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<document id="DF21D8B368806E42C14CE8916F2D5370" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFE3FFB8FCEE7334DA8DFB62" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Tetrabothrus pubescens Bernhauer 1915" docType="treatment" docVersion="11" lastPageNumber="96" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="95" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:title id="FB4B3FE8FCF3B969F0F3D86F1FFACBBB">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<treatment id="766F7C36FFE3FFB8FCEE7334DA8DFB62" ID-DOI="http://doi.org/10.5281/zenodo.5462729" ID-GBIF-Taxon="192602764" ID-Zenodo-Dep="5462729" LSID="urn:lsid:plazi:treatment:766F7C36FFE3FFB8FCEE7334DA8DFB62" httpUri="http://treatment.plazi.org/id/766F7C36FFE3FFB8FCEE7334DA8DFB62" lastPageId="83" lastPageNumber="96" pageId="82" pageNumber="95">
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<paragraph id="FE79CD20FFE3FFB9FCEE7334DD63F9AD" blockId="82.[869,1307,1555,1615]" box="[869,1307,1555,1582]" pageId="82" pageNumber="95">
<heading id="A5317A4CFFE3FFB9FCEE7334DD63F9AD" box="[869,1307,1555,1582]" centered="true" fontSize="11" level="2" pageId="82" pageNumber="95" reason="3">
<taxonomicName id="39C6B6A3FFE3FFB9FCEE7334DD63F9AD" ID-CoL="55L2R" authority="BERNHAUER, 1915" authorityName="Bernhauer" authorityYear="1915" box="[869,1307,1555,1582]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="95" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE3FFB9FCEE7334DC23F9AD" box="[869,1115,1555,1582]" italics="true" pageId="82" pageNumber="95">Tetrabothrus pubescens</emphasis>
BERNHAUER, 1915
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FE79CD20FFE3FFB9FCEE731EDB9BF9CC" blockId="82.[869,1307,1555,1615]" box="[869,995,1593,1615]" pageId="82" pageNumber="95">
(
<figureCitation id="66FDD1A5FFE3FFB9FCE1731EDBA6F9CC" box="[874,990,1593,1615]" captionStart="Figs 144159" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[189,1436,232,1940]" captionTargetId="figure-141@16.[189,1436,232,1948]" captionText="Figs 144159. Amaurodera thailandensis (144145), A. fasciata (146), A. meorum (147), A. spinans (148), A. varicollis (149), A. disparicollis, male (150), A. disparicollis, female (151), Tetrabothrus nilgiricus (152153), T. pubescens, syntype (154156), T. collucatus (157158), and T. neoguineensis, holotype (159): forebody (144152, 154, 157, 159); abdomen (153, 156); antenna (155, 158). Scale bars: 144154, 156157, 159: 1.0 mm; 155, 158: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421192" httpUri="https://zenodo.org/record/6421192/files/figure.png" pageId="82" pageNumber="95">Figs 154156</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE3FFB9FCB4734EDD35F902" box="[831,1357,1641,1665]" pageId="82" pageNumber="95" type="reference_group">
<paragraph id="FE79CD20FFE3FFB9FCB4734EDD35F902" blockId="82.[831,1357,1641,1665]" box="[831,1357,1641,1665]" pageId="82" pageNumber="95">
<treatmentCitationGroup id="DED6EA0EFFE3FFB9FCB4734EDD35F902" box="[831,1357,1641,1665]" pageId="82" pageNumber="95">
<treatmentCitation id="7F67EB31FFE3FFB9FCB4734EDD40F902" author="BERNHAUER, M." box="[831,1336,1641,1665]" page="241" pageId="82" pageNumber="95" year="1915">
<taxonomicName id="39C6B6A3FFE3FFB9FCB4734EDD40F902" ID-CoL="55L2R" authority="BERNHAUER, 1915 b: 241" authorityName="Bernhauer" authorityPageNumber="241" authorityYear="1915" box="[831,1336,1641,1665]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="95" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE3FFB9FCB4734EDC4AF902" box="[831,1074,1641,1665]" italics="true" pageId="82" pageNumber="95">Tetrabothrus pubescens</emphasis>
<bibRefCitation id="9A57B0D1FFE3FFB9FBB3734EDD40F902" author="BERNHAUER, M." box="[1080,1336,1641,1665]" pageId="82" pageNumber="95" pagination="213 - 243" refId="ref76974" refString="BERNHAUER, M. 1915 b: Neue Staphyliniden aus Java und Sumatra (7. Beitrag zur indo-malayischen Staphylinidenfauna). - Tijdschrift voor Entomologie 58: 213 - 243." type="journal article" year="1915">
BERNHAUER, 1915b:
<specimenCount id="E8C006A9FFE3FFB9FA99734EDD40F902" box="[1298,1336,1641,1665]" pageId="82" pageNumber="95" type="generic">241</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE3FFB9FCB4738EDBD1F8C2" pageId="82" pageNumber="95" type="materials_examined">
<paragraph id="FE79CD20FFE3FFB9FCB4738EDBD1F8C2" blockId="82.[831,1437,1705,1857]" pageId="82" pageNumber="95">
<emphasis id="CCB21132FFE3FFB9FCB4738EDC30F942" bold="true" box="[831,1096,1705,1729]" pageId="82" pageNumber="95">Type material examined</emphasis>
:
<materialsCitation id="4EAEC77DFFE3FFB9FBDE738EDBDDF8C2" ID-GBIF-Occurrence="3455118321" collectionCode="FMNH" collectorName="M. Bernhauer Collection" location="Java" pageId="82" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Java" typeStatus="syntype">
<typeStatus id="217D7382FFE3FFB9FBDE738EDCD2F942" box="[1109,1194,1705,1729]" pageId="82" pageNumber="95" type="syntype">Syntype</typeStatus>
<specimenCount id="E8C006A9FFE3FFB9FB3B738DDCBDF942" box="[1200,1221,1706,1729]" pageId="82" pageNumber="95" type="female"></specimenCount>
: “Banjoewangi,
<collectingRegion id="3C0203C2FFE3FFB9FAFA738DDDE4F942" box="[1393,1436,1706,1729]" country="Indonesia" name="Jawa Timur" pageId="82" pageNumber="95">Java</collectingRegion>
1910, Mac Gillavry /
<taxonomicName id="39C6B6A3FFE3FFB9FBA973EEDD2AF962" authority="Brh" authorityName="Bernhauer" authorityYear="1915" box="[1058,1362,1737,1761]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="95" phylum="Arthropoda" rank="species" species="pubescens">Tetrabothrus pubescens Brh</taxonomicName>
,
<typeStatus id="217D7382FFE3FFB9FAD773EDDDE4F961" box="[1372,1436,1738,1762]" pageId="82" pageNumber="95">Typus</typeStatus>
/ Chicago NHMus,
<collectorName id="5333A8F6FFE3FFB9FB9C73CDDD5AF882" box="[1047,1314,1769,1793]" pageId="82" pageNumber="95">M. Bernhauer Collection</collectorName>
/
<typeStatus id="217D7382FFE3FFB9FAB773CEDDE5F882" box="[1340,1437,1769,1793]" pageId="82" pageNumber="95" type="syntype">Syntypus</typeStatus>
<taxonomicName id="39C6B6A3FFE3FFB9FCC9722EDCC4F8A2" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1915" box="[834,1212,1801,1825]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="95" phylum="Arthropoda" rank="species" species="pubescens">Tetrabothrus pubescens Bernhauer</taxonomicName>
, rev. V.
<bibRefCitation id="9A57B0D1FFE3FFB9FA84722EDDEBF8A2" author="ASSING, V." box="[1295,1427,1801,1825]" pageId="82" pageNumber="95" pagination="87 - 175" refId="ref76876" refString="ASSING, V. 2016: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">Assing 2016</bibRefCitation>
” (
<collectionCode id="98D755E5FFE3FFB9FCC2720EDBE5F8C2" box="[841,925,1833,1857]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="82" pageNumber="95" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFE3FFB8FCB4724EDA8DFB62" lastPageId="83" lastPageNumber="96" pageId="82" pageNumber="95" type="discussion">
<paragraph id="FE79CD20FFE3FFB8FCB4724ED9F9FEA2" blockId="82.[831,1436,1897,2049]" lastBlockId="83.[149,757,233,1249]" lastPageId="83" lastPageNumber="96" pageId="82" pageNumber="95">
<emphasis id="CCB21132FFE3FFB9FCB4724EDBD6F802" bold="true" box="[831,942,1897,1921]" pageId="82" pageNumber="95">Comment</emphasis>
: The original description is based on an unspecified number of
<typeStatus id="217D7382FFE3FFB9FBB672ADDCE1F821" box="[1085,1177,1930,1954]" pageId="82" pageNumber="95" type="syntype">syntypes</typeStatus>
from “
<collectingRegion id="3C0203C2FFE3FFB9FB6C72AEDD6EF822" box="[1255,1302,1929,1953]" country="Indonesia" name="Jawa Timur" pageId="82" pageNumber="95">Java</collectingRegion>
: Bajoewangi (Mac Gillavry), Preanger (
<collectorName id="5333A8F6FFE3FFB9FBFB728EDC96F842" box="[1136,1262,1961,1985]" pageId="82" pageNumber="95">P.F. Sijthoff</collectorName>
)” (
<bibRefCitation id="9A57B0D1FFE3FFB9FA91728EDBFCF862" author="BERNHAUER, M." pageId="82" pageNumber="95" pagination="213 - 243" refId="ref76974" refString="BERNHAUER, M. 1915 b: Neue Staphyliniden aus Java und Sumatra (7. Beitrag zur indo-malayischen Staphylinidenfauna). - Tijdschrift voor Entomologie 58: 213 - 243." type="journal article" year="1915">BERNHAUER 1915b</bibRefCitation>
). The sole
<typeStatus id="217D7382FFE3FFB9FB8672EDDC27F861" box="[1037,1119,1994,2018]" pageId="82" pageNumber="95" type="syntype">syntype</typeStatus>
located in the BERNHAUER collection is a female and consequently not designated as the
<typeStatus id="217D7382FFE2FFB8FF4975CED95EFE82" box="[194,294,233,257]" pageId="83" pageNumber="96" type="lectotype">lectotype</typeStatus>
. The depository of the type material from Preanger is unknown.
</paragraph>
<paragraph id="FE79CD20FFE2FFB8FF1C740EDA9FFDC2" blockId="83.[149,757,233,1249]" pageId="83" pageNumber="96">
The external characters (
<figureCitation id="66FDD1A5FFE2FFB8FE38740EDA3BFEC2" box="[435,579,297,321]" captionStart="Figs 144159" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[189,1436,232,1940]" captionTargetId="figure-141@16.[189,1436,232,1948]" captionText="Figs 144159. Amaurodera thailandensis (144145), A. fasciata (146), A. meorum (147), A. spinans (148), A. varicollis (149), A. disparicollis, male (150), A. disparicollis, female (151), Tetrabothrus nilgiricus (152153), T. pubescens, syntype (154156), T. collucatus (157158), and T. neoguineensis, holotype (159): forebody (144152, 154, 157, 159); abdomen (153, 156); antenna (155, 158). Scale bars: 144154, 156157, 159: 1.0 mm; 155, 158: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421192" httpUri="https://zenodo.org/record/6421192/files/figure.png" pageId="83" pageNumber="96">Figs 154156</figureCitation>
) are practically identical to those of the
<typeStatus id="217D7382FFE2FFB8FE27746EDA6BFEE2" box="[428,531,329,353]" pageId="83" pageNumber="96" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="39C6B6A3FFE2FFB8FDB6746DDADAFEE2" authorityName="Cameron" authorityYear="1939" box="[573,674,329,353]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE2FFB8FDB6746DDADAFEE2" box="[573,674,329,353]" italics="true" pageId="83" pageNumber="96">T. indicus</emphasis>
</taxonomicName>
and of
<taxonomicName id="39C6B6A3FFE2FFB8FF1C744DD9E6FE02" authority="PACE, 2014" authorityName="PACE" authorityYear="2014" box="[151,414,361,385]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE2FFB8FF1C744DD968FE02" box="[151,272,361,385]" italics="true" pageId="83" pageNumber="96">T. femoralis</emphasis>
<bibRefCitation id="9A57B0D1FFE2FFB8FEAB744ED9E6FE02" author="PACE, R." box="[288,414,361,385]" pageId="83" pageNumber="96" pagination="727 - 794" refId="ref78194" refString="PACE, R. 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Linzer Biologische Beitrage 46 (1): 727 - 794." type="journal article" year="2014">PACE, 2014</bibRefCitation>
</taxonomicName>
(Borneo:
<collectingRegion id="3C0203C2FFE2FFB8FD90744EDA18FE02" box="[539,608,361,385]" country="Malaysia" name="Sabah" pageId="83" pageNumber="96">Sabah</collectingRegion>
). Coloration: body reddish with the elytra, except for the base, darker; antennae uniformly reddish; legs yellow with the apices of all femora black. Tergite VII without non-setiferous punctation anteriorly. Males from
<collectingRegion id="3C0203C2FFE2FFB8FD8974CDDA55FD82" box="[514,557,490,513]" country="Indonesia" name="Jawa Timur" pageId="83" pageNumber="96">Java</collectingRegion>
would be required for a reliable identification and for clarifying if
<taxonomicName id="39C6B6A3FFE2FFB8FD1B772DDA8CFDA2" authorityName="Cameron" authorityYear="1939" box="[656,756,521,545]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE2FFB8FD1B772DDA8CFDA2" box="[656,756,521,545]" italics="true" pageId="83" pageNumber="96">T. indicus</emphasis>
</taxonomicName>
and/or
<taxonomicName id="39C6B6A3FFE2FFB8FF6E770DD926FDC2" authorityName="PACE" authorityYear="2014" box="[229,350,553,577]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE2FFB8FF6E770DD926FDC2" box="[229,350,553,577]" italics="true" pageId="83" pageNumber="96">T. femoralis</emphasis>
</taxonomicName>
are junior synonyms of
<taxonomicName id="39C6B6A3FFE2FFB8FDEB770DDA99FDC2" authorityName="Bernhauer" authorityYear="1915" box="[608,737,553,577]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE2FFB8FDEB770DDA99FDC2" box="[608,737,553,577]" italics="true" pageId="83" pageNumber="96">T. pubescens</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="FE79CD20FFE2FFB8FF1C776DDA62FBA2" blockId="83.[149,757,233,1249]" pageId="83" pageNumber="96">
Not only the external characters, but also the lateral aspect of the median lobe of the aedeagus of
<taxonomicName id="39C6B6A3FFE2FFB8FD2C774DD8C0FD22" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE2FFB8FD2C774DD8C0FD22" italics="true" pageId="83" pageNumber="96">T. indicus</emphasis>
</taxonomicName>
(
<figureCitation id="66FDD1A5FFE2FFB8FF4677AED918FD22" box="[205,352,649,673]" captionStart="Figs 420441" captionStartId="30.[189,229,1972,1994]" captionTargetBox="[189,1427,232,1968]" captionTargetId="figure-139@30.[189,1436,232,1971]" captionText="Figs 420441. Amaurodera meorum (420422), A. disparicollis (423429), A. spinans (430431), A. varicollis (432434), Tetrabothrus nilgiricus (435436), T. indicus (437439; 439: lectotype), andT. inflexus fromLaos (440441): spermatheca (420422, 429, 434); male pronotum (423424); female pronotum (425426); median lobe of aedeagus in lateral and in ventral view (427428, 430433, 435437, 440); ventral process of aedeagus (438439, 441). Scale bars: 423426: 1.0 mm; 420422, 427441: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421224" httpUri="https://zenodo.org/record/6421224/files/figure.png" pageId="83" pageNumber="96">Figs 437439</figureCitation>
),
<taxonomicName id="39C6B6A3FFE2FFB8FEF277ADDA72FD22" authorityName="PACE" authorityYear="1998" box="[377,522,650,673]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="rougemonti">
<emphasis id="CCB21132FFE2FFB8FEF277ADDA72FD22" box="[377,522,650,673]" italics="true" pageId="83" pageNumber="96">T. rougemonti</emphasis>
</taxonomicName>
(
<bibRefCitation id="9A57B0D1FFE2FFB8FDAA77AEDAE2FD22" author="PACE, R." box="[545,666,649,673]" pageId="83" pageNumber="96" pagination="911 - 982" refId="ref77804" refString="PACE, R. 1998: Aleocharinae della Cina: Parte IV (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 105 (4): 911 - 982." type="journal article" year="1998">PACE 1998</bibRefCitation>
: figures 211212),
<taxonomicName id="39C6B6A3FFE2FFB8FE83778DD9F9FD42" authorityName="PACE" authorityYear="2014" box="[264,385,681,705]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE2FFB8FE83778DD9F9FD42" box="[264,385,681,705]" italics="true" pageId="83" pageNumber="96">T. femoralis</emphasis>
</taxonomicName>
(
<bibRefCitation id="9A57B0D1FFE2FFB8FE1A778EDA7BFD42" author="PACE, R." box="[401,515,681,705]" pageId="83" pageNumber="96" pagination="727 - 794" refId="ref78194" refString="PACE, R. 2014: Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Linzer Biologische Beitrage 46 (1): 727 - 794." type="journal article" year="2014">PACE 2014</bibRefCitation>
: figures 120121), and
<taxonomicName id="39C6B6A3FFE2FFB8FF1C77EDD97CFD62" authorityName="ASSING" authorityYear="2015" box="[151,260,713,737]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="inflexus">
<emphasis id="CCB21132FFE2FFB8FF1C77EDD97CFD62" box="[151,260,713,737]" italics="true" pageId="83" pageNumber="96">T. inflexus</emphasis>
</taxonomicName>
(
<figureCitation id="66FDD1A5FFE2FFB8FE9D77EED9DFFD62" box="[278,423,713,737]" captionStart="Figs 420441" captionStartId="30.[189,229,1972,1994]" captionTargetBox="[189,1427,232,1968]" captionTargetId="figure-139@30.[189,1436,232,1971]" captionText="Figs 420441. Amaurodera meorum (420422), A. disparicollis (423429), A. spinans (430431), A. varicollis (432434), Tetrabothrus nilgiricus (435436), T. indicus (437439; 439: lectotype), andT. inflexus fromLaos (440441): spermatheca (420422, 429, 434); male pronotum (423424); female pronotum (425426); median lobe of aedeagus in lateral and in ventral view (427428, 430433, 435437, 440); ventral process of aedeagus (438439, 441). Scale bars: 423426: 1.0 mm; 420422, 427441: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421224" httpUri="https://zenodo.org/record/6421224/files/figure.png" pageId="83" pageNumber="96">Figs 440441</figureCitation>
) are highly similar. However, based on currently available material, these species seem to be distinguished by the ventral aspect of the ventral process of the aedeagus (broad and strongly sinuate in
<taxonomicName id="39C6B6A3FFE2FFB8FF1C766DD950FCE2" authorityName="PACE" authorityYear="1998" box="[151,296,842,865]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="rougemonti">
<emphasis id="CCB21132FFE2FFB8FF1C766DD950FCE2" box="[151,296,842,865]" italics="true" pageId="83" pageNumber="96">T. rougemonti</emphasis>
</taxonomicName>
and
<taxonomicName id="39C6B6A3FFE2FFB8FEEE766DD9A6FCE2" authorityName="PACE" authorityYear="2014" box="[357,478,841,865]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE2FFB8FEEE766DD9A6FCE2" box="[357,478,841,865]" italics="true" pageId="83" pageNumber="96">T. femoralis</emphasis>
</taxonomicName>
; narrow, small, and with sinuate lateral margins in
<taxonomicName id="39C6B6A3FFE2FFB8FE32764DDA5DFC02" authorityName="ASSING" authorityYear="2015" box="[441,549,873,897]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="inflexus">
<emphasis id="CCB21132FFE2FFB8FE32764DDA5DFC02" box="[441,549,873,897]" italics="true" pageId="83" pageNumber="96">T. inflexus</emphasis>
</taxonomicName>
; weakly sinuate in
<taxonomicName id="39C6B6A3FFE2FFB8FF1C76ADD884FC22" authorityName="Cameron" authorityYear="1939" box="[151,252,905,929]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE2FFB8FF1C76ADD884FC22" box="[151,252,905,929]" italics="true" pageId="83" pageNumber="96">T. indicus</emphasis>
</taxonomicName>
). Moreover,
<taxonomicName id="39C6B6A3FFE2FFB8FEF576ADD992FC22" authorityName="ASSING" authorityYear="2015" box="[382,490,905,929]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="inflexus">
<emphasis id="CCB21132FFE2FFB8FEF576ADD992FC22" box="[382,490,905,929]" italics="true" pageId="83" pageNumber="96">T. inflexus</emphasis>
</taxonomicName>
is distinguished from the other species by generally darker coloration of the body and particularly by blackish-brown antennomeres IVX (more or less uniformly reddish in
<taxonomicName id="39C6B6A3FFE2FFB8FD9376CDDAE1FB82" authorityName="Bernhauer" authorityYear="1915" box="[536,665,1001,1025]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE2FFB8FD9376CDDAE1FB82" box="[536,665,1001,1025]" italics="true" pageId="83" pageNumber="96">T. pubescens</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FFE2FFB8FD2C76CDD8C0FBA2" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="CCB21132FFE2FFB8FD2C76CDD8C0FBA2" italics="true" pageId="83" pageNumber="96">T. indicus</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FFE2FFB8FF4E712DD92EFBA2" authorityName="PACE" authorityYear="1998" box="[197,342,1034,1057]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="rougemonti">
<emphasis id="CCB21132FFE2FFB8FF4E712DD92EFBA2" box="[197,342,1034,1057]" italics="true" pageId="83" pageNumber="96">T. rougemonti</emphasis>
</taxonomicName>
, and
<taxonomicName id="39C6B6A3FFE2FFB8FE1B712DDA71FBA2" authorityName="PACE" authorityYear="2014" box="[400,521,1033,1057]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="CCB21132FFE2FFB8FE1B712DDA71FBA2" box="[400,521,1033,1057]" italics="true" pageId="83" pageNumber="96">T. femoralis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="FE79CD20FFE2FFB8FF1C710EDA8DFB62" blockId="83.[149,757,233,1249]" pageId="83" pageNumber="96">
Aside from Java,
<taxonomicName id="39C6B6A3FFE2FFB8FEC9710DD9BCFBC2" authorityName="Bernhauer" authorityYear="1915" box="[322,452,1065,1089]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE2FFB8FEC9710DD9BCFBC2" box="[322,452,1065,1089]" italics="true" pageId="83" pageNumber="96">T. pubescens</emphasis>
</taxonomicName>
has been recorded from Java, Sumatra, and from the
<collectingCountry id="86D18DB0FFE2FFB8FE1C716EDA6AFBE2" box="[407,530,1097,1121]" name="Philippines" pageId="83" pageNumber="96">Philippines</collectingCountry>
. However, the reference specimens from the
<collectingCountry id="86D18DB0FFE2FFB8FE21714EDA5AFB02" box="[426,546,1129,1153]" name="Philippines" pageId="83" pageNumber="96">Philippines</collectingCountry>
are not conspecific with the
<typeStatus id="217D7382FFE2FFB8FF7671ADD953FB21" box="[253,299,1162,1186]" pageId="83" pageNumber="96">type</typeStatus>
material (see the section on
<taxonomicName id="39C6B6A3FFE2FFB8FDE471AED898FB42" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="CCB21132FFE2FFB8FDE471AEDA8CFB22" box="[623,756,1161,1185]" italics="true" pageId="83" pageNumber="96">Tetrabothrus</emphasis>
spec. 2
</taxonomicName>
) and no material from Sumatra has been revised. At present,
<taxonomicName id="39C6B6A3FFE2FFB8FE9A71EDD9EAFB62" authorityName="Bernhauer" authorityYear="1915" box="[273,402,1225,1249]" class="Insecta" family="Staphylinidae" genus="Tetrabothrus" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="96" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="CCB21132FFE2FFB8FE9A71EDD9EAFB62" box="[273,402,1225,1249]" italics="true" pageId="83" pageNumber="96">T. pubescens</emphasis>
</taxonomicName>
is reliably known only from Java.
</paragraph>
</subSubSection>
</treatment>
</document>