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<document id="5A85DCDD6D39276C8CAD2A7B4833AA93" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFE8FFB2FF3677F4DCFCFB82" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Tetrabothrus punctiventris Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="102" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="102" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:title id="D4241423E9DEC39E07B3AC2DCC3587E1">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="D48701B1D4D668F9D57CA9BF05BFF54B">Assing, Volker</mods:namePart>
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<mods:date id="7666E9F73F495D1C8E0535C799DA173B">2016</mods:date>
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<heading id="A5317A4CFFE8FFB2FF3677F4DA41FD6D" box="[189,569,723,750]" centered="true" fontSize="11" level="2" pageId="89" pageNumber="102" reason="3">
<taxonomicName id="39C6B6A3FFE8FFB2FF3677F4D9B2FD6D" ID-CoL="55L2T" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[189,458,723,750]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="punctiventris" status="sp. nov.">
<emphasis id="CCB21132FFE8FFB2FF3677F4D9B2FD6D" box="[189,458,723,750]" italics="true" pageId="89" pageNumber="102">Tetrabothrus punctiventris</emphasis>
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</paragraph>
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<subSubSection id="B6DC9EABFFE8FFB2FF3677DED943FC8C" box="[189,315,761,783]" pageId="89" pageNumber="102" type="description">
<paragraph id="FE79CD20FFE8FFB2FF3677DED943FC8C" blockId="89.[189,569,723,783]" box="[189,315,761,783]" pageId="89" pageNumber="102">
(
<figureCitation id="66FDD1A5FFE8FFB2FF4A77DED94DFC8C" box="[193,309,761,783]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Figs 163166</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE8FFB2FF1C760ED9C8FC22" pageId="89" pageNumber="102" type="materials_examined">
<paragraph id="FE79CD20FFE8FFB2FF1C760ED9C8FC22" blockId="89.[151,757,809,929]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FF1C760ED948FCC2" bold="true" box="[151,304,809,833]" pageId="89" pageNumber="102">Type material</emphasis>
:
<materialsCitation id="4EAEC77DFFE8FFB2FEB4760ED9C8FC22" ID-GBIF-Occurrence="3455118317" collectingDate="1987-04-24" collectionCode="MHNG" collectorName="Burckhardt-Lobl" elevation="1600" location="Mt. Kinabalu" pageId="89" pageNumber="102" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="217D7382FFE8FFB2FEB4760ED9DAFCC2" box="[319,418,809,833]" pageId="89" pageNumber="102" type="holotype">Holotype</typeStatus>
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[teneral]: “
<collectingRegion id="3C0203C2FFE8FFB2FDB0760EDA05FCC2" box="[571,637,809,833]" country="Malaysia" name="Sabah" pageId="89" pageNumber="102">Sabah</collectingRegion>
,
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,
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<elevation id="75EB2A13FFE8FFB2FF68766ED90EFCE2" box="[227,374,841,865]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="89" pageNumber="102" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">15501650 m</elevation>
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,
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<collectingDate id="9A3C1208FFE8FFB2FE19766EDA7DFCE2" box="[402,517,841,865]" pageId="89" pageNumber="102" value="1987-04-24">24.IV.1987</collectingDate>
</date>
,
<collectorName id="5333A8F6FFE8FFB2FDAA766EDAACFCE2" box="[545,724,841,865]" pageId="89" pageNumber="102">Burckhardt-Löbl</collectorName>
/ Holotypus ♂
<taxonomicName id="39C6B6A3FFE8FFB2FECF764EDA1EFC02" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[324,614,873,897]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="punctiventris" status="sp. nov.">
<emphasis id="CCB21132FFE8FFB2FECF764EDA1EFC02" box="[324,614,873,897]" italics="true" pageId="89" pageNumber="102">Tetrabothrus punctiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FFE8FFB2FDF1764DDACBFC02" box="[634,691,874,897]" pageId="89" pageNumber="102" rank="species">sp. n.</taxonomicNameLabel>
, det. V.
<bibRefCitation id="9A57B0D1FFE8FFB2FF3976AED94DFC22" author="ASSING, V." box="[178,309,905,929]" pageId="89" pageNumber="102" pagination="87 - 175" refId="ref76876" refString="ASSING, V. 2016: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">Assing 2016</bibRefCitation>
” (
<collectionCode id="98D755E5FFE8FFB2FEC076AED9DCFC22" box="[331,420,905,929]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="89" pageNumber="102" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE8FFB2FF1C76EED901FBA2" pageId="89" pageNumber="102" type="discussion">
<paragraph id="FE79CD20FFE8FFB2FF1C76EED901FBA2" blockId="89.[151,756,969,1057]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FF1C76EED97EFC62" bold="true" box="[151,262,969,993]" pageId="89" pageNumber="102">Comment</emphasis>
: The specific epithet (adjective) alludes to the dense and distinctive non-setiferous punctation on tergites VII and VIII.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE8FFB2FF1C716EDCDBFCE2" pageId="89" pageNumber="102" type="description">
<paragraph id="FE79CD20FFE8FFB2FF1C716EDA2AFA82" blockId="89.[151,757,1097,2049]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FF1C716ED961FBE2" bold="true" box="[151,281,1097,1121]" pageId="89" pageNumber="102">Description</emphasis>
: Body length
<quantity id="393E60C5FFE8FFB2FE4B716EDA6BFBE2" box="[448,531,1097,1121]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="89" pageNumber="102" unit="mm" value="5.4">5.4 mm</quantity>
; length of forebody
<quantity id="393E60C5FFE8FFB2FF1C714ED893FB02" box="[151,235,1129,1153]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="89" pageNumber="102" unit="mm" value="2.2">2.2 mm</quantity>
. Coloration: body pale-brown, with the posterior margins of the abdominal segments yellowish; legs pale-yellowish; antennae bicoloured, with antennomeres IVVIII dark-brown, IIII yellowish, and IXXI dark-yellowish; maxillary palpi yellowish.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FF1C702ED8B5FA62" blockId="89.[151,757,1097,2049]" pageId="89" pageNumber="102">
Head (
<figureCitation id="66FDD1A5FFE8FFB2FF55702ED94EFAA2" box="[222,310,1289,1313]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 163</figureCitation>
) strongly transverse, 1.3 times as broad as long, with weakly marked posterior angles, extensively impunctate in the middle; punctation in lateral dorsal portions fine and rather dense. Eyes large, approximately as long as distance from posterior margin of eye to posterior constriction of head. Antenna (
<figureCitation id="66FDD1A5FFE8FFB2FDBD708EDAF6FA42" box="[566,654,1449,1473]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 164</figureCitation>
)
<quantity id="393E60C5FFE8FFB2FD29708EDA8CFA42" box="[674,756,1449,1473]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="89" pageNumber="102" unit="mm" value="1.4">1.4 mm</quantity>
long.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FF1C70CDDA1EF902" blockId="89.[151,757,1097,2049]" pageId="89" pageNumber="102">
Pronotum (
<figureCitation id="66FDD1A5FFE8FFB2FE9270CED909F982" box="[281,369,1513,1537]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 163</figureCitation>
) 1.15 times as broad as long and 1.08 times as broad as head, somewhat flattened in postero-median portion of disc; punctation fine and rather dense, except along the impunctate midline; pubescence long and depressed to suberect.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FF1C73AEDA6DF882" blockId="89.[151,757,1097,2049]" pageId="89" pageNumber="102">
Elytra (
<figureCitation id="66FDD1A5FFE8FFB2FF6D73AED945F922" box="[230,317,1673,1697]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 163</figureCitation>
) long, approximately 1.1 times as long as pronotum; punctation fine and rather dense. Hind wings fully developed. Metatarsomere as long as the combined length of metatarsomeres II and III.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FF1C722EDB80FEA2" blockId="89.[151,757,1097,2049]" lastBlockId="89.[792,1399,233,481]" pageId="89" pageNumber="102">
Abdomen (
<figureCitation id="66FDD1A5FFE8FFB2FE9B722ED910F8A2" box="[272,360,1801,1825]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 165</figureCitation>
) narrower than elytra and with deep anterior impressions on tergites IIIVI; tergites IIIVI each with four setiferous punctures at posterior margins; tergite VII (
<figureCitation id="66FDD1A5FFE8FFB2FE9C724ED917F802" box="[279,367,1897,1921]" captionStart="Figs 160180" captionStartId="17.[151,191,1972,1994]" captionTargetBox="[151,1397,232,1954]" captionTargetId="figure-153@17.[151,1398,232,1964]" captionTargetPageId="17" captionText="Figs 160180. Tetrabothrus borneensis, holotype (160162), T. punctiventris (163166), T. sulawesicus (167169), Zyras illecebrosus, paratype (170172), Z. gibbus, holotype (173177), and Z. quasar, holotype (178180): forebody (160, 163, 167, 170, 173, 178); antenna (161, 164, 168, 171, 174, 179); abdomen (162, 165, 169, 172, 175, 180); tergite VIII (166); tergite III (176); tergites VII VIII (177). Scale bars: 160165, 167175, 178180: 1.0 mm; 166, 176177: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421194" httpUri="https://zenodo.org/record/6421194/files/figure.png" pageId="89" pageNumber="102">Fig. 166</figureCitation>
) with coarse and moderately dense non-setiferous punctation in anterior three-fourths, posterior margin with palisade fringe; tergite VIII with very dense and moderately coarse non-setiferous punctation in anterior three-fourths (much denser and less coarse than that of tergite VII), posteriorly with a pair of setiferous punctures.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FC91740DDC2FFE62" blockId="89.[792,1399,233,481]" pageId="89" pageNumber="102">
<specimenCount id="E8C006A9FFE8FFB2FC91740DDB57FEC2" box="[794,815,298,321]" pageId="89" pageNumber="102" type="male"></specimenCount>
: posterior margin of sternite VIII produced, broadly truncate in the middle; median lobe of aedeagus (
<figureCitation id="66FDD1A5FFE8FFB2FCAA744EDBD5FE02" box="[801,941,361,385]" captionStart="Figs 442458" captionStartId="31.[151,191,1971,1993]" captionTargetBox="[155,1398,240,1950]" captionTargetId="figure-119@31.[151,1398,232,1971]" captionText="Figs 442458. Tetrabothrus collucatus (442443), T. borneensis (444449), T. punctiventris (450451), T. sulawesicus (452454), Zyras illecebrosus, paratype (455456), and Z. quasar, holotype (457458): median lobe of aedeagus in lateral and in ventral view (442445, 450453, 455458); apico-ventral portion of median lobe in lateral view (446448); male sternite VIII (449, 454). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421226" httpUri="https://zenodo.org/record/6421226/files/figure.png" pageId="89" pageNumber="102">Fig. 450451</figureCitation>
)
<quantity id="393E60C5FFE8FFB2FC49744EDC58FE02" box="[962,1056,361,385]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="89" pageNumber="102" unit="mm" value="0.53">0.53 mm</quantity>
long, with pronounced crista apicalis; ventral process subapically angled in lateral view; paramere nearly
<quantity id="393E60C5FFE8FFB2FC47748EDC67FE42" box="[972,1055,425,449]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="89" pageNumber="102" unit="mm" value="0.7">0.7 mm</quantity>
long, its apical lobe of moderate length and somewhat conical.
</paragraph>
<paragraph id="FE79CD20FFE8FFB2FC91772EDCDBFCE2" blockId="89.[794,1400,521,865]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FC91772EDB94FDA2" bold="true" box="[794,1004,521,545]" pageId="89" pageNumber="102">Comparative notes</emphasis>
:
<taxonomicName id="39C6B6A3FFE8FFB2FC72772EDD77FDA2" authorityName="Assing" authorityYear="2016" box="[1017,1295,521,545]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="CCB21132FFE8FFB2FC72772EDD77FDA2" box="[1017,1295,521,545]" italics="true" pageId="89" pageNumber="102">Tetrabothrus punctiventris</emphasis>
</taxonomicName>
is distinguished from its congeners particularly by the dense and extensive non-setiferous punctation of tergites VII and VIII, the medially depressed pronotum, the coloration of the antennae, as well as by the shape of the aedeagus, above all the pronounced crista apicalis. It additionally differs from
<taxonomicName id="39C6B6A3FFE8FFB2FC2F77EDDC38FD62" authorityName="PACE" authorityYear="2013" box="[932,1088,713,737]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="89" pageNumber="102" phylum="Arthropoda" rank="species" species="malaysianus">
<emphasis id="CCB21132FFE8FFB2FC2F77EDDC38FD62" box="[932,1088,713,737]" italics="true" pageId="89" pageNumber="102">T. malaysianus</emphasis>
</taxonomicName>
, the only other species with non-setiferous punctation on tergite VII known from Borneo, by the uniformly yellowish legs, the much paler coloration of the body, and the medially truncate posterior margin of the male sternite VIII.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE8FFB2FC9176AEDCFCFB82" pageId="89" pageNumber="102" type="distribution">
<paragraph id="FE79CD20FFE8FFB2FC9176AEDCFCFB82" blockId="89.[794,1399,905,1025]" pageId="89" pageNumber="102">
<emphasis id="CCB21132FFE8FFB2FC9176AEDCF4FC22" bold="true" box="[794,1164,905,929]" pageId="89" pageNumber="102">Distribution and natural history</emphasis>
: The type locality is situated in northern Borneo (
<collectingCountry id="86D18DB0FFE8FFB2FBF3768EDCA3FC42" box="[1144,1243,937,961]" name="Malaysia" pageId="89" pageNumber="102">Malaysia</collectingCountry>
:
<collectingRegion id="3C0203C2FFE8FFB2FB65768EDD4BFC42" box="[1262,1331,937,961]" country="Malaysia" name="Sabah" pageId="89" pageNumber="102">Sabah</collectingRegion>
). The slightly teneral
<typeStatus id="217D7382FFE8FFB2FC4F76EEDC5AFC62" box="[964,1058,969,993]" pageId="89" pageNumber="102" type="holotype">holotype</typeStatus>
was collected at an altitude of
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, probably by sifting.
</paragraph>
</subSubSection>
</treatment>
</document>