287 lines
33 KiB
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287 lines
33 KiB
XML
<document ID-CLB-Dataset="9622" ID-DOI="http://dx.doi.org/10.3897/zookeys.987.53648" ID-GBIF-Dataset="e14abbdb-e0a3-4c35-ad98-1cfd0cee46b5" ID-PMC="PMC7666096" ID-Pensoft-Pub="1313-2970-987-135" ID-Pensoft-UUID="A304608B226C5889BE8AF26E618669A3" ID-PubMed="33223888" ID-ZooBank="861F9C146E6D4222A41FFC56F978C2E8" ModsDocID="1313-2970-987-135" checkinTime="1604753978193" checkinUser="pensoft" docAuthor="Inoue, Shota, Nomura, Shuhei & Yin, Zi-Wei" docDate="2020" docId="190D674F79B358378011AF4CAE51FF2B" docLanguage="en" docName="ZooKeys 987: 135-156" docOrigin="ZooKeys 987" docSource="http://dx.doi.org/10.3897/zookeys.987.53648" docTitle="Pseudophanias excavatus Inoue, Nomura & Yin 2020, sp. nov." docType="treatment" docUuid="E9C17AEF-CEA2-45DB-A952-52905F0943BE" docUuidSource="ZooBank" docVersion="6" id="A304608B226C5889BE8AF26E618669A3" lastPageNumber="135" masterDocId="A304608B226C5889BE8AF26E618669A3" masterDocTitle="Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="156" masterPageNumber="135" pageNumber="135" updateTime="1732761297298" updateUser="ExternalLinkService">
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<mods:mods id="78BCCFD10DA63DFA905B7A94F79C9033" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="571E577FA569E6B350DA535958D59D8B">
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<mods:title id="2EB7AFB11196DD85E3E8DD7518FE191D">Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0C1D103101FAAC4AAE359C81F673FB63" type="personal">
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<mods:roleTerm id="8783C0825CD6C3DE3E9F927C1199821E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4411424A89D68D0B23657A63522E854C">Inoue, Shota</mods:namePart>
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<mods:affiliation id="60A0560A067FE903640363CBE494B2BF">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819 - 0395, Japan & The Kyushu University Museum, Fukuoka 812 - 8581, Japan</mods:affiliation>
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<mods:nameIdentifier id="1B977F7B8C673F1BF3B885936587F8C8" type="orcid">https://orcid.org/0000-0002-7005-6531</mods:nameIdentifier>
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<mods:nameIdentifier id="2CC44F3414042C53EC02E4552E7D288E" type="email">pselaphineman@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="5D1E37D429373F0900561A6C71656F06" type="personal">
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<mods:roleTerm id="3B18FBA0CDBF7421CB72FD29C15D80AD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="807F0FACD008211C60D0E8B2E08A38DD">Nomura, Shuhei</mods:namePart>
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<mods:affiliation id="46A1A7AB239D0DA6472496FE9D4C4602">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki 305 - 0005, Japan</mods:affiliation>
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</mods:name>
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<mods:name id="9AF49B92DC45107D1FB4249DD996AC23" type="personal">
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<mods:roleTerm id="C2A11810CA54D3BC59E137636939A4C5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C62B3112056BD1CA8B9D48C6412A8B8">Yin, Zi-Wei</mods:namePart>
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<mods:affiliation id="246DC0A0CC4C3FD8E358B57F1CD2A621">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai 200234, China</mods:affiliation>
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<mods:typeOfResource id="48B696F753D361F6BC564A91E92A76E4">text</mods:typeOfResource>
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<mods:title id="D089E9DE329D8C8C0B49B4BEF3230241">ZooKeys</mods:title>
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<mods:date id="BD8BC2389B77F5F1F83751846166A7D5">2020</mods:date>
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<mods:detail id="8C7925616A85B4868AE8957B18C4154F" type="volume">
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<mods:number id="E1AA81CA7F21EEFA1388C8DE72748229">987</mods:number>
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<mods:start id="AB7FD22BEBA32500903553D6DDC135E3">135</mods:start>
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<mods:url id="9C79C33B213762456ED6BCACBAF4CB4F">http://dx.doi.org/10.3897/zookeys.987.53648</mods:url>
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<mods:classification id="2596BFFB68C3768C17874D1582BC1A4F">journal article</mods:classification>
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<mods:identifier id="03DEDA8D42120E8F4747945E839B4F3E" type="DOI">http://dx.doi.org/10.3897/zookeys.987.53648</mods:identifier>
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<mods:identifier id="7A802325F38A0B3BA76BA59F488BCBF5" type="Pensoft-Pub">1313-2970-987-135</mods:identifier>
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<mods:identifier id="13464075A1594B0792BB06E0A727F57E" type="ZooBank">861F9C146E6D4222A41FFC56F978C2E8</mods:identifier>
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<treatment id="190D674F79B358378011AF4CAE51FF2B" ID-GBIF-Taxon="168821845" LSID="urn:lsid:zoobank.org:act:E9C17AEF-CEA2-45DB-A952-52905F0943BE" httpUri="http://treatment.plazi.org/id/190D674F79B358378011AF4CAE51FF2B" lastPageNumber="135" pageId="0" pageNumber="135" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Pselaphinae">
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<subSubSection id="48EF3952A04C0A8965A4484D93B46DD1" pageId="0" pageNumber="135" type="nomenclature">
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<paragraph id="928EC4D3825748B8ECEA7B94A5042F30" pageId="0" pageNumber="135">
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<taxonomicName id="6A2F3700F06BEE71780F98CDD7DAEF00" ID-CoL="8Y8SY" LSID="190D674F-79B3-5837-8011-AF4CAE51FF2B" authority="Inoue, Nomura & Yin" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus" status="sp. nov.">Pseudophanias excavatus Inoue, Nomura & Yin</taxonomicName>
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<taxonomicNameLabel id="B19B1D9DA7AC09E54C6179B7FD2EE85E" pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
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<figureCitation id="2A372C2C0A7B3FA7D36372AB31F8E86A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">Figs 9</figureCitation>
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<figureCitation id="6767A48C855A15E6374E2487E5615D95" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">, 10</figureCitation>
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<figureCitation id="9113CD9269C826196C0C5325718C5054" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">, 11</figureCitation>
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<figureCitation id="3D0D760776E4ADDB016907163558233D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male antenna of Pseudophanias excavatus by SEM images A lateral view B ventral view C antennomere 11 in ventral view D antennomeres 5 - 10 in ventral view E special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (A, B); 0.1 mm (C, D); 0.01 mm (E)." figureDoi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135">, 12</figureCitation>
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<figureCitation id="53A2A92AE6E674D038780EC1D5C760A0" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">, 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F88707B36F0E214D4F1552DC8DB6FB20" pageId="0" pageNumber="135" type="type material">
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<paragraph id="12E99027397A5FD82FF464415FD7459D" pageId="0" pageNumber="135">Type material.</paragraph>
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<paragraph id="D0C46992A382E3F24AF1B53DB6516CDA" pageId="0" pageNumber="135">
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<emphasis id="2ADBD3F1769AF2BD56B644DF5A834BC1" bold="true" italics="true" pageId="0" pageNumber="135">Holotype</emphasis>
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(NSMT): ♂, "Tengshih (1400 m, litter) / Kaosiung Hsien / [M-Taiwan] /
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<normalizedToken id="4EC1DE9F2D510813EBEB59BB75360560" originalValue="台湾高雄縣藤枝">台湾高雄縣藤枝</normalizedToken>
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/ 20-22. iv. 2001, H. Sugaya leg. // HOLOTYPE (red) /♂,
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<taxonomicName id="B97B925C303A8BE2CBE954AD53CA7C81" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
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<emphasis id="D6E97F8BCD2B739866CCF75002AF0C78" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020"
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<emphasis id="1757BBEA914EBD563C04ACC8B75C4F68" bold="true" italics="true" pageId="0" pageNumber="135">Paratypes</emphasis>
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: (NSMT, KUM, NMNS) 1 ♂, 3 ♀, same data as holotype; 6 ♂, 1 ♀, same data as holotype, but 29-30 IV 2001; 1 ♂, 2 ♀, same data as holotype, but 30 IV 2001. Each paratype pinned with the following label: "PARATYPE (yellow) / ♂ (or ♀),
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<taxonomicName id="3363835998E84EBDDA98CDD5B7C20C3F" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
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<emphasis id="07D73DE31904541582F80D9AEE535976" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
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</taxonomicName>
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sp. nov., / det. Inoue, Nomura & Yin, 2020".
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</paragraph>
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</subSubSection>
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<subSubSection id="6F41518663F1155C291646F7F26CDD52" pageId="0" pageNumber="135" type="diagnosis">
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<paragraph id="5639B1B6F7B09D19FD867817C2BAAA3F" pageId="0" pageNumber="135">Diagnosis.</paragraph>
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<paragraph id="8D04BA59FF23797DB0EA28D39BD80DC3" pageId="0" pageNumber="135">
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This species is readily distinguished from other members of
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<taxonomicName id="55F88003ADEAB5E4F605CA427E11D9AC" authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
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<emphasis id="4FEB889EBD6D5F5481F0AD57483A8A2B" italics="true" pageId="0" pageNumber="135">Pseudophanias</emphasis>
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</taxonomicName>
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by the clasping formed antennae in the male, frontal sulcus indistinct, and the rounded pronotum.
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</paragraph>
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</subSubSection>
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<subSubSection id="078BC2131A23ABC1BA45BCEEEE9915F9" pageId="0" pageNumber="135" type="description">
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<paragraph id="A6B4FDBE44E236181D08D22F57E230E2" pageId="0" pageNumber="135">Description.</paragraph>
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<paragraph id="709A119A46ED14DA16CF1987E40CAFA1" pageId="0" pageNumber="135">
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<emphasis id="70C5DB3A2A3B6227DA3D710B3E90E564" bold="true" pageId="0" pageNumber="135">Male</emphasis>
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(Figs
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<figureCitation id="3647193B5A87D98001CB53E4FCA811D8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">9A</figureCitation>
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,
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<figureCitation id="3609825B8235FB890F0D54A1102D4032" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10A</figureCitation>
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). Body length 2.21-2.48 mm. Dorsal surface with dense setae.
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</paragraph>
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<paragraph id="452B9CF7DEA00DE6F1AF9F296E80C5E9" pageId="0" pageNumber="135">
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<emphasis id="400C788182FB6FFD1A5CEB046B56463E" bold="true" italics="true" pageId="0" pageNumber="135">Head</emphasis>
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(Fig.
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<figureCitation id="41A89AE0EB338CB34BB2330502737F1F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11A</figureCitation>
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) as long as wide, HL 0.45-0.50 mm, HW 0.43-0.50 mm, densely punctate, with dense, long setae; frontal rostrum broad, with frontal fovea nude; antennal tubercles distinct; vertex flat, with pair of glabrous foveae; eyes prominent, small; occiput with dense setae; postocular margin two times longer than eyes; small areas just posterior to U-shaped setose sulci finely punctate.
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<emphasis id="5BAD14C97777BE0D864DC28D1AACF9D7" bold="true" italics="true" pageId="0" pageNumber="135">Antennae</emphasis>
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(Figs
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<figureCitation id="8D090EB0721CA144B934971856337C65" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10B</figureCitation>
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,
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<figureCitation id="C88B7B0B94E48D10C188ABE2EE8B7FF6" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male antenna of Pseudophanias excavatus by SEM images A lateral view B ventral view C antennomere 11 in ventral view D antennomeres 5 - 10 in ventral view E special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (A, B); 0.1 mm (C, D); 0.01 mm (E)." figureDoi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135">12A-E</figureCitation>
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) strongly modified, 0.98-1.12 mm in length; antennomeres 1 elongate, as long as 2-4 combined; 2-4 each transverse, successively shorter; 5-11 strongly excavated on ventral side, with tufts of setae on ventral surface, modified to form clasping, each excavated on ventral surface; 5-9 each two times wider than 4, each transverse; 9 longer than 8; 10 as long as 9, asymmetrical; outer side strongly produced ventrally in 9; 11 enlarged, with glabrous areas on ventral side. Maxillary palpi (Fig.
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<figureCitation id="C3BF064F36E1E19672FB3DCF01B9B95E" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11B</figureCitation>
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) symmetrical; palpomeres 1 minute; palpomeres 2 elongate, narrowed in basal halves; palpomeres 3 small, widest at apices; palpomeres 4 fusiform.
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<emphasis id="0DFEF5119FCA75A357BD18DA338F9BEA" bold="true" italics="true" pageId="0" pageNumber="135">Pronotum</emphasis>
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(Fig.
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<figureCitation id="CE441D46CD6F2B2C9EFE3864D153CD0B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11C</figureCitation>
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) about as long as wide, PL 0.47-0.55 mm, PW 0.50-0.55 mm, broadly rounded, widest at middle, finely punctate on dorsal surface, with a median and pair of lateral antebasal foveae; antebasal area strongly punctate.
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<emphasis id="6F3188EE9271CAE91EEA3476EE87460E" bold="true" italics="true" pageId="0" pageNumber="135">Metaventrite</emphasis>
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(Figs
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<figureCitation id="D6032B6F70D7AFA18053DF6A2B251DFF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10D</figureCitation>
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,
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<figureCitation id="59516AF1F308DE499C58A7CD00409E7B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11D</figureCitation>
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) finely punctate, strongly convex, but area just above metaventral apex roundly impressed; that impression 2/3 as long as metaventral length, occupying 1/4 metaventral width; anterior margin of that impression sharply rounded, distinct.
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<emphasis id="E1A45EC25A08727289668B20A06EA11D" bold="true" italics="true" pageId="0" pageNumber="135">Elytra</emphasis>
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(Fig.
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<figureCitation id="EC324AD729296DAF9B01F90490F9E694" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11E</figureCitation>
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) much wider than long, EL 0.57-0.65 mm, EW 0.90-0.98 mm, trapezoidal, finely punctate, each elytron with two basal fovea; discal stria shallow, extending from basal fovea placed middle to posterior 1/3.
|
|
<emphasis id="A67CF5BB07D89CFCCAA5DBEAD1DF98F2" bold="true" pageId="0" pageNumber="135">
|
|
<emphasis id="B51FC62E1EFF1F853E13003A3B12A065" italics="true" pageId="0" pageNumber="135">Legs</emphasis>
|
|
.
|
|
</emphasis>
|
|
All legs moderately short; each femora broadest near middle; protibiae, mesotibiae with dense yellow setae at apices; protibiae, metatibiae moderately straight; mesotibiae slightly arcuate at apical fourth; tarsi (Fig.
|
|
<figureCitation id="B95CC8B80E51C05B0046C0DED06506FC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11G</figureCitation>
|
|
) each with tarsomere 2 half as long as tarsomere 3; tarsal claws (Fig.
|
|
<figureCitation id="74014D088340DAA2664073F0BDB2D751" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11H</figureCitation>
|
|
) asymmetrical; anterior claws long, posterior claws thin, short.
|
|
<emphasis id="1132C1F082EA0CD267C1A67F53681565" bold="true" italics="true" pageId="0" pageNumber="135">Abdomen</emphasis>
|
|
(Fig.
|
|
<figureCitation id="122731712C2FF9B3FB8BE3E40A22AC59" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. SEM images of Pseudophanias excavatus, male A head B maxillary palp C pronotum D metaventrite E elytra F abdomen G protarsus H protarsal claws. Scale bars: 0.2 mm (A, C-F); 0.1 mm (B, G); 0.05 mm (H)." figureDoi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135">11F</figureCitation>
|
|
) much wider than long, widest at tergite IV, AL 0.64-0.81 mm, AW 0.97-1.05 mm, lacking discal carinae; tergite IV longest, twice as long as V, with setose depression at base. Tergite and sternite VIII as in Fig.
|
|
<figureCitation id="F2A789F4868A8A7FDA10ACCA2669D11B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">13D, E</figureCitation>
|
|
.
|
|
<emphasis id="30DDA1CBDE0F2AF3374947BE8B4171DD" bold="true" italics="true" pageId="0" pageNumber="135">Aedeagus</emphasis>
|
|
(Fig.
|
|
<figureCitation id="F94519D96D9D420CC8E0A41E7DAEABD7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male genitalia of Pseudophanias excavatus A ventral view B lateral view C dorsal view D tergite VIII E sternite VIII. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135">13A-C</figureCitation>
|
|
) 0.60-0.63 mm in length, well-sclerotized, slightly asymmetrical in dorsal and ventral view, tubular in lateral view; parameres symmetrical, extremely elongate, reaching apical third, each with five setae at apex; median lobe roundly curved, C-shaped in lateral view; apical part widely opened, narrowed towards basal part to connect ovoidal dorsal diaphragm; apical lobe extending downward, curved to form S-shaped in lateral view, widely opened at base, strongly produced laterally at apex; endophallus indistinct.
|
|
</paragraph>
|
|
<paragraph id="7B1FD023BCB40BC367350C1C08D6395E" pageId="0" pageNumber="135">
|
|
<emphasis id="AB3FA1CC60CFEDC2E0970DB6C88DEB34" bold="true" pageId="0" pageNumber="135">Female</emphasis>
|
|
(Fig.
|
|
<figureCitation id="F6C9AB2104264736DB536C09DB807CD1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Dorsal habitus of Pseudophanias excavatus A male B female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135">9B</figureCitation>
|
|
). BL 2.21-2.27 mm; HL 0.44-0.48 mm; HW 0.44-0.48 mm; PL 0.47-0.55 mm; PW 0.46-0.51 mm; EL 0.56-0.63 mm; EW 0.90-0.94 mm; AL 0.67-0.70 mm; AW 0.94-0.96 mm. Antennae (Fig.
|
|
<figureCitation id="8A17647D3F24B9D30D4BB7A6FF6E1132" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Morphological details of Pseudophanias excavatus A habitus in lateral view B male antenna C female antenna D male metaventrite. Scale bars: 1.0 mm (A); 0.5 mm (D); 0.2 mm (B, C)." figureDoi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135">10C</figureCitation>
|
|
) with antennomeres 11 simple, successively widened towards apices; antennomeres 2-10 each transverse; 11 ovoid, largest. Metaventrite convex, lacking metaventral impression.
|
|
</paragraph>
|
|
<caption id="3BCC0603A84D5F80D1BC94C77FB47FD6" doi="10.3897/zookeys.987.53648.figure9" httpUri="https://binary.pensoft.net/fig/472464" pageId="0" pageNumber="135" start="Figure 9" startId="F9">
|
|
<paragraph id="CC4C91C48E5617CB9CF6B288A89D8E91" pageId="0" pageNumber="135">
|
|
<emphasis id="CBE2DD73FC428C8A9FB0677C739EDB90" bold="true" pageId="0" pageNumber="135">Figure 9.</emphasis>
|
|
Dorsal habitus of
|
|
<taxonomicName id="840B02C03CA09C3233D325FEF99465BC" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis id="D53A088895BD98045FE6277FF42F8679" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="67645893C2416A4C2458F187B1313454" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
male
|
|
<emphasis id="E751A7CBF7FE1D34FD407FF0371250F2" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
female. Scale bar: 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="462EAF6AEDD06E0C5CAAB5E859DD3104" doi="10.3897/zookeys.987.53648.figure10" httpUri="https://binary.pensoft.net/fig/472465" pageId="0" pageNumber="135" start="Figure 10" startId="F10">
|
|
<paragraph id="98C9082C00A7CAE71A3AC8F3EDC14C22" pageId="0" pageNumber="135">
|
|
<emphasis id="45401A7941CF0E2A2A528B6BA936EB41" bold="true" pageId="0" pageNumber="135">Figure 10.</emphasis>
|
|
Morphological details of
|
|
<taxonomicName id="70C11A09151ACA1870124861BEA9BF5B" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis id="BF44D67893C7F111385F9C81E8352EDA" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="CB54A64756E4D46CCBBCCAB6D12E632E" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
habitus in lateral view
|
|
<emphasis id="F7AB6995304E078A73B11573D182C985" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
male antenna
|
|
<emphasis id="0C55BEDDFD2B820443C110913FC5DE56" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
female antenna
|
|
<emphasis id="688A9174D95BB590759D951D51F2F41D" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
male metaventrite. Scale bars: 1.0 mm (
|
|
<emphasis id="4859D2BB697E57B2E4047DAC518A783D" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
); 0.5 mm (
|
|
<emphasis id="02B6FB9469320B4FF05F1DC98821B54D" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
); 0.2 mm (
|
|
<emphasis id="864FE7356EF300C551CCF8EBD130868F" bold="true" pageId="0" pageNumber="135">B, C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="0CA60E6C5FA1A57677C07EC7AF7EA96D" doi="10.3897/zookeys.987.53648.figure11" httpUri="https://binary.pensoft.net/fig/472466" pageId="0" pageNumber="135" start="Figure 11" startId="F11">
|
|
<paragraph id="4FB156E6F2B2E365D7BF2FC746F99F99" pageId="0" pageNumber="135">
|
|
<emphasis id="01E3E149DCCFFE24B2F854FD25DE9C82" bold="true" pageId="0" pageNumber="135">Figure 11.</emphasis>
|
|
SEM images of
|
|
<taxonomicName id="F0B09590ED9738B156FE1CE6F4B358C9" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis id="115339188387B6F47D8B7EEDFBA06D84" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
, male
|
|
<emphasis id="E3968FA91B3C480B274038B3C9DD44A3" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
head
|
|
<emphasis id="68B34BE0F1BF060D265598560BEA9D46" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
maxillary palp
|
|
<emphasis id="7F8A5AA66D3E62EA3668F48CDCF6614D" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
pronotum
|
|
<emphasis id="B5BD7655F54F6A09EE15CA0B65CDA39B" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
metaventrite
|
|
<emphasis id="66EB033103DC2F01F4388C7E9E6D0C15" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
elytra
|
|
<emphasis id="CCF59F72F6A3EFB2CAA6DC6C3A248710" bold="true" pageId="0" pageNumber="135">F</emphasis>
|
|
abdomen
|
|
<emphasis id="FD30BD613E520202673E7D2755BC70D1" bold="true" pageId="0" pageNumber="135">G</emphasis>
|
|
protarsus
|
|
<emphasis id="269672966DFFEB95D6F9950AAB0CDF8E" bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
protarsal claws. Scale bars: 0.2 mm (
|
|
<emphasis id="0E6237A598C9AC31240816696C64BCC2" bold="true" pageId="0" pageNumber="135">A, C-F</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="599AABDE7EE7F59C1D30CBE8F1A9F4CC" bold="true" pageId="0" pageNumber="135">B, G</emphasis>
|
|
); 0.05 mm (
|
|
<emphasis id="AB7290DD4678C63069D3672C10312221" bold="true" pageId="0" pageNumber="135">H</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="00DD5C45C0294C330AC9CF151D241B53" doi="10.3897/zookeys.987.53648.figure12" httpUri="https://binary.pensoft.net/fig/472467" pageId="0" pageNumber="135" start="Figure 12" startId="F12">
|
|
<paragraph id="019C1A06DFD29D6B8B7B1D1A4D96BF8F" pageId="0" pageNumber="135">
|
|
<emphasis id="1587229150C934A513077CE7FE92FB0A" bold="true" pageId="0" pageNumber="135">Figure 12.</emphasis>
|
|
Male antenna of
|
|
<taxonomicName id="8C19E1384607F0149DFD57703D5D976B" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis id="64A8AC1F4C3AA7932CE3BF52A41E1EB9" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
by SEM images
|
|
<emphasis id="7E4314AD23012BC24865AAC8C4B11D35" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
lateral view
|
|
<emphasis id="9F3E822AE6748591914B9AEDDC0FDB9B" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
ventral view
|
|
<emphasis id="B82DBFBA8EABFCB39459AD0C39F30090" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
antennomere 11 in ventral view
|
|
<emphasis id="36B9777679C0F7C3580FE5652329D760" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
antennomeres 5-10 in ventral view
|
|
<emphasis id="F7EA689E00C79B29C441205E06C2CAD6" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
special setae on antennomere VIII in ventral view. Scale bars: 0.2 mm (
|
|
<emphasis id="E082D697F26DF0FAFC9F8D8982A1F903" bold="true" pageId="0" pageNumber="135">A, B</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis id="F355BF83B46ABFA297E67C885286B462" bold="true" pageId="0" pageNumber="135">C, D</emphasis>
|
|
); 0.01 mm (
|
|
<emphasis id="C842B1F500F92E30E5EDFA730765DD73" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="2EF8434DE4B0EF213493B2CB93DC35BA" doi="10.3897/zookeys.987.53648.figure13" httpUri="https://binary.pensoft.net/fig/472468" pageId="0" pageNumber="135" start="Figure 13" startId="F13">
|
|
<paragraph id="3F93B9ACF0BC8F6FC40B22C094219E07" pageId="0" pageNumber="135">
|
|
<emphasis id="8E5E077FEF27470C0B54638BAD7F3565" bold="true" pageId="0" pageNumber="135">Figure 13.</emphasis>
|
|
Male genitalia of
|
|
<taxonomicName id="0A9D5948E2F05AD397BDFBA04BDD11DB" authorityName="Inoue, Nomura & Yin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias excavatus" order="Coleoptera" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="excavatus">
|
|
<emphasis id="5C4C05F88DEA93B18668C9089F34BD85" italics="true" pageId="0" pageNumber="135">Pseudophanias excavatus</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="8850024C9DF6718E58F54F413A48378F" bold="true" pageId="0" pageNumber="135">A</emphasis>
|
|
ventral view
|
|
<emphasis id="C3E915A9D901C895B173F7EECF91CA90" bold="true" pageId="0" pageNumber="135">B</emphasis>
|
|
lateral view
|
|
<emphasis id="92FB2AFD34980A58A88CB4BFFCB2E6E8" bold="true" pageId="0" pageNumber="135">C</emphasis>
|
|
dorsal view
|
|
<emphasis id="30B5CE486021166D08BE4CC5902697F0" bold="true" pageId="0" pageNumber="135">D</emphasis>
|
|
tergite VIII
|
|
<emphasis id="1CA6AF7E0594E1BE6EBA7545D15DE5FE" bold="true" pageId="0" pageNumber="135">E</emphasis>
|
|
sternite VIII. Scale bars: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="0EF767E0AFBEB65100212507FDAC2312" pageId="0" pageNumber="135" type="etymology">
|
|
<paragraph id="44EDDAC39A040D5E7780BC3BA7CA8A4C" pageId="0" pageNumber="135">Etymology.</paragraph>
|
|
<paragraph id="CB0974FDBFABD437E693E74484832AF0" pageId="0" pageNumber="135">The specific epithet refers to the strongly excavated antennae in the male of the new species.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FB5DE6A693B69F06F50EBF373645ADF9" pageId="0" pageNumber="135" type="distribution">
|
|
<paragraph id="DFFC12EA05D906672F528EE958BAA37C" pageId="0" pageNumber="135">Distribution.</paragraph>
|
|
<paragraph id="8A90DA09DC1C2F4F2DE0CFD76496AB5C" pageId="0" pageNumber="135">China (Taiwan).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BB7A11D6F42346E7FB6F663C033BFB15" pageId="0" pageNumber="135" type="biology_ecology">
|
|
<paragraph id="64197940B2368E2224ECE02610EE789D" pageId="0" pageNumber="135">Biology.</paragraph>
|
|
<paragraph id="A901B3C0BF8C6B564BDD7F62DBB2E09B" pageId="0" pageNumber="135">This species was collected from leaf litter.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |