>
) with distal margin almost straight, about 1/3 length of lateral lobes (
<emphasis bold="true" pageId="5" pageNumber="24">ll</emphasis>
); median carina (
<emphasis bold="true" pageId="5" pageNumber="24">mc</emphasis>
)(13C, 14C) extended distad, or not, beyond anterior margin of lateral lobes; pit organs (
<emphasis bold="true" pageId="5" pageNumber="24">po</emphasis>
) distinct, opened through oval orifices in basal part of mentum or on mental-submental suture; palpomere 3 (
<emphasis bold="true" pageId="5" pageNumber="24">lp3</emphasis>
) subequal in length to palpomere 2, palpomere 2 (
<emphasis bold="true" pageId="5" pageNumber="24">lp2</emphasis>
) bisetose; ligula (
<figureCitation captionStart="Figures 13–21" captionStartId="F4" captionText="Figures 13 - 21. Labium ofClivinini (ventral aspects): A mentum and palpi B ligula C mental carina, lateral aspect. 13 - 14 Genus Semiardistomis: 13 S. glabratus (Putzeys) 14 Semiardistomis labialis (Chaudoir) 15 Ardistomis fasciolatus Putzeys 16 Aspidoglossa mexicana (Chaudoir) 17 Clivina dentipes Dejean 18 Oxydrepanus rufus (Putzeys) 19 Schizogenius arimao Darlington 20 Ancus excavaticeps (Putzeys) 21 Nyctosyles planicollis (Reiche). Legend: as apical setae of glossal sclerite gs glossal sclerite lp 2 labial palpomere 2 lp 3 labial palpomere 3 ll lateral lobes mc median carina m mentum mt mental tooth pas preapical seta of glossal sclerite pg paraglossa. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19662" pageId="5" pageNumber="24">Figs 13B, 14B</figureCitation>
) with glossal sclerite (
<emphasis bold="true" pageId="5" pageNumber="24">gs</emphasis>
) notably wide, its anterior margin with acuminate median projection and two long setae (
<emphasis bold="true" pageId="5" pageNumber="24">as</emphasis>
), two additional small
<pageBreakToken pageId="6" pageNumber="25" start="start">setae</pageBreakToken>
(
<emphasis bold="true" pageId="6" pageNumber="25">pas</emphasis>
) located preapically; paraglossae (
<emphasis bold="true" pageId="6" pageNumber="25">pg</emphasis>
) divergent from glossal sclerite and covered with microtrichia dorsally, lobes thin and acuminate, extended distally beyond distal margin of glossal sclerite, apices serrulate.
</paragraph>
<paragraph pageId="6" pageNumber="25">
<emphasis bold="true" pageId="6" pageNumber="25">Pronotum.</emphasis>
Ovate or cordate, anterior transverse and median longitudinal impressions distinct, proepisternum visible from above, lateral border not extended to base in some specimens. Anterior transverse and median longitudinal impressions distinct. Proepipleura visible from above.
</paragraph>
<paragraph pageId="6" pageNumber="25">
<emphasis bold="true" pageId="6" pageNumber="25">Elytra.</emphasis>
Oval, convex; striae complete or obliterated to various degrees or absent (stria 1 always complete), punctate or not, punctures shallowly to deeply impressed. Preapical epipleural plica (
<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="6" pageNumber="25">Fig. 22</figureCitation>
,
<emphasis bold="true" pageId="6" pageNumber="25">ep</emphasis>
) displaced outward, interrupting outline of elytra.
</paragraph>
<paragraph pageId="6" pageNumber="25">
<emphasis bold="true" pageId="6" pageNumber="25">Hind wings.</emphasis>
Macropterous or brachypterous.
</paragraph>
<paragraph pageId="6" pageNumber="25">
<emphasis bold="true" pageId="6" pageNumber="25">Underside</emphasis>
(
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="6" pageNumber="25">Fig. 2</figureCitation>
). Metasternum with a row of setiferous punctures along margin of coxal cavity, median contact with abdomen dividing metacoxae. Suture between abdominal sterna III and IV obliterated at middle. Pleurite VII with small projection(
<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="6" pageNumber="25">Fig. 22</figureCitation>
,
<emphasis bold="true" pageId="6" pageNumber="25">app</emphasis>
) fitting into elytral epipleural plica (
<emphasis bold="true" pageId="6" pageNumber="25">ep</emphasis>
,).
</paragraph>
<paragraph pageId="7" pageNumber="26">
<emphasis bold="true" pageId="7" pageNumber="26">
<pageBreakToken pageId="7" pageNumber="26" start="start">Legs</pageBreakToken>
.
</emphasis>
Slender, protibia (
<figureCitation captionStart="Figures 22–24" captionStartId="F5" captionText="Figures 22 - 24. 22 Body extremity (lateral aspect) of Semiardistomis labialis (Chaudoir) 23 - 24 Protarsus (ventral aspect) of Semiardistomis 23 Semiardistomis labialis (Chaudoir) 24 Semiardistomis glabratus (Putzeys). Legend: app abdominal epipleural projectionof abdominal sternum VII ep elytral plica. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19663" pageId="7" pageNumber="26">Figs 23, 24</figureCitation>
) with basal half of ventral surface with a group of setae, protarsomeres in males slightly dilated.
</paragraph>
<paragraph pageId="7" pageNumber="26">
<emphasis bold="true" pageId="7" pageNumber="26">Male genitalia</emphasis>
(
<figureCitation captionStart="Figures 25–26" captionStartId="F6" captionText="Figures 25 - 26. Male genitalia of genus Semiardistomis: A phallus and parameres in dorsal aspect B phallus and endophallus everted in lateral aspect 25 Semiardistomis labialis (Chaudoir) 26 Semiardistomis glabratus (Putzeys). Legend: ba basal apophysis bo basal opening bp basal projection bs basal sclerite ll left lobe rl right lobe. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19664" pageId="7" pageNumber="26">Figs 25, 26</figureCitation>
). Phallus lightly sclerotized; basal bulb with basal orifice (
<emphasis bold="true" pageId="7" pageNumber="26">bo</emphasis>
) oval, wide; basal projection (
<emphasis bold="true" pageId="7" pageNumber="26">bp</emphasis>
) distinct, apical margin curved; basal apophysis (
<emphasis bold="true" pageId="7" pageNumber="26">ba</emphasis>
) moderate; median portion tubular; apical portion formed by two lamellae (
<emphasis bold="true" pageId="7" pageNumber="26">rl</emphasis>
and
<emphasis bold="true" pageId="7" pageNumber="26">ll</emphasis>
). Endophallus pubescent in apical portion; basal sclerite (
<emphasis bold="true" pageId="7" pageNumber="26">bs</emphasis>
) clearly developed. Parameres subequal in length, the right one wider, both with 4-5 apical setae.
</paragraph>
<paragraph lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26">
<emphasis bold="true" pageId="7" pageNumber="26">Female genitalia</emphasis>
(
<figureCitation captionStart="Figures 27–30" captionStartId="F7" captionText="Figures 27 - 30. Female genitalia (ventral aspect). of Ardistomina 27 Semiardistomis glabratus (Putzeys) 28 Semiardistomis labialis (Chaudoir) 29 Ardistomis fasciolatus Putzeys 30 Aspidoglossa mexicana (Chaudoir). Legend: bc bursa copulatrix gc gonocoxa (gonocoxite 1 and 2, fused) lt laterotergite mt mediotergite sp spermatheca spd spermathecal duct spg spermathecal gland spgd spermathecal gland duct. Scale bar 0.1 mm." httpUri="https://binary.pensoft.net/fig/19665" pageId="7" pageNumber="26">Figs 27, 28</figureCitation>
). Ovipositor: gonocoxa (
<emphasis bold="true" pageId="7" pageNumber="26">gc</emphasis>
) unsegmented, very slender, with a pair of setae at apex, surface with micropores; laterotergites (
<emphasis bold="true" pageId="7" pageNumber="26">lt</emphasis>
) thinly sclerotized, flat and asetose, sclerotized border thin and folded in basal portion with a small anterior part; mediotergite (
<emphasis bold="true" pageId="7" pageNumber="26">mt</emphasis>
) wide, its width delimited by external borders
<pageBreakToken pageId="8" pageNumber="27" start="start">of</pageBreakToken>
laterotergites. Reproductive tract: bursa copulatrix (
<emphasis bold="true" pageId="8" pageNumber="27">bc</emphasis>
) elongate; spermathecal duct (
<emphasis bold="true" pageId="8" pageNumber="27">spd</emphasis>
) =distance from apex of bursa copulatrix to insertion point of spermathecal gland duct (
<emphasis bold="true" pageId="8" pageNumber="27">spgd</emphasis>
) relatively short or long, spermathecal gland duct elongate; spermatheca (
<emphasis bold="true" pageId="8" pageNumber="27">sp</emphasis>
) various in thickness, length, and conformation; see description of this structure for each of the two species-groups.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27" type="morphological notes">
<paragraph pageId="8" pageNumber="27">Morphological notes.</paragraph>
<paragraph lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27">
<emphasis bold="true" pageId="8" pageNumber="27">Cuticular sculpture:</emphasis>
The absence of a submarginal band of microsculpture on the proepisternum has been used to differentiate
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="8" pageNumber="27">Semiardistomis</emphasis>
</taxonomicName>
<pageBreakToken pageId="9" pageNumber="28" start="start">from</pageBreakToken>
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Ardistomis</emphasis>
</taxonomicName>
but the character state is not diagnostic since the band has been lost in at least two lineages of
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Ardistomis</emphasis>
</taxonomicName>
(Valdes, 2009).
</paragraph>
<paragraph pageId="9" pageNumber="28">The microsculpture on the elytral disc has a diagnostic value, the absence of microsculpture being the apotypic state.</paragraph>
<paragraph pageId="9" pageNumber="28">
<emphasis bold="true" pageId="9" pageNumber="28">Vestiture:</emphasis>
Hirsuteness is a common phenomenon in this genus. In the two species groups, this apotypic character state appears, its origin independently developed. In the
<emphasis italics="true" pageId="9" pageNumber="28">puncticollis</emphasis>
group, three species have hirsute structures:
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis puncticollis</emphasis>
</taxonomicName>
from southern U.S.A. (supraorbital area of the head capsule, pronotal disc, elytral disc, profemur and abdominal sternum VII),
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="pilosellus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis pilosellus</emphasis>
</taxonomicName>
(Kult) from southern South America (elytral disc and profemur) and
<taxonomicName baseAuthorityName="van Emden" baseAuthorityYear="1949" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="subglabra">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis subglabra</emphasis>
</taxonomicName>
(van Emden) from South America (profemur). In the
<emphasis italics="true" pageId="9" pageNumber="28">labialis</emphasis>
group, two species have hirsute structures:
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="viridis">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis viridis</emphasis>
</taxonomicName>
from southern U.S.A. (pronotal disc and elytral disc) and
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis propinquus</emphasis>
</taxonomicName>
(Putzeys) from Mexico (elytral disc),
</paragraph>
<paragraph pageId="9" pageNumber="28">
<emphasis bold="true" pageId="9" pageNumber="28">Elytra:</emphasis>
In this genus, I regard the state of having the striae complete and impunctate as plesiotypic, being present in both species groups.
</paragraph>
<paragraph pageId="9" pageNumber="28">
<emphasis bold="true" pageId="9" pageNumber="28">Hind wings:</emphasis>
Macropterous except
<taxonomicName baseAuthorityName="Fleutiaux &amp; Salle" baseAuthorityYear="1889" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="laevistriatus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis laevistriatus</emphasis>
</taxonomicName>
brachypterous.
<bibRefCitation author="Nichols, SW" journalOrPublisher="Doctoral dissertation, Cornell University, Ithaca, New York" pageId="34" pageNumber="53" refId="B29" refString="Nichols, SW, 1988a. Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Doctoral dissertation, Cornell University, Ithaca, New York, 393 pp." title="Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae)." year="1988 a">Nichols (1988a)</bibRefCitation>
also reported that
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis puncticollis</emphasis>
</taxonomicName>
is brachypterous.
</paragraph>
<paragraph pageId="9" pageNumber="28">
<emphasis bold="true" pageId="9" pageNumber="28">Legs:</emphasis>
Protarsomeres in males so slightly dilated that this feature is useless for sex recognition.
</paragraph>
<paragraph pageId="9" pageNumber="28">
<emphasis bold="true" pageId="9" pageNumber="28">Male genitalia:</emphasis>
Phallic structure enhances definition of the two species groups of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis</emphasis>
</taxonomicName>
. See details, provided in the species-group diagnoses. The differences in the phallic structures are too small to separate the species within each group, a situation also known in other clivinine genera, such as
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Schizogenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Schizogenius</emphasis>
</taxonomicName>
Putzeys (see
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Quaestiones entomologicae" pageId="34" pageNumber="53" pagination="131 - 348" refId="B37" refString="Whitehead, DR, 1972. Classification, phylogeny, and zoogeography of Schizogenius Putzeys (Coleoptera: Carabidae: Scaritini). Quaestiones entomologicae 8: 131 - 348" title="Classification, phylogeny, and zoogeography of Schizogenius Putzeys (Coleoptera: Carabidae: Scaritini)." volume="8" year="1972">Whitehead 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="28" type="larvae">
<paragraph pageId="9" pageNumber="28">Larvae.</paragraph>
<paragraph pageId="9" pageNumber="28">
The first instar of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis</emphasis>
</taxonomicName>
was described by
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Zootaxa" pageId="34" pageNumber="53" pagination="1 - 29" refId="B7" refString="Bousquet, Y, 2006. Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico. Zootaxa 1308: 1 - 29" title="Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico." volume="1308" year="2006">Bousquet (2006)</bibRefCitation>
. It was compared with
<taxonomicName authorityName="Putzeys" authorityYear="1846" class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Ardistomis</emphasis>
</taxonomicName>
larva by Valdes (2007) who pointed out two notable synapotypic features: absence of the coronal suture and a segmented maxillary palpomere 4.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="28" type="classification">
<paragraph pageId="9" pageNumber="28">Classification.</paragraph>
<paragraph pageId="9" pageNumber="28">
Based primarily on male and female genitalic features, the species of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis</emphasis>
</taxonomicName>
are arranged in two species-groups designated as the
<emphasis italics="true" pageId="9" pageNumber="28">puncticollis</emphasis>
and
<emphasis italics="true" pageId="9" pageNumber="28">labialis</emphasis>
group, respectively.
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="29" pageId="9" pageNumber="28" type="criteria for species definition">
<paragraph pageId="9" pageNumber="28">Criteria for species definition.</paragraph>
<paragraph pageId="9" pageNumber="28">
Of the 20 known species of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis</emphasis>
</taxonomicName>
only those occurring in U.S.A. (
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="viridis">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis viridis</emphasis>
</taxonomicName>
(Say) and
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="9" pageNumber="28">Semiardistomis puncticollis</emphasis>
</taxonomicName>
(Dejean)) are represented by large series of specimens in collections (
<bibRefCitation author="Nichols, SW" journalOrPublisher="Doctoral dissertation, Cornell University, Ithaca, New York" pageId="34" pageNumber="53" refId="B29" refString="Nichols, SW, 1988a. Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Doctoral dissertation, Cornell University, Ithaca, New York, 393 pp." title="Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae)." year="1988 a">Nichols 1988a</bibRefCitation>
;
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Zootaxa" pageId="34" pageNumber="53" pagination="1 - 29" refId="B7" refString="Bousquet, Y, 2006. Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico. Zootaxa 1308: 1 - 29" title="Review of the species of Ardistomina (Coleoptera: Carabidae: Clivinini) in America north of Mexico." volume="1308" year="2006">Bousquet 2006</bibRefCitation>
). The species inhabiting Central and South America and the West Indies are represented in collections by few specimens which give only an incomplete picture of the ranges of the species.
</paragraph>
<paragraph lastPageId="10" lastPageNumber="29" pageId="9" pageNumber="28">
Members of this genus are difficult to identify in part because of a shortage of useful external features, and in part because the male and female genitalia, so useful in distinguishing between species groups, are not useful within each group. Moreover, some of the characters traditionally used to separate the species have been found in large samples to vary extensively. Some samples are a mixture of combinations from extreme patterns considered as different species; being impossible to demonstrate if those populations are hybrids or simply polymorphs, while in other putative species, allopatric populations exhibit no evident differences. In the
<emphasis italics="true" pageId="9" pageNumber="28">labialis</emphasis>
group in southern South America, the species limits are especially difficult to assess probably because the group is young and composed of species with a great power of dispersal. Without direct evidence of the state of gene flow between known populations, it is almost impossible to testify in favor of the reproductive isolation of each species described or in other cases to designate subspecies. I will follow a criterion
<pageBreakToken pageId="10" pageNumber="29" start="start">with</pageBreakToken>
a practical value, so maybe for some species, taxonomic determination will denote unintentionally a combination of extreme characters shared by more than one biological species.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="29" type="geographical distribution">
<paragraph pageId="10" pageNumber="29">Geographical distribution.</paragraph>
<paragraph pageId="10" pageNumber="29">
(
<figureCitation captionStart="Figure 57" captionStartId="F19" captionText="Figure 57. Distribution map of members of the puncticollis species-group." httpUri="https://binary.pensoft.net/fig/19677" pageId="10" pageNumber="29">Figs 57</figureCitation>
-
<figureCitation captionStart="Figure 59" captionStartId="F21" captionText="Figure 59. Distribution map of members of the labialis species-group in Central America." httpUri="https://binary.pensoft.net/fig/19679" pageId="10" pageNumber="29">59</figureCitation>
)The range of this genus is co-extensive with the range of the
<taxonomicName authorityName="Putzeys" authorityYear="1867" genus="Ardistomina" lsidName="" pageId="10" pageNumber="29" rank="genus">Ardistomina</taxonomicName>
.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/19659" pageId="10" pageNumber="29" start="Figures 1–2" startId="F1">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 1-2.</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir).
<emphasis bold="true" pageId="10" pageNumber="29">1</emphasis>
Dorsal aspect
<emphasis bold="true" pageId="10" pageNumber="29">2</emphasis>
Ventral aspect. Legend:
<emphasis bold="true" pageId="10" pageNumber="29">a1 a2 a3</emphasis>
antennomeres 1-3, respectively. Scale bar 1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19660" pageId="10" pageNumber="29" start="Figures 3–5" startId="F2">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 3-5.</emphasis>
<emphasis bold="true" pageId="10" pageNumber="29">3-4</emphasis>
Labrum (dorsal aspect)
<emphasis bold="true" pageId="10" pageNumber="29">3</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis glabratus</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">4</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">5</emphasis>
RightMaxilla (dorsal aspect) of
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir). Legend: Labrum:
<emphasis bold="true" pageId="10" pageNumber="29">ds</emphasis>
dorsal seta
<emphasis bold="true" pageId="10" pageNumber="29">es</emphasis>
epipharyngeal seta
<emphasis bold="true" pageId="10" pageNumber="29">lp</emphasis>
labral dentiform projection. Maxilla:
<emphasis bold="true" pageId="10" pageNumber="29">c</emphasis>
cardo
<emphasis bold="true" pageId="10" pageNumber="29">g1</emphasis>
galeomere 1
<emphasis bold="true" pageId="10" pageNumber="29">g2</emphasis>
galeomere 2
<emphasis bold="true" pageId="10" pageNumber="29">lac</emphasis>
lacinia
<emphasis bold="true" pageId="10" pageNumber="29">mp2 mp3 mp4</emphasis>
maxillary palpomeres 2-4 respectively
<emphasis bold="true" pageId="10" pageNumber="29">pf</emphasis>
palpifer
<emphasis bold="true" pageId="10" pageNumber="29">s</emphasis>
stipes. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19661" pageId="10" pageNumber="29" start="Figures 6–12" startId="F3">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 6-12.</emphasis>
Mandibles ofArdistomina
<emphasis bold="true" pageId="10" pageNumber="29">,</emphasis>
<taxonomicName authorityName="Rafinesque" authorityYear="1815" genus="Clivinina" lsidName="" pageId="10" pageNumber="29" rank="genus">Clivinina</taxonomicName>
and
<taxonomicName genus="Dyschiriina" lsidName="" pageId="10" pageNumber="29" rank="genus">Dyschiriina</taxonomicName>
with length/width ratios:
<emphasis bold="true" pageId="10" pageNumber="29">A</emphasis>
left, dorsal aspect
<emphasis bold="true" pageId="10" pageNumber="29">B</emphasis>
right, dorsal aspect
<emphasis bold="true" pageId="10" pageNumber="29">C</emphasis>
left, ventral aspect and
<emphasis bold="true" pageId="10" pageNumber="29">D</emphasis>
right, ventral aspect)
<emphasis bold="true" pageId="10" pageNumber="29">6</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">7</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="fasciolatus">
<emphasis italics="true" pageId="10" pageNumber="29">Ardistomis fasciolatus</emphasis>
</taxonomicName>
Putzeys
<emphasis bold="true" pageId="10" pageNumber="29">8</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Aspidoglossa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis italics="true" pageId="10" pageNumber="29">Aspidoglossa mexicana</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">9</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Oxydrepanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="10" pageNumber="29">Oxydrepanus rufus</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">10</emphasis>
<taxonomicName authorityName="Dejean" authorityYear="1825" class="Hexapoda" family="Carabidae" genus="Clivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis italics="true" pageId="10" pageNumber="29">Clivina dentipes</emphasis>
</taxonomicName>
Dejean
<emphasis bold="true" pageId="10" pageNumber="29">11</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1934" class="Hexapoda" family="Carabidae" genus="Schizogenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="arimao">
<emphasis italics="true" pageId="10" pageNumber="29">Schizogenius arimao</emphasis>
</taxonomicName>
Darlington
<emphasis bold="true" pageId="10" pageNumber="29">12</emphasis>
<taxonomicName baseAuthorityName="Bousquet" baseAuthorityYear="1988" class="Insecta" family="Carabidae" genus="Dyschiriodes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="larochellei">
<emphasis italics="true" pageId="10" pageNumber="29">Dyschiriodes larochellei</emphasis>
</taxonomicName>
Bousquet. Legend: Occlusal teeth:
<emphasis bold="true" pageId="10" pageNumber="29">m</emphasis>
molar
<emphasis bold="true" pageId="10" pageNumber="29">pm</emphasis>
premolar
<emphasis bold="true" pageId="10" pageNumber="29">rt</emphasis>
retinacular
<emphasis bold="true" pageId="10" pageNumber="29">tt</emphasis>
terebral. Ventral surface:
<emphasis bold="true" pageId="10" pageNumber="29">vg</emphasis>
ventral groove
<emphasis bold="true" pageId="10" pageNumber="29">mt</emphasis>
microtrichia. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19662" pageId="10" pageNumber="29" start="Figures 13–21" startId="F4">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 13-21.</emphasis>
Labium ofClivinini (ventral aspects):
<emphasis bold="true" pageId="10" pageNumber="29">A</emphasis>
mentum and palpi
<emphasis bold="true" pageId="10" pageNumber="29">B</emphasis>
ligula
<emphasis bold="true" pageId="10" pageNumber="29">C</emphasis>
mental carina, lateral aspect.
<emphasis bold="true" pageId="10" pageNumber="29">13-14</emphasis>
Genus
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="10" pageNumber="29">13</emphasis>
S. glabratus (Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">14</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">15</emphasis>
<emphasis italics="true" pageId="10" pageNumber="29">Ardistomis fasciolatus</emphasis>
Putzeys
<emphasis bold="true" pageId="10" pageNumber="29">16</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Aspidoglossa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis italics="true" pageId="10" pageNumber="29">Aspidoglossa mexicana</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">17</emphasis>
<taxonomicName authorityName="Dejean" authorityYear="1825" class="Hexapoda" family="Carabidae" genus="Clivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis italics="true" pageId="10" pageNumber="29">Clivina dentipes</emphasis>
</taxonomicName>
Dejean
<emphasis bold="true" pageId="10" pageNumber="29">18</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Oxydrepanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="10" pageNumber="29">Oxydrepanus rufus</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">19</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1934" class="Hexapoda" family="Carabidae" genus="Schizogenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="arimao">
<emphasis italics="true" pageId="10" pageNumber="29">Schizogenius arimao</emphasis>
</taxonomicName>
Darlington
<emphasis bold="true" pageId="10" pageNumber="29">20</emphasis>
<taxonomicName authorityName="Putzeys" authorityYear="1867" class="Insecta" family="Carabidae" genus="Ancus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="excavaticeps">
<emphasis italics="true" pageId="10" pageNumber="29">Ancus excavaticeps</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">21</emphasis>
<taxonomicName baseAuthorityName="Reiche" baseAuthorityYear="1843" class="Hexapoda" family="Carabidae" genus="Nyctosyles" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="planicollis">
<emphasis italics="true" pageId="10" pageNumber="29">Nyctosyles planicollis</emphasis>
</taxonomicName>
(Reiche). Legend:
<emphasis bold="true" pageId="10" pageNumber="29">as</emphasis>
apical setae of glossal sclerite
<emphasis bold="true" pageId="10" pageNumber="29">gs</emphasis>
glossal sclerite
<emphasis bold="true" pageId="10" pageNumber="29">lp2</emphasis>
labial palpomere 2
<emphasis bold="true" pageId="10" pageNumber="29">lp3</emphasis>
labial palpomere 3
<emphasis bold="true" pageId="10" pageNumber="29">ll</emphasis>
lateral lobes
<emphasis bold="true" pageId="10" pageNumber="29">mc</emphasis>
median carina
<emphasis bold="true" pageId="10" pageNumber="29">m</emphasis>
mentum
<emphasis bold="true" pageId="10" pageNumber="29">mt</emphasis>
mental tooth
<emphasis bold="true" pageId="10" pageNumber="29">pas</emphasis>
preapical seta of glossal sclerite
<emphasis bold="true" pageId="10" pageNumber="29">pg</emphasis>
paraglossa. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19663" pageId="10" pageNumber="29" start="Figures 22–24" startId="F5">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 22-24.</emphasis>
<emphasis bold="true" pageId="10" pageNumber="29">22</emphasis>
Body extremity (lateral aspect) of
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">23-24</emphasis>
Protarsus (ventral aspect) of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="10" pageNumber="29">23</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">24</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis glabratus</emphasis>
</taxonomicName>
(Putzeys). Legend:
<emphasis bold="true" pageId="10" pageNumber="29">app</emphasis>
abdominal epipleural projectionof abdominal sternum VII
<emphasis bold="true" pageId="10" pageNumber="29">ep</emphasis>
elytral plica. Scale bar 1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19664" pageId="10" pageNumber="29" start="Figures 25–26" startId="F6">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 25-26.</emphasis>
Male genitalia of genus
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="10" pageNumber="29">A</emphasis>
phallus and parameres in dorsal aspect
<emphasis bold="true" pageId="10" pageNumber="29">B</emphasis>
phallus and endophallus everted in lateral aspect
<emphasis bold="true" pageId="10" pageNumber="29">25</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">26</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis glabratus</emphasis>
</taxonomicName>
(Putzeys). Legend:
<emphasis bold="true" pageId="10" pageNumber="29">ba</emphasis>
basal apophysis
<emphasis bold="true" pageId="10" pageNumber="29">bo</emphasis>
basal opening
<emphasis bold="true" pageId="10" pageNumber="29">bp</emphasis>
basal projection
<emphasis bold="true" pageId="10" pageNumber="29">bs</emphasis>
basal sclerite
<emphasis bold="true" pageId="10" pageNumber="29">ll</emphasis>
left lobe
<emphasis bold="true" pageId="10" pageNumber="29">rl</emphasis>
right lobe. Scale bar 0.1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19665" pageId="10" pageNumber="29" start="Figures 27–30" startId="F7">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 27-30.</emphasis>
Female genitalia (ventral aspect). of
<taxonomicName authorityName="Putzeys" authorityYear="1867" genus="Ardistomina" lsidName="" pageId="10" pageNumber="29" rank="genus">Ardistomina</taxonomicName>
<emphasis bold="true" pageId="10" pageNumber="29">27</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis glabratus</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">28</emphasis>
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
(Chaudoir)
<emphasis bold="true" pageId="10" pageNumber="29">29</emphasis>
<taxonomicName class="Hexapoda" family="Carabidae" genus="Ardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="fasciolatus">
<emphasis italics="true" pageId="10" pageNumber="29">Ardistomis fasciolatus</emphasis>
</taxonomicName>
Putzeys
<emphasis bold="true" pageId="10" pageNumber="29">30</emphasis>
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Aspidoglossa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis italics="true" pageId="10" pageNumber="29">Aspidoglossa mexicana</emphasis>
</taxonomicName>
(Chaudoir). Legend:
<emphasis bold="true" pageId="10" pageNumber="29">bc</emphasis>
bursa copulatrix
<emphasis bold="true" pageId="10" pageNumber="29">gc</emphasis>
gonocoxa (gonocoxite 1 and 2, fused)
<emphasis bold="true" pageId="10" pageNumber="29">lt</emphasis>
laterotergite
<emphasis bold="true" pageId="10" pageNumber="29">mt</emphasis>
mediotergite
<emphasis bold="true" pageId="10" pageNumber="29">sp</emphasis>
spermatheca
<emphasis bold="true" pageId="10" pageNumber="29">spd</emphasis>
spermathecal duct
<emphasis bold="true" pageId="10" pageNumber="29">spg</emphasis>
spermathecal gland
<emphasis bold="true" pageId="10" pageNumber="29">spgd</emphasis>
spermathecal gland duct.Scale bar 0.1 mm.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/19666" pageId="10" pageNumber="29" start="Figures 31–36" startId="F8">
<paragraph pageId="10" pageNumber="29">
<emphasis bold="true" pageId="10" pageNumber="29">Figures 31-36.</emphasis>
Female genitalia (dorsal aspect) and gonocoxites /laterotergites (lateral aspect) of
<taxonomicName authorityName="Rafinesque" authorityYear="1815" genus="Clivinina" lsidName="" pageId="10" pageNumber="29" rank="genus">Clivinina</taxonomicName>
<emphasis bold="true" pageId="10" pageNumber="29">31</emphasis>
<taxonomicName authorityName="Dejean" authorityYear="1825" class="Hexapoda" family="Carabidae" genus="Clivina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis italics="true" pageId="10" pageNumber="29">Clivina dentipes</emphasis>
</taxonomicName>
Dejean
<emphasis bold="true" pageId="10" pageNumber="29">32</emphasis>
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Oxydrepanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="rufus">
<emphasis italics="true" pageId="10" pageNumber="29">Oxydrepanus rufus</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">33</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1934" class="Hexapoda" family="Carabidae" genus="Schizogenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="arimao">
<emphasis italics="true" pageId="10" pageNumber="29">Schizogenius arimao</emphasis>
</taxonomicName>
Darlington
<emphasis bold="true" pageId="10" pageNumber="29">34</emphasis>
<taxonomicName baseAuthorityName="Reiche" baseAuthorityYear="1843" class="Hexapoda" family="Carabidae" genus="Nyctosyles" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="planicollis">
<emphasis italics="true" pageId="10" pageNumber="29">Nyctosyles planicollis</emphasis>
</taxonomicName>
(Reiche)
<emphasis bold="true" pageId="10" pageNumber="29">35</emphasis>
<taxonomicName authorityName="Putzeys" authorityYear="1867" class="Insecta" family="Carabidae" genus="Ancus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="excavaticeps">
<emphasis italics="true" pageId="10" pageNumber="29">Ancus excavaticeps</emphasis>
</taxonomicName>
(Putzeys)
<emphasis bold="true" pageId="10" pageNumber="29">36</emphasis>
<taxonomicName authorityName="Burmeister" authorityYear="1875" class="Insecta" family="Carabidae" genus="Obadius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="insignis">
<emphasis italics="true" pageId="10" pageNumber="29">Obadius insignis</emphasis>
</taxonomicName>
Burmeister. Legend: See legend for
<emphasis bold="true" pageId="10" pageNumber="29">Figs</emphasis>
<emphasis bold="true" pageId="10" pageNumber="29">27-30</emphasis>
, except gonocoxite 1 (
<emphasis bold="true" pageId="10" pageNumber="29">gc1</emphasis>
) and. gonocoxite 2 (
<emphasis bold="true" pageId="10" pageNumber="29">gc2</emphasis>
) Scale bar 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="30" pageId="10" pageNumber="29" type="key to described species of semiardistomis, based on adult characters">
<paragraph pageId="10" pageNumber="29">
Key to described species of
<taxonomicName authorityName="Kult" authorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis</emphasis>
</taxonomicName>
, based on adult characters
</paragraph>
<paragraph lastPageId="11" lastPageNumber="30" pageId="10" pageNumber="29">
<table inLine="true" lastPageId="11" lastPageNumber="30" pageId="10" pageNumber="29">
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">1</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral disc plurisetose</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">2</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">1'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral disc with setiferous punctures in interval 3, or sometimes also in interval 5</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">5</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">2(1)</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Profemur plurisetose</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">3</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">2'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Profemur with the usual setae</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">4</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">3(2)</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Pronotal disc plurisetose (along margin and on disc). Habitus as in
<figureCitation captionStart="Figures 47–48" captionStartId="F13" captionText="Figures 47 - 48. 47 Semiardistomis laevistriatus (Fleutiaux &amp; Salle) 48 Semiardistomis puncticollis (Dejean). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19671" pageId="10" pageNumber="29">Fig. 48</figureCitation>
. USA
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="puncticollis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis puncticollis</emphasis>
</taxonomicName>
(Dejean
<emphasis italics="true" pageId="10" pageNumber="29">)</emphasis>
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">3'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Pronotal disc with the usual 2 pairs of marginal setae. Habitus as in
<figureCitation captionStart="Figures 41–44" captionStartId="F11" captionText="Figures 41 - 44. 41 - 43 Semiardistomis glabratus (Putzeys) 41 exemplar from Peru, Loreto 42 exemplar from Argentina, Misiones 43 exemplar from Uruguay, Montevideo 44 Semiardistomis pilosellus (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19669" pageId="10" pageNumber="29">Fig. 44</figureCitation>
. South America
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="pilosellus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis pilosellus</emphasis>
</taxonomicName>
(Kult)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">4(2')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Pronotal disc plurisetose along margin. Habitus as in
<figureCitation captionStart="Figures 54–55" captionStartId="F17" captionText="Figures 54 - 55. 54 Semiardistomis propinquus (Putzeys) 55 Semiardistomis viridis (Say). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19675" pageId="10" pageNumber="29">Fig. 55</figureCitation>
. USA
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Say" baseAuthorityYear="1823" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="viridis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis viridis</emphasis>
</taxonomicName>
(Say)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">4'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Pronotal disc with the usual 2 pairs of marginal setae. Habitus as in
<figureCitation captionStart="Figures 54–55" captionStartId="F17" captionText="Figures 54 - 55. 54 Semiardistomis propinquus (Putzeys) 55 Semiardistomis viridis (Say). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19675" pageId="10" pageNumber="29">Fig. 54</figureCitation>
. Mexico
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="propinquus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis propinquus</emphasis>
</taxonomicName>
(Putzeys)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">5(1')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Elytron with two setiferous punctures on interval 3; humerus rounded; brachypterous. Habitus as in
<figureCitation captionStart="Figures 47–48" captionStartId="F13" captionText="Figures 47 - 48. 47 Semiardistomis laevistriatus (Fleutiaux &amp; Salle) 48 Semiardistomis puncticollis (Dejean). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19671" pageId="10" pageNumber="29">Fig. 47</figureCitation>
Guadeloupe
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Fleutiaux &amp; Salle" baseAuthorityYear="1889" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="laevistriatus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis laevistriatus</emphasis>
</taxonomicName>
(Fleutiaux &amp;
<normalizedToken originalValue="Sallé">Salle</normalizedToken>
<emphasis italics="true" pageId="10" pageNumber="29">)</emphasis>
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">5'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytron with three or more setiferous punctures on interval 3; macropterous</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">6</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">6(5')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytron with three setiferous punctures on interval 3</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">7</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">6'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytron with more than three setiferous punctures on interval 3</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">12</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">7(6)</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Elytral striae impunctate. Habitus as in
<figureCitation captionStart="Figures 51–52" captionStartId="F15" captionText="Figures 51 - 52. 51 Semiardistomis cyaneolimbatus (Chevrolat) 52 Semiardistomis pallipes (Dejean). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19673" pageId="10" pageNumber="29">Fig. 52</figureCitation>
. South America
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis pallipes</emphasis>
</taxonomicName>
(Dejean)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">7'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral striae punctate</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">8</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">8(7')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral disc with isodiametric mesh pattern</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">9</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">8'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral disc smooth</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">10</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">9(8)</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral striae impressed only in basal fifth, punctures deep along three-quarters of elytra, surface completely covered with isodiametric mesh pattern. South America</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="deletus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis deletus</emphasis>
</taxonomicName>
(Putzeys)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">9'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral striae obliterated in apical third, punctures moderately deep, surface with isodiametric mesh pattern, evanescent in central area of elytral disc in some specimens. South America</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis flavipes</emphasis>
</taxonomicName>
(Dejean)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">10(8')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Elytral striae obliterated in apical fifth, punctures moderately deep. Habitus as in
<figureCitation captionStart="Figures 51–52" captionStartId="F15" captionText="Figures 51 - 52. 51 Semiardistomis cyaneolimbatus (Chevrolat) 52 Semiardistomis pallipes (Dejean). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19673" pageId="10" pageNumber="29">Fig. 51</figureCitation>
. West Indies
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Chevrolat" baseAuthorityYear="1863" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="cyaneolimbatus">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis cyaneolimbatus</emphasis>
</taxonomicName>
(Chevrolat)
</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">10'</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">Elytral striae impressed only in basal fifth</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">11</td>
</tr>
<tr pageId="10" pageNumber="29">
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">11(10')</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
Elytral punctures deeply impressed in basal two-thirds of elytra. Habitus as in
<figureCitation captionStart="Figures 1–2" captionStartId="F1" captionText="Figures 1 - 2. Semiardistomis labialis (Chaudoir). 1 Dorsal aspect 2 Ventral aspect. Legend: a 1 a 2 a 3 antennomeres 1 - 3, respectively. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19659" pageId="10" pageNumber="29">Figs 1, 2</figureCitation>
. Central America
</td>
<td colspan="1" pageId="10" pageNumber="29" rowspan="1">
<taxonomicName baseAuthorityName="Chaudoir" baseAuthorityYear="1837" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="labialis">
<emphasis italics="true" pageId="10" pageNumber="29">Semiardistomis labialis</emphasis>
</taxonomicName>
(Chaudoir)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<pageBreakToken pageId="11" pageNumber="30" start="start">11</pageBreakToken>
'
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytral punctures moderately impressed in basal third of elytra. Southern South America</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="semipunctatus">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis semipunctatus</emphasis>
</taxonomicName>
(Dejean)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">12(6')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytral disc surface completely covered with isodiametric mesh pattern. Habitus as in
<figureCitation captionStart="Figures 45–46" captionStartId="F12" captionText="Figures 45 - 46. 45 Semiardistomis jedlickai (Kult) 46 Semiardistomis maindroni (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19670" pageId="11" pageNumber="30">Fig. 46</figureCitation>
. South America
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis maindroni</emphasis>
</taxonomicName>
(Kult)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">12'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytral disc surface smooth</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">13</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">13(12')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytral striae complete; impunctate</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">14</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">13'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytral striae incomplete</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">17</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">14(13)</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Front between antennal lobes with transverse rugae. Habitus as in
<figureCitation captionStart="Figures 49–50" captionStartId="F14" captionText="Figures 49 - 50.. 47 Semiardistomis major sp. n. 50 Semiardistomis rugosus (Putzeys). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19672" pageId="11" pageNumber="30">Fig. 50</figureCitation>
. Southern Brazil
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis rugosus</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">14'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Front between antennal lobes smooth</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">15</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">15(14')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytron with five setiferous punctures on interval 3; body size more than 7 mm. Habitus as in
<figureCitation captionStart="Figures 49–50" captionStartId="F14" captionText="Figures 49 - 50.. 47 Semiardistomis major sp. n. 50 Semiardistomis rugosus (Putzeys). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19672" pageId="11" pageNumber="30">Fig. 49</figureCitation>
. Peru
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName authorityName="Valdés" authorityYear="2012" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="major">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis major</emphasis>
</taxonomicName>
n. sp.
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">15'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytron with four setiferous punctures on interval 3; body size less than 6 mm</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">16</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">16(15')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Pronotum markedly cordate, 1.25 times wider than long; humeri square, sides subparallel. Habitus as in
<figureCitation captionStart="Figures 37–38" captionStartId="F9" captionText="Figures 37 - 38. 37 Semiardistomis cordicollis (Putzeys) 38 Semiardistomis exspectatus sp. n. Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19667" pageId="11" pageNumber="30">Fig. 37</figureCitation>
. Colombia
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1846" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="cordicollis">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis cordicollis</emphasis>
</taxonomicName>
(Putzeys)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">16'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Pronotum subcordate, 1.14 times wider than long; shoulders rounded. Habitus as in
<figureCitation captionStart="Figures 37–38" captionStartId="F9" captionText="Figures 37 - 38. 37 Semiardistomis cordicollis (Putzeys) 38 Semiardistomis exspectatus sp. n. Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19667" pageId="11" pageNumber="30">Fig. 38</figureCitation>
. South America.
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName authorityName="Valdés" authorityYear="2012" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="exspectatus">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis exspectatus</emphasis>
</taxonomicName>
n. sp.
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">17(13')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytron with eight setiferous punctures in interval 3</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">18</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">17'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">Elytron with four setiferous punctures in interval 3</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">19</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">18(17)</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytron with three setiferous punctures in interval 5. Habitus as in
<figureCitation captionStart="Figures 39–40" captionStartId="F10" captionText="Figures 39 - 40. 39 Semiardistomis darlingtoni (Kult) (teneral) 40 Semiardistomis subglabra (van Emden). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19668" pageId="11" pageNumber="30">Fig. 39</figureCitation>
. Brazil
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis darlingtoni</emphasis>
</taxonomicName>
(Kult)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">18'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytron with interval 5 asetose. Habitus as in
<figureCitation captionStart="Figures 39–40" captionStartId="F10" captionText="Figures 39 - 40. 39 Semiardistomis darlingtoni (Kult) (teneral) 40 Semiardistomis subglabra (van Emden). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19668" pageId="11" pageNumber="30">Fig. 40</figureCitation>
. South America
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="van Emden" baseAuthorityYear="1949" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="subglabra">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis subglabra</emphasis>
</taxonomicName>
(van Emden)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">19(17')</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytral striae 3,4,5 and 6 impressed in basal half of disc, stria 2 present. Body length around 4 mm. Habitus as in
<figureCitation captionStart="Figures 45–46" captionStartId="F12" captionText="Figures 45 - 46. 45 Semiardistomis jedlickai (Kult) 46 Semiardistomis maindroni (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19670" pageId="11" pageNumber="30">Fig. 45</figureCitation>
. South America
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Kult" baseAuthorityYear="1950" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="jedlickai">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis jedlickai</emphasis>
</taxonomicName>
(Kult)
</td>
</tr>
<tr pageId="11" pageNumber="30">
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">19'</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
Elytral striae 3, 4, 5 and 6 impressed in basal third of disc, stria 2 absent. Body length around 6 mm. Habitus as in
<figureCitation captionStart="Figures 41–44" captionStartId="F11" captionText="Figures 41 - 44. 41 - 43 Semiardistomis glabratus (Putzeys) 41 exemplar from Peru, Loreto 42 exemplar from Argentina, Misiones 43 exemplar from Uruguay, Montevideo 44 Semiardistomis pilosellus (Kult). Dorsal aspect. Scale bar 1 mm." httpUri="https://binary.pensoft.net/fig/19669" pageId="11" pageNumber="30">Fig. 41</figureCitation>
. South America
</td>
<td colspan="1" pageId="11" pageNumber="30" rowspan="1">
<taxonomicName baseAuthorityName="Putzeys" baseAuthorityYear="1866" class="Hexapoda" family="Carabidae" genus="Semiardistomis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis italics="true" pageId="11" pageNumber="30">Semiardistomis glabratus</emphasis>
</taxonomicName>
(Putzeys)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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