treatments-xml/data/76/6F/7C/766F7C36FF97FFCCFCEE7714D9A7F882.xml
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<document id="868D03C9C17074F4DA3734C53706A6AE" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FF97FFCCFCEE7714D9A7F882" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Orphnebius dilatatus Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="52" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="51" updateTime="1699235685730" updateUser="ExternalLinkService">
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<mods:title id="1A32E3345BFB62E2B597B5EECE29D298">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="AF8967B6D2EBF5C0440BE0871852E8B7">Assing, Volker</mods:namePart>
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<mods:title id="363CE8D8B616C34C29748A682FBB8626">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="763EEF55A9AA2DDCA6864220B328363C">
<mods:date id="1321F9D9BD8F8C3BBB1A04EBA46A21FF">2016</mods:date>
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<mods:number id="E4A289EA136ADCD470AA8E3D783ECEE8">2016-12-20</mods:number>
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<mods:identifier id="0948E520F07EC910CB56910A0C9A1C80" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
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<treatment id="766F7C36FF97FFCCFCEE7714D9A7F882" ID-DOI="http://doi.org/10.5281/zenodo.5903414" ID-GBIF-Taxon="192602828" ID-Zenodo-Dep="5903414" LSID="urn:lsid:plazi:treatment:766F7C36FF97FFCCFCEE7714D9A7F882" httpUri="http://treatment.plazi.org/id/766F7C36FF97FFCCFCEE7714D9A7F882" lastPageId="39" lastPageNumber="52" pageId="38" pageNumber="51">
<subSubSection id="B6DC9EABFF97FFCDFCEE7714DCD3FDCD" box="[869,1195,563,590]" pageId="38" pageNumber="51" type="nomenclature">
<paragraph id="FE79CD20FF97FFCDFCEE7714DCD3FDCD" blockId="38.[869,1195,563,623]" box="[869,1195,563,590]" pageId="38" pageNumber="51">
<heading id="A5317A4CFF97FFCDFCEE7714DCD3FDCD" box="[869,1195,563,590]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="51" reason="3">
<taxonomicName id="39C6B6A3FF97FFCDFCEE7714DC43FDCD" ID-CoL="8Y79Y" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[869,1083,563,590]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="51" phylum="Arthropoda" rank="species" species="dilatatus" status="sp. nov.">
<emphasis id="CCB21132FF97FFCDFCEE7714DC43FDCD" box="[869,1083,563,590]" italics="true" pageId="38" pageNumber="51">Orphnebius dilatatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FF97FFCDFBC97714DCD3FDCD" box="[1090,1195,563,590]" pageId="38" pageNumber="51" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF97FFCDFCEE777EDC39FDEC" box="[869,1089,601,623]" pageId="38" pageNumber="51" type="description">
<paragraph id="FE79CD20FF97FFCDFCEE777EDC39FDEC" blockId="38.[869,1195,563,623]" box="[869,1089,601,623]" pageId="38" pageNumber="51">
(
<figureCitation id="66FDD1A5FF97FFCDFCE1777EDBD2FDEC" box="[874,938,601,623]" captionStart="Figs 126" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 126. Antenna (121) and forebody (2226) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 2226: 1.0 mm; 121: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="38" pageNumber="51">Figs 15</figureCitation>
,
<figureCitation id="66FDD1A5FF97FFCDFC39777EDB91FDEC" box="[946,1001,601,623]" captionStart="Figs 2739" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 2739. Forebody (2734), abdomen (3537), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 2737: 1.0 mm; 3849: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="38" pageNumber="51">28, 36</figureCitation>
,
<figureCitation id="66FDD1A5FF97FFCDFC7A777EDC45FDEC" box="[1009,1085,601,623]" captionStart="Figs 216235" captionStartId="20.[189,229,2001,2023]" captionTargetBox="[195,1430,239,1958]" captionTargetId="figure-128@20.[189,1436,232,1971]" captionText="Figs 216235.Orphnebiusfuscapicalis(216219),O. nigrapicalis(220222),O. bakeri(223225),O. breviceps (226228),O. dilatatus (229232), and O. extensus (233235): median lobe of aedeagus in lateral and in ventral view (216217, 220221, 223, 226227, 229230, 233234); paramere (218, 222, 224, 228, 231, 235); apex of paramere (219); spermatheca (225, 232). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421200" httpUri="https://zenodo.org/record/6421200/files/figure.png" pageId="38" pageNumber="51">229232</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF97FFCDFCB477AEDB9DFC02" pageId="38" pageNumber="51" type="materials_examined">
<paragraph id="FE79CD20FF97FFCDFCB477AEDCFBFC82" blockId="38.[830,1436,649,897]" pageId="38" pageNumber="51">
<emphasis id="CCB21132FF97FFCDFCB477AEDBA7FD22" bold="true" box="[831,991,649,673]" pageId="38" pageNumber="51">Type material</emphasis>
:
<materialsCitation id="4EAEC77DFF97FFCDFC7E77AEDCFBFC82" ID-GBIF-Occurrence="3455118445" collectingDate="2004-05-19" collectingDateMax="2004-05-26" collectingDateMin="2004-05-19" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1150" latitude="21.35" location="Ban Sano Mai" longLatPrecision="1264" longitude="102.05" pageId="38" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Phongsaly prov." typeStatus="holotype">
<typeStatus id="217D7382FF97FFCDFC7E77AEDC20FD22" box="[1013,1112,649,673]" pageId="38" pageNumber="51" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C006A9FF97FFCDFBE377ADDC05FD22" box="[1128,1149,650,673]" pageId="38" pageNumber="51" type="male"></specimenCount>
: “
<collectingCountry id="86D18DB0FF97FFCDFB1777AEDCA9FD22" box="[1180,1233,649,673]" name="Laos" pageId="38" pageNumber="51">Laos</collectingCountry>
,
<collectingRegion id="3C0203C2FF97FFCDFB6F77AEDDE1FD22" box="[1252,1433,649,673]" country="Laos" name="Phongsali" pageId="38" pageNumber="51">Phongsaly prov.</collectingRegion>
,
<geoCoordinate id="9BF2ABE7FF97FFCDFCB4778EDBECFD42" box="[831,916,681,705]" degrees="21" direction="north" minutes="21" orientation="latitude" pageId="38" pageNumber="51" precision="925" value="21.35">21°21'N</geoCoordinate>
<geoCoordinate id="9BF2ABE7FF97FFCDFC12778EDB8FFD42" box="[921,1015,681,705]" degrees="102" direction="east" minutes="03" orientation="longitude" pageId="38" pageNumber="51" precision="925" value="102.05">102°03'E</geoCoordinate>
,
<location id="FB199BFBFF97FFCDFB8B778DDCECFD42" LSID="urn:lsid:plazi:treatment:766F7C36FF97FFCCFCEE7714D9A7F882:FB199BFBFF97FFCDFB8B778DDCECFD42" box="[1024,1172,681,705]" country="Laos" latitude="21.35" longLatPrecision="1264" longitude="102.05" name="Ban Sano Mai" pageId="38" pageNumber="51" stateProvince="Phongsaly prov.">Ban Sano Mai</location>
,
<date id="8A78EBE0FF97FFCDFB17778EDD57FD42" box="[1180,1327,681,705]" pageId="38" pageNumber="51" value="2004-05-19" valueMax="2004-05-26" valueMin="2004-05-19">
<collectingDate id="9A3C1208FF97FFCDFB17778EDD57FD42" box="[1180,1327,681,705]" pageId="38" pageNumber="51" value="2004-05-19" valueMax="2004-05-26" valueMin="2004-05-19">19.26.v.2004</collectingDate>
</date>
, ~
<quantity id="393E60C5FF97FFCDFACE778EDDE0FD42" box="[1349,1432,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="38" pageNumber="51" unit="m" value="1150.0">
<elevation id="75EB2A13FF97FFCDFACE778EDDE0FD42" box="[1349,1432,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="38" pageNumber="51" unit="m" value="1150.0">1150 m</elevation>
</quantity>
,
<collectorName id="5333A8F6FF97FFCDFCB577EEDBD3FD62" box="[830,939,713,737]" pageId="38" pageNumber="51">Vít Kubáň</collectorName>
leg. / Holotypus ♂
<taxonomicName id="39C6B6A3FF97FFCDFBF677EEDD20FD62" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[1149,1368,713,737]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="51" phylum="Arthropoda" rank="species" species="dilatatus" status="sp. nov.">
<emphasis id="CCB21132FF97FFCDFBF677EEDD20FD62" box="[1149,1368,713,737]" italics="true" pageId="38" pageNumber="51">Orphnebius dilatatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FF97FFCDFAD577EDDDEFFD62" box="[1374,1431,714,737]" pageId="38" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="0BA521E4FF97FFCDFCE677CDDBB7FC82" box="[877,975,745,769]" pageId="38" pageNumber="51">V. Assing</determinerName>
2015” (
<collectionCode id="98D755E5FF97FFCDFBAA77CEDC0FFC82" box="[1057,1143,745,769]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="38" pageNumber="51">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="FE79CD20FF97FFCDFCB4762DDB9DFC02" blockId="38.[830,1436,649,897]" pageId="38" pageNumber="51">
<materialsCitation id="4EAEC77DFF97FFCDFCB4762DDB03FCC2" ID-GBIF-Occurrence="3455118508" collectingDate="2004-05-19" collectingDateMax="2004-05-26" collectingDateMin="2004-05-19" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1150" latitude="21.35" location="Ban Sano Mai" longLatPrecision="1264" longitude="102.05" pageId="38" pageNumber="51" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phongsali" typeStatus="paratype">
<typeStatus id="217D7382FF97FFCDFCB4762DDBD1FCA1" box="[831,937,778,802]" pageId="38" pageNumber="51" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="E8C006A9FF97FFCDFC30762EDB98FCA2" box="[955,992,777,801]" pageId="38" pageNumber="51" type="male">1 ♂</specimenCount>
,
<specimenCount id="E8C006A9FF97FFCDFC7F762EDC61FCA2" box="[1012,1049,777,801]" pageId="38" pageNumber="51" type="female">1 ♀</specimenCount>
: same data as holotype (
<collectionCode id="98D755E5FF97FFCDFACE762EDDE0FCA2" box="[1349,1432,777,801]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="38" pageNumber="51">NHMB</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="4EAEC77DFF97FFCDFC02760EDB99FC02" ID-GBIF-Occurrence="3455118542" collectingDate="2002-05-11" collectingDateMax="2002-05-21" collectingDateMin="2002-05-11" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" elevation="750" latitude="19.583334" location="Laos-N" longLatPrecision="1271" longitude="101.96667" pageId="38" pageNumber="51" specimenCount="2" specimenCount-male="2" stateProvince="Louangphabang" typeStatus="paratype">
<specimenCount id="E8C006A9FF97FFCDFC02760EDBBAFCC2" box="[905,962,809,833]" pageId="38" pageNumber="51" type="male">2 ♂♂</specimenCount>
: “
<location id="FB199BFBFF97FFCDFC50760EDC53FCC2" LSID="urn:lsid:plazi:treatment:766F7C36FF97FFCCFCEE7714D9A7F882:FB199BFBFF97FFCDFC50760EDC53FCC2" box="[987,1067,809,833]" country="Laos" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Laos-N" pageId="38" pageNumber="51" stateProvince="Louangphabang">Laos-N</location>
(
<location id="FB199BFBFF97FFCDFBB0760EDC8FFCC2" LSID="urn:lsid:plazi:treatment:766F7C36FF97FFCCFCEE7714D9A7F882:FB199BFBFF97FFCDFBB0760EDC8FFCC2" box="[1083,1271,809,833]" country="Laos" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Louangphrabang" pageId="38" pageNumber="51" stateProvince="Louangphabang">Louangphrabang</location>
),
<date id="8A78EBE0FF97FFCDFA80760EDDE0FCC2" box="[1291,1432,809,833]" pageId="38" pageNumber="51" value="2002-05-11" valueMax="2002-05-21" valueMin="2002-05-11">
<collectingDate id="9A3C1208FF97FFCDFA80760EDDE0FCC2" box="[1291,1432,809,833]" pageId="38" pageNumber="51" value="2002-05-11" valueMax="2002-05-21" valueMin="2002-05-11">1121.v.2002</collectingDate>
</date>
,
<geoCoordinate id="9BF2ABE7FF97FFCDFCB4766EDBEEFCE2" box="[831,918,841,865]" degrees="19" direction="north" minutes="35" orientation="latitude" pageId="38" pageNumber="51" precision="925" value="19.583334">19°35'N</geoCoordinate>
,
<geoCoordinate id="9BF2ABE7FF97FFCDFC2A766EDC78FCE2" box="[929,1024,841,865]" degrees="101" direction="east" minutes="58" orientation="longitude" pageId="38" pageNumber="51" precision="925" value="101.96667">101°58'E</geoCoordinate>
,
<location id="FB199BFBFF97FFCDFB81766EDCECFCE2" LSID="urn:lsid:plazi:treatment:766F7C36FF97FFCCFCEE7714D9A7F882:FB199BFBFF97FFCDFB81766EDCECFCE2" box="[1034,1172,841,865]" country="Laos" latitude="19.583334" longLatPrecision="1271" longitude="101.96667" name="Thong Khan" pageId="38" pageNumber="51" stateProvince="Louangphabang">Thong Khan</location>
, ~
<quantity id="393E60C5FF97FFCDFB27766EDC8BFCE2" box="[1196,1267,841,865]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="38" pageNumber="51" unit="m" value="750.0">
<elevation id="75EB2A13FF97FFCDFB27766EDC8BFCE2" box="[1196,1267,841,865]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="38" pageNumber="51" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<collectorName id="5333A8F6FF97FFCDFB76766EDD12FCE2" box="[1277,1386,841,865]" pageId="38" pageNumber="51">Vít Kubáň</collectorName>
leg.” (
<collectionCode id="98D755E5FF97FFCDFCCC764EDBE2FC02" box="[839,922,873,897]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="38" pageNumber="51">NHMB</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF97FFCDFCB4768EDBD4FB82" pageId="38" pageNumber="51" type="etymology">
<paragraph id="FE79CD20FF97FFCDFCB4768EDBD4FB82" blockId="38.[831,1437,937,1025]" pageId="38" pageNumber="51">
<emphasis id="CCB21132FF97FFCDFCB4768EDBCEFC42" bold="true" box="[831,950,937,961]" pageId="38" pageNumber="51">Etymology</emphasis>
: The specific epithet (Latin, adjective) alludes to the medially distinctly dilated condylite of the paramere.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF97FFCCFCB4710ED939FA62" lastPageId="39" lastPageNumber="52" pageId="38" pageNumber="51" type="description">
<paragraph id="FE79CD20FF97FFCDFCB4710EDC5AFA82" blockId="38.[830,1438,1065,2049]" pageId="38" pageNumber="51">
<emphasis id="CCB21132FF97FFCDFCB4710EDBB9FBC2" bold="true" box="[831,961,1065,1089]" pageId="38" pageNumber="51">Description</emphasis>
: Body size very variable; body length 3.35.0 mm; length of forebody 1.32.0 mm. Coloration: forebody black; abdomen red; legs with blackish-brown femora, brown tibiae, and reddish tarsi; antennae blackish with antennomeres IIIV at least partly dark-reddish; maxillary palpi brown to dark-brown with the apical palpomere yellowish.
</paragraph>
<paragraph id="FE79CD20FF97FFCDFCB4702EDD65F902" blockId="38.[830,1438,1065,2049]" pageId="38" pageNumber="51">
Head (
<figureCitation id="66FDD1A5FF97FFCDFC1A702EDBA4FAA2" box="[913,988,1289,1313]" captionStart="Figs 2739" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 2739. Forebody (2734), abdomen (3537), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 2737: 1.0 mm; 3849: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="38" pageNumber="51">Fig. 28</figureCitation>
) conspicuously large, strongly wedgeshaped, broadest posteriorly, 1.51.6 times as broad as long; anterior margin of clypeus very broad and convexly produced in the middle; punctation extremely fine and sparse. Eyes moderately large and distinctly bulging, situated in posterior portion of head. Antenna (
<figureCitation id="66FDD1A5FF97FFCDFAC1708EDDECFA42" box="[1354,1428,1449,1473]" captionStart="Figs 126" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 126. Antenna (121) and forebody (2226) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 2226: 1.0 mm; 121: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="38" pageNumber="51">Fig. 15</figureCitation>
)
<quantity id="393E60C5FF97FFCDFCB470EEDBC7FA62" box="[831,959,1481,1505]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="38" pageNumber="51" unit="mm" value="1.45" valueMax="1.7" valueMin="1.2">1.21.7 mm</quantity>
long; antennomere I conspicuously elongate, approximately as long as combined length of antennomeres IIIV; antennomeres IIIIV distinctly, VX weakly flattened; IVX transverse and of gradually increasing width; X approximately 1.5 times as broad as long; XI nearly as long as combined length of VIIIX.
</paragraph>
<paragraph id="FE79CD20FF97FFCDFCB473ADDD6AF802" blockId="38.[830,1438,1065,2049]" pageId="38" pageNumber="51">
Pronotum (
<figureCitation id="66FDD1A5FF97FFCDFC3073AEDC7EF922" box="[955,1030,1673,1697]" captionStart="Figs 2739" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 2739. Forebody (2734), abdomen (3537), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 2737: 1.0 mm; 3849: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="38" pageNumber="51">Fig. 28</figureCitation>
) 1.521.59 times as broad as long and 0.820.95 times as broad as head, smaller and narrower in relation to head in larger than in smaller specimens, moderately convex in cross-section; lateral and posterior margins together nearly forming a semi-circle, posterior angles completely obsolete; disc with a median pair of punctures, otherwise impunctate; lateral margins anteriorly with three long, erect, stout black setae.
</paragraph>
<paragraph id="FE79CD20FF97FFCDFCB472AEDC05F782" blockId="38.[830,1438,1065,2049]" pageId="38" pageNumber="51">
Elytra (
<figureCitation id="66FDD1A5FF97FFCDFC1172AEDB9CF822" box="[922,996,1929,1953]" captionStart="Figs 2739" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 2739. Forebody (2734), abdomen (3537), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 2737: 1.0 mm; 3849: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="38" pageNumber="51">Fig. 28</figureCitation>
) approximately 0.8 times as long as pronotum; punctation sparse and fine. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II and III.
</paragraph>
<paragraph id="FE79CD20FF96FFCCFF1C75CED982FDA2" blockId="39.[150,757,233,929]" pageId="39" pageNumber="52">
Abdomen (
<figureCitation id="66FDD1A5FF96FFCCFE9E75CED919FE82" box="[277,353,233,257]" captionStart="Figs 2739" captionStartId="10.[189,229,2001,2023]" captionTargetBox="[189,1436,232,1963]" captionTargetId="figure-121@10.[189,1436,232,1971]" captionTargetPageId="10" captionText="Figs 2739. Forebody (2734), abdomen (3537), mesotibia (38), and protibia (39) of Orphnebius spp. of the O. hauseri group: fuscapicalis (27, 35), dilatatus (28, 36), latitibialis (29, 38), breviceps (30), extensus (31), serratus (32, 39), integer (33, 37), bicuspis, Laos (34). Scale bars: 2737: 1.0 mm; 3849: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421175" httpUri="https://zenodo.org/record/6421175/files/figure.png" pageId="39" pageNumber="52">Fig. 36</figureCitation>
): tergites IIIVI with a lateral setiferous puncture on either side, VI with four additional punctures at posterior margin; tergite VII primarily with dense longitudinal striae and additional oblong non-setiferous punctures in posterior two-thirds, near posterior margin with some setiferous punctures, posterior margin with palisade fringe; tergite VIII with two transverse series of long black setae posteriorly, posterior margin convex, in the middle truncate; sternite VIII with broadly convex posterior margin.
</paragraph>
<paragraph id="FE79CD20FF96FFCCFF1C770DD904FCA2" blockId="39.[150,757,233,929]" pageId="39" pageNumber="52">
<specimenCount id="E8C006A9FF96FFCCFF1C770DD8D4FDC2" box="[151,172,554,577]" pageId="39" pageNumber="52" type="male"></specimenCount>
: segments IXX distinctly modified (of the
<taxonomicName id="39C6B6A3FF96FFCCFD02770EDA8DFDC2" authorityName="Eppelsheim" authorityYear="1895" box="[649,757,553,577]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="CCB21132FF96FFCCFD02770EDA8DFDC2" box="[649,757,553,577]" italics="true" pageId="39" pageNumber="52">O. hauseri</emphasis>
</taxonomicName>
<typeStatus id="217D7382FF96FFCCFF1C776ED8B1FDE2" box="[151,201,585,609]" pageId="39" pageNumber="52">type</typeStatus>
), with dense and long pubescence; median lobe of aedeagus (
<figureCitation id="66FDD1A5FF96FFCCFE86774ED9E6FD02" box="[269,414,617,641]" captionStart="Figs 216235" captionStartId="20.[189,229,2001,2023]" captionTargetBox="[195,1430,239,1958]" captionTargetId="figure-128@20.[189,1436,232,1971]" captionText="Figs 216235.Orphnebiusfuscapicalis(216219),O. nigrapicalis(220222),O. bakeri(223225),O. breviceps (226228),O. dilatatus (229232), and O. extensus (233235): median lobe of aedeagus in lateral and in ventral view (216217, 220221, 223, 226227, 229230, 233234); paramere (218, 222, 224, 228, 231, 235); apex of paramere (219); spermatheca (225, 232). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421200" httpUri="https://zenodo.org/record/6421200/files/figure.png" pageId="39" pageNumber="52">Figs 229230</figureCitation>
)
<quantity id="393E60C5FF96FFCCFE38774EDA34FD02" box="[435,588,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="8.299999999999999" metricValueMin="6.0" pageId="39" pageNumber="52" unit="mm" value="0.715" valueMax="0.83" valueMin="0.6">0.600.83 mm</quantity>
long, strongly sclerotized, and of very distinctive shape; ventral process strongly curved in lateral view, apically concave in ventral view; paramere (
<figureCitation id="66FDD1A5FF96FFCCFEDE77EED9D6FD62" box="[341,430,713,737]" captionStart="Figs 216235" captionStartId="20.[189,229,2001,2023]" captionTargetBox="[195,1430,239,1958]" captionTargetId="figure-128@20.[189,1436,232,1971]" captionText="Figs 216235.Orphnebiusfuscapicalis(216219),O. nigrapicalis(220222),O. bakeri(223225),O. breviceps (226228),O. dilatatus (229232), and O. extensus (233235): median lobe of aedeagus in lateral and in ventral view (216217, 220221, 223, 226227, 229230, 233234); paramere (218, 222, 224, 228, 231, 235); apex of paramere (219); spermatheca (225, 232). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421200" httpUri="https://zenodo.org/record/6421200/files/figure.png" pageId="39" pageNumber="52">Fig. 231</figureCitation>
) of highly distinctive shape: paramerite and condylite very slender, condylite strongly dilated in the middle.
</paragraph>
<paragraph id="FE79CD20FF96FFCCFF1C760DD987FC22" blockId="39.[150,757,233,929]" pageId="39" pageNumber="52">
<specimenCount id="E8C006A9FF96FFCCFF1C760DD8D4FCC2" box="[151,172,810,833]" pageId="39" pageNumber="52" type="female"></specimenCount>
: segments IXX distinctly modified (of the
<taxonomicName id="39C6B6A3FF96FFCCFD02760EDA8DFCC2" authorityName="Eppelsheim" authorityYear="1895" box="[649,757,809,833]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="CCB21132FF96FFCCFD02760EDA8DFCC2" box="[649,757,809,833]" italics="true" pageId="39" pageNumber="52">O. hauseri</emphasis>
</taxonomicName>
<typeStatus id="217D7382FF96FFCCFF1C766ED8B1FCE2" box="[151,201,841,865]" pageId="39" pageNumber="52">type</typeStatus>
), with dense and long pubescence; spermathecal capsule (
<figureCitation id="66FDD1A5FF96FFCCFF73764ED937FC02" box="[248,335,873,897]" captionStart="Figs 216235" captionStartId="20.[189,229,2001,2023]" captionTargetBox="[195,1430,239,1958]" captionTargetId="figure-128@20.[189,1436,232,1971]" captionText="Figs 216235.Orphnebiusfuscapicalis(216219),O. nigrapicalis(220222),O. bakeri(223225),O. breviceps (226228),O. dilatatus (229232), and O. extensus (233235): median lobe of aedeagus in lateral and in ventral view (216217, 220221, 223, 226227, 229230, 233234); paramere (218, 222, 224, 228, 231, 235); apex of paramere (219); spermatheca (225, 232). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421200" httpUri="https://zenodo.org/record/6421200/files/figure.png" pageId="39" pageNumber="52">Fig. 232</figureCitation>
) with long and slender distal portion and with coiled proximal portion.
</paragraph>
<paragraph id="FE79CD20FF96FFCCFF1C76EED913FBE2" blockId="39.[151,756,969,1121]" pageId="39" pageNumber="52">
<emphasis id="CCB21132FF96FFCCFF1C76EED9E1FC62" bold="true" box="[151,409,969,993]" pageId="39" pageNumber="52">Intraspecific variation</emphasis>
: This species is subject to pronounced intraspecific variation of body size. Other variable characters (shape and relative size of the head, length of the antennae, size of the aedeagus, etc.) are related to body size.
</paragraph>
<paragraph id="FE79CD20FF96FFCCFF1C71AED939FA62" blockId="39.[151,757,1161,1505]" pageId="39" pageNumber="52">
<emphasis id="CCB21132FF96FFCCFF1C71AED90AFB22" bold="true" box="[151,370,1161,1185]" pageId="39" pageNumber="52">Comparative notes</emphasis>
: Based on the modifications of the abdominal segments IXX and on the general morphology,
<taxonomicName id="39C6B6A3FF96FFCCFEAE71EED9E7FB62" authorityName="Assing" authorityYear="2016" box="[293,415,1225,1249]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="CCB21132FF96FFCCFEAE71EED9E7FB62" box="[293,415,1225,1249]" italics="true" pageId="39" pageNumber="52">O. dilatatus</emphasis>
</taxonomicName>
belongs to the
<taxonomicName id="39C6B6A3FF96FFCCFDB671EEDAD1FB62" authorityName="Eppelsheim" authorityYear="1895" box="[573,681,1225,1249]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="CCB21132FF96FFCCFDB671EEDAD1FB62" box="[573,681,1225,1249]" italics="true" pageId="39" pageNumber="52">O. hauseri</emphasis>
</taxonomicName>
group. As can be inferred from the similarly modified head and pronotum, it is closely allied to
<taxonomicName id="39C6B6A3FF96FFCCFE71702ED8B2FAC2" authority="CAMERON, 1940" authorityName="CAMERON" authorityYear="1940" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="CCB21132FF96FFCCFE71702EDA0FFAA2" box="[506,631,1289,1313]" italics="true" pageId="39" pageNumber="52">O. breviceps</emphasis>
CAMERON, 1940
</taxonomicName>
(
<collectingCountry id="86D18DB0FF96FFCCFF54700ED93BFAC2" box="[223,323,1321,1345]" name="Thailand" pageId="39" pageNumber="52">Thailand</collectingCountry>
; body length
<quantity id="393E60C5FF96FFCCFE62700EDA13FAC2" box="[489,619,1321,1345]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="39" pageNumber="52" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 mm</quantity>
),
<taxonomicName id="39C6B6A3FF96FFCCFD0E700ED93FFAE2" authority="PACE, 2000" authorityName="PACE" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="semivorax">
<emphasis id="CCB21132FF96FFCCFD0E700ED8C2FAE2" italics="true" pageId="39" pageNumber="52">O. semivorax</emphasis>
<bibRefCitation id="9A57B0D1FF96FFCCFF42706ED93FFAE2" author="PACE, R." box="[201,327,1353,1377]" pageId="39" pageNumber="52" pagination="39 - 86" refId="ref77836" refString="PACE, R. 2000: Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE, 2000</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="86D18DB0FF96FFCCFED5706ED9BAFAE2" box="[350,450,1353,1377]" name="Thailand" pageId="39" pageNumber="52">Thailand</collectingCountry>
; male unknown;
<quantity id="393E60C5FF96FFCCFD18706EDA90FAE2" box="[659,744,1353,1377]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="39" pageNumber="52" unit="mm" value="2.3">2.3 mm</quantity>
), and
<taxonomicName id="39C6B6A3FF96FFCCFF41704ED9BAFA02" authority="PACE, 2004" authorityName="PACE" authorityYear="2004" box="[202,450,1385,1409]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="CCB21132FF96FFCCFF41704ED944FA02" box="[202,316,1385,1409]" italics="true" pageId="39" pageNumber="52">O. incertus</emphasis>
<bibRefCitation id="9A57B0D1FF96FFCCFECC704ED9BAFA02" author="PACE, R." box="[327,450,1385,1409]" pageId="39" pageNumber="52" pagination="243 - 302" refId="ref77907" refString="PACE, R. 2004: Aleocharinae di Thailandia, Cambogia, Laos e Malaysia (Coleoptera, Staphylinidae). - Belgian Journal of Entomology 6: 243 - 302." type="journal article" year="2004">PACE, 2004</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="86D18DB0FF96FFCCFE5E704EDA41FA02" box="[469,569,1385,1409]" name="Thailand" pageId="39" pageNumber="52">Thailand</collectingCountry>
; male unknown;
<quantity id="393E60C5FF96FFCCFF1C70AED895FA22" box="[151,237,1417,1441]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="52" unit="mm" value="3.5">3.5 mm</quantity>
). It is distinguished from all of them by larger body size and (except
<taxonomicName id="39C6B6A3FF96FFCCFE00708EDA6AFA42" authorityName="PACE" authorityYear="2000" box="[395,530,1449,1473]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="semivorax">
<emphasis id="CCB21132FF96FFCCFE00708EDA6AFA42" box="[395,530,1449,1473]" italics="true" pageId="39" pageNumber="52">O. semivorax</emphasis>
</taxonomicName>
) different coloration of the forebody.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFF96FFCCFF1C732ED9A7F882" pageId="39" pageNumber="52" type="distribution">
<paragraph id="FE79CD20FF96FFCCFF1C732ED9A7F882" blockId="39.[151,756,1545,1793]" pageId="39" pageNumber="52">
<emphasis id="CCB21132FF96FFCCFF1C732EDA7DF9A2" bold="true" box="[151,517,1545,1569]" pageId="39" pageNumber="52">Distribution and natural history</emphasis>
: The specimens were collected in two localities in
<collectingRegion id="3C0203C2FF96FFCCFE4D730EDA4CF9C2" box="[454,564,1577,1601]" country="Laos" name="Phongsali" pageId="39" pageNumber="52">Phongsaly</collectingRegion>
and
<collectingRegion id="3C0203C2FF96FFCCFDE3730ED8B3F9E2" country="Laos" name="Louangphabang" pageId="39" pageNumber="52">Louangphrabang</collectingRegion>
provinces, North
<collectingCountry id="86D18DB0FF96FFCCFE01736DD9C6F9E2" box="[394,446,1610,1633]" name="Laos" pageId="39" pageNumber="52">Laos</collectingCountry>
, at altitudes of approximately 750 and
<quantity id="393E60C5FF96FFCCFF70734ED935F902" box="[251,333,1641,1665]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="52" unit="m" value="1500.0">1500 m</quantity>
. In the
<typeStatus id="217D7382FF96FFCCFE20734DD9A1F901" box="[427,473,1642,1666]" pageId="39" pageNumber="52">type</typeStatus>
locality they were found together with
<taxonomicName id="39C6B6A3FF96FFCCFEBD73AED9C5F922" authorityName="Assing" authorityYear="2016" box="[310,445,1673,1697]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="CCB21132FF96FFCCFEBD73AED9C5F922" box="[310,445,1673,1697]" italics="true" pageId="39" pageNumber="52">O. cultellatus</emphasis>
</taxonomicName>
and
<taxonomicName id="39C6B6A3FF96FFCCFE7773AEDA15F922" authorityName="Assing" authorityYear="2016" box="[508,621,1673,1697]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF96FFCCFE7773AEDA15F922" box="[508,621,1673,1697]" italics="true" pageId="39" pageNumber="52">O. serratus</emphasis>
</taxonomicName>
, and in the locality in
<collectingRegion id="3C0203C2FF96FFCCFE94738ED9ADF942" box="[287,469,1705,1729]" country="Laos" name="Louangphabang" pageId="39" pageNumber="52">Louangphrabang</collectingRegion>
together with
<taxonomicName id="39C6B6A3FF96FFCCFD1F738ED8CFF962" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="extensus">
<emphasis id="CCB21132FF96FFCCFD1F738ED8CFF962" italics="true" pageId="39" pageNumber="52">O. extensus</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FF96FFCCFF4673EED947F962" authorityName="Assing" authorityYear="2016" box="[205,319,1737,1761]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="CCB21132FF96FFCCFF4673EED947F962" box="[205,319,1737,1761]" italics="true" pageId="39" pageNumber="52">O. serratus</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FF96FFCCFEDF73EED9A3F962" authorityName="Assing" authorityYear="2016" box="[340,475,1737,1761]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="latitibialis">
<emphasis id="CCB21132FF96FFCCFEDF73EED9A3F962" box="[340,475,1737,1761]" italics="true" pageId="39" pageNumber="52">O. latitibialis</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FF96FFCCFE7A73EEDA19F962" box="[497,609,1737,1761]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="CCB21132FF96FFCCFE7A73EEDA19F962" box="[497,609,1737,1761]" italics="true" pageId="39" pageNumber="52">O. bicuspis</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FF96FFCCFDF373EEDA96F962" authorityName="Assing" authorityYear="2016" box="[632,750,1737,1761]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="retunsus">
<emphasis id="CCB21132FF96FFCCFDF373EEDA96F962" box="[632,750,1737,1761]" italics="true" pageId="39" pageNumber="52">O. retunsus</emphasis>
</taxonomicName>
,
<taxonomicName id="39C6B6A3FF96FFCCFF1C73CED967F882" authorityName="Assing" authorityYear="2016" box="[151,287,1769,1793]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="cultellatus">
<emphasis id="CCB21132FF96FFCCFF1C73CED967F882" box="[151,287,1769,1793]" italics="true" pageId="39" pageNumber="52">O. cultellatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="39C6B6A3FF96FFCCFED273CED9A1F882" authorityName="Assing" authorityYear="2016" box="[345,473,1769,1793]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="52" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="CCB21132FF96FFCCFED273CED9A1F882" box="[345,473,1769,1793]" italics="true" pageId="39" pageNumber="52">O. carinatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>