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<document id="72B875E48239A3B8B1A02BCE1818A6EE" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFECFFB5FCB47174DCBDFC42" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) wunderlei Assing 2016, spec. nov." docType="treatment" docVersion="11" lastPageNumber="107" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="106" updateTime="1699235685730" updateUser="ExternalLinkService">
<mods:mods id="E8B83DF59083FED219DD4432313361A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8D5D9CEBAFE72A067AD2D195B8CF176C">
<mods:title id="546F454D7BD80A6F9A59833764F7F20E">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="EE5DC3FD5B8CFD85C7B16549BC3FEC66" type="personal">
<mods:role id="C893DCB6E76A0FED9F04469DA8AABC79">
<mods:roleTerm id="348CC19CE2BFBB781165D72809B5EC9C">Author</mods:roleTerm>
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<mods:namePart id="73A7D8B0CC7F0BD60D7545D463794FB1">Assing, Volker</mods:namePart>
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<mods:title id="68CE3A4DEE4474DBC08B942B93C3645A">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="848291E4E27EE5F6201A8324FB984642">
<mods:date id="0C8FAD4022AE9C8C30B3CD87F86F1D70">2016</mods:date>
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<mods:number id="673AB600189AC85E606A1246EF202132">2016-12-20</mods:number>
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<mods:detail id="BA42A29F145C03FC6F70B749218688F9" type="volume">
<mods:number id="6BE98B450A2FC9FC13F1E08807623A3B">66</mods:number>
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<mods:number id="77ACAF00B8AF2C64A0962DA6C28F4D08">1</mods:number>
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<mods:url id="DD5C471F3D8F0380A03AB11297DE5BDB">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="40B38F9C0E667BC3B6BF46221541B20F">journal article</mods:classification>
<mods:identifier id="95FFB872343465C54F649A5A65C47DEE" type="CLB-Dataset">2419</mods:identifier>
<mods:identifier id="B9438A1FC903CCB34AB82EF540508BC8" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
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</mods:mods>
<treatment id="766F7C36FFECFFB5FCB47174DCBDFC42" ID-DOI="http://doi.org/10.5281/zenodo.5879539" ID-GBIF-Taxon="192602812" ID-Zenodo-Dep="5879539" LSID="urn:lsid:plazi:treatment:766F7C36FFECFFB5FCB47174DCBDFC42" httpUri="http://treatment.plazi.org/id/766F7C36FFECFFB5FCB47174DCBDFC42" lastPageId="94" lastPageNumber="107" pageId="93" pageNumber="106">
<subSubSection id="B6DC9EABFFECFFB6FCB47174DCE5FBED" box="[831,1181,1107,1134]" pageId="93" pageNumber="106" type="nomenclature">
<paragraph id="FE79CD20FFECFFB6FCB47174DCE5FBED" blockId="93.[831,1181,1107,1167]" box="[831,1181,1107,1134]" pageId="93" pageNumber="106">
<heading id="A5317A4CFFECFFB6FCB47174DCE5FBED" box="[831,1181,1107,1134]" centered="true" fontSize="11" level="2" pageId="93" pageNumber="106" reason="3">
<taxonomicName id="39C6B6A3FFECFFB6FCB47174DC56FBED" ID-CoL="5DDGH" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[831,1070,1107,1134]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="106" phylum="Arthropoda" rank="species" species="wunderlei" status="sp. nov." subGenus="Zyras">
<emphasis id="CCB21132FFECFFB6FCB47174DC56FBED" box="[831,1070,1107,1134]" italics="true" pageId="93" pageNumber="106">Zyras (Zyras) wunderlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FFECFFB6FBBF7174DCE5FBED" box="[1076,1181,1107,1134]" pageId="93" pageNumber="106" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFECFFB6FCB4715EDC6BFB0C" box="[831,1043,1145,1167]" pageId="93" pageNumber="106" type="description">
<paragraph id="FE79CD20FFECFFB6FCB4715EDC6BFB0C" blockId="93.[831,1181,1107,1167]" box="[831,1043,1145,1167]" pageId="93" pageNumber="106">
(
<figureCitation id="66FDD1A5FFECFFB6FCCF715EDBC2FB0C" box="[836,954,1145,1167]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="93" pageNumber="106">Figs 184–186</figureCitation>
,
<figureCitation id="66FDD1A5FFECFFB6FC49715EDC75FB0C" box="[962,1037,1145,1167]" captionStart="Figs 459–471" captionStartId="32.[189,229,1646,1668]" captionTargetBox="[197,1433,236,1628]" captionTargetId="figure-119@32.[189,1436,232,1638]" captionTargetPageId="32" captionText="Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421228" httpUri="https://zenodo.org/record/6421228/files/figure.png" pageId="93" pageNumber="106">461–465</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFECFFB6FC91718EDC68FAA2" pageId="93" pageNumber="106" type="materials_examined">
<paragraph id="FE79CD20FFECFFB6FC91718EDC68FAA2" blockId="93.[794,1399,1193,1313]" pageId="93" pageNumber="106">
<emphasis id="CCB21132FFECFFB6FC91718EDBCFFB42" bold="true" box="[794,951,1193,1217]" pageId="93" pageNumber="106">Type material</emphasis>
:
<materialsCitation id="4EAEC77DFFECFFB6FC40718EDC68FAA2" ID-GBIF-Occurrence="3455118528" collectingDate="1994-06-14" collectionCode="cAss" collectorName=". Wunderle" country="Indonesia" county="Central" determinerName="V. Assing" elevation="700" location="Central Bali" municipality="Pupuan" pageId="93" pageNumber="106" specimenCount="1" specimenCount-male="1" stateProvince="Bali" typeStatus="holotype">
<typeStatus id="217D7382FFECFFB6FC40718EDC56FB42" box="[971,1070,1193,1217]" pageId="93" pageNumber="106" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C006A9FFECFFB6FBB0718DDC28FB42" box="[1083,1104,1194,1217]" pageId="93" pageNumber="106" type="male">♂</specimenCount>
: “
<collectingCountry id="86D18DB0FFECFFB6FBE6718EDCA1FB42" box="[1133,1241,1193,1217]" name="Indonesia" pageId="93" pageNumber="106">Indonesia</collectingCountry>
,
<location id="FB199BFBFFECFFB6FB62718EDD0BFB42" LSID="urn:lsid:plazi:treatment:766F7C36FFECFFB5FCB47174DCBDFC42:FB199BFBFFECFFB6FB62718EDD0BFB42" box="[1257,1395,1193,1217]" country="Indonesia" county="Central" municipality="Pupuan" name="Central Bali" pageId="93" pageNumber="106" stateProvince="Bali">Central Bali</location>
,
<collectingMunicipality id="1E1D575AFFECFFB6FC9171EDDB17FB62" box="[794,879,1226,1249]" pageId="93" pageNumber="106">Pupuan</collectingMunicipality>
,
<location id="FB199BFBFFECFFB6FC0771EEDB84FB62" LSID="urn:lsid:plazi:treatment:766F7C36FFECFFB5FCB47174DCBDFC42:FB199BFBFFECFFB6FC0771EEDB84FB62" box="[908,1020,1225,1249]" country="Indonesia" county="Central" municipality="Pupuan" name="Wasserfall" pageId="93" pageNumber="106" stateProvince="Bali">Wasserfall</location>
,
<quantity id="393E60C5FFECFFB6FB9271EEDC25FB62" box="[1049,1117,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="93" pageNumber="106" unit="m" value="700.0">
<elevation id="75EB2A13FFECFFB6FB9271EEDC25FB62" box="[1049,1117,1225,1249]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="93" pageNumber="106" unit="m" value="700.0">700 m</elevation>
</quantity>
, Uferschotter,
<date id="8A78EBE0FFECFFB6FA9671EEDD0FFB62" box="[1309,1399,1225,1249]" pageId="93" pageNumber="106" value="94-06-14">
<collectingDate id="9A3C1208FFECFFB6FA9671EEDD0FFB62" box="[1309,1399,1225,1249]" pageId="93" pageNumber="106" value="94-06-14">14.06.94</collectingDate>
</date>
P
<collectorName id="5333A8F6FFECFFB6FCA371CDDBE2FA82" box="[808,922,1257,1281]" pageId="93" pageNumber="106">. Wunderle</collectorName>
/ Holotypus ♂
<taxonomicName id="39C6B6A3FFECFFB6FBDF71CDDD79FA82" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[1108,1281,1257,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="93" pageNumber="106" phylum="Arthropoda" rank="species" species="wunderlei" status="sp. nov.">
<emphasis id="CCB21132FFECFFB6FBDF71CDDD79FA82" box="[1108,1281,1257,1281]" italics="true" pageId="93" pageNumber="106">Zyras wunderlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FFECFFB6FA8771CDDD3CFA82" box="[1292,1348,1258,1281]" pageId="93" pageNumber="106" rank="species">sp. n.</taxonomicNameLabel>
det.
<determinerName id="0BA521E4FFECFFB6FC91702DDB04FAA2" box="[794,892,1289,1313]" pageId="93" pageNumber="106">V. Assing</determinerName>
2015” (
<collectionCode id="98D755E5FFECFFB6FC45702EDC7DFAA2" box="[974,1029,1289,1313]" pageId="93" pageNumber="106">cAss</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFECFFB6FC91706EDCCCFA22" pageId="93" pageNumber="106" type="etymology">
<paragraph id="FE79CD20FFECFFB6FC91706EDCCCFA22" blockId="93.[794,1398,1353,1441]" pageId="93" pageNumber="106">
<emphasis id="CCB21132FFECFFB6FC91706EDBE9FAE2" bold="true" box="[794,913,1353,1377]" pageId="93" pageNumber="106">Etymology</emphasis>
: This species is dedicated to my friend and long-time field companion Paul Wunderle (Mönchengladbach), who collected the
<typeStatus id="217D7382FFECFFB6FBDB70AEDCC8FA22" box="[1104,1200,1417,1441]" pageId="93" pageNumber="106" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFECFFB5FC9170EEDCC9FCC2" lastPageId="94" lastPageNumber="107" pageId="93" pageNumber="106" type="description">
<paragraph id="FE79CD20FFECFFB6FC9170EEDCEBF962" blockId="93.[791,1400,1481,2081]" pageId="93" pageNumber="106">
<emphasis id="CCB21132FFECFFB6FC9170EEDBE4FA62" bold="true" box="[794,924,1481,1505]" pageId="93" pageNumber="106">Description</emphasis>
: Body length
<quantity id="393E60C5FFECFFB6FBC970EEDCEEFA62" box="[1090,1174,1481,1505]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="93" pageNumber="106" unit="mm" value="6.8">6.8 mm</quantity>
; length of forebody
<quantity id="393E60C5FFECFFB6FC9170CEDB15F982" box="[794,877,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="93" pageNumber="106" unit="mm" value="3.3">3.3 mm</quantity>
. Coloration: head and pronotum dark-brown; elytra reddish-brown; abdomen dark-brown, with segments VIII–X and the posterior portion of tergite VII paler; legs pale-yellowish; antennae brown, with antennomeres I–II and the base of III reddish and antennomeres IX–XI pale-yellowish (except for the infuscate base of IX); maxillary palpi dark-yellowish with the terminal palpomere pale-yellowish.
</paragraph>
<paragraph id="FE79CD20FFECFFB6FC9173CEDC2FF7A2" blockId="93.[791,1400,1481,2081]" pageId="93" pageNumber="106">
Head (
<figureCitation id="66FDD1A5FFECFFB6FCE873CEDBC2F882" box="[867,954,1769,1793]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="93" pageNumber="106">Fig. 184</figureCitation>
) distinctly transverse, with rather coarse and dense punctation, except for a small patch in median portion, and with rather dense, long, erect, and dark pubescence. Eyes somewhat longer than postocular region in dorsal view. Antenna (
<figureCitation id="66FDD1A5FFECFFB6FB95724EDC00F802" box="[1054,1144,1897,1921]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="93" pageNumber="106">Fig. 185</figureCitation>
)
<quantity id="393E60C5FFECFFB6FB0F724EDCAEF802" box="[1156,1238,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="93" pageNumber="106" unit="mm" value="2.3">2.3 mm</quantity>
long; antennomeres IV approximately as long as broad, V indistinctly transverse, VI–X of increasing width and increasingly transverse, X slightly more than half as broad as long, and XI of conical shape and short, distinctly shorter than the combined length of IX and X.
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FF3675CDDB7DFE02" blockId="94.[188,795,233,1601]" pageId="94" pageNumber="107">
Pronotum (
<figureCitation id="66FDD1A5FFEFFFB5FEB575CED9EDFE82" box="[318,405,233,257]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="94" pageNumber="107">Fig. 184</figureCitation>
) 1.16 times as broad as long, and 1.22 times as broad as head, rather weakly convex in crosssection; lateral margins weakly sinuate in posterior half (dorsal view); punctation dense, moderately coarse, and rather shallow; pubescence dense, long, erect, and dark.
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FF3674AED982FDC2" blockId="94.[188,795,233,1601]" pageId="94" pageNumber="107">
Elytra (
<figureCitation id="66FDD1A5FFEFFFB5FE8674AED91DFE22" box="[269,357,393,417]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="94" pageNumber="107">Fig. 184</figureCitation>
) 0.78 times as long as pronotum; punctation defined, very dense, and moderately coarse; interstices much narrower than diameter of punctures; pubescence dense, long, erect, and pale-brown. Hind wings fully developed. Metatarsomere I nearly as long as the combined length of II–IV.
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FF36776ED9F8FAE2" blockId="94.[188,795,233,1601]" pageId="94" pageNumber="107">
Abdomen (
<figureCitation id="66FDD1A5FFEFFFB5FEB2776ED9E9FDE2" box="[313,401,585,609]" captionStart="Figs 181–196" captionStartId="18.[189,229,1971,1993]" captionTargetBox="[193,1430,232,1956]" captionTargetId="figure-136@18.[189,1436,232,1956]" captionTargetPageId="18" captionText="Figs 181–196. Zyrasporrectus(181–183),Z.wunderlei(184–186),Pedinopleurusnotabilis (187–189),andAenictoides derivatus (190– 196): forebody (181, 184, 187, 190); antenna (182, 185, 188, 192); abdomen (183, 186, 194); tergites IX–X (189); forebody in lateral view (191); thorax in ventral view (193); abdomen in lateral view (195); abdomen in ventral view (196). Scale bars: 181–188, 190–196: 1.0 mm; 189: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421196" httpUri="https://zenodo.org/record/6421196/files/figure.png" pageId="94" pageNumber="107">Fig. 186</figureCitation>
) broad, slightly broader than elytra and with moderately deep anterior impressions on tergites III–V; all sternites with dense and long, semierect to erect brown pubescence; anterior impressions of tergites III–V each with a transverse series of sparse and fine non-setiferous punctures; tergites III–IV with numerous setiferous punctures bearing long and semierect dark setae at posterior margin and in lateral portions, with scattered setiferous punctures on disc, and with interspersed micropunctation; tergite V with setiferous punctures bearing long and semi-erect dark setae at posterior margin and on disc, and with interspersed micropunctation; tergite VI with a transverse band of numerous and moderately coarse non-setiferous punctures anteriorly, and with setiferous punctures bearing long and semi-erect setae in posterior two-thirds; tergite VII with a broad transverse band of numerous and moderately coarse non-setiferous punctures in anterior third and with setiferous punctures not arranged in distinct series in posterior two-thirds; tergite VIII with numerous dark long setae in posterior two-fifths; posterior margin of tergite VII with palisade fringe; tergite VIII (
<figureCitation id="66FDD1A5FFEFFFB5FEDB702ED9D1FAA2" box="[336,425,1289,1313]" captionStart="Figs 459–471" captionStartId="32.[189,229,1646,1668]" captionTargetBox="[197,1433,236,1628]" captionTargetId="figure-119@32.[189,1436,232,1638]" captionTargetPageId="32" captionText="Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421228" httpUri="https://zenodo.org/record/6421228/files/figure.png" pageId="94" pageNumber="107">Fig. 464</figureCitation>
) with dense long pubescences posteriorly, posterior margin with small and shallow median concavity.
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FF36704DD986F9C2" blockId="94.[188,795,233,1601]" pageId="94" pageNumber="107">
<specimenCount id="E8C006A9FFEFFFB5FF36704DD8AAFA02" box="[189,210,1386,1409]" pageId="94" pageNumber="107" type="male">♂</specimenCount>
: sternite VIII (
<figureCitation id="66FDD1A5FFEFFFB5FEFA704ED9B0FA02" box="[369,456,1385,1409]" captionStart="Figs 459–471" captionStartId="32.[189,229,1646,1668]" captionTargetBox="[197,1433,236,1628]" captionTargetId="figure-119@32.[189,1436,232,1638]" captionTargetPageId="32" captionText="Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421228" httpUri="https://zenodo.org/record/6421228/files/figure.png" pageId="94" pageNumber="107">Fig. 465</figureCitation>
) much longer than tergite VIII, with dense and long pubescence in posterior half, its posterior margin convex; median lobe of aedeagus (
<figureCitation id="66FDD1A5FFEFFFB5FF4F70EED92DFA62" box="[196,341,1481,1505]" captionStart="Figs 459–471" captionStartId="32.[189,229,1646,1668]" captionTargetBox="[197,1433,236,1628]" captionTargetId="figure-119@32.[189,1436,232,1638]" captionTargetPageId="32" captionText="Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421228" httpUri="https://zenodo.org/record/6421228/files/figure.png" pageId="94" pageNumber="107">Figs 461–462</figureCitation>
) 1.0 mm long; ventral process rather slender and apically acute; paramere (
<figureCitation id="66FDD1A5FFEFFFB5FD9570CEDA0DF982" box="[542,629,1513,1537]" captionStart="Figs 459–471" captionStartId="32.[189,229,1646,1668]" captionTargetBox="[197,1433,236,1628]" captionTargetId="figure-119@32.[189,1436,232,1638]" captionTargetPageId="32" captionText="Figs 459–471. Zyras porrectus (459–460), Z. wunderlei (461–465), and Aenictoides derivata (466–471): median lobe of aedeagus in lateral and in ventral view (459–462, 468–469); paramere (463, 470); male tergite VIII (464); male sternite VIII (465); maxilla (466); labium (467); apical lobe of paramere (471). Scale bars: 459–466, 468–470: 0.2 mm; 467, 471: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421228" httpUri="https://zenodo.org/record/6421228/files/figure.png" pageId="94" pageNumber="107">Fig. 463</figureCitation>
) approximately as long as median lobe and with very short and apically obliquely truncate apical lobe.
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FF36734EDA28F782" blockId="94.[188,794,1641,2049]" pageId="94" pageNumber="107">
<emphasis id="CCB21132FFEFFFB5FF36734EDA40F902" bold="true" box="[189,568,1641,1665]" pageId="94" pageNumber="107">Comparative notes and comment</emphasis>
: As can be inferred from external (robust body; broad abdomen; dense punctation with long and erect pubescence) and the male sexual characters (apical lobe of paramere very short),
<taxonomicName id="39C6B6A3FFEFFFB5FF3673CDD939F882" authorityName="Assing" authorityYear="2016" box="[189,321,1769,1793]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="wunderlei">
<emphasis id="CCB21132FFEFFFB5FF3673CDD939F882" box="[189,321,1769,1793]" italics="true" pageId="94" pageNumber="107">Z. wunderlei</emphasis>
</taxonomicName>
belongs to the group of species allied to
<taxonomicName id="39C6B6A3FFEFFFB5FF36722DD9ACF8A2" authority="(KRAATZ, 1859)" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[189,468,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="CCB21132FFEFFFB5FF36722DD96FF8A2" box="[189,279,1801,1825]" italics="true" pageId="94" pageNumber="107">Z. hirtus</emphasis>
(KRAATZ, 1859)
</taxonomicName>
, which, aside from
<taxonomicName id="39C6B6A3FFEFFFB5FD31722DDB6CF8A2" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[698,788,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="CCB21132FFEFFFB5FD31722DDB6CF8A2" box="[698,788,1801,1825]" italics="true" pageId="94" pageNumber="107">Z. hirtus</emphasis>
</taxonomicName>
, includes several named and unnamed species from the southern East Palaearctic and the Oriental regions, such as
<taxonomicName id="39C6B6A3FFEFFFB5FF6B724DDA2AF802" authority="SCHEERPELTZ, 1965" authorityName="SCHEERPELTZ" authorityYear="1965" box="[224,594,1897,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="setosivestis">
<emphasis id="CCB21132FFEFFFB5FF6B724DD913F802" box="[224,363,1898,1921]" italics="true" pageId="94" pageNumber="107">Z. setosivestis</emphasis>
SCHEERPELTZ, 1965
</taxonomicName>
(
<collectingCountry id="86D18DB0FFEFFFB5FDEC724EDAACF802" box="[615,724,1897,1921]" name="Myanmar" pageId="94" pageNumber="107">Myanmar</collectingCountry>
; male unknown),
<taxonomicName id="39C6B6A3FFEFFFB5FEB772ADDA11F822" authority="(DVORAK, 1996)" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[316,617,1929,1957]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FEB772ADD9D6F822" box="[316,430,1929,1953]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
(
<bibRefCitation id="9A57B0D1FFEFFFB5FE4B72ADDA19F822" author="DVORAK, M." box="[448,609,1929,1957]" pageId="94" pageNumber="107" pagination="1 - 15" refId="ref77273" refString="DVORAK, M. 1996: Einige neue myrmekophile oder termitophile Arten des Tribus Zyrasini aus Sudasien (Coleoptera, Staphylinidae, Aleocharinae). - Coleoptera Schwanfelder Coleopterologische Mitteilungen 23: 1 - 15." type="journal article" year="1996">DVOŘÁK, 1996</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="86D18DB0FFEFFFB5FDF272AEDAA1F822" box="[633,729,1929,1953]" name="Vietnam" pageId="94" pageNumber="107">Vietnam</collectingCountry>
; male unknown), and
<taxonomicName id="39C6B6A3FFEFFFB5FEE4728DDABCF842" authority="ASSING, 2016" authorityName="ASSING" authorityYear="2016" box="[367,708,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FEE4728DD9B0F842" box="[367,456,1961,1985]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
<bibRefCitation id="9A57B0D1FFEFFFB5FE59728EDA1BF842" author="ASSING, V." box="[466,611,1961,1985]" pageId="94" pageNumber="107" pagination="87 - 175" refId="ref76876" refString="ASSING, V. 2016: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING, 2016</bibRefCitation>
(
<collectingCountry id="86D18DB0FFEFFFB5FDFE728EDAC4F842" box="[629,700,1961,1985]" name="China" pageId="94" pageNumber="107">China</collectingCountry>
)
</taxonomicName>
. In size and habitus,
<taxonomicName id="39C6B6A3FFEFFFB5FEDB72EDD9ACF862" authorityName="Assing" authorityYear="2016" box="[336,468,1993,2017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="wunderlei">
<emphasis id="CCB21132FFEFFFB5FEDB72EDD9ACF862" box="[336,468,1993,2017]" italics="true" pageId="94" pageNumber="107">Z. wunderlei</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="39C6B6A3FFEFFFB5FDFF72EDDA9EF862" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[628,742,1993,2017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FDFF72EDDA9EF862" box="[628,742,1993,2017]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
and
<taxonomicName id="39C6B6A3FFEFFFB5FF3672CDD96EF782" authorityName="ASSING" authorityYear="2016" box="[189,278,2025,2049]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FF3672CDD96EF782" box="[189,278,2025,2049]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
, but distinguished as follows:
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FCB475CDDC51FDA2" blockId="94.[831,1437,233,833]" pageId="94" pageNumber="107">
– from
<taxonomicName id="39C6B6A3FFEFFFB5FC1E75CDDC7FFE82" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[917,1031,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FC1E75CDDC7FFE82" box="[917,1031,233,257]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
by much paler legs (
<taxonomicName id="39C6B6A3FFEFFFB5FB6A75CDDD2BFE82" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[1249,1363,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FB6A75CDDD2BFE82" box="[1249,1363,233,257]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
: apical halves of femora dark-brown), paler apical antennomeres IX–XI (
<taxonomicName id="39C6B6A3FFEFFFB5FBA0740DDCE5FEC2" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[1067,1181,297,321]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FBA0740DDCE5FEC2" box="[1067,1181,297,321]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
: antennomeres IX–X dark-brown, XI reddish), longer and less strongly incrassate antennae with less transverse antennomeres V–X (
<taxonomicName id="39C6B6A3FFEFFFB5FC7274ADDC13FE22" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[1017,1131,393,417]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FC7274ADDC13FE22" box="[1017,1131,393,417]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
: antenna approximately 2.0 mm long), a larger and more transverse pronotum (
<taxonomicName id="39C6B6A3FFEFFFB5FCE874EDDBADFE62" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[867,981,457,481]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FCE874EDDBADFE62" box="[867,981,457,481]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
: pronotum 1.1 times as broad as long), and by the distinctly finer punctation of the abdominal tergites VI and VII;
</paragraph>
<paragraph id="FE79CD20FFEFFFB5FCB4770DDCC9FCC2" blockId="94.[831,1437,233,833]" pageId="94" pageNumber="107">
– from
<taxonomicName id="39C6B6A3FFEFFFB5FC19770DDB93FDC2" authorityName="ASSING" authorityYear="2016" box="[914,1003,553,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FC19770DDB93FDC2" box="[914,1003,553,577]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
bymuchpalerlegs (
<taxonomicName id="39C6B6A3FFEFFFB5FB3C770DDD68FDC2" authorityName="ASSING" authorityYear="2016" box="[1207,1296,553,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FB3C770DDD68FDC2" box="[1207,1296,553,577]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
: apicalhalves of femora dark-brown), the coloration of the antennae (
<taxonomicName id="39C6B6A3FFEFFFB5FCE8774DDBC4FD02" authorityName="ASSING" authorityYear="2016" box="[867,956,617,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FCE8774DDBC4FD02" box="[867,956,617,641]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
: antennomeres I–X black, XI yellow), coarser and denser punctation of the head, the median concavity of the posterior margin of tergite VIII (absent in
<taxonomicName id="39C6B6A3FFEFFFB5FCD177EDDBCBFD62" authorityName="ASSING" authorityYear="2016" box="[858,947,713,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FCD177EDDBCBFD62" box="[858,947,713,737]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
), and by the distinctly larger aedeagus with an apically less strongly bent ventral process. For illustrations of
<taxonomicName id="39C6B6A3FFEFFFB5FC3C762DDC51FCA2" baseAuthorityName="DVORAK" baseAuthorityYear="1996" box="[951,1065,777,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="funestus">
<emphasis id="CCB21132FFEFFFB5FC3C762DDC51FCA2" box="[951,1065,777,801]" italics="true" pageId="94" pageNumber="107">Z. funestus</emphasis>
</taxonomicName>
and
<taxonomicName id="39C6B6A3FFEFFFB5FBEF762DDCC5FCA2" authorityName="ASSING" authorityYear="2016" box="[1124,1213,777,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="107" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="CCB21132FFEFFFB5FBEF762DDCC5FCA2" box="[1124,1213,777,801]" italics="true" pageId="94" pageNumber="107">Z. flexus</emphasis>
</taxonomicName>
see
<bibRefCitation id="9A57B0D1FFEFFFB5FB7B762EDDE4FCA2" author="ASSING, V." box="[1264,1436,777,801]" pageId="94" pageNumber="107" pagination="1257 - 1282" refId="ref76789" refString="ASSING, V. 2015 d: On the Lomechusini fauna of Vietnam (Coleoptera: Staphylinidae: Aleocharinae). - Linzer Biologische Beitrage 47 (2): 1257 - 1282." type="journal article" year="2015">ASSING (2015d)</bibRefCitation>
and
<bibRefCitation id="9A57B0D1FFEFFFB5FC03760EDC5CFCC2" author="ASSING, V." box="[904,1060,809,833]" pageId="94" pageNumber="107" pagination="87 - 175" refId="ref76876" refString="ASSING, V. 2016: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016)</bibRefCitation>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFEFFFB5FCB4764EDCBDFC42" pageId="94" pageNumber="107" type="distribution">
<paragraph id="FE79CD20FFEFFFB5FCB4764EDCBDFC42" blockId="94.[831,1437,873,961]" pageId="94" pageNumber="107">
<emphasis id="CCB21132FFEFFFB5FCB4764EDCCAFC02" bold="true" box="[831,1202,873,897]" pageId="94" pageNumber="107">Distribution and natural history</emphasis>
: The type locality is situated in central
<collectingRegion id="3C0203C2FFEFFFB5FB9176AEDC3EFC22" box="[1050,1094,905,929]" country="Indonesia" name="Bali" pageId="94" pageNumber="107">Bali</collectingRegion>
,
<collectingCountry id="86D18DB0FFEFFFB5FBDD76AEDCBAFC22" box="[1110,1218,905,929]" name="Indonesia" pageId="94" pageNumber="107">Indonesia</collectingCountry>
. The
<typeStatus id="217D7382FFEFFFB5FA8176AEDD10FC22" box="[1290,1384,905,929]" pageId="94" pageNumber="107" type="holotype">holotype</typeStatus>
was collected from scree near a waterfall.
</paragraph>
</subSubSection>
</treatment>
</document>
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