131 lines
15 KiB
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131 lines
15 KiB
XML
<document id="24EF1972A0049BFC239A30BBCEEDCFD7" ID="10.11646/zootaxa.4125.1.1" ID-CLB-Dataset="38807" ID-DOI="10.11646/zootaxa.4125.1.1" ID-GBIF-Dataset="1af13aec-3c54-4a92-abdf-e4e29b97e82f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271704" ID-ZooBank="118F4865-D89E-45EA-A210-8D61946CC37F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465971881907" checkinUser="plazi" docAuthor="Matz, Jess & Brower, Andrew V. Z." docDate="2016" docId="03F187D7FFB18446FF11F920FB9CB8B5" docLanguage="en" docName="zootaxa.4125.1.1.pdf" docOrigin="Zootaxa 4125 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pamperis Heimlich 1959" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="FFC8FFAFFF8B847DFF86FFA0FFD9B918" masterDocTitle="The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes" masterLastPageNumber="108" masterPageNumber="1" pageNumber="59" updateTime="1698662999642" updateUser="ExternalLinkService">
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<mods:mods id="5AF510514F944E6B8A589B43D47B7B25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9648E884CEED5AD8199365F6D331195D">
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<mods:title id="3C354E843DC6BE704FE03E391F459F8D">The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes</mods:title>
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<mods:name id="B41AA99DBBC6AB9BD36E2A517E29CF1E" type="personal">
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<mods:roleTerm id="3C78ACF0A4A0FFEE81870DFFFCF2F177">Author</mods:roleTerm>
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<mods:namePart id="6EEEB3724FBE1DD1E97AD6C2BE23BCC1">Matz, Jess</mods:namePart>
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<mods:name id="1CCB7B68BAE70B498E24381D0E3B0BFF" type="personal">
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<mods:roleTerm id="29F5B78F0AF91D26B2799322F5CA91A3">Author</mods:roleTerm>
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<mods:namePart id="A4905A834641C148A34A4F72301E7622">Brower, Andrew V. Z.</mods:namePart>
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<mods:typeOfResource id="67E64ADDC258B8409E6C27EDA3DAC210">text</mods:typeOfResource>
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<mods:title id="D1B2A5B9A51E801F0AB09878590FF4BB">Zootaxa</mods:title>
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<mods:part id="72E4EDF7A5E553D9376FEB12F1AA4AE0">
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<mods:date id="47A5DB4F14E1CD1AF3787E389ABD04C2">2016</mods:date>
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<mods:detail id="B3CB4C823A92755F6521179D0C056892" type="volume">
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<mods:number id="684C2AF7A3D7E8CAD33F1D0D794EC5F8">4125</mods:number>
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<mods:number id="86C471EBE063883E6C27CA22D7F80E20">1</mods:number>
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<mods:classification id="675AE1A4F8897CB95A7E98BD07AF2EE4">journal article</mods:classification>
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<mods:identifier id="92B9120E1D1A248BB3C24290A66C5BFF" type="CLB-Dataset">38807</mods:identifier>
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<mods:identifier id="D8264A642F3354129C86698E3F111953" type="DOI">10.11646/zootaxa.4125.1.1</mods:identifier>
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<treatment id="03F187D7FFB18446FF11F920FB9CB8B5" ID-DOI="http://doi.org/10.5281/zenodo.6070082" ID-GBIF-Taxon="120828508" ID-Zenodo-Dep="6070082" LSID="urn:lsid:plazi:treatment:03F187D7FFB18446FF11F920FB9CB8B5" httpUri="http://treatment.plazi.org/id/03F187D7FFB18446FF11F920FB9CB8B5" lastPageId="59" lastPageNumber="59" pageId="58" pageNumber="59">
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<subSubSection id="C342654AFFB18447FF11F920FE15BF82" box="[151,460,1664,1690]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="8BE736C1FFB18447FF11F920FE15BF82" blockId="58.[151,460,1664,1690]" box="[151,460,1664,1690]" pageId="58" pageNumber="59">
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<heading id="D0AF81ADFFB18447FF11F920FE15BF82" bold="true" box="[151,460,1664,1690]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<taxonomicName id="4C584D42FFB18447FF11F920FE15BF82" ID-CoL="92HRF" authority="Heimlich, 1959" authorityName="Heimlich" authorityYear="1959" box="[151,460,1664,1690]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB18447FF11F920FE15BF82" bold="true" box="[151,460,1664,1690]" pageId="58" pageNumber="59">
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<emphasis id="B92CEAD3FFB18447FF11F920FED0BF82" bold="true" box="[151,265,1664,1690]" italics="true" pageId="58" pageNumber="59">Pamperis</emphasis>
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<bibRefCitation id="EFC94B30FFB18447FE96F920FE15BF82" author="Heimlich" box="[272,460,1664,1690]" pageId="58" pageNumber="59" refString="Heimlich, W. (1959) Eine neue satyride aus Chile. Entomologische Zeitschrift, 69 (16), 173 - 179." type="journal article" year="1959">Heimlich, 1959</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFB18447FF11F966FD47BFC4" box="[151,670,1734,1757]" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8BE736C1FFB18447FF11F966FD47BFC4" blockId="58.[151,670,1734,1757]" box="[151,670,1734,1757]" pageId="58" pageNumber="59">
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<emphasis id="B92CEAD3FFB18447FF11F966FEFCBFC4" bold="true" box="[151,293,1734,1756]" pageId="58" pageNumber="59">
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<typeStatus id="54E38863FFB18447FF11F966FF15BFC4" box="[151,204,1734,1756]" pageId="58" pageNumber="59">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C584D42FFB18447FEADF968FD47BFC4" authority="Heimlich, 1959" authorityName="Heimlich" authorityYear="1959" box="[299,670,1734,1757]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="poaoeneis">
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<emphasis id="B92CEAD3FFB18447FEADF968FE23BFC5" box="[299,506,1736,1757]" italics="true" pageId="58" pageNumber="59">Pamperis poaoeneis</emphasis>
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<bibRefCitation id="EFC94B30FFB18447FD86F966FD47BFC4" author="Heimlich" box="[512,670,1734,1756]" pageId="58" pageNumber="59" refString="Heimlich, W. (1959) Eine neue satyride aus Chile. Entomologische Zeitschrift, 69 (16), 173 - 179." type="journal article" year="1959">Heimlich, 1959</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFB18447FF11F8A9FC76BE72" pageId="58" pageNumber="59" type="diagnosis">
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<paragraph id="8BE736C1FFB18447FF11F8A9FC76BE72" blockId="58.[151,1436,1801,2006]" pageId="58" pageNumber="59">
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<emphasis id="B92CEAD3FFB18447FF11F8A9FED3BE3A" bold="true" box="[151,266,1801,1826]" pageId="58" pageNumber="59">Diagnosis</emphasis>
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. A dark gray butterfly with no forewing ocelli, and extremely aberrant wing venation. Forewing R1 has a separate origin at the anterior end of the discal cell and fuses with the subcosta, while the remaining radial veins branch as follows; R5, R2, R3+R4. in the hindwing, vein A3 is absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C342654AFFB18446FF41F8D5FB9CB8B5" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="discussion">
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<paragraph id="8BE736C1FFB18446FF41F8D5FB9CB8B5" blockId="58.[151,1436,1801,2006]" lastBlockId="59.[151,1436,151,429]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
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<emphasis id="B92CEAD3FFB18447FF41F8D5FEEDBE96" bold="true" box="[199,308,1909,1934]" pageId="58" pageNumber="59">Remarks</emphasis>
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. In the original description, Heimlich associated
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<taxonomicName id="4C584D42FFB18447FCF2F8D6FB85BE95" box="[884,1116,1910,1933]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="poaoeneis">
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<emphasis id="B92CEAD3FFB18447FCF2F8D6FB85BE95" box="[884,1116,1910,1933]" italics="true" pageId="58" pageNumber="59">Pamperis poaoeneis</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="4C584D42FFB18447FB25F8D6FA98BE95" box="[1187,1345,1910,1933]" class="Insecta" family="Nymphalidae" genus="Cosmosatyrus" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB18447FB25F8D6FA98BE95" box="[1187,1345,1910,1933]" italics="true" pageId="58" pageNumber="59">Cosmosatyrus</emphasis>
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</taxonomicName>
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, but he considered the resemblance insufficient for placing in that genus, and created the monotypic genus
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<taxonomicName id="4C584D42FFB18447FA81F83AFAAABEA9" box="[1287,1395,1946,1969]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB18447FA81F83AFAAABEA9" box="[1287,1395,1946,1969]" italics="true" pageId="58" pageNumber="59">Pamperis</emphasis>
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</taxonomicName>
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for this unusual species based primarily on peculiarities in the wing venation.
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<bibRefCitation id="EFC94B30FFB18447FB8EF81DFB7DBECE" author="Miller" box="[1032,1188,1981,2006]" pageId="58" pageNumber="59" refString="Miller, L. D. (1968) The higher classification, phylogeny and zoogeography of the Satyridae (Lepidoptera). Memoirs of the American Entomological Society, 24, 1 - 174. http: // dx. doi. org / 10.1130 / MEM 110 - p 1" type="journal article" year="1968">Miller (1968)</bibRefCitation>
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further set
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<taxonomicName id="4C584D42FFB18446FAB7F81EFED0B9A8" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" lastPageId="59" lastPageNumber="60" order="Lepidoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="poaoeneis">
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<emphasis id="B92CEAD3FFB18446FAB7F81EFED0B9A8" italics="true" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">Pamperis poaoeneis</emphasis>
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</taxonomicName>
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apart as a genus of uncertain position, again due to its unusual venation, noting that "the veins in the apical portion of the forewing are arranged like no known satyrid."
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<bibRefCitation id="EFC94B30FFB08446FC43FF1DFB22B9CC" author="Pyrcz" box="[965,1275,188,213]" pageId="59" pageNumber="60" refString="Pyrcz, T. W. & Wojtusiak, J. (2010) A new species of Argyrophorus Blanchard from northern Peru and considerations on the value of wing venation as a source of synapomorphies in some temperate Neotropical Satyrinae (Lepidoptera: Nymphalidae). Genus, 21, 605 - 613." type="journal article" year="2010">Pyrcz & Wojtusiak (2010)</bibRefCitation>
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synonymized
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<taxonomicName id="4C584D42FFB08446FF11FF41FEDAB9E0" box="[151,259,225,248]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB08446FF11FF41FEDAB9E0" box="[151,259,225,248]" italics="true" pageId="59" pageNumber="60">Pamperis</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="4C584D42FFB08446FEC3FF7FFE39B9E0" box="[325,480,223,248]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB08446FEC3FF7FFE39B9E0" box="[325,480,223,248]" italics="true" pageId="59" pageNumber="60">Argyrophorus</emphasis>
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</taxonomicName>
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, but noted its unusual venation, the absence of an M1-M2 ventral forewing ocellus, and antennae only partially covered in scales as features that set it apart: in other words, it is very little like
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<taxonomicName id="4C584D42FFB08446FF11FE87FEEBB858" box="[151,306,295,320]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB08446FF11FE87FEEBB858" box="[151,306,295,320]" italics="true" pageId="59" pageNumber="60">Argyrophorus</emphasis>
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</taxonomicName>
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at all.
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<bibRefCitation id="EFC94B30FFB08446FE0AFE87FDF1B858" author="Miller" box="[396,552,295,320]" pageId="59" pageNumber="60" refString="Miller, L. D. (1968) The higher classification, phylogeny and zoogeography of the Satyridae (Lepidoptera). Memoirs of the American Entomological Society, 24, 1 - 174. http: // dx. doi. org / 10.1130 / MEM 110 - p 1" type="journal article" year="1968">Miller (1968)</bibRefCitation>
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also noted, "Until these structures, as well as others not considered in the original description, have been studied, it will be quite impossible to assign this genus to its proper position among the
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<taxonomicName id="4C584D42FFB08446FF40FECFFEEFB890" box="[198,310,367,392]" class="Insecta" family="Nymphalidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="family">Satyridae</taxonomicName>
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." Molecular data confirm what morphology suggests: that
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<taxonomicName id="4C584D42FFB08446FC77FED1FB02B890" box="[1009,1243,369,392]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="poaoeneis">
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<emphasis id="B92CEAD3FFB08446FC77FED1FB02B890" box="[1009,1243,369,392]" italics="true" pageId="59" pageNumber="60">Pamperis poaoeneis</emphasis>
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</taxonomicName>
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is distinct from
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<taxonomicName id="4C584D42FFB08446FF11FE34FEEBB8B5" box="[151,306,404,429]" class="Insecta" family="Nymphalidae" genus="Argyrophorus" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB08446FF11FE34FEEBB8B5" box="[151,306,404,429]" italics="true" pageId="59" pageNumber="60">Argyrophorus</emphasis>
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</taxonomicName>
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or any other taxon.
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<taxonomicName id="4C584D42FFB08446FD95FE35FDA7B8B4" box="[531,638,405,428]" class="Insecta" family="Nymphalidae" genus="Pamperis" kingdom="Animalia" order="Lepidoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B92CEAD3FFB08446FD95FE35FDA7B8B4" box="[531,638,405,428]" italics="true" pageId="59" pageNumber="60">Pamperis</emphasis>
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</taxonomicName>
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, then, remains a valid monotypic genus.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |