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<document id="DFE13463D725ED3D21876809998BCBE9" ID-DOI="10.11646/zootaxa.5512.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849159" ID-ZooBank="45911D23-1EED-4DE4-987A-F806B309AD79" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727440331145" checkinUser="plazi" docAuthor="Shamshev, Igor V. &amp; Barták, Miroslav" docDate="2024" docId="756887E7017F6702FF031C25FD7B8FDC" docLanguage="en" docName="zootaxa.5512.4.3.pdf" docOrigin="Zootaxa 5512 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5512.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhamphomyia (Holoclera) vikhrevi Shamshev &amp; Barták, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="526" masterDocId="8951FF9F0173670CFF941A2DFFA18C31" masterDocTitle="New species and records of the Rhamphomyia subgenus Holoclera (Diptera, Empididae) from the Palaearctic Region" masterLastPageNumber="530" masterPageNumber="512" pageNumber="524" updateTime="1727441476738" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="32597AC4B4822C3D983C9FCCE98A72F8">New species and records of the Rhamphomyia subgenus Holoclera (Diptera, Empididae) from the Palaearctic Region</mods:title>
</mods:titleInfo>
<mods:name id="DB516B1479FA1207AEB3F457F8E6E083" type="personal">
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<mods:namePart id="1B57974BFD1CCC95A88FAAA798BCA673">Shamshev, Igor V.</mods:namePart>
<mods:nameIdentifier id="746B77F205E5843936C3A07DCC327AD0" type="ORCID">0000-0003-3575-4243</mods:nameIdentifier>
<mods:affiliation id="9BBD1B8D0172F78753C7C141193A3B1E">All-Russian Institute of Plant Protection, St. Petersburg - Pushkin, Russia</mods:affiliation>
<mods:nameIdentifier id="7ABDB4997C060B9841EC637CD46F6657" type="email">shamshev@mail.ru</mods:nameIdentifier>
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<mods:namePart id="45558440A64FEC00117ECD3105FF54D8">Barták, Miroslav</mods:namePart>
<mods:affiliation id="06FC44E17930910BFE3B90E322B28EBB">Czech University of Life Sciences Prague, Kamýcká 129, 165 00 Praha-Suchdol, Czech Republic</mods:affiliation>
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<mods:title id="D85194D08EC0CB4FBA021AB1C38E4A0E">Zootaxa</mods:title>
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<mods:date id="2B09EB9AECB06723F765FB32282E6EA7">2024</mods:date>
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<mods:number id="16D2E7A055C4CBEF521FFB36265E8633">2024-09-23</mods:number>
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<mods:number id="7EA46D9B8FC11F54DFDC84D03F52B118">5512</mods:number>
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<mods:classification id="B35807F7FA1AE032F97FB25655763CC4">journal article</mods:classification>
<mods:identifier id="662FC011FFAFF2ADA6C6F3874808E0F5" type="DOI">10.11646/zootaxa.5512.4.3</mods:identifier>
<mods:identifier id="CD4799FAC961202670BFB3E357E48FF1" type="ISSN">1175-5326</mods:identifier>
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<treatment id="756887E7017F6702FF031C25FD7B8FDC" ID-DOI="http://doi.org/10.5281/zenodo.13849166" ID-Zenodo-Dep="13849166" LSID="urn:lsid:plazi:treatment:756887E7017F6702FF031C25FD7B8FDC" httpUri="http://treatment.plazi.org/id/756887E7017F6702FF031C25FD7B8FDC" lastPageId="14" lastPageNumber="526" pageId="12" pageNumber="524">
<subSubSection id="B5DB657A017F6700FF031C25FD0B8A12" box="[151,682,1544,1571]" pageId="12" pageNumber="524" type="nomenclature">
<paragraph id="FD7E36F1017F6700FF031C25FD0B8A12" blockId="12.[151,682,1544,1607]" box="[151,682,1544,1571]" pageId="12" pageNumber="524">
<heading id="A636819D017F6700FF031C25FD0B8A12" bold="true" box="[151,682,1544,1571]" fontSize="11" level="1" pageId="12" pageNumber="524" reason="1">
<emphasis id="CFB5EAE3017F6700FF031C25FD0B8A12" bold="true" box="[151,682,1544,1571]" pageId="12" pageNumber="524">
<taxonomicName id="3AC14D72017F6700FF031C25FDE78A12" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[151,582,1544,1571]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="524" phylum="Arthropoda" rank="species" species="vikhrevi" status="sp. nov." subGenus="Holoclera">
<emphasis id="CFB5EAE3017F6700FF031C25FEE88A12" bold="true" box="[151,329,1544,1571]" italics="true" pageId="12" pageNumber="524">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE3017F6700FECE1C25FE728A12" bold="true" box="[346,467,1544,1571]" italics="true" pageId="12" pageNumber="524">Holoclera</emphasis>
)
<emphasis id="CFB5EAE3017F6700FE701C25FDE78A12" bold="true" box="[484,582,1544,1571]" italics="true" pageId="12" pageNumber="524">vikhrevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4865798017F6700FDD91C24FD0B8A12" box="[589,682,1545,1571]" pageId="12" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017F6700FF031C01FF4A8A76" box="[151,235,1580,1607]" pageId="12" pageNumber="524" type="description">
<paragraph id="FD7E36F1017F6700FF031C01FF4A8A76" blockId="12.[151,682,1544,1607]" box="[151,235,1580,1607]" pageId="12" pageNumber="524">
(
<figureCitation id="65FA2A74017F6700FF0B1C01FF428A76" box="[159,227,1580,1607]" captionStart="FIGURE 4" captionStartId="13.[152,255,1373,1397]" captionTargetBox="[201,1374,273,1349]" captionTargetId="figure-90@13.[201,1386,273,1349]" captionTargetPageId="13" captionText="FIGURE 4. Rhamphomyia (Holoclera) vikhrevi sp. nov., male. A. Habitus, lateral view. B, C. postabdomen, lateral view. Abbreviations: cerc—cercus; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bars: A—1 mm, C—0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849171" httpUri="https://zenodo.org/record/13849171/files/figure.png" pageId="12" pageNumber="524">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017F6700FF031C58FD188ACA" pageId="12" pageNumber="524" type="materials_examined">
<paragraph id="FD7E36F1017F6700FF031C58FD188ACA" blockId="12.[151,1437,1652,2039]" pageId="12" pageNumber="524">
<emphasis id="CFB5EAE3017F6700FF031C58FEE98ABF" bold="true" box="[151,328,1652,1679]" pageId="12" pageNumber="524">Type material.</emphasis>
<materialsCitation id="4DA93CAC017F6700FEC61C59FED88A82" collectingDate="2012-05-17" collectionCode="ZMMU" collectorName="N. Vikhrev" country="Morocco" elevation="2000" latitude="31.227" location="Oukaimeden" longLatPrecision="72" longitude="-7.812" pageId="12" pageNumber="524" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="CFB5EAE3017F6700FEC61C59FE598ABF" bold="true" box="[338,504,1652,1678]" pageId="12" pageNumber="524">
<typeStatus id="227A8853017F6700FEC61C59FE598ABF" box="[338,504,1652,1678]" pageId="12" pageNumber="524" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="EBC7FD78017F6700FD961C58FDB88ABE" box="[514,537,1653,1679]" count="1" pageId="12" pageNumber="524" type="male">♂</specimenCount>
, labelled:
<emphasis id="CFB5EAE3017F6700FD071C59FC908ABF" bold="true" box="[659,817,1652,1678]" pageId="12" pageNumber="524">
<collectingCountry id="85D67661017F6700FD071C59FC908ABF" box="[659,817,1652,1678]" name="Morocco" pageId="12" pageNumber="524">MOROCCO</collectingCountry>
</emphasis>
,
<location id="F81E602A017F6700FCD61C59FC708ABF" LSID="urn:lsid:plazi:treatment:756887E7017F6702FF031C25FD7B8FDC:F81E602A017F6700FCD61C59FC708ABF" box="[834,977,1652,1678]" country="Morocco" latitude="31.227" longLatPrecision="72" longitude="-7.812" name="Oukaimeden" pageId="12" pageNumber="524">Oukaimeden</location>
<quantity id="3A399B14017F6700FC4F1C58FB948ABE" box="[987,1077,1653,1679]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="524" unit="m" value="2000.0">
<elevation id="76ECD1C2017F6700FC4F1C58FB948ABE" box="[987,1077,1653,1679]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="524" unit="m" value="2000.0">2000 m</elevation>
</quantity>
,
<geoCoordinate id="98F55036017F6700FBD01C58FB098ABE" box="[1092,1192,1653,1679]" degrees="31.227" direction="north" orientation="latitude" pageId="12" pageNumber="524" precision="55" value="31.227">31.227N</geoCoordinate>
,
<geoCoordinate id="98F55036017F6700FB221C58FAB18ABE" box="[1206,1296,1653,1679]" degrees="7.812" direction="west" orientation="longitude" pageId="12" pageNumber="524" precision="55" value="-7.812">7.812W</geoCoordinate>
,
<collectorName id="50345327017F6700FA8A1C58FA3D8ABF" box="[1310,1436,1652,1679]" pageId="12" pageNumber="524">N. Vikhrev</collectorName>
<date id="897F1031017F6700FF031CB4FEA48A82" box="[151,261,1689,1715]" pageId="12" pageNumber="524" value="2012-05-17">
<collectingDate id="993BE9D9017F6700FF031CB4FEA48A82" box="[151,261,1689,1715]" pageId="12" pageNumber="524" value="2012-05-17">17.v.2012</collectingDate>
</date>
(
<collectionCode id="9BD0AE34017F6700FE871CB4FED08A82" box="[275,369,1689,1715]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="524" type="Museum">ZMMU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4DA93CAC017F6700FE171CB5FE9A8AE6" collectingDate="2012-05-12" collectionCode="CULSP" collectorName="N. Vikhrev" country="Morocco" elevation="2050" latitude="31.26" location="Aguelmouss" longLatPrecision="72" longitude="-7.411" pageId="12" pageNumber="524" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="CFB5EAE3017F6700FE171CB5FD428A82" bold="true" box="[387,739,1688,1715]" pageId="12" pageNumber="524">
<typeStatus id="227A8853017F6700FE171CB5FD8E8A83" box="[387,559,1688,1714]" pageId="12" pageNumber="524" type="paratype">PARATYPES</typeStatus>
.
<collectingCountry id="85D67661017F6700FDA91CB5FD7A8A83" box="[573,731,1688,1714]" name="Morocco" pageId="12" pageNumber="524">MOROCCO</collectingCountry>
.
</emphasis>
<location id="F81E602A017F6700FD7C1CB5FCD58A83" LSID="urn:lsid:plazi:treatment:756887E7017F6702FF031C25FD7B8FDC:F81E602A017F6700FD7C1CB5FCD58A83" box="[744,884,1688,1714]" country="Morocco" latitude="31.26" longLatPrecision="72" longitude="-7.411" name="Aguelmouss" pageId="12" pageNumber="524">Aguelmouss</location>
<quantity id="3A399B14017F6700FCEE1CB5FC718A82" box="[890,976,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="524" unit="m" value="2050.0">
<elevation id="76ECD1C2017F6700FCEE1CB5FC718A82" box="[890,976,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="524" unit="m" value="2050.0">2050 m</elevation>
</quantity>
,
<geoCoordinate id="98F55036017F6700FC4F1CB5FBE18A83" box="[987,1088,1688,1714]" degrees="31.260" direction="north" orientation="latitude" pageId="12" pageNumber="524" precision="55" value="31.26">31.260N</geoCoordinate>
,
<geoCoordinate id="98F55036017F6700FBDE1CB4FB038A82" box="[1098,1186,1689,1715]" degrees="7.411" direction="west" orientation="longitude" pageId="12" pageNumber="524" precision="55" value="-7.411">7.411W</geoCoordinate>
,
<collectorName id="50345327017F6700FB391CB4FA898A83" box="[1197,1320,1688,1715]" pageId="12" pageNumber="524">N. Vikhrev</collectorName>
<date id="897F1031017F6700FABA1CB4FA3D8A82" box="[1326,1436,1689,1715]" pageId="12" pageNumber="524" value="2012-05-12">
<collectingDate id="993BE9D9017F6700FABA1CB4FA3D8A82" box="[1326,1436,1689,1715]" pageId="12" pageNumber="524" value="2012-05-12">12.v.2012</collectingDate>
</date>
(
<specimenCount id="EBC7FD78017F6700FF361C90FF6D8AE6" box="[162,204,1725,1751]" count="1" pageId="12" pageNumber="524" type="male">1 ♂</specimenCount>
,
<collectionCode id="9BD0AE34017F6700FF431C90FE928AE6" box="[215,307,1725,1751]" pageId="12" pageNumber="524">CULSP</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="4DA93CAC017F6700FED11C90FD148ACA" collectingDate="1993-05-08" collectionCode="CULSP, ZMMU, ZISP, CNC" collectorName="G. Shewell" country="Morocco" elevation="1740" latitude="31.135" location="Imlil" longLatPrecision="72" longitude="-7.919" municipality="High Atlas" pageId="12" pageNumber="524" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="paratype">
same data as holotype (
<specimenCount id="EBC7FD78017F6700FDDE1C90FDD38AE6" box="[586,626,1725,1751]" count="1" pageId="12" pageNumber="524" type="female">1 ♀</specimenCount>
<collectionCode id="9BD0AE34017F6700FDEC1C90FD728AE6" box="[632,723,1725,1751]" pageId="12" pageNumber="524">CULSP</collectionCode>
;
<specimenCount id="EBC7FD78017F6700FD491C90FCA88AE6" box="[733,777,1725,1751]" count="1" pageId="12" pageNumber="524" type="male">1 ♂</specimenCount>
,
<specimenCount id="EBC7FD78017F6700FC801C90FC9C8AE6" box="[788,829,1725,1751]" count="2" pageId="12" pageNumber="524" type="female">2 ♀</specimenCount>
<collectionCode id="9BD0AE34017F6700FCD01C90FC3E8AE6" box="[836,927,1725,1751]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="12" pageNumber="524" type="Museum">ZMMU</collectionCode>
;
<specimenCount id="EBC7FD78017F6700FC3D1C90FC738AE6" box="[937,978,1725,1751]" count="1" pageId="12" pageNumber="524" type="male">1 ♂</specimenCount>
<collectionCode id="9BD0AE34017F6700FC4C1C90FBB78AE6" box="[984,1046,1725,1751]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="12" pageNumber="524">ZISP</collectionCode>
);
<location id="F81E602A017F6700FBBD1C91FBFE8AE7" LSID="urn:lsid:plazi:treatment:756887E7017F6702FF031C25FD7B8FDC:F81E602A017F6700FBBD1C91FBFE8AE7" box="[1065,1119,1724,1750]" country="Morocco" latitude="31.135" longLatPrecision="72" longitude="-7.919" municipality="High Atlas" name="Imlil" pageId="12" pageNumber="524">Imlil</location>
[
<geoCoordinate id="98F55036017F6700FBF81C91FB7F8AE6" box="[1132,1246,1724,1751]" degrees="31.135" direction="north" orientation="latitude" pageId="12" pageNumber="524" precision="55" value="31.135">31.135°N</geoCoordinate>
,
<geoCoordinate id="98F55036017F6700FB7D1C90FAF28AE6" box="[1257,1363,1725,1751]" degrees="7.919" direction="west" orientation="longitude" pageId="12" pageNumber="524" precision="55" value="-7.919">7.919°W</geoCoordinate>
],
<collectingMunicipality id="1D1AAC8B017F6700FAF01C91FF748ACB" pageId="12" pageNumber="524">High Atlas</collectingMunicipality>
,
<quantity id="3A399B14017F6700FF741CCCFE968ACA" box="[224,311,1761,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="12" pageNumber="524" unit="m" value="1740.0">
<elevation id="76ECD1C2017F6700FF741CCCFE968ACA" box="[224,311,1761,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="12" pageNumber="524" unit="m" value="1740.0">1740 m</elevation>
</quantity>
,
<date id="897F1031017F6700FED71CCCFE048ACA" box="[323,421,1761,1787]" pageId="12" pageNumber="524" value="1993-05-08">
<collectingDate id="993BE9D9017F6700FED71CCCFE048ACA" box="[323,421,1761,1787]" pageId="12" pageNumber="524" value="1993-05-08">8.v.1993</collectingDate>
</date>
,
<collectorName id="50345327017F6700FE251CCCFD8C8ACB" box="[433,557,1760,1787]" pageId="12" pageNumber="524">G. Shewell</collectorName>
(
<specimenCount id="EBC7FD78017F6700FDD41CCCFDC98ACA" box="[576,616,1761,1787]" count="1" pageId="12" pageNumber="524" type="male">1 ♂</specimenCount>
<collectionCode id="9BD0AE34017F6700FDFB1CCCFD0C8ACA" box="[623,685,1761,1787]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="12" pageNumber="524">CNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017F6700FF531D29FD258B56" pageId="12" pageNumber="524" type="diagnosis">
<paragraph id="FD7E36F1017F6700FF531D29FD258B56" blockId="12.[151,1437,1652,2039]" pageId="12" pageNumber="524">
<emphasis id="CFB5EAE3017F6700FF531D29FEE38B2F" bold="true" box="[199,322,1796,1822]" pageId="12" pageNumber="524">Diagnosis.</emphasis>
Black species of the
<taxonomicName id="3AC14D72017F6700FDA51D28FCA48B2F" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[561,773,1797,1823]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="524" phylum="Arthropoda" rank="species" species="nigripennis" subGenus="Holoclera">
<emphasis id="CFB5EAE3017F6700FDA51D28FDE38B2F" box="[561,578,1797,1822]" italics="true" pageId="12" pageNumber="524">R</emphasis>
. (
<emphasis id="CFB5EAE3017F6700FDCD1D28FDD48B2F" box="[601,629,1797,1822]" italics="true" pageId="12" pageNumber="524">H.</emphasis>
)
<emphasis id="CFB5EAE3017F6700FD111D28FCA48B2F" box="[645,773,1797,1822]" italics="true" pageId="12" pageNumber="524">nigripennis</emphasis>
</taxonomicName>
group with black palpus, uniserial dorsocentrals, fore coxa paler than pleura, supra-alar seta absent, cercus almost straight dorsally (not deeply concave), short phallus, and hypandrium with two strong and long setae.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017F6702FF531D5DFD738DC4" lastPageId="14" lastPageNumber="526" pageId="12" pageNumber="524" type="description">
<paragraph id="FD7E36F1017F6701FF531D5DFAE68CC8" blockId="12.[151,1437,1652,2039]" lastBlockId="13.[151,1437,151,249]" lastPageId="13" lastPageNumber="525" pageId="12" pageNumber="524">
<emphasis id="CFB5EAE3017F6700FF531D5DFE008BBA" bold="true" box="[199,417,1904,1931]" pageId="12" pageNumber="524">Description. Male</emphasis>
(
<figureCitation id="65FA2A74017F6700FE201D5DFDB38BBA" box="[436,530,1904,1931]" captionStart="FIGURE 4" captionStartId="13.[152,255,1373,1397]" captionTargetBox="[201,1374,273,1349]" captionTargetId="figure-90@13.[201,1386,273,1349]" captionTargetPageId="13" captionText="FIGURE 4. Rhamphomyia (Holoclera) vikhrevi sp. nov., male. A. Habitus, lateral view. B, C. postabdomen, lateral view. Abbreviations: cerc—cercus; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bars: A—1 mm, C—0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849171" httpUri="https://zenodo.org/record/13849171/files/figure.png" pageId="12" pageNumber="524">Fig. 4A</figureCitation>
). Body 4.0–
<quantity id="3A399B14017F6700FD301D5CFD5F8BBA" box="[676,766,1905,1931]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="524" unit="mm" value="4.1">4.1 mm</quantity>
, wing
<quantity id="3A399B14017F6700FCC51D5CFC7E8BBA" box="[849,991,1905,1931]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="12" pageNumber="524" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 mm</quantity>
. Head brown, entirely pruinose, black setose, eyes holoptic. Frons confined to small triangle above antennae, without setulae. Dorsal half of eye with greatly enlarged ommatidia. Ocellar triangle prominent, pruinose; ocellar setae fine and approx.
<quantity id="3A399B14017F6700FB671D94FAFB8BE2" box="[1267,1370,1977,2003]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="524" unit="mm" value="0.2">0.20 mm</quantity>
long. Occiput sparsely covered with black setae as long as ocellars or shorter, postocular row incomplete. Face nearly
<quantity id="3A399B14017F6701FAFF1DF1FF628C80" lastPageId="13" lastPageNumber="525" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="524" unit="mm" value="0.15">0.15 mm</quantity>
wide, bare. Clypeus pruinose, gena very narrow. Antenna black, both basal segments short setose; postpedicel nearly 3X longer than basal width; stylus nearly 1.5X shorter than postpedicel. Labrum black, shiny, slightly longer than head height; labellum narrow with rather long setae. Palpus black, short, with several very short setae.
</paragraph>
<caption id="A9BE6679017E6701FF0C1F70FAF789A9" ID-DOI="http://doi.org/10.5281/zenodo.13849171" ID-Zenodo-Dep="13849171" httpUri="https://zenodo.org/record/13849171/files/figure.png" pageId="13" pageNumber="525" startId="13.[152,255,1373,1397]" targetBox="[201,1374,273,1349]" targetPageId="13" targetType="figure">
<paragraph id="FD7E36F1017E6701FF0C1F70FAF789A9" blockId="13.[151,1437,1373,1433]" pageId="13" pageNumber="525">
<emphasis id="CFB5EAE3017E6701FF0C1F70FEBA8944" bold="true" box="[152,283,1373,1397]" pageId="13" pageNumber="525">FIGURE 4.</emphasis>
<taxonomicName id="3AC14D72017E6701FEB01F70FD308944" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[292,657,1373,1397]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="525" phylum="Arthropoda" rank="species" species="vikhrevi" status="sp. nov." subGenus="Holoclera">
<emphasis id="CFB5EAE3017E6701FEB01F70FE178944" box="[292,438,1373,1397]" italics="true" pageId="13" pageNumber="525">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE3017E6701FE5C1F70FD8E8944" box="[456,559,1373,1397]" italics="true" pageId="13" pageNumber="525">Holoclera</emphasis>
)
<emphasis id="CFB5EAE3017E6701FDD41F70FD308944" box="[576,657,1373,1397]" italics="true" pageId="13" pageNumber="525">vikhrevi</emphasis>
</taxonomicName>
<emphasis id="CFB5EAE3017E6701FD0E1F70FD4C8944" bold="true" box="[666,749,1373,1397]" pageId="13" pageNumber="525">
<taxonomicNameLabel id="D4865798017E6701FD0E1F70FD4C8944" box="[666,749,1373,1397]" pageId="13" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="CFB5EAE3017E6701FCAA1F70FCF68944" bold="true" box="[830,855,1373,1397]" pageId="13" pageNumber="525">A.</emphasis>
Habitus, lateral view.
<emphasis id="CFB5EAE3017E6701FBD11F70FBDF8944" bold="true" box="[1093,1150,1373,1397]" pageId="13" pageNumber="525">B, C.</emphasis>
postabdomen, lateral view. Abbreviations: cerc—cercus; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bars: A—1 mm, C—0.1 mm.
</paragraph>
</caption>
<paragraph id="FD7E36F1017E6701FF531FC9FB238ACA" blockId="13.[151,1437,1508,2039]" pageId="13" pageNumber="525">
Thorax brown in ground-colour. Mesoscutum faintly pruinose; in anterodorsal view brownish grey and with two narrow dark vittae between dorsocentrals and virtual acrostichals; in dorsal view uniformly brownish. All thoracic setae black. Antepronotum with several short setae on each side. Proepisternum with several short and fine setulae. Prosternum and upper part of proepisternum bare; postpronotal lobe with 1 long, strong and several much shorter setulae. Mesonotal chaetotaxy: acrostichals absent; dorsocentrals uniserial, 3–4 anterior pairs approx.
<quantity id="3A399B14017E6701FAFF1C58FF628A82" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="525" unit="mm" value="0.2">0.20 mm</quantity>
long and oriented posteriorly as usually, posterior 2–3 pairs oriented anteriorly; 1 long presutural supra-alar; notopleuron with 3 long setae and 1–2 smaller setae in anterior part; postsutural supra-alar (including prealar) region without setae; 1 long and 1 short postalar; 2 pairs of scutellars. Laterotergite black setose.
</paragraph>
<paragraph id="FD7E36F1017E6701FF531D28FD0C8BC7" blockId="13.[151,1437,1508,2039]" pageId="13" pageNumber="525">
Legs colour: fore coxa and trochanter yellow to light brownish yellow, much lighter than pleura; mid and hind coxae and trochanters brown; femora and tibiae yellowish brown, hind tibia at apex and tarsi brown. All legs sparsely pruinose, black setose. Fore femur with anterior row of spine-like setae (approx.
<quantity id="3A399B14017E6701FBE11D60FB768B56" box="[1141,1239,1869,1895]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="525" unit="mm" value="0.03">0.03 mm</quantity>
long). All femora and fore tibiae without prominent setae except preapicals. Mid tibia with 0–1 scarcely distinguishable setae both dorsally and ventrally. Hind tibia with several setae ventrally shorter than tibia depth and with several setae dorsally somewhat longer than tibia depth. Tarsi of all legs slender and long; short setose. Seta in comb at tip of hind tibia very short, not longer than setae forming comb.
</paragraph>
<paragraph id="FD7E36F1017D6702FF531ABAFA3A8CE4" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">Wing brownish tinged, veins brown, axillary angle right to slightly obtuse; costal seta short. Measurements: M2/d ratio = 1.3–1.9, CuA1 ratio = 3.9–4.7, lw:ww ratio = 2.6–2.8. Halter brow, calypter brown with dark fringes.</paragraph>
<paragraph id="FD7E36F1017D6702FF531AF2FE1B8D70" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">Abdomen brown, subshiny, black setose. Lateral marginal setae on tergites 2–3 as long as corresponding segments, remaining tergites with somewhat shorter setae; sternites with long setae, those on segments 3–7 with long pair of median setae.</paragraph>
<paragraph id="FD7E36F1017D6702FF531B66FC298DB8" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">
Hypopygium as in
<figureCitation id="65FA2A74017D6702FE361B66FDBC8D54" box="[418,541,331,357]" captionStart="FIGURE 4" captionStartId="13.[152,255,1373,1397]" captionTargetBox="[201,1374,273,1349]" captionTargetId="figure-90@13.[201,1386,273,1349]" captionTargetPageId="13" captionText="FIGURE 4. Rhamphomyia (Holoclera) vikhrevi sp. nov., male. A. Habitus, lateral view. B, C. postabdomen, lateral view. Abbreviations: cerc—cercus; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bars: A—1 mm, C—0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849171" httpUri="https://zenodo.org/record/13849171/files/figure.png" pageId="14" pageNumber="526">Fig. 4B, C</figureCitation>
. Hypandrium with two long and strong setae; epandrium elongate and without apical spines; cercus slightly concave dorsally, phallus rather short.
</paragraph>
<paragraph id="FD7E36F1017D6702FF531BBEFD738DC4" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">
<emphasis id="CFB5EAE3017D6702FF531BBEFE848D9C" bold="true" box="[199,293,403,429]" pageId="14" pageNumber="526">Female.</emphasis>
Body
<quantity id="3A399B14017D6702FEE71BBFFE6C8D9C" box="[371,461,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="526" unit="mm" value="4.5">4.5 mm</quantity>
, wing
<quantity id="3A399B14017D6702FD8E1BBEFDD48D9C" box="[538,629,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="526" unit="mm" value="4.9">4.9 mm</quantity>
. Dichoptic, all ommatidia almost equal in size, frons pruinose, approx.
<quantity id="3A399B14017D6702FF031B9AFF5F8DE0" box="[151,254,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="526" unit="mm" value="0.1">0.10 mm</quantity>
broad with 3–4 black setae subequally long as frons width. Otherwise very similar to male, only hind femur with prominent subapical anteroventral seta.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017D6702FF531BD2FB2D8E0C" pageId="14" pageNumber="526" type="etymology">
<paragraph id="FD7E36F1017D6702FF531BD2FB2D8E0C" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">
<emphasis id="CFB5EAE3017D6702FF531BD2FEEF8E28" bold="true" box="[199,334,511,537]" pageId="14" pageNumber="526">Etymology.</emphasis>
The species epithet,
<taxonomicName id="3AC14D72017D6702FDD61BD2FD3A8E28" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[578,667,511,537]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="vikhrevi">
<emphasis id="CFB5EAE3017D6702FDD61BD2FD3A8E28" box="[578,667,511,537]" italics="true" pageId="14" pageNumber="526">vikhrevi</emphasis>
</taxonomicName>
, is a Latin genitive patronym to honour of Russian dipterist Nikita Vikhrev (
<collectingRegion id="3F05F813017D6702FE94180EFEC78E0C" box="[256,358,547,573]" country="Russia" name="Moscow" pageId="14" pageNumber="526">Moscow</collectingRegion>
), who kindly provided his very interesting specimens used in this study.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017D6702FF53186AFB278FF8" pageId="14" pageNumber="526" type="discussion">
<paragraph id="FD7E36F1017D6702FF53186AFE438FB0" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">
<emphasis id="CFB5EAE3017D6702FF53186AFE9A8E50" bold="true" box="[199,315,583,609]" pageId="14" pageNumber="526">Remarks.</emphasis>
<taxonomicName id="3AC14D72017D6702FED0186AFDDC8E50" authorityName="Shamshev &amp; Barták" authorityYear="2024" box="[324,637,583,609]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="vikhrevi" status="sp. nov." subGenus="Holoclera">
<emphasis id="CFB5EAE3017D6702FED0186AFE478E50" box="[324,486,583,609]" italics="true" pageId="14" pageNumber="526">Rhamphomyia</emphasis>
(
<emphasis id="CFB5EAE3017D6702FE63186AFDB28E51" box="[503,531,583,608]" italics="true" pageId="14" pageNumber="526">H.</emphasis>
)
<emphasis id="CFB5EAE3017D6702FDB0186AFDDC8E50" box="[548,637,583,609]" italics="true" pageId="14" pageNumber="526">vikhrevi</emphasis>
</taxonomicName>
<emphasis id="CFB5EAE3017D6702FD12186AFD408E50" bold="true" box="[646,737,583,609]" pageId="14" pageNumber="526">
<taxonomicNameLabel id="D4865798017D6702FD12186AFD408E50" box="[646,737,583,609]" pageId="14" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<emphasis id="CFB5EAE3017D6702FCED186AFC2B8E51" box="[889,906,583,608]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FC0E186AFACC8E50" authority="Bartak &amp; Kubik. However" authorityName="Bartak &amp; Kubik. However" box="[922,1389,583,609]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="helvetica" subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FC37186AFC168E51" box="[931,951,583,608]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FC44186AFB948E50" box="[976,1077,583,609]" italics="true" pageId="14" pageNumber="526">helvetica</emphasis>
Barták &amp; Kubík. However
</taxonomicName>
, the new species differs from the latter species in the shape of genitalia (compare fig.
<quantity id="3A399B14017D6702FBBD1846FBF48EB4" box="[1065,1109,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="526" unit="in" value="3.0">3 in</quantity>
<bibRefCitation id="99504B00017D6702FBC81846FAC28EB4" author="Bartak, M. &amp; Kubik, S." box="[1116,1379,619,645]" pageId="14" pageNumber="526" pagination="385 - 407" refId="ref21248" refString="Bartak, M. &amp; Kubik, S. (2012) A review of the Palaearctic species of Rhamphomyia subgenus Holoclera (Diptera: Empididae) with description of 5 new species. Revue Suisse de Zoologie, 119 (3), 385 - 407. https: // doi. org / 10.5962 / bhl. part. 150199" type="journal article" year="2012">Barták &amp; Kubík (2012)</bibRefCitation>
with
<figureCitation id="65FA2A74017D6702FF0318A2FF4E8E98" box="[151,239,655,681]" captionStart="FIGURE 4" captionStartId="13.[152,255,1373,1397]" captionTargetBox="[201,1374,273,1349]" captionTargetId="figure-90@13.[201,1386,273,1349]" captionTargetPageId="13" captionText="FIGURE 4. Rhamphomyia (Holoclera) vikhrevi sp. nov., male. A. Habitus, lateral view. B, C. postabdomen, lateral view. Abbreviations: cerc—cercus; epand—epandrium; hypd—hypandrium; ph—phallus. Scale bars: A—1 mm, C—0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13849171" httpUri="https://zenodo.org/record/13849171/files/figure.png" pageId="14" pageNumber="526">Fig. 4C</figureCitation>
). In
<emphasis id="CFB5EAE3017D6702FEB518A2FE938E99" box="[289,306,655,680]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FED418A2FE788E98" box="[320,473,655,681]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="helvetica" subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FEDD18A2FEFC8E99" box="[329,349,655,680]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FEE018A2FE788E98" box="[372,473,655,681]" italics="true" pageId="14" pageNumber="526">helvetica</emphasis>
</taxonomicName>
, the hypandrium is differently setose and the phallus is much narrower than in
<emphasis id="CFB5EAE3017D6702FADB18A2FAC18E99" box="[1359,1376,655,680]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FAFA18A2FF518EFC" authorityName="Shamshev &amp; Barták" authorityYear="2024" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="vikhrevi" status="sp. nov." subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FAEC18A2FA2D8E99" box="[1400,1420,655,680]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FF03189EFF518EFC" box="[151,240,691,717]" italics="true" pageId="14" pageNumber="526">vikhrevi</emphasis>
</taxonomicName>
<emphasis id="CFB5EAE3017D6702FF63189EFEF08EFC" bold="true" box="[247,337,691,717]" pageId="14" pageNumber="526">
<taxonomicNameLabel id="D4865798017D6702FF63189EFEF08EFC" box="[247,337,691,717]" pageId="14" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In addition, the shape of the cercus in the new species is different, being slightly concave dorsally rather than straight in
<emphasis id="CFB5EAE3017D6702FE1918FAFE3F8EC1" box="[397,414,727,752]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FE3818FAFDE78EC0" box="[428,582,727,753]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="helvetica" subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FE2118FAFE688EC1" box="[437,457,727,752]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FE7518FAFDE78EC0" box="[481,582,727,753]" italics="true" pageId="14" pageNumber="526">helvetica</emphasis>
</taxonomicName>
. Similarly as in
<taxonomicName id="3AC14D72017D6702FD6918FAFC718EC1" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[765,976,727,753]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="524" phylum="Arthropoda" rank="species" species="nigripennis" subGenus="Holoclera">
<emphasis id="CFB5EAE3017D6702FD6918FAFCAF8EC1" box="[765,782,727,752]" italics="true" pageId="14" pageNumber="526">R</emphasis>
. (
<emphasis id="CFB5EAE3017D6702FCB118FAFC988EC1" box="[805,825,727,752]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FCC418FAFC718EC1" box="[848,976,727,752]" italics="true" pageId="14" pageNumber="526">nigripennis</emphasis>
</taxonomicName>
, the male cerci of the new species have a distinct basal swelling (absent in
<emphasis id="CFB5EAE3017D6702FD9A18D6FDBE8F25" box="[526,543,763,788]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FDB918D6FD668F24" box="[557,711,763,789]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="helvetica" subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FDA218D6FDEB8F25" box="[566,586,763,788]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FDF618D6FD668F24" box="[610,711,763,789]" italics="true" pageId="14" pageNumber="526">helvetica</emphasis>
</taxonomicName>
). Also, the female is similar to
<emphasis id="CFB5EAE3017D6702FBBE18D6FB9A8F25" box="[1066,1083,763,788]" italics="true" pageId="14" pageNumber="526">R</emphasis>
.
<taxonomicName id="3AC14D72017D6702FBDE18D6FB458F24" box="[1098,1252,763,789]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="species" species="helvetica" subGenus="Holoclera">
(
<emphasis id="CFB5EAE3017D6702FBC718D6FBC68F25" box="[1107,1127,763,788]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FBEB18D6FB458F24" box="[1151,1252,763,789]" italics="true" pageId="14" pageNumber="526">helvetica</emphasis>
</taxonomicName>
, however, it has a subapical anteroventral seta on the hind femur similar to
<emphasis id="CFB5EAE3017D6702FCBE1932FC9A8F09" box="[810,827,799,824]" italics="true" pageId="14" pageNumber="526">R</emphasis>
. (
<emphasis id="CFB5EAE3017D6702FCC71932FCC68F09" box="[851,871,799,824]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FCEB1932FC7D8F09" box="[895,988,799,824]" italics="true" pageId="14" pageNumber="526">tenuipes</emphasis>
. Moreover, the above described species is markedly larger than the remaining species of the
<taxonomicName id="3AC14D72017D6702FD7E196EFC618F6D" baseAuthorityName="Fabricius" baseAuthorityYear="1794" box="[746,960,835,861]" class="Insecta" family="Empididae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="524" phylum="Arthropoda" rank="species" species="nigripennis" subGenus="Holoclera">
<emphasis id="CFB5EAE3017D6702FD7E196EFD5A8F6D" box="[746,763,835,860]" italics="true" pageId="14" pageNumber="526">R</emphasis>
. (
<emphasis id="CFB5EAE3017D6702FC87196EFC868F6D" box="[787,807,835,860]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FCD4196EFC618F6D" box="[832,960,835,860]" italics="true" pageId="14" pageNumber="526">nigripennis</emphasis>
</taxonomicName>
group (wing length of all other species of this group is less than
<quantity id="3A399B14017D6702FE05194AFE778FB0" box="[401,470,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="526" unit="mm" value="4.0">4 mm</quantity>
).
</paragraph>
<paragraph id="FD7E36F1017D6702FF5319A6FB278FF8" blockId="14.[151,1437,151,1005]" pageId="14" pageNumber="526">
Prior to our study, only one species of
<emphasis id="CFB5EAE3017D6702FD1E19A6FD038F95" box="[650,674,907,932]" italics="true" pageId="14" pageNumber="526">R.</emphasis>
(
<taxonomicName id="3AC14D72017D6702FD2119A6FC868F94" authorityName="Schiner" authorityYear="1860" box="[693,807,907,933]" class="Insecta" family="Empididae" genus="Holoclera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="CFB5EAE3017D6702FD2119A6FC868F94" box="[693,807,907,933]" italics="true" pageId="14" pageNumber="526">Holoclera</emphasis>
</taxonomicName>
) was reported from
<collectingCountry id="85D67661017D6702FB8119A6FBDC8F94" box="[1045,1149,907,933]" name="Morocco" pageId="14" pageNumber="526">Morocco</collectingCountry>
, namely
<emphasis id="CFB5EAE3017D6702FB7219A6FB568F95" box="[1254,1271,907,932]" italics="true" pageId="14" pageNumber="526">R</emphasis>
. (
<emphasis id="CFB5EAE3017D6702FA8519A6FA848F95" box="[1297,1317,907,932]" italics="true" pageId="14" pageNumber="526">H</emphasis>
.)
<emphasis id="CFB5EAE3017D6702FAAB19A6FA3D8F95" box="[1343,1436,907,932]" italics="true" pageId="14" pageNumber="526">tenuipes</emphasis>
Becker (
<bibRefCitation id="99504B00017D6702FF631982FE6D8FF8" author="Kettani, K. &amp; Ebejer, M. J. &amp; Ackland, D. M. &amp; Bachli, G. &amp; Barraclough, D. &amp; Bartak, M. &amp; Carles-Tolra, M. &amp; Cerny, M. &amp; Cerretti, P. &amp; Chandler, P. &amp; Dakki, M. &amp; Daugeron, C. &amp; De Jong, H. &amp; Dils, J. &amp; Disney, H. &amp; Droz, B. &amp; Evenhuis, N. &amp; Gatt, P. &amp; Graciolli, G. &amp; Grichanov, I. Y. &amp; Haenni, J-P. &amp; Hauser, M. &amp; Himmi, O. &amp; Macgowan, I. &amp; Mathieu, B. &amp; Mouna, M. &amp; Munari, L. &amp; Nartshuk, E. P. &amp; Negrobov, O. P. &amp; Oosterbroek, P. &amp; Pape, T. &amp; Pont, A. C. &amp; Popov, G. V. &amp; Rognes, K. &amp; Skuhrava, M. &amp; Skuhravy, V. &amp; Speight, M. &amp; Tomasovic, G. &amp; Trari, B. &amp; Tschorsnig, H-P. &amp; Vala, J-C. &amp; von Tschirnhaus, M. &amp; Wagner, R. &amp; Whitmore, D. &amp; Woznica, A. J. &amp; Zatwarnicki, T. &amp; Zwick, P." box="[247,460,943,969]" pageId="14" pageNumber="526" pagination="1 - 466" refId="ref21806" refString="Kettani, K., Ebejer, M. J., Ackland, D. M., Bachli, G., Barraclough, D., Bartak, M., Carles-Tolra, M., Cerny, M., Cerretti, P., Chandler, P., Dakki, M., Daugeron, C., De Jong, H., Dils, J., Disney, H., Droz, B., Evenhuis, N., Gatt, P., Graciolli, G., Grichanov, I. Y., Haenni, J-P., Hauser, M., Himmi, O., Macgowan, I., Mathieu, B., Mouna, M., Munari, L., Nartshuk, E. P., Negrobov, O. P., Oosterbroek, P., Pape, T., Pont, A. C., Popov, G. V., Rognes, K., Skuhrava, M., Skuhravy, V., Speight, M., Tomasovic, G., Trari, B., Tschorsnig, H-P., Vala, J-C., von Tschirnhaus, M., Wagner, R., Whitmore, D., Woznica, A. J., Zatwarnicki, T. &amp; Zwick, P. (2022) Catalogue of the Diptera (Insecta) of Morocco - an annotated checklist, with distributions and a bibliography. ZooKeys, 1094, 1 - 466. https: // doi. org / 10.3897 / zookeys. 1094.62644" type="journal article" year="2022">
Kettani
<emphasis id="CFB5EAE3017D6702FEC41982FE208FF8" box="[336,385,943,969]" italics="true" pageId="14" pageNumber="526">et al</emphasis>
. 2022
</bibRefCitation>
). Currently, the subgenus includes two species from
<collectingCountry id="85D67661017D6702FB8D1982FB238FF8" box="[1049,1154,943,969]" name="Morocco" pageId="14" pageNumber="526">Morocco</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="B5DB657A017D6702FF5319FFFD7B8FDC" box="[199,730,978,1005]" pageId="14" pageNumber="526" type="distribution">
<paragraph id="FD7E36F1017D6702FF5319FFFD7B8FDC" blockId="14.[151,1437,151,1005]" box="[199,730,978,1005]" pageId="14" pageNumber="526">
<emphasis id="CFB5EAE3017D6702FF5319FFFEC08FDD" bold="true" box="[199,353,978,1004]" pageId="14" pageNumber="526">Distribution.</emphasis>
<collectingCountry id="85D67661017D6702FEF319FEFE6E8FDC" box="[359,463,979,1005]" name="Morocco" pageId="14" pageNumber="526">Morocco</collectingCountry>
, High Atlas Mountains.
</paragraph>
</subSubSection>
</treatment>
</document>
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