206 lines
29 KiB
XML
206 lines
29 KiB
XML
<document id="40F219C674D5C1921B6318BC5779BE37" ID-CLB-Dataset="35024" ID-DOI="http://dx.doi.org/10.3897/zookeys.608.9427" ID-GBIF-Dataset="f37895dc-7316-48e6-ac3e-901aa5f79eeb" ID-PMC="PMC4982377" ID-Pensoft-Pub="1313-2970-608-1" ID-PubMed="27559303" ID-ZooBank="F865473C03374FD2915A0E3DD2299E66" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-608-1" ModsDocOrigin="ZooKeys 608" ModsDocTitle="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)" checkinTime="1470408253360" checkinUser="pensoft" docAuthor="Borowiec, Marek L." docDate="2016" docId="D3DDB12DF2E357CA6D4D1B54BC6F25E4" docLanguage="en" docName="ZooKeys 608: 1-280" docOrigin="ZooKeys 608" docSource="http://dx.doi.org/10.3897/zookeys.608.9427" docTitle="Tanipone Bolton & Fisher 2012" docType="treatment" docVersion="10" lastPageNumber="180" masterDocId="D563FF93FFEAD163FF9A3E6FFFDDF937" masterDocTitle="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)" masterLastPageNumber="280" masterPageNumber="1" pageNumber="178" updateTime="1732481141242" updateUser="ExternalLinkService">
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<mods:title id="CD8B2AFE669C6D0C62C4E3F0039888C4">Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="DADE3F9868943EEFF930C392437A78DC">Borowiec, Marek L.</mods:namePart>
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<mods:date id="C13936E344169920BD89191689CF12A6">2016</mods:date>
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<mods:classification id="F059B989D9FC129CDA6336E4BCCE5531">journal article</mods:classification>
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<mods:identifier id="D72C88560020B259AD99AD262F1B0D4D" type="DOI">http://dx.doi.org/10.3897/zookeys.608.9427</mods:identifier>
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<treatment id="D3DDB12DF2E357CA6D4D1B54BC6F25E4" ID-GBIF-Taxon="127875730" LSID="urn:lsid:plazi:treatment:D3DDB12DF2E357CA6D4D1B54BC6F25E4" httpUri="http://treatment.plazi.org/id/D3DDB12DF2E357CA6D4D1B54BC6F25E4" lastPageId="179" lastPageNumber="180" pageId="177" pageNumber="178" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Aenictogitoninae">
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<subSubSection id="4EA8CB1AA3384194710556E8D9EC9D51" pageId="177" pageNumber="178" type="multiple">
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<paragraph id="F0C7B5F78DFDB3CC3F3575EE18C79D40" pageId="177" pageNumber="178">
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<pageBreakToken id="055E044A77E081F9D7762460BBC0E00C" pageId="177" pageNumber="178" start="start">Taxon</pageBreakToken>
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classification Animalia Hymenoptera Formicidae
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</paragraph>
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</subSubSection>
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<subSubSection id="DA12174202510AA584D9AEF501200169" pageId="177" pageNumber="178" type="nomenclature">
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<paragraph id="C09882A119373ED8B8D1BDB86CDF64A2" pageId="177" pageNumber="178">
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<taxonomicName id="C7BED4BBB223E82872A4F8DBFF772AAE" ID-CoL="CMXKD" authority="Bolton & Fisher, 2012" authorityName="Bolton & Fisher" authorityYear="2012" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone Bolton & Fisher, 2012</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="4C78528B2123023CC998054DE82B66F9" pageId="177" pageNumber="178" type="type-species">
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<paragraph id="82915652F24831075D40E033577BE85F" pageId="177" pageNumber="178">Type-species.</paragraph>
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<paragraph id="BC925D6BB846A14B916ED58DB96F9B65" pageId="177" pageNumber="178">
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<taxonomicName id="2591D17E35CCA143B01454194DC70B5E" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone hirsuta" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="hirsuta">Tanipone hirsuta</taxonomicName>
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, by original designation.
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</paragraph>
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<paragraph id="89B358DFB1D8A70EF9412B31D3C838EB" pageId="177" pageNumber="178">
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<taxonomicName id="C641C17F64FB881A71D073CECC7000CC" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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is a small genus with unknown biology, endemic to arid and semi-arid habitats of Madagascar.
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</paragraph>
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</subSubSection>
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<subSubSection id="8DEE3BA679D02EE1754DECB8ECDB320F" pageId="177" pageNumber="178" type="diagnosis">
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<paragraph id="7DE6DD141E918B0C092D44A4E97283D2" pageId="177" pageNumber="178">Diagnosis.</paragraph>
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<paragraph id="057F9E421B33364F366237D10BA68CCB" pageId="177" pageNumber="178">
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Worker.
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<taxonomicName id="3868DD8DE184978460D822964560A68F" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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workers are distinctive ants with large eyes and ocelli, very long palps and unique glandular patches on abdominal segment III. The latter are present in all species except one (
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<taxonomicName id="2302F5C92366AC4B4DEE57B2F5DD9A1A" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone aglandula" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="aglandula">Tanipone aglandula</taxonomicName>
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). The body coloration is black or bicolored reddish and black, with a light band or two light spots present at the posterior edge of abdominal segment III. The workers of
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<taxonomicName id="C16A4A46C6C2AAC684E9D49713B668CB" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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lack a conspicuous mid tibial spur. Other genera without the spur include
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<taxonomicName id="61916DB057691F17EA96E828B6628729" class="Insecta" family="Formicidae" genus="Simopone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simopone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Simopone</taxonomicName>
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and
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<taxonomicName id="2E7F702265C43CB8DE8A1B92B119FD06" class="Insecta" family="Formicidae" genus="Vicinopone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vicinopone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Vicinopone</taxonomicName>
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. Additionally, in certain species of
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<taxonomicName id="D49D10557E7A50FF4B5799123ACB3709" class="Insecta" family="Formicidae" genus="Lioponera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lioponera" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Lioponera</taxonomicName>
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the mid tibial spur may be reduced and not easily discernible.
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<taxonomicName id="633800C7692C1CAAF1C3EFAF4D3D420F" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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workers can be easily distinguished from all three lineages by remarkably long palps that are always visible on preserved specimens, reaching the occipital foramen.
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</paragraph>
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<paragraph id="5B243C313F38BD6FE9ABC3F0120894F9" pageId="177" pageNumber="178">
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Male. The male of
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<taxonomicName id="81DB7A04B1B7F2697F3DDE8D2FC51CA8" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="177" pageNumber="178" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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is also easily distinguished from all other dorylines by the long maxillary palps, almost always extruded and reaching occipital foramen. The wing venation is variously developed but the submarginal cell (SMC) is open or closed by a faint 2rs-m cross-vein. There are no notauli and no spurs on middle tibiae.
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</paragraph>
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</subSubSection>
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<subSubSection id="D59093F9DEE11F3DB652DB6963523F09" lastPageId="179" lastPageNumber="180" pageId="177" pageNumber="178" type="description">
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<paragraph id="68A5498B7308166715154E0DF5397203" pageId="177" pageNumber="178">Description.</paragraph>
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<paragraph id="A4AD88160073A3A5996B7FCDAD49122E" lastPageId="178" lastPageNumber="179" pageId="177" pageNumber="178">
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Worker.Head: Antennae with 12 segments. Apical antennal segment not enlarged, not broader and longer than two preceding segments combined. Clypeus without cuticular apron. Lateroclypeal teeth absent.
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<taxonomicName id="D92E4006E704BF0211B17CA46908825F" class="Insecta" genus="Parafrontal" lsidName="Parafrontal ridges" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="ridges">Parafrontal ridges</taxonomicName>
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reduced. Torulo-posttorular complex vertical. Antennal scrobes absent. Labrum with median notch or concavity. Proximal face of stipes projecting beyond inner margin of sclerite, concealing prementum when mouthparts fully closed. Maxillary palps 6-segmented. Labial palps 4-segmented. Mandibles triangular, edentate. Eyes present, composed of more than 20 ommatidia. Ocelli present. Head capsule with differentiated vertical posterior surface above occipital foramen. Ventrolateral margins of head without lamella or ridge extending towards mandibles and beyond carina surrounding occipital foramen. Posterior head corners dorsolaterally immarginate. Carina surrounding occipital foramen ventrally present. Mesosoma: Pronotal flange not separated from collar by distinct ridge. Promesonotal connection with Pronotomesopleural suture completely fused.
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<taxonomicName id="872693200EABC5441A4B9EBA1BAF723F" class="Insecta" genus="Pronotomesopleural" lsidName="Pronotomesopleural suture" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="suture">Pronotomesopleural suture</taxonomicName>
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visible, unfused up to notal surface.
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<taxonomicName id="A1C9D8EFFDA8991179ACD99F3FDC7495" class="Insecta" genus="Mesometapleural" lsidName="Mesometapleural groove" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="groove">Mesometapleural groove</taxonomicName>
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deeply impressed, conspicuous. Transverse
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<taxonomicName id="93731373AF841F40F1A398E89EEA44E1" class="Insecta" lsidName="groove" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="groove">groove</taxonomicName>
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dividing mesopleuron present. Pleural endophragmal pit concavity present. Mesosoma dorsolaterally immarginate. Metanotal depression or
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<taxonomicName id="287B0D1A880DF9345EFDA0712EDC047B" class="Insecta" lsidName="groove" pageId="177" pageNumber="178" phylum="Arthropoda" rank="species" species="groove">groove</taxonomicName>
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on mesosoma absent or a weakly impressed line. Propodeal spiracle situated low on sclerite. Propodeal declivity with distinct dorsal edge or margin and rectangular in posterior view. Metapleural gland without bulla visible through cuticle. Propodeal lobes present, well developed. Metasoma: Petiole anterodorsally marginate, dorsolaterally immarginate, and laterally above spiracle marginate. Helcium in relation to tergosternal Pronotomesopleural suture placed at posttergite and axial. Prora simple, not delimited by carina. Spiracle openings of abdominal segments
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<normalizedToken id="8061B6CB1254F902354AED0099C27077" originalValue="IV–VI">IV-VI</normalizedToken>
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circular. Abdominal segment III anterodorsally immarginate and dorsolaterally immarginate. Abdominal segment III more than half
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<pageBreakToken id="5F77A815AC070A3D00BA1CA4D924AE68" pageId="178" pageNumber="179" start="start">size</pageBreakToken>
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of succeeding segment IV, which is weakly constricted at presegmental portion (uninodal waist). Girdling constriction of segment IV present, i.e. pre- and postsclerites distinct. Cinctus of abdominal segment IV gutter-like, not sculptured. Cinctus of abdominal segment IV gutter-like and smooth or cross-ribbed. Abdominal segment IV not conspicuously largest segment. Abdominal tergite IV not folding over sternite, and anterior portions of sternite and tergite equally well visible in lateral view. Girdling constriction between pre- and posttergites of abdominal segments V and VI absent. Girdling constriction between pre- and poststernites of abdominal segments V and VI present or absent. Pygidium large, with weakly impressed medial field, and armed with few modified setae restricted to apex. Hypopygium unarmed. Legs: Mid tibia without spurs. Hind tibia with single pectinate spur. Hind basitarsus not widening distally, circular in cross-section. Posterior flange of hind coxa not produced as raised lamella. Metatibial gland present as oval patch of whitish cuticle. Metabasitarsal gland absent. Hind pretarsal claws each armed with a tooth. Polymorphism: Monomorphic.
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</paragraph>
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<paragraph id="847EE4922DDD151C02718F8376BBF2C7" lastPageId="179" lastPageNumber="180" pageId="178" pageNumber="179">
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Male.Head: Antennae with 13 segments. Clypeus without cuticular apron.
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<taxonomicName id="2EBAD05E78167FA3B27D722DEFF85735" class="Insecta" genus="Parafrontal" lsidName="Parafrontal ridges" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="ridges">Parafrontal ridges</taxonomicName>
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absent. Torulo-posttorular complex vertical. Maxillary palps 6-segmented. Labial palps 4-segmented. Mandibles triangular, edentate. Ventrolateral margins of head without lamella or ridge extending towards mandibles and beyond carina surrounding occipital foramen. Carina surrounding occipital foramen ventrally present. Mesosoma: Pronotal flange separated from collar by distinct ridge or not.
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<taxonomicName id="CF069E2C42714FCB4A04B384D8BE1B47" class="Insecta" genus="Notauli" lsidName="Notauli" pageId="178" pageNumber="179" phylum="Arthropoda" rank="genus">Notauli</taxonomicName>
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absent. Transverse
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<taxonomicName id="836A29C2AC4F864A3B73F2A17DE882A7" class="Insecta" lsidName="groove" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="groove">groove</taxonomicName>
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dividing mesopleuron present. Propodeal declivity with distinct dorsal edge or margin. Metapleural gland opening absent. Propodeal lobes present. Metasoma: Petiole anterodorsally immarginate, dorsolaterally immarginate, and laterally above spiracle marginate. Helcium in relation to tergosternal Pronotomesopleural suture placed at posttergite and axial. Prora forming a simple U-shaped margin or U-shaped protrusion. Spiracle openings of abdominal segments
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<normalizedToken id="13A45CB2E7CD505351FEB9C7709698E5" originalValue="IV–VI">IV-VI</normalizedToken>
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circular. Abdominal segment III more than half size of succeeding segment IV; latter weakly constricted at presegmental portion (uninodal waist). Girdling constriction of segment IV present, i.e. pre- and postsclerites distinct. Cinctus of abdominal segment IV gutter-like and cross-ribbed. Girdling constriction between pre- and postsclerites of abdominal segments V and VI absent. Abdominal segment IV not conspicuously largest segment. Abdominal sternite VII simple. Abdominal sternite IX distally armed with two spines, with lateral apodemes about as long as medial apodeme, directed anteriorly (towards head). Genitalia: Cupula long relative to rest of genital capsule and shorter ventrally than dorsally. Basimere broadly fused to telomere, with no sulcus trace at junction, and ventrally with left and right arms separated. Telomere gradually tapering toward apex.
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<taxonomicName id="2ADDDACB5F0ECF28DEDA50375A3EB5A9" class="Insecta" family="Mytilidae" genus="Volsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Volsella" order="Mytilida" pageId="178" pageNumber="179" phylum="Arthropoda" rank="genus">Volsella</taxonomicName>
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gradually tapering toward apex. Penisvalva laterally compressed, rounded at apex. Legs: Mid tibia without spurs. Hind tibia with single pectinate spur. Posterior flange of hind coxa not produced as raised lamella. Metatibial gland absent. Metabasitarsal glands absent. Hind pretarsal claws simple. Hind pretarsal claws each armed with a tooth. Wings: Tegula present, broad, demiovate in shape. Vein C in fore wing absent. Pterostigma broad. Abscissa R·f3 absent. Abscissae Rs·f2-3 absent. Cross-vein 2r-rs absent, present and forming base of 'free stigmal
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<normalizedToken id="BD576AB08A6A3E7690A2EABD4A75F62A" originalValue="vein’">vein'</normalizedToken>
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(2r-rs&Rs·f4-5) in absence of Rs·f3 and 2rs-m, or present and
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<pageBreakToken id="30F724B7D1005F1938512E8A65BA23E8" pageId="179" pageNumber="180" start="start">connected</pageBreakToken>
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to Rs·f2-3&Rs·f4. Abscissae Rs·f4-5 absent or differentiated into Rs·f4 and Rs·f5 by 2rs-m. Abscissa M·f2 in fore wing absent or contiguous with Rs+M. Abscissa M·f4 in fore wing absent or present, reaching wing margin. Cross-vein 1m-cu in fore wing absent or present. Cross-vein cu-a in fore wing present, arising from M+Cu and proximal to M·f1. Vein Cu in fore wing present, with only Cu1 branch prominent. Vein A in fore wing with only abscissa A·f1 present or with abscissae A·f1 and A·f2 present. Vein C in hind wing absent. Vein R in hind wing absent. Vein Sc+R in hind wing present. Abscissa Rs·f1 in hind wing present, as long as 1rs-m or longer than 1rs-m. Abscissa Rs·f2 in hind wing absent or present, reaching wing margin. Cross-vein 1rs-m in hind wing absent or present, about as long as M·f1. Vein M+Cu in hind wing present. Abscissa M·f1 in hind wing present. Abscissa M·f2 in hind wing absent. Cross-vein cu-a in hind wing absent or present. Vein Cu in hind wing present. Vein A in hind wing abscissa A·f1 or with abscissae A·f1 and A·f2 present.
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</paragraph>
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<paragraph id="4B403E5D24F063E9080EC3C1E0FFD975" pageId="179" pageNumber="180">
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Gyne. Presumably extremely ergatoid in all species; specimens identified as putative gynes differ from workers only in sculpturation. See
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<bibRefCitation id="A4F79254FE4E68EDF927987C604A4C26" author="Bolton, B" journalOrPublisher="Zootaxa" pageId="191" pageNumber="192" pagination="1 - 101" title="Taxonomy of the cerapachyine ant genera Simopone Forel, Vicinopone gen. n. and Tanipone gen. n. (Hymenoptera: Formicidae)." volume="3283" year="2012">Bolton and Fisher (2012)</bibRefCitation>
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for a discussion.
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</paragraph>
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<paragraph id="6214A0207102ADCFFD56B357DBB4BBD5" pageId="179" pageNumber="180">Larva. Not described. Cocoons absent.</paragraph>
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</subSubSection>
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<subSubSection id="D1604B3798C3481BA9C2433D77D3E71B" pageId="179" pageNumber="180" type="distribution">
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<paragraph id="9DC9A7FA6B1C1C4172ACDC0075298C0D" pageId="179" pageNumber="180">Distribution.</paragraph>
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<paragraph id="4CA33A3240875C5A6C8250FD1734E349" pageId="179" pageNumber="180">Endemic to Madagascar.</paragraph>
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</subSubSection>
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<subSubSection id="09CE0E1525B285B7B352B6BEB4272E41" pageId="179" pageNumber="180" type="taxonomy and phylogeny">
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<paragraph id="02B954C645E6162DD9A554A15E43B1CC" pageId="179" pageNumber="180">Taxonomy and phylogeny.</paragraph>
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<paragraph id="D1506D63FF977226FFB98F83D7D4B5FA" pageId="179" pageNumber="180">
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There are ten
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<taxonomicName id="B7CA8393740FB47974251671C2D4642E" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
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species currently known, all confined to Madagascar (
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<bibRefCitation id="83CFABCB64D53314379C99CC5ABC1B36" author="Bolton, B" journalOrPublisher="Zootaxa" pageId="191" pageNumber="192" pagination="1 - 101" title="Taxonomy of the cerapachyine ant genera Simopone Forel, Vicinopone gen. n. and Tanipone gen. n. (Hymenoptera: Formicidae)." volume="3283" year="2012">Bolton and Fisher 2012</bibRefCitation>
|
||
). The position of this lineage is not known, and internal phylogeny has never been investigated, although
|
||
<bibRefCitation id="91FEC7F5CF1C807188B009A42DA787FE" author="Bolton, B" journalOrPublisher="Zootaxa" pageId="191" pageNumber="192" pagination="1 - 101" title="Taxonomy of the cerapachyine ant genera Simopone Forel, Vicinopone gen. n. and Tanipone gen. n. (Hymenoptera: Formicidae)." volume="3283" year="2012">Bolton and Fisher (2012)</bibRefCitation>
|
||
assigned the species to three species-groups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C73159A7DD77EA5BBF2AE3DB4ECC46BE" pageId="179" pageNumber="180" type="biology_ecology">
|
||
<paragraph id="0564FC8266A7552A4974653E16752430" pageId="179" pageNumber="180">Biology.</paragraph>
|
||
<paragraph id="186C8BD3B45B5FBF854D033FF6DA3684" pageId="179" pageNumber="180">
|
||
The known specimens have been collected mostly in a variety of drier habitats in Madagascar, including dry tropical forest, savannah, and spiny forest. Most workers were collected on low vegetation, on the ground or in rot holes on tree trunks. The sole nest sample of
|
||
<taxonomicName id="C40353BD4CD9D3B5BE2162BBA9428B59" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
|
||
(
|
||
<taxonomicName id="0EB490AA87546A6709C7FE94659DC0C5" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone zona" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="zona">Tanipone zona</taxonomicName>
|
||
) was collected under a stone. Nothing is known about feeding habits of this lineage. The putative queen specimens are ergatoid. Based on nest collections where larvae of different sizes and pupae were collected together, known for
|
||
<taxonomicName id="258C304BA4F0875828B33F9A1E9D9822" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone hirsuta" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="hirsuta">Tanipone hirsuta</taxonomicName>
|
||
,
|
||
<taxonomicName id="9E8B3236514F61854D88B98D6396E31C" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone subpilosa" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="subpilosa">Tanipone subpilosa</taxonomicName>
|
||
, and
|
||
<taxonomicName id="F52B71D83B1ED8340C82FEEF94DC4DC6" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone zona" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="zona">Tanipone zona</taxonomicName>
|
||
, brood development appears not synchronized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D81B18A58F7DDD63873CDDE94FF868CE" pageId="179" pageNumber="180" type="species of tanipone">
|
||
<paragraph id="160542FB71022B171E6F95C4C65645D7" pageId="179" pageNumber="180">
|
||
Species of
|
||
<taxonomicName id="E4BE3CDDC0755808B153A7E2FC7C16D2" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="genus">Tanipone</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="B8A69089F3FFE168E60E05BB9225B197" pageId="179" pageNumber="180">
|
||
<taxonomicName id="1CBBAABEC5B57C7D432D1B8016B8B977" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone aglandula" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="aglandula">Tanipone aglandula</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="67B9689B8A4909506EBEEC468ABDAB1E" pageId="179" pageNumber="180">
|
||
<taxonomicName id="A43346C733798786C9AD7C6D34423AB3" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone aversa" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="aversa">Tanipone aversa</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="CB6289D5866BEDB2F86A0C9F693709BE" pageId="179" pageNumber="180">
|
||
<taxonomicName id="8A0C599F40AAAE3888A4EA02E6DA5F47" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone cognata" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="cognata">Tanipone cognata</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="219612C6E0C5B9BE6D221DB984E03734" pageId="179" pageNumber="180">
|
||
<taxonomicName id="D2E2E8F5E806BBC20AC98122C04F8AE2" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone hirsuta" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="hirsuta">Tanipone hirsuta</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="9475A5D1EE421D7CA628E8A2BD1C4CD9" pageId="179" pageNumber="180">
|
||
<taxonomicName id="CE8A1BDBC0E7E6BEFB13305D0C91F960" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone maculata" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="maculata">Tanipone maculata</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="0E8AF76F389F800B88305EC519A44DE9" pageId="179" pageNumber="180">
|
||
<taxonomicName id="7C52D9086786136AD0FEDE2F01FB090F" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone pilosa" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="pilosa">Tanipone pilosa</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="4F099C6C96A471E55FA1F029AAC7A1C2" pageId="179" pageNumber="180">
|
||
<taxonomicName id="2F44A517253B55F0BCC7CF3F318BC4ED" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone scelesta" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="scelesta">Tanipone scelesta</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="3E121827846B5F187CD421B1A9ADCE1D" pageId="179" pageNumber="180">
|
||
<taxonomicName id="7263C0C7D8BAB49FD3AD4CA13729AEE8" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone subpilosa" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="subpilosa">Tanipone subpilosa</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="80EF2AD4BB9881901054635DD8907158" pageId="179" pageNumber="180">
|
||
<taxonomicName id="2CEE873C1E562A327C328A18A9DECA1E" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone varia" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="varia">Tanipone varia</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
<paragraph id="4E515FBD0BA1134B471D268823057CB0" pageId="179" pageNumber="180">
|
||
<taxonomicName id="069C0C3DB843E8081A5ACC4E4C1EE51F" class="Insecta" family="Formicidae" genus="Tanipone" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tanipone zona" order="Hymenoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="zona">Tanipone zona</taxonomicName>
|
||
Bolton and Fisher, 2012: Madagascar
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |