treatments-xml/data/03/EC/95/03EC95531F046B00FF01F896FCED5406.xml

<document id="97517922826BA66062EAE807B352487E" ID-DOI="10.11646/zootaxa.5374.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10159019" ID-ZooBank="72C9EC0B-F3D1-48F1-9A4C-C9F887C45327" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1700471180442" checkinUser="plazi" docAuthor="Santos, Laisa S., Roseno, Rafaella S., Solé, Mirco &amp; Dias, Iuri Ribeiro" docDate="2023" docId="03EC95531F046B00FF01F896FCED5406" docLanguage="en" docName="zootaxa.5374.4.4.pdf" docOrigin="Zootaxa 5374 (4)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.4/52304" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllodytes iuna Santos &amp; Roseno &amp; Solé &amp; Dias 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="526" masterDocId="FFD5ED2B1F056B07FF96FFC0FFA15112" masterDocTitle="Another new species (and it’s not over yet) of Phyllodytes Wagler, 1930 (Anura, Hylidae) from the Atlantic Forest of southern Bahia, northeastern Brazil" masterLastPageNumber="532" masterPageNumber="519" pageNumber="520" updateTime="1701116728914" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D5B040471F838BABCAFD319F290FB7CC">Another new species (and it’s not over yet) of Phyllodytes Wagler, 1930 (Anura, Hylidae) from the Atlantic Forest of southern Bahia, northeastern Brazil</mods:title>
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<mods:namePart id="66A91BFC0607F7C08C16FC88DB7DC249">Santos, Laisa S.</mods:namePart>
<mods:affiliation id="F3DCA911D6EB3E4ABE79F5F19ECCCC5C">Programa de Pós-Graduação em Zoologia, Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brasil</mods:affiliation>
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<mods:namePart id="621F855F26853FDB6B90A14D2E11A012">Roseno, Rafaella S.</mods:namePart>
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<mods:affiliation id="F909E553F2D7A3F2CD3E9B3DF8AE707B">Programa de Pós-Graduação em Zoologia, Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brasil &amp; raf. iroseno 96 @ gmail. com; https: // orcid. org / 0000 - 0002 - 8503 - 5530</mods:affiliation>
<mods:nameIdentifier id="B9D60F2BBB7F6440B60F151008A285B3" type="email">raf.iroseno96@gmail.com</mods:nameIdentifier>
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<mods:namePart id="88D2034E61F1FF6B12AF0FC40AF476FB">Solé, Mirco</mods:namePart>
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<mods:affiliation id="C30C76B06921593ECAD9C398357D9456">Programa de Pós-Graduação em Zoologia, Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, Ilhéus, Bahia, Brasil &amp; Herpetology Section, Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D- 53113 Bonn, Germany msole @ uesc. br; https: // orcid. org / 0000 - 0001 - 7881 - 6227</mods:affiliation>
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<mods:namePart id="DEC1A9158A0AE22EFEBA3FED3667EB24">Dias, Iuri Ribeiro</mods:namePart>
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<mods:date id="59FF053C76B23395F9E2DD33697F17AE">2023</mods:date>
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<heading id="D0B293291F046B06FF01F896FE645663" bold="true" box="[151,453,1878,1905]" fontSize="11" level="1" pageId="1" pageNumber="520" reason="1">
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<taxonomicName id="4C455FC61F046B06FF01F896FEC15663" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[151,352,1878,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="1" pageNumber="520" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F046B06FF01F896FEC15663" bold="true" box="[151,352,1878,1905]" italics="true" pageId="1" pageNumber="520">Phyllodytes iuna</emphasis>
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<taxonomicNameLabel id="A202452C1F046B06FEFEF897FE645663" box="[360,453,1879,1905]" pageId="1" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BFA24451F046B06FF01F85BFDAE56A1" blockId="1.[151,527,1947,1971]" box="[151,527,1947,1971]" pageId="1" pageNumber="520">
<taxonomicName id="4C455FC61F046B06FF01F85BFEE056A1" box="[151,321,1947,1971]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="1" pageNumber="520" phylum="Chordata" rank="species" species="undefined-3">
<emphasis id="B931F8571F046B06FF01F85BFEAC56A1" box="[151,269,1947,1971]" italics="true" pageId="1" pageNumber="520">Phyllodytes</emphasis>
sp. 3
</taxonomicName>
— 
<bibRefCitation id="EFD459B41F046B06FECAF85BFDAE56A1" author="Blotto, B. L. &amp; Lyra, M. L. &amp; Cardoso, M. C. S. &amp; Rodrigues, M. T. &amp; Dias, I. R. &amp; Marciano-Jr, E. &amp; Vechio, F. D. &amp; Orrico, V. G. D. &amp; Assis, C. L. &amp; Lantyer-Silva, A. S. F. &amp; Rutherford, M. G. &amp; Gagliardi-Urrutia, G. &amp; Sole, M. &amp; Baldo, D. &amp; Nunes, I. &amp; Cajade, R. &amp; Torres, A. &amp; Grant, T. &amp; Jungfer, K. H. &amp; Silva, H. R. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[348,527,1947,1971]" pageId="1" pageNumber="520" pagination="36 - 72" refId="ref6375" refString="Blotto, B. L., Lyra, M. L., Cardoso, M. C. S., Rodrigues, M. T., Dias, I. R., Marciano-Jr, E., Vechio, F. D., Orrico, V. G. D., Brand &quot; o, R. A., Assis, C. L., Lantyer-Silva, A. S. F., Rutherford, M. G., Gagliardi-Urrutia, G., Sole, M., Baldo, D., Nunes, I., Cajade, R., Torres, A., Grant, T., Jungfer, K. H., Silva, H. R., Haddad, C. F. B. &amp; Faivovich, J. (2020) The phylogeny of the Casqueheaded Treefrogs (Hylidae: Hylinae: Lophyohylini). Cladistics, 37 (1), 36 - 72. https: // doi. org / 10.1111 / cla. 12409" type="journal article" year="2020">
Blotto 
<emphasis id="B931F8571F046B06FE35F85CFE7756A1" box="[419,470,1947,1971]" italics="true" pageId="1" pageNumber="520">et al.</emphasis>
2020
</bibRefCitation>
</paragraph>
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<emphasis id="B931F8571F076B05FF01FF57FEA551A3" bold="true" box="[151,260,151,177]" pageId="2" pageNumber="521">
<typeStatus id="54FE9AE71F076B05FF01FF57FEA551A3" box="[151,260,151,177]" pageId="2" pageNumber="521" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<specimenCode id="DBE38C3E1F076B05FE83FF57FE7B51A3" box="[277,474,150,177]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="521">MZUESC 18950</specimenCode>
, an adult male (
<figureCitation id="137E38C01F076B05FD01FF57FD4651A3" box="[663,743,151,177]" captionStart="FIGURE 1" captionStartId="3.[152,255,1541,1565]" captionTargetBox="[170,1417,813,1517]" captionTargetId="figure-305@3.[170,1417,813,1517]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Phyllodytes iuna sp. nov. (MZUESC 18950). (A) Dorsal and (B) ventral view of the body. Scale bar = 10 mm" figureDoi="http://doi.org/10.5281/zenodo.10159023" httpUri="https://zenodo.org/record/10159023/files/figure.png" pageId="2" pageNumber="521">Figs. 1</figureCitation>
and 
<figureCitation id="137E38C01F076B05FCB5FF57FC9351A3" box="[803,818,151,177]" captionStart="FIGURE 2" captionStartId="4.[152,255,1357,1381]" captionTargetBox="[170,1416,309,1334]" captionTargetId="figure-92@4.[170,1416,309,1334]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Phyllodytes iuna sp. nov. (MZUESC 18950). (A) Dorsal view of head; (B) Lateral view of head (right side); (C) Palmar view of left hand; (D) Plantar view of left foot. Scale bar = 3 mm" figureDoi="http://doi.org/10.5281/zenodo.10159025" httpUri="https://zenodo.org/record/10159025/files/figure.png" pageId="2" pageNumber="521">2</figureCitation>
) from 
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(
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, 
<geoCoordinate id="EE7142821F076B05FE99FF7BFE2551C7" box="[271,388,187,213]" degrees="39.71880" direction="west" orientation="longitude" pageId="2" pageNumber="521" precision="1" value="-39.7188">-39.71880</geoCoordinate>
, 
<elevation id="0068C3761F076B05FE19FF7AFDB251C7" box="[399,531,186,213]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="2" pageNumber="521" unit="m" value="675.0">
<quantity id="4CBD89A01F076B05FE19FF7AFE7651C7" box="[399,471,186,213]" metricMagnitude="2" metricUnit="m" metricValue="6.75" pageId="2" pageNumber="521" unit="m" value="675.0">675 m</quantity>
a.s.l.
</elevation>
), municipality of 
<collectingMunicipality id="6B9EBE3F1F076B05FD4BFF7BFC7B51C7" box="[733,986,187,213]" pageId="2" pageNumber="521">Wenceslau Guimarães</collectingMunicipality>
, 
<collectingRegion id="4981EAA71F076B05FC73FF7BFB8851C7" box="[997,1065,187,213]" country="Brazil" name="Bahia" pageId="2" pageNumber="521">Bahia</collectingRegion>
, 
<collectingCountry id="F35264D51F076B05FBA3FF7BFBDD51C7" box="[1077,1148,187,213]" name="Brazil" pageId="2" pageNumber="521">Brazil</collectingCountry>
, collected by 
<collectorName id="26B041931F076B05FA8EFF7BFA3D51C7" box="[1304,1436,187,213]" pageId="2" pageNumber="521">Iuri R. Dias</collectorName>
and 
<collectorName id="26B041931F076B05FF51FF1FFE3E51EB" box="[199,415,223,249]" pageId="2" pageNumber="521">Rafael O. de Abreu</collectorName>
on 
<date id="FFFB02851F076B05FE5FFF1FFDF351EB" box="[457,594,223,249]" pageId="2" pageNumber="521" value="2003-02">
<collectingDate id="EFBFFB6D1F076B05FE5FFF1FFDF351EB" box="[457,594,223,249]" pageId="2" pageNumber="521" value="2003-02">03 February</collectingDate>
</date>
, 2015.
</materialsCitation>
</paragraph>
<paragraph id="8BFA24451F076B05FF51FEC3FA215053" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<materialsCitation id="3B2D2E181F076B05FF51FEC3FADD5053" collectionCode="MZUESC" collectorName="Marcos F. Vila Nova" location="Vila Nova in May" pageId="2" pageNumber="521" specimenCode="MZUESC18952" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" typeStatus="paratopotype">
<emphasis id="B931F8571F076B05FF51FEC3FED5500F" bold="true" box="[199,372,259,285]" pageId="2" pageNumber="521">
<typeStatus id="54FE9AE71F076B05FF51FEC3FED5500F" box="[199,372,259,285]" pageId="2" pageNumber="521" type="paratopotype">Paratopotypes</typeStatus>
</emphasis>
. 
<specimenCode id="DBE38C3E1F076B05FE15FEC2FD9C500F" box="[387,573,258,285]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="521">MZUESC18952</specimenCode>
, 
<specimenCount id="9D43EFCC1F076B05FDDCFEC3FD23500F" box="[586,642,259,285]" pageId="2" pageNumber="521" type="adult">adult</specimenCount>
<specimenCount id="9D43EFCC1F076B05FD1CFEC3FD61500F" box="[650,704,259,285]" pageId="2" pageNumber="521" type="male">male</specimenCount>
collected with the holotype; 
<specimenCode id="DBE38C3E1F076B05FB9BFEC3FB70500F" box="[1037,1233,259,285]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="521">MZUESC 23000</specimenCode>
; 23002, 
<specimenCount id="9D43EFCC1F076B05FAA4FEC3FA3D500F" box="[1330,1436,259,285]" count="2" pageId="2" pageNumber="521" type="adult">two adult</specimenCount>
males, and 
<specimenCode id="DBE38C3E1F076B05FE80FEE7FE795053" box="[278,472,295,321]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="2" pageNumber="521">MZUESC 23001</specimenCode>
; 23003, 
<specimenCount id="9D43EFCC1F076B05FDA1FEE7FD015053" box="[567,672,295,321]" count="2" pageId="2" pageNumber="521" type="adult">two adult</specimenCount>
females, collected by 
<collectorName id="26B041931F076B05FC0DFEE7FB255053" box="[923,1156,295,321]" pageId="2" pageNumber="521">Marcos F. Vila Nova</collectorName>
in May, 2018 (
<figureCitation id="137E38C01F076B05FAB8FEE7FAD35053" box="[1326,1394,295,321]" captionStart="FIGURE 3" captionStartId="5.[152,255,655,679]" captionTargetBox="[151,1435,181,631]" captionTargetId="figure-19@5.[151,1435,181,631]" captionTargetPageId="5" captionText="FIGURE 3. Paratopotypes of Phyllodytes iuna sp. nov. MZUESC 18952 (A and F); MZUESC 23000 (B and G); MZUESC 23001 (C and H); MZUESC 23002 (D and I); MZUESC 23003 (E and J). Scale bar = 10 m" figureDoi="http://doi.org/10.5281/zenodo.10159027" httpUri="https://zenodo.org/record/10159027/files/figure.png" pageId="2" pageNumber="521">Fig. 3</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFA24451F076B05FF51FE8AFD2E50BF" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<emphasis id="B931F8571F076B05FF51FE8AFE0E5077" bold="true" box="[199,431,330,357]" pageId="2" pageNumber="521">Generic Placement.</emphasis>
The new species is assigned to the genus 
<taxonomicName id="4C455FC61F076B05FCEFFE8BFC5D5077" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[889,1020,331,357]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="genus">
<emphasis id="B931F8571F076B05FCEFFE8BFC5D5077" box="[889,1020,331,357]" italics="true" pageId="2" pageNumber="521">Phyllodytes</emphasis>
</taxonomicName>
based on the occurrence of odontoids in the lower jaw. In addition, molecular data support the placement of 
<taxonomicName id="4C455FC61F076B05FC2DFEAFFB9F509B" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[955,1086,367,393]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="B931F8571F076B05FC2DFEAFFB9F509B" box="[955,1086,367,393]" italics="true" pageId="2" pageNumber="521">Phyllodytes</emphasis>
</taxonomicName>
<emphasis id="B931F8571F076B05FBD0FEAFFB00509B" bold="true" box="[1094,1185,367,393]" pageId="2" pageNumber="521">
<taxonomicNameLabel id="A202452C1F076B05FBD0FEAFFB00509B" box="[1094,1185,367,393]" pageId="2" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as a sister taxon of 
<taxonomicName id="4C455FC61F076B05FA1EFEAFFEB950BF" authorityName="Peixoto &amp; Cruz" authorityYear="1988" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B931F8571F076B05FA1EFEAFFEB950BF" italics="true" pageId="2" pageNumber="521">P. brevirostris</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C455FC61F076B05FEA0FE53FD2A50BF" authority="(Blotto et al. 2020)" baseAuthorityName="Blotto" baseAuthorityYear="2020" box="[310,651,403,429]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B931F8571F076B05FEA0FE53FE0B50BF" box="[310,426,403,429]" italics="true" pageId="2" pageNumber="521">P. edelmoi</emphasis>
(
<bibRefCitation id="EFD459B41F076B05FE2EFE53FD2250BF" author="Blotto, B. L. &amp; Lyra, M. L. &amp; Cardoso, M. C. S. &amp; Rodrigues, M. T. &amp; Dias, I. R. &amp; Marciano-Jr, E. &amp; Vechio, F. D. &amp; Orrico, V. G. D. &amp; Assis, C. L. &amp; Lantyer-Silva, A. S. F. &amp; Rutherford, M. G. &amp; Gagliardi-Urrutia, G. &amp; Sole, M. &amp; Baldo, D. &amp; Nunes, I. &amp; Cajade, R. &amp; Torres, A. &amp; Grant, T. &amp; Jungfer, K. H. &amp; Silva, H. R. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[440,643,403,429]" pageId="2" pageNumber="521" pagination="36 - 72" refId="ref6375" refString="Blotto, B. L., Lyra, M. L., Cardoso, M. C. S., Rodrigues, M. T., Dias, I. R., Marciano-Jr, E., Vechio, F. D., Orrico, V. G. D., Brand &quot; o, R. A., Assis, C. L., Lantyer-Silva, A. S. F., Rutherford, M. G., Gagliardi-Urrutia, G., Sole, M., Baldo, D., Nunes, I., Cajade, R., Torres, A., Grant, T., Jungfer, K. H., Silva, H. R., Haddad, C. F. B. &amp; Faivovich, J. (2020) The phylogeny of the Casqueheaded Treefrogs (Hylidae: Hylinae: Lophyohylini). Cladistics, 37 (1), 36 - 72. https: // doi. org / 10.1111 / cla. 12409" type="journal article" year="2020">
Blotto 
<emphasis id="B931F8571F076B05FD91FE53FD9950BF" box="[519,568,403,429]" italics="true" pageId="2" pageNumber="521">et al</emphasis>
. 2020
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F77CE1F076B05FF51FE77FBFB55FB" pageId="2" pageNumber="521" type="etymology">
<paragraph id="8BFA24451F076B05FF51FE77FE525207" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<emphasis id="B931F8571F076B05FF51FE77FEE950C3" bold="true" box="[199,328,439,465]" pageId="2" pageNumber="521">Etymology</emphasis>
. The specific epithet 
<taxonomicName id="4C455FC61F076B05FDD2FE77FD2350C3" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[580,642,439,465]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">“iuna</taxonomicName>
” alludes to a berimbau riff (a musical percussion instrument) played during a Capoeira bout (called “jogo”; “game”). The ‘iuna’ riff represents a rite of passage in Capoeira, marking the moment of transition from student to teacher. When this riff is being played, the game is reserved for teachers and masters. It is also played during posthumous homages to honor the skill of the deceased Capoeira player (capoeirista). Capoeira is an Afro-Brazilian cultural expression, developed by enslaved Africans in 
<collectingCountry id="F35264D51F076B05FB77FD87FA845373" box="[1249,1317,583,609]" name="Brazil" pageId="2" pageNumber="521">Brazil</collectingCountry>
during the colonial era as a means of self-expression, resistance, and community-building. It is recognized as Cultural Heritage of 
<collectingCountry id="F35264D51F076B05FF20FD4FFF5B53BB" box="[182,250,655,681]" name="Brazil" pageId="2" pageNumber="521">Brazil</collectingCountry>
and Intangible Cultural Heritage of Humanity by UNESCO, symbolizing the black resistance during the slavery period in the country. By adopting the name 
<taxonomicName id="4C455FC61F076B05FD6BFD73FC9A53DF" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[765,827,691,717]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">“iuna</taxonomicName>
” to honor Capoeira, we are also celebrating the rich Afro-Brazilian heritage and acknowledging the legacy left by Capoeira masters in the resistance, consolidation, and perpetuation of this martial art.
</paragraph>
<paragraph id="8BFA24451F076B05FF51FCDEFBFB55FB" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<emphasis id="B931F8571F076B05FF51FCDEFE3B522A" bold="true" box="[199,410,798,824]" pageId="2" pageNumber="521">Characterization.</emphasis>
(1) Medium sized adult males, SVL in males 
<quantity id="4CBD89A01F076B05FC36FCDEFBE2522B" box="[928,1091,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" metricValueMax="2.52" metricValueMin="2.2199999999999998" pageId="2" pageNumber="521" unit="mm" value="23.7" valueMax="25.2" valueMin="22.2">22.2–25.2 mm</quantity>
(n = 4); SVL in females 24.0– 
<quantity id="4CBD89A01F076B05FF01FC82FF5D524F" box="[151,252,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="2" pageNumber="521" unit="mm" value="25.9">25.9 mm</quantity>
(n = 2); (2) slender body; (3) head as long as wide (HL 98.7 % of HW); (4) truncated snout with apical tubercle in dorsal view and protuberant in lateral view; (5) nostrils with rounded contours positioned close to the tip of the snout, directed dorsolaterally; (6) 
<emphasis id="B931F8571F076B05FD1AFC4BFCEB52B7" box="[652,842,907,933]" italics="true" pageId="2" pageNumber="521">canthus rostralis</emphasis>
rounded; (7) loreal region concave and straight; (8) tympanum evident; (9) tympanic ring evident; (10) well-marked supratympanic fold from the back of the eyes to the insertion of the arm; (11) prominent eyes, positioned anterolaterally; (12) single sub-gular vocal sac; (13) horizontal pupil; (14) two anterior large, and additional smaller (2–6) subequal odontoids on each side of the mandible; (15) lateral margin of forearms with inconspicuous outer tubercles; (16) palmar tubercle developed and elongated; (17) single and prominent tubercle near the tibiotarsal articulation; (18) brownish yellow coloration; (19) dorsum of body exhibits a coloration with a diffuse band-like pattern from the interocular region to the posterior region of the body; (20) dorsum of arms and legs with small and irregular brown blotches; (21) dark strip from the tip of the snout to the middle of the body; (22) smooth dorsal skin; (23) uniform cream ventral coloration; (24) ventral surface with two rows of central tubercles, with adjacent tubercles bordering without a specific pattern.
</paragraph>
</subSubSection>
<subSubSection id="C35F77CE1F076B00FF51FB32FCED5406" lastPageId="7" lastPageNumber="526" pageId="2" pageNumber="521" type="description">
<paragraph id="8BFA24451F076B05FF51FB32FB9C54F7" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<emphasis id="B931F8571F076B05FF51FB32FD9B541E" bold="true" box="[199,570,1266,1293]" pageId="2" pageNumber="521">Comparison with other species</emphasis>
. Character states for the other species are shown in parenthesis. 
<taxonomicName id="4C455FC61F076B05FA8CFB33FF685422" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FA8CFB33FF685422" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is recovered as the sister taxon of 
<taxonomicName id="4C455FC61F076B05FDCDFAD7FD5B5423" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[603,762,1303,1329]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B931F8571F076B05FDCDFAD7FD5B5423" box="[603,762,1303,1329]" italics="true" pageId="2" pageNumber="521">P. brevirostris</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C455FC61F076B05FC8DFAD7FB205423" authority="(Blotto et al., 2020)" baseAuthorityName="Blotto" baseAuthorityYear="2020" box="[795,1153,1303,1329]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B931F8571F076B05FC8DFAD7FC335423" box="[795,914,1303,1329]" italics="true" pageId="2" pageNumber="521">P. edelmoi</emphasis>
(
<bibRefCitation id="EFD459B41F076B05FC34FAD7FBD85423" author="Blotto, B. L. &amp; Lyra, M. L. &amp; Cardoso, M. C. S. &amp; Rodrigues, M. T. &amp; Dias, I. R. &amp; Marciano-Jr, E. &amp; Vechio, F. D. &amp; Orrico, V. G. D. &amp; Assis, C. L. &amp; Lantyer-Silva, A. S. F. &amp; Rutherford, M. G. &amp; Gagliardi-Urrutia, G. &amp; Sole, M. &amp; Baldo, D. &amp; Nunes, I. &amp; Cajade, R. &amp; Torres, A. &amp; Grant, T. &amp; Jungfer, K. H. &amp; Silva, H. R. &amp; Haddad, C. F. B. &amp; Faivovich, J." box="[930,1145,1303,1329]" pageId="2" pageNumber="521" pagination="36 - 72" refId="ref6375" refString="Blotto, B. L., Lyra, M. L., Cardoso, M. C. S., Rodrigues, M. T., Dias, I. R., Marciano-Jr, E., Vechio, F. D., Orrico, V. G. D., Brand &quot; o, R. A., Assis, C. L., Lantyer-Silva, A. S. F., Rutherford, M. G., Gagliardi-Urrutia, G., Sole, M., Baldo, D., Nunes, I., Cajade, R., Torres, A., Grant, T., Jungfer, K. H., Silva, H. R., Haddad, C. F. B. &amp; Faivovich, J. (2020) The phylogeny of the Casqueheaded Treefrogs (Hylidae: Hylinae: Lophyohylini). Cladistics, 37 (1), 36 - 72. https: // doi. org / 10.1111 / cla. 12409" type="journal article" year="2020">
Blotto 
<emphasis id="B931F8571F076B05FC64FAD7FB845423" box="[1010,1061,1303,1329]" italics="true" pageId="2" pageNumber="521">et al</emphasis>
., 2020
</bibRefCitation>
)
</taxonomicName>
. 
<taxonomicName id="4C455FC61F076B05FB1BFAD7FAEA5422" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1165,1355,1303,1329]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FB1BFAD7FAEA5422" box="[1165,1355,1303,1329]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
differs from them by having a head that is as wide as it is long, with HL comprising 98.7% of HW (wider than longer; HL 84.0% of HW), truncated snout in dorsal view and protruding in lateral view (rounded snout in dorsal and lateral view), presence of an apical tubercle on the snout (absent), dark stripe between the tip of the snout and the eye, on the 
<emphasis id="B931F8571F076B05FF54FA67FEDE54D3" box="[194,383,1447,1473]" italics="true" pageId="2" pageNumber="521">canthus rostralis</emphasis>
(absent), dorsal coloration with diffuse dark stripes/blotches (immaculate pattern) and single tubercle near the tibiotarsal articulation (several tubercles along the tarsus) (
<figureCitation id="137E38C01F076B05FC7CFA0BFB9154F7" box="[1002,1072,1483,1509]" captionStart="FIGURE 4" captionStartId="5.[152,255,1889,1913]" captionTargetBox="[170,1417,874,1865]" captionTargetId="figure-110@5.[170,1417,874,1865]" captionTargetPageId="5" captionText="FIGURE 4. Variation pattern of the dorsal coloration of P. iuna sp. nov. and its sister clade P. edelmoi and P. brevirostris. Paratopotypes of Phyllodytes iuna sp. nov. (MZUESC 18952, A and MZUESC 23001, B, Photos by Rafael O. de Abreu and Marcos Vila Nova, respectively); Phyllodytes edelmoi (C, Reserva Biológica de Saltinho, Tamandaré—PE, Photo by Pedro Ivo Simıes) and Phyllodytes brevirostris (D, Cruz do Espírito Santo—PB, Photo by Washington Luiz S. Vieira)." figureDoi="http://doi.org/10.5281/zenodo.10159029" httpUri="https://zenodo.org/record/10159029/files/figure.png" pageId="2" pageNumber="521">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BFA24451F076B05FF51FA2FFEC55687" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="521">
<taxonomicName id="4C455FC61F076B05FF51FA2FFE29571A" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[199,392,1519,1545]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FF51FA2FFE29571A" box="[199,392,1519,1545]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C455FC61F076B05FD0EFA2FFCF8571B" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[664,857,1519,1545]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F076B05FD0EFA2FFCF8571B" box="[664,857,1519,1545]" italics="true" pageId="2" pageNumber="521">P. melanomystax</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FCFAFA2FFB9F571A" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[876,1086,1519,1544]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F076B05FCFAFA2FFB9F571A" box="[876,1086,1519,1544]" italics="true" pageId="2" pageNumber="521">P. megatympanum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FBC4FA2FFB6B571A" box="[1106,1226,1519,1544]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F076B05FBC4FA2FFB6B571A" box="[1106,1226,1519,1544]" italics="true" pageId="2" pageNumber="521">P. magnus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FB4BFA2FFAD5571B" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[1245,1396,1519,1545]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F076B05FB4BFA2FFAD5571B" box="[1245,1396,1519,1545]" italics="true" pageId="2" pageNumber="521">P. maculosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FA1EFA2FFF51573F" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B931F8571F076B05FA1EFA2FFF51573F" italics="true" pageId="2" pageNumber="521">P. luteolus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FE96F9D3FE00573F" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[256,417,1555,1581]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F076B05FE96F9D3FE00573F" box="[256,417,1555,1581]" italics="true" pageId="2" pageNumber="521">P. gyrinaethes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FE27F9D3FD8A573F" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[433,555,1555,1581]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F076B05FE27F9D3FD8A573F" box="[433,555,1555,1581]" italics="true" pageId="2" pageNumber="521">P. kautskyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F076B05FDF0F9D3FD79573F" authorityName="Voros, Dias &amp; Sole" authorityYear="2017" box="[614,728,1555,1581]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B931F8571F076B05FDF0F9D3FD79573F" box="[614,728,1555,1581]" italics="true" pageId="2" pageNumber="521">P. amadoi</emphasis>
</taxonomicName>
by having a head as long as it is wide (wider than long); for having a medium body size (
<quantity id="4CBD89A01F076B05FE42F9F6FDD95743" box="[468,632,1590,1617]" metricMagnitude="-2" metricUnit="m" metricValue="2.405" metricValueMax="2.59" metricValueMin="2.2199999999999998" pageId="2" pageNumber="521" unit="mm" value="24.049999999999997" valueMax="25.9" valueMin="22.2">22.2–25.9 mm</quantity>
SVL), 
<taxonomicName id="4C455FC61F076B05FD5FF9F7FC255742" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[713,900,1591,1617]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FD5FF9F7FC255742" box="[713,900,1591,1617]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C455FC61F076B05FBECF9F7FB4A5742" box="[1146,1259,1591,1616]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F076B05FBECF9F7FB4A5742" box="[1146,1259,1591,1616]" italics="true" pageId="2" pageNumber="521">P. magnus</emphasis>
</taxonomicName>
(
<quantity id="4CBD89A01F076B05FB6FF9F7FA3C5743" box="[1273,1437,1591,1617]" metricMagnitude="-2" metricUnit="m" metricValue="3.875" metricValueMax="4.11" metricValueMin="3.6399999999999997" pageId="2" pageNumber="521" unit="mm" value="38.75" valueMax="41.1" valueMin="36.4">36.4–41.1 mm</quantity>
SVL), 
<taxonomicName id="4C455FC61F076B05FF75F99BFEFA5767" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[227,347,1627,1653]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F076B05FF75F99BFEFA5767" box="[227,347,1627,1653]" italics="true" pageId="2" pageNumber="521">P. kautskyi</emphasis>
</taxonomicName>
(
<quantity id="4CBD89A01F076B05FEFFF99AFDAF5767" box="[361,526,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.35" metricValueMin="3.65" pageId="2" pageNumber="521" unit="mm" value="40.0" valueMax="43.5" valueMin="36.5">36.5–43.5 mm</quantity>
SVL) and 
<taxonomicName id="4C455FC61F076B05FD1FF99BFCBA5767" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[649,795,1627,1653]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F076B05FD1FF99BFCBA5767" box="[649,795,1627,1653]" italics="true" pageId="2" pageNumber="521">P. maculosus</emphasis>
</taxonomicName>
(
<quantity id="4CBD89A01F076B05FCBFF99AFC6F5767" box="[809,974,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.35" metricValueMin="3.95" pageId="2" pageNumber="521" unit="mm" value="41.5" valueMax="43.5" valueMin="39.5">39.5–43.5 mm</quantity>
SVL); the truncated snout in dorsal view distinguishes 
<taxonomicName id="4C455FC61F076B05FEA6F9BFFE4B578A" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[304,490,1663,1689]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FEA6F9BFFE4B578A" box="[304,490,1663,1689]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C455FC61F076B05FDBBF9BFFD3A578B" authorityName="Voros, Dias &amp; Sole" authorityYear="2017" box="[557,667,1663,1689]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B931F8571F076B05FDBBF9BFFD3A578B" box="[557,667,1663,1689]" italics="true" pageId="2" pageNumber="521">P. amadoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FD3EF9BFFCE7578B" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[680,838,1663,1689]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F076B05FD3EF9BFFCE7578B" box="[680,838,1663,1689]" italics="true" pageId="2" pageNumber="521">P. gyrinaethes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FCC5F9BFFC42578B" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[851,995,1663,1689]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F076B05FCC5F9BFFC42578B" box="[851,995,1663,1689]" italics="true" pageId="2" pageNumber="521">P. maculosus</emphasis>
</taxonomicName>
(rounded) and from 
<taxonomicName id="4C455FC61F076B05FB5FF9BFFA34578A" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[1225,1429,1663,1688]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F076B05FB5FF9BFFA34578A" box="[1225,1429,1663,1688]" italics="true" pageId="2" pageNumber="521">P. megatympanum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FF01F963FEAE57AF" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[151,271,1699,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F076B05FF01F963FEAE57AF" box="[151,271,1699,1725]" italics="true" pageId="2" pageNumber="521">P. kautskyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F076B05FED3F963FE4357AE" authorityName="Bokermann" authorityYear="1966" box="[325,482,1699,1724]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B931F8571F076B05FED3F963FE4357AE" box="[325,482,1699,1724]" italics="true" pageId="2" pageNumber="521">P. acuminatus</emphasis>
</taxonomicName>
(acute); in lateral view, the snout of 
<taxonomicName id="4C455FC61F076B05FCE8F963FB9B57AE" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[894,1082,1699,1725]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FCE8F963FB9B57AE" box="[894,1082,1699,1725]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is protuberant, which separates it from 
<taxonomicName id="4C455FC61F076B05FF7CF907FEF957F3" authorityName="Voros, Dias &amp; Sole" authorityYear="2017" box="[234,344,1735,1761]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B931F8571F076B05FF7CF907FEF957F3" box="[234,344,1735,1761]" italics="true" pageId="2" pageNumber="521">P. amadoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F076B05FEF3F907FE5757F3" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[357,502,1735,1761]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F076B05FEF3F907FE5757F3" box="[357,502,1735,1761]" italics="true" pageId="2" pageNumber="521">P. maculosus</emphasis>
</taxonomicName>
(vertical) and 
<taxonomicName id="4C455FC61F076B05FD0EF907FCF357F3" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[664,850,1735,1761]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F076B05FD0EF907FCF357F3" box="[664,850,1735,1761]" italics="true" pageId="2" pageNumber="521">P. melanomystax</emphasis>
</taxonomicName>
(rounded); the presence of a tubercle at the tip of the snout separates 
<taxonomicName id="4C455FC61F076B05FEDDF92BFDA95616" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[331,520,1771,1797]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FEDDF92BFDA95616" box="[331,520,1771,1797]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C455FC61F076B05FDD9F92BFD465616" authorityName="Orrico, Dias &amp; Marciano" authorityYear="2018" box="[591,743,1771,1796]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="praeceptor">
<emphasis id="B931F8571F076B05FDD9F92BFD465616" box="[591,743,1771,1796]" italics="true" pageId="2" pageNumber="521">P. praeceptor</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F076B05FCB6F92BFC7C5617" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[800,989,1771,1797]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F076B05FCB6F92BFC7C5617" box="[800,989,1771,1797]" italics="true" pageId="2" pageNumber="521">P. melanomystax</emphasis>
</taxonomicName>
(absent); the tympanum of 
<taxonomicName id="4C455FC61F076B05FA8CF92BFF68563A" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FA8CF92BFF68563A" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is evident, distinguishing it from 
<taxonomicName id="4C455FC61F076B05FDD9F8CFFD51563B" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[591,752,1807,1833]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F076B05FDD9F8CFFD51563B" box="[591,752,1807,1833]" italics="true" pageId="2" pageNumber="521">P. gyrinaethes</emphasis>
</taxonomicName>
(hidden); the tympanum size of 
<taxonomicName id="4C455FC61F076B05FBFEF8CFFA87563A" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1128,1318,1807,1833]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FBFEF8CFFA87563A" box="[1128,1318,1807,1833]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
is smaller (&lt;7% of the SVL) compared to 
<taxonomicName id="4C455FC61F076B05FD87F8F3FD41565E" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[529,736,1843,1868]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F076B05FD87F8F3FD41565E" box="[529,736,1843,1868]" italics="true" pageId="2" pageNumber="521">P. megatympanum</emphasis>
</taxonomicName>
(7.73% of the SVL) and 
<taxonomicName id="4C455FC61F076B05FB9EF8F3FB06565E" authorityName="Bokermann" authorityYear="1966" box="[1032,1191,1843,1868]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B931F8571F076B05FB9EF8F3FB06565E" box="[1032,1191,1843,1868]" italics="true" pageId="2" pageNumber="521">P. acuminatus</emphasis>
</taxonomicName>
(8.16% of the SVL); 
<taxonomicName id="4C455FC61F076B05FF01F897FEF65662" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[151,343,1879,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FF01F897FEF65662" box="[151,343,1879,1905]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
has a well-marked supratympanic fold, distinguishing it from 
<taxonomicName id="4C455FC61F076B05FBA2F897FB775663" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[1076,1238,1879,1905]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F076B05FBA2F897FB775663" box="[1076,1238,1879,1905]" italics="true" pageId="2" pageNumber="521">P. gyrinaethes</emphasis>
</taxonomicName>
(discreet) and 
<taxonomicName id="4C455FC61F076B05FA1EF897FEA95687" baseAuthorityName="Peters" baseAuthorityYear="1873" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B931F8571F076B05FA1EF897FEA95687" italics="true" pageId="2" pageNumber="521">P. wuchereri</emphasis>
</taxonomicName>
(weak).
</paragraph>
<paragraph id="8BFA24451F076B04FF51F85FFB875077" blockId="2.[151,1437,150,2013]" lastBlockId="3.[151,1437,151,789]" lastPageId="3" lastPageNumber="522" pageId="2" pageNumber="521">
The presence of inconspicuous tubercles on the lateral margin of the forearm distinguishes 
<taxonomicName id="4C455FC61F076B05FB3EF85FFAC056AA" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1192,1377,1951,1977]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FB3EF85FFAC056AA" box="[1192,1377,1951,1977]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C455FC61F076B05FF01F803FE9756CE" authorityName="Bokermann" authorityYear="1966" box="[151,310,1987,2012]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B931F8571F076B05FF01F803FE9756CE" box="[151,310,1987,2012]" italics="true" pageId="2" pageNumber="521">P. acuminatus</emphasis>
</taxonomicName>
(one well-developed), 
<taxonomicName id="4C455FC61F076B05FDD6F803FD5C56CF" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[576,765,1987,2013]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F076B05FDD6F803FD5C56CF" box="[576,765,1987,2013]" italics="true" pageId="2" pageNumber="521">P. melanomystax</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F076B05FCAEF803FC4556CF" authorityName="Bokermann" authorityYear="1966" box="[824,996,1987,2013]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B931F8571F076B05FCAEF803FC4556CF" box="[824,996,1987,2013]" italics="true" pageId="2" pageNumber="521">P. tuberculosus</emphasis>
</taxonomicName>
(three very evident); 
<taxonomicName id="4C455FC61F076B05FB49F803FA3D56CE" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1247,1436,1987,2013]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="2" pageNumber="521" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F076B05FB49F803FA3D56CE" box="[1247,1436,1987,2013]" italics="true" pageId="2" pageNumber="521">Phyllodytes iuna</emphasis>
</taxonomicName>
has a round and simple subarticular tubercle, differentiating it from 
<taxonomicName id="4C455FC61F066B04FC0AFF57FB8E51A3" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[924,1071,151,177]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F066B04FC0AFF57FB8E51A3" box="[924,1071,151,177]" italics="true" pageId="3" pageNumber="522">P. maculosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F066B04FBFEFF57FB4051A3" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[1128,1249,151,177]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F066B04FBFEFF57FB4051A3" box="[1128,1249,151,177]" italics="true" pageId="3" pageNumber="522">P. kautskyi</emphasis>
</taxonomicName>
(bifid elongated tubercles); the palmar tubercle of 
<taxonomicName id="4C455FC61F066B04FDB7FF7BFD4151C6" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[545,736,187,213]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F066B04FDB7FF7BFD4151C6" box="[545,736,187,213]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
is developed and elongated diverging from 
<taxonomicName id="4C455FC61F066B04FB74FF7BFAD751C7" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[1250,1398,187,213]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F066B04FB74FF7BFAD751C7" box="[1250,1398,187,213]" italics="true" pageId="3" pageNumber="522">P. maculosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FA1EFF7BFEA951EB" baseAuthorityName="Peters" baseAuthorityYear="1873" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B931F8571F066B04FA1EFF7BFEA951EB" italics="true" pageId="3" pageNumber="522">P. wuchereri</emphasis>
</taxonomicName>
(ovoid), 
<taxonomicName id="4C455FC61F066B04FEE6FF1FFDAE51EB" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[368,527,223,249]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F066B04FEE6FF1FFDAE51EB" box="[368,527,223,249]" italics="true" pageId="3" pageNumber="522">P. gyrinaethes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FD8BFF1FFD3151EA" box="[541,656,223,248]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F066B04FD8BFF1FFD3151EA" box="[541,656,223,248]" italics="true" pageId="3" pageNumber="522">P. magnus</emphasis>
</taxonomicName>
(round), 
<taxonomicName id="4C455FC61F066B04FD6DFF1FFC6951EA" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[763,968,223,248]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F066B04FD6DFF1FFC6951EA" box="[763,968,223,248]" italics="true" pageId="3" pageNumber="522">P. megatympanum</emphasis>
</taxonomicName>
(triangular), 
<taxonomicName id="4C455FC61F066B04FBC8FF1FFABB51EB" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[1118,1306,223,249]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F066B04FBC8FF1FFABB51EB" box="[1118,1306,223,249]" italics="true" pageId="3" pageNumber="522">P. melanomystax</emphasis>
</taxonomicName>
(indistinct) and 
<taxonomicName id="4C455FC61F066B04FF52FEC3FEEB500E" authorityName="Caramaschi &amp; Peixoto" authorityYear="2004" box="[196,330,259,284]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B931F8571F066B04FF52FEC3FEEB500E" box="[196,330,259,284]" italics="true" pageId="3" pageNumber="522">P. punctatus</emphasis>
</taxonomicName>
(elliptical); 
<taxonomicName id="4C455FC61F066B04FE59FEC3FD29500E" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[463,648,259,285]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F066B04FE59FEC3FD29500E" box="[463,648,259,285]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
presents a single tubercle near the tibiotarsal articulation, differentiating it from 
<taxonomicName id="4C455FC61F066B04FF7BFEE7FE2B5052" authorityName="Bokermann" authorityYear="1966" box="[237,394,295,320]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B931F8571F066B04FF7BFEE7FE2B5052" box="[237,394,295,320]" italics="true" pageId="3" pageNumber="522">P. acuminatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FE0FFEE7FDB35053" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[409,530,295,321]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F066B04FE0FFEE7FDB35053" box="[409,530,295,321]" italics="true" pageId="3" pageNumber="522">P. kautskyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FDB6FEE7FD7C5053" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[544,733,295,321]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F066B04FDB6FEE7FD7C5053" box="[544,733,295,321]" italics="true" pageId="3" pageNumber="522">P. melanomystax</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FD7AFEE7FCC05053" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[748,865,295,321]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B931F8571F066B04FD7AFEE7FCC05053" box="[748,865,295,321]" italics="true" pageId="3" pageNumber="522">P. luteolus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FCE6FEE7FC425052" box="[880,995,295,320]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F066B04FCE6FEE7FC425052" box="[880,995,295,320]" italics="true" pageId="3" pageNumber="522">P. magnus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FC64FEE7FB1E5052" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[1010,1215,295,320]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F066B04FC64FEE7FB1E5052" box="[1010,1215,295,320]" italics="true" pageId="3" pageNumber="522">P. megatympanum</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FB58FEE7FAD85053" authorityName="Bokermann" authorityYear="1966" box="[1230,1401,295,321]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B931F8571F066B04FB58FEE7FAD85053" box="[1230,1401,295,321]" italics="true" pageId="3" pageNumber="522">P. tuberculosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FA1EFEE7FEA95077" baseAuthorityName="Peters" baseAuthorityYear="1873" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B931F8571F066B04FA1EFEE7FEA95077" italics="true" pageId="3" pageNumber="522">P. wuchereri</emphasis>
</taxonomicName>
(several tubercles along the tarsus), and 
<taxonomicName id="4C455FC61F066B04FD47FE8BFCCE5077" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[721,879,331,357]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F066B04FD47FE8BFCCE5077" box="[721,879,331,357]" italics="true" pageId="3" pageNumber="522">P. gyrinaethes</emphasis>
</taxonomicName>
(two tubercles).
</paragraph>
<paragraph id="8BFA24451F066B04FF51FEAFFED55397" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="522">
<taxonomicName id="4C455FC61F066B04FF51FEAFFE28509A" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[199,393,367,393]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F066B04FF51FEAFFE28509A" box="[199,393,367,393]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
has eye stripes extending from the tip of the snout to halfway down the body, which distinguishes it from 
<taxonomicName id="4C455FC61F066B04FE06FE53FDA650BE" box="[400,519,403,428]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F066B04FE06FE53FDA650BE" box="[400,519,403,428]" italics="true" pageId="3" pageNumber="522">P. magnus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F066B04FDD3FE53FD4950BF" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[581,744,403,429]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F066B04FDD3FE53FD4950BF" box="[581,744,403,429]" italics="true" pageId="3" pageNumber="522">P. gyrinaethes</emphasis>
</taxonomicName>
(stripes absent); the dorsum of 
<taxonomicName id="4C455FC61F066B04FBF4FE53FA8050BE" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1122,1313,403,429]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F066B04FBF4FE53FA8050BE" box="[1122,1313,403,429]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
exhibits a coloration forming a diffuse band-like pattern extending to the posterior region of the body, which differs from 
<taxonomicName id="4C455FC61F066B04FA1EFE77FF5250E7" authorityName="Peixoto &amp; Cruz" authorityYear="1988" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F066B04FA1EFE77FF5250E7" italics="true" pageId="3" pageNumber="522">P. kautskyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FE94FE1BFED450E6" box="[258,373,475,500]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F066B04FE94FE1BFED450E6" box="[258,373,475,500]" italics="true" pageId="3" pageNumber="522">P. magnus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F066B04FE38FE1BFDDD50E6" authorityName="Marciano, Lantyer-Silva &amp; Sole" authorityYear="2017" box="[430,636,475,500]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="megatympanum">
<emphasis id="B931F8571F066B04FE38FE1BFDDD50E6" box="[430,636,475,500]" italics="true" pageId="3" pageNumber="522">P. megatympanum</emphasis>
</taxonomicName>
(immaculate pattern), 
<emphasis id="B931F8571F066B04FCE8FE1BFACD50E7" box="[894,1388,475,501]" italics="true" pageId="3" pageNumber="522">
<taxonomicName id="4C455FC61F066B04FCE8FE1BFBBE50E6" authorityName="Bokermann" authorityYear="1966" box="[894,1055,475,500]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="acuminatus">P. acuminatus</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FBBDFE1BFB0350E7" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[1067,1186,475,501]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="luteolus">P. luteolus</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FB39FE1BFACD50E7" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[1199,1388,475,501]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="melanomystax">P. melanomystax</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C455FC61F066B04FF01FE3FFEE1530B" authorityName="Bokermann" authorityYear="1966" box="[151,320,511,537]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B931F8571F066B04FF01FE3FFEE1530B" box="[151,320,511,537]" italics="true" pageId="3" pageNumber="522">P. tuberculosus</emphasis>
</taxonomicName>
(immaculate pattern or with small spots), 
<emphasis id="B931F8571F066B04FC80FE3FFB8D530A" box="[790,1068,511,537]" italics="true" pageId="3" pageNumber="522">
<taxonomicName id="4C455FC61F066B04FC80FE3FFC29530B" authorityName="Voros, Dias &amp; Sole" authorityYear="2017" box="[790,904,511,537]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="amadoi">P. amadoi</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FC04FE3FFB89530A" authorityName="Orrico, Dias &amp; Marciano" authorityYear="2018" box="[914,1064,511,536]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="praeceptor">P. praeceptor</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C455FC61F066B04FBF7FE3FFB48530A" authorityName="Caramaschi &amp; Peixoto" authorityYear="2004" box="[1121,1257,511,536]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B931F8571F066B04FBF7FE3FFB48530A" box="[1121,1257,511,536]" italics="true" pageId="3" pageNumber="522">P. punctatus</emphasis>
</taxonomicName>
(small irregular spots), 
<taxonomicName id="4C455FC61F066B04FF7EFDE3FE27532F" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[232,390,547,573]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F066B04FF7EFDE3FE27532F" box="[232,390,547,573]" italics="true" pageId="3" pageNumber="522">P. gyrinaethes</emphasis>
</taxonomicName>
(marbled pattern), 
<taxonomicName id="4C455FC61F066B04FDCBFDE3FD4F532F" authorityName="Cruz, Feio &amp; Cardoso" authorityYear="2007" box="[605,750,547,573]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B931F8571F066B04FDCBFDE3FD4F532F" box="[605,750,547,573]" italics="true" pageId="3" pageNumber="522">P. maculosus</emphasis>
</taxonomicName>
(maculate pattern, but see Novaes-e-Fagundes &amp; Solé 2021), and 
<taxonomicName id="4C455FC61F066B04FF5EFD87FEF65373" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[200,343,583,609]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B931F8571F066B04FF5EFD87FEF65373" box="[200,343,583,609]" italics="true" pageId="3" pageNumber="522">P. wuchereri</emphasis>
</taxonomicName>
(two dorsolateral white stripes outlined by a dark brown to black line with or without irregular patches in dorsum).
</paragraph>
<paragraph id="8BFA24451F066B04FF51FD4FFD665207" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="522">
<taxonomicName id="4C455FC61F066B04FF51FD4FFE2653BA" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[199,391,655,681]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F066B04FF51FD4FFE2653BA" box="[199,391,655,681]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
has a belly with two rows of central tubercles with adjacent tubercles bordering without showing a pattern, which separates it from 
<taxonomicName id="4C455FC61F066B04FD15FD73FD5753DE" box="[643,758,691,716]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="magnus">
<emphasis id="B931F8571F066B04FD15FD73FD5753DE" box="[643,758,691,716]" italics="true" pageId="3" pageNumber="522">P. magnus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FC90FD73FCDE53DF" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[774,895,691,717]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="kautskyi">
<emphasis id="B931F8571F066B04FC90FD73FCDE53DF" box="[774,895,691,717]" italics="true" pageId="3" pageNumber="522">P. kautskyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FC18FD73FBEB53DF" authorityName="Caramaschi, Silva &amp; Britto-Pereira" authorityYear="1992" box="[910,1098,691,717]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="melanomystax">
<emphasis id="B931F8571F066B04FC18FD73FBEB53DF" box="[910,1098,691,717]" italics="true" pageId="3" pageNumber="522">P. melanomystax</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FBCCFD73FB5B53DF" authorityName="Peixoto, Caramaschi &amp; Freire" authorityYear="2003" box="[1114,1274,691,717]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="gyrinaethes">
<emphasis id="B931F8571F066B04FBCCFD73FB5B53DF" box="[1114,1274,691,717]" italics="true" pageId="3" pageNumber="522">P. gyrinaethes</emphasis>
</taxonomicName>
(tubercles not distinct), from 
<taxonomicName id="4C455FC61F066B04FED7FD17FE7E53E2" authorityName="Bokermann" authorityYear="1966" box="[321,479,727,752]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="B931F8571F066B04FED7FD17FE7E53E2" box="[321,479,727,752]" italics="true" pageId="3" pageNumber="522">P. acuminatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FE78FD17FD3853E3" authorityName="Bokermann" authorityYear="1966" box="[494,665,727,753]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="tuberculosus">
<emphasis id="B931F8571F066B04FE78FD17FD3853E3" box="[494,665,727,753]" italics="true" pageId="3" pageNumber="522">P. tuberculosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FD3EFD17FCB853E3" authorityName="Voros, Dias &amp; Sole" authorityYear="2017" box="[680,793,727,753]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="amadoi">
<emphasis id="B931F8571F066B04FD3EFD17FCB853E3" box="[680,793,727,753]" italics="true" pageId="3" pageNumber="522">P. amadoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FCBEFD17FC3F53E3" baseAuthorityName="Wied-Neuwied" baseAuthorityYear="1821" box="[808,926,727,753]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="luteolus">
<emphasis id="B931F8571F066B04FCBEFD17FC3F53E3" box="[808,926,727,753]" italics="true" pageId="3" pageNumber="522">P. luteolus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C455FC61F066B04FC3BFD17FB9A53E3" baseAuthorityName="Peters" baseAuthorityYear="1873" box="[941,1083,727,753]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="wuchereri">
<emphasis id="B931F8571F066B04FC3BFD17FB9A53E3" box="[941,1083,727,753]" italics="true" pageId="3" pageNumber="522">P. wuchereri</emphasis>
</taxonomicName>
(six rows of rounded tubercles laterally merging with the granules on the flanks).
</paragraph>
<caption id="DF3A74CD1F066B04FF0EF9C5FF535752" ID-DOI="http://doi.org/10.5281/zenodo.10159023" ID-Zenodo-Dep="10159023" httpUri="https://zenodo.org/record/10159023/files/figure.png" pageId="3" pageNumber="522" startId="3.[152,255,1541,1565]" targetBox="[170,1417,813,1517]" targetPageId="3" targetType="figure">
<paragraph id="8BFA24451F066B04FF0EF9C5FF535752" blockId="3.[151,1436,1541,1601]" pageId="3" pageNumber="522">
<emphasis id="B931F8571F066B04FF0EF9C5FEB8570F" bold="true" box="[152,281,1541,1565]" pageId="3" pageNumber="522">FIGURE 1.</emphasis>
Holotype of 
<taxonomicName id="4C455FC61F066B04FE36F9C5FDEB570F" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[416,586,1541,1565]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="3" pageNumber="522" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F066B04FE36F9C5FDEB570F" box="[416,586,1541,1565]" italics="true" pageId="3" pageNumber="522">Phyllodytes iuna</emphasis>
</taxonomicName>
<emphasis id="B931F8571F066B04FDC6F9C5FD00570F" bold="true" box="[592,673,1541,1565]" pageId="3" pageNumber="522">
<taxonomicNameLabel id="A202452C1F066B04FDC6F9C5FD00570F" box="[592,673,1541,1565]" pageId="3" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZUESC 18950). (A) Dorsal and (B) ventral view of the body. Scale bar = 10 mm
</paragraph>
</caption>
<paragraph id="8BFA24451F066B03FF51F953FC64500F" blockId="3.[151,1436,1683,2034]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="523" pageId="3" pageNumber="522">
<emphasis id="B931F8571F066B04FF51F953FDB957BF" bold="true" box="[199,536,1683,1709]" pageId="3" pageNumber="522">
Description of the 
<typeStatus id="54FE9AE71F066B04FE3CF953FDB257BF" box="[426,531,1683,1709]" pageId="3" pageNumber="522" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male in good condition, with a piece of muscle removed from the right thigh for molecular analysis. Detailed measurements are in 
<tableCitation id="C6C711FE1F066B04FCBAF977FC2157C3" box="[812,896,1719,1745]" captionStart="TABLE 1" captionStartId="6.[152,244,512,536]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of the type series of Phyllodytes iuna sp. nov." httpUri="http://table.plazi.org/id/DF3A74CD1F036B01FF0EFDC0FC40530B" pageId="3" pageNumber="522" tableUuid="DF3A74CD1F036B01FF0EFDC0FC40530B">Table 1</tableCitation>
. Head is as long as it is wide (HL 97.4% of HW; HW 35.5% of SVL; HL 34.6% of SVL); snout truncate with apical tubercle in dorsal view and projected slightly forward in lateral view (
<figureCitation id="137E38C01F066B04FE38F93FFE54560B" box="[430,501,1791,1817]" captionStart="FIGURE 2" captionStartId="4.[152,255,1357,1381]" captionTargetBox="[170,1416,309,1334]" captionTargetId="figure-92@4.[170,1416,309,1334]" captionTargetPageId="4" captionText="FIGURE 2. Holotype of Phyllodytes iuna sp. nov. (MZUESC 18950). (A) Dorsal view of head; (B) Lateral view of head (right side); (C) Palmar view of left hand; (D) Plantar view of left foot. Scale bar = 3 mm" figureDoi="http://doi.org/10.5281/zenodo.10159025" httpUri="https://zenodo.org/record/10159025/files/figure.png" pageId="3" pageNumber="522">Fig. 2</figureCitation>
); nostrils with rounded contours, positioned close to the tip of the snout, directed dorsolaterally; distance between nostrils smaller than the eye-nose distance (IND 57.2% of NDE); large eyes (ED 34.4% of HL; 33.5% of HW; 126.2% of NDE), prominent, positioned anterolaterally; 
<emphasis id="B931F8571F066B04FBDBF887FAA95673" box="[1101,1288,1863,1889]" italics="true" pageId="3" pageNumber="522">canthus rostralis</emphasis>
round; loreal region concave and straight; tympanum evident, relatively large (TD 19.4% of HL), nearly circular; diameter of the tympanum is larger than the distance between nostrils (125% of IND) and smaller than the eye-narrow distance (TD 71.4% of NDE), eye diameter (TD 57.6% of ED), and interorbital distance (TD 28.3% of IOD); diameter of the tympanum is slightly smaller than the width of the discs of the third toe (TD/DF3 = 96.6%) and fourth toe (TD/4TD = 89.9%); tympanic ring evident; the supratympanic fold is well marked from the back of the eyes to near the insertion of the arm and has an arched outline; single vocal sac; vomerine teeth forming two slightly arched series, close to each other arranged horizontally between and behind the coanas; two anterior large, and additional smaller (2–3) subequal odontoids on each side of the mandible; horizontal pupil.
</paragraph>
<caption id="DF3A74CD1F016B03FF0EFA8DFC41549B" ID-DOI="http://doi.org/10.5281/zenodo.10159025" ID-Zenodo-Dep="10159025" httpUri="https://zenodo.org/record/10159025/files/figure.png" pageId="4" pageNumber="523" startId="4.[152,255,1357,1381]" targetBox="[170,1416,309,1334]" targetPageId="4" targetType="figure">
<paragraph id="8BFA24451F016B03FF0EFA8DFC41549B" blockId="4.[151,1436,1357,1417]" pageId="4" pageNumber="523">
<emphasis id="B931F8571F016B03FF0EFA8DFEB95477" bold="true" box="[152,280,1357,1381]" pageId="4" pageNumber="523">FIGURE 2.</emphasis>
Holotype of 
<taxonomicName id="4C455FC61F016B03FE0AFA8DFDE45477" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[412,581,1357,1381]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="4" pageNumber="523" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F016B03FE0AFA8DFDE45477" box="[412,581,1357,1381]" italics="true" pageId="4" pageNumber="523">Phyllodytes iuna</emphasis>
</taxonomicName>
<emphasis id="B931F8571F016B03FDDCFA8DFD3B5477" bold="true" box="[586,666,1357,1381]" pageId="4" pageNumber="523">
<taxonomicNameLabel id="A202452C1F016B03FDDCFA8DFD3B5477" box="[586,666,1357,1381]" pageId="4" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZUESC 18950). (A) Dorsal view of head; (B) Lateral view of head (right side); (C) Palmar view of left hand; (D) Plantar view of left foot. Scale bar = 3 mm
</paragraph>
</caption>
<paragraph id="8BFA24451F016B03FF51FA78FC4E5770" blockId="4.[151,1437,1463,1958]" pageId="4" pageNumber="523">Forearm with outer four tubercles, with the most evident ones near the hand. Hands large (HAL 91.5 % of HL), subarticular tubercles simple and round; supernumerary tubercles absent; palmar tubercle elongate and ovoid; fringe between fingers absent; fingers in increasing order of size I &lt;IV &lt;II &lt;III; unpigmented nuptial pad on the internal base of finger I, without dark nuptial excrescence, but showing a different color pattern than the surrounding one with dark spots; hand membrane webbing I 2 - 2 - II 2 + - 2 - III 2 - - 2 + IV.</paragraph>
<paragraph id="8BFA24451F016B03FF51F9ACFC71564C" blockId="4.[151,1437,1463,1958]" pageId="4" pageNumber="523">Long hind limbs (THL+TBL+TAL+FL equal 180.3% of SVL), thigh is shorter than tibia (THL 89.7% of TBL; THL 49.3% of SVL; TBL 55.8% of SVL); total of the length of the thigh and tibia greater than the SVL (THL+TBL 104.5% of the SVL); a single tubercle on the inner posteroventral margin of the tarsus, near the junction with the tibia. Tarsus shorter than foot (TAL 66.1% of FL). Foot length shorter than thigh and tibia length (FL 91.2% of THL; FL 81% of TBL). Plantar surface with poorly developed supernumerary tubercles; large elongated internal metatarsal tubercle; small and round external metatarsal tubercle; toes without membranes between the I, II and III toes; foot membrane webbing I 2 - - 2 + II 2 + - 3 III 1 + + 3 IV 1 - 1 + V.</paragraph>
<paragraph id="8BFA24451F016B03FF51F8A7FC6A56B7" blockId="4.[151,1437,1463,1958]" pageId="4" pageNumber="523">Dorsal surfaces smooth; calcar appendix and supracloacal crest absent. Ventral surface of thighs with tubercles; ventral tubercles reaching to pectoral; mesal pericloacal tubercles absent.</paragraph>
<caption id="DF3A74CD1F006B02FF0EFD4FFB9553D9" ID-DOI="http://doi.org/10.5281/zenodo.10159027" ID-Zenodo-Dep="10159027" httpUri="https://zenodo.org/record/10159027/files/figure.png" pageId="5" pageNumber="524" startId="5.[152,255,655,679]" targetBox="[151,1435,181,631]" targetPageId="5" targetType="figure">
<paragraph id="8BFA24451F006B02FF0EFD4FFB9553D9" blockId="5.[151,1437,655,715]" pageId="5" pageNumber="524">
<emphasis id="B931F8571F006B02FF0EFD4FFEBB53B5" bold="true" box="[152,282,655,679]" pageId="5" pageNumber="524">FIGURE 3.</emphasis>
Paratopotypes of 
<taxonomicName id="4C455FC61F006B02FE43FD4FFD2153B5" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[469,640,655,679]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F006B02FE43FD4FFD2153B5" box="[469,640,655,679]" italics="true" pageId="5" pageNumber="524">Phyllodytes iuna</emphasis>
</taxonomicName>
<emphasis id="B931F8571F006B02FD1EFD4FFD7B53B5" bold="true" box="[648,730,655,679]" pageId="5" pageNumber="524">
<taxonomicNameLabel id="A202452C1F006B02FD1EFD4FFD7B53B5" box="[648,730,655,679]" pageId="5" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
MZUESC 18952 (A and F); MZUESC 23000 (B and G); MZUESC 23001 (C and H); MZUESC 23002 (D and I); MZUESC 23003 (E and J). Scale bar = 10 m
</paragraph>
</caption>
<paragraph id="8BFA24451F006B02FF51FD30FC2A5240" blockId="5.[151,1437,752,851]" pageId="5" pageNumber="524">
<emphasis id="B931F8571F006B02FF51FD30FDDB5218" bold="true" box="[199,634,752,778]" pageId="5" pageNumber="524">
Color of the 
<typeStatus id="54FE9AE71F006B02FECDFD30FE635218" box="[347,450,752,778]" pageId="5" pageNumber="524" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
. In preservative it has a yellowish-brown coloration, a dorsal coloration presenting a diffuse band-like pattern extending halfway down the body and a dark band extending from the eyes to halfway down the body; the entire ventral surface is cream colored.
</paragraph>
<caption id="DF3A74CD1F006B02FF0EF8A1FAEC56F4" ID-DOI="http://doi.org/10.5281/zenodo.10159029" ID-Zenodo-Dep="10159029" httpUri="https://zenodo.org/record/10159029/files/figure.png" pageId="5" pageNumber="524" startId="5.[152,255,1889,1913]" targetBox="[170,1417,874,1865]" targetPageId="5" targetType="figure">
<paragraph id="8BFA24451F006B02FF0EF8A1FAEC56F4" blockId="5.[151,1436,1889,2022]" pageId="5" pageNumber="524">
<emphasis id="B931F8571F006B02FF0EF8A1FEB9566B" bold="true" box="[152,280,1889,1913]" pageId="5" pageNumber="524">FIGURE 4.</emphasis>
Variation pattern of the dorsal coloration of 
<taxonomicName id="4C455FC61F006B02FD4AF8A2FC80566B" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[732,801,1890,1913]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F006B02FD4AF8A2FC80566B" box="[732,801,1890,1913]" italics="true" pageId="5" pageNumber="524">P. iuna</emphasis>
</taxonomicName>
<emphasis id="B931F8571F006B02FCBEF8A1FCD9566B" bold="true" box="[808,888,1889,1913]" pageId="5" pageNumber="524">
<taxonomicNameLabel id="A202452C1F006B02FCBEF8A1FCD9566B" box="[808,888,1889,1913]" pageId="5" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and its sister clade 
<taxonomicName id="4C455FC61F006B02FBD6F8A2FB08566B" box="[1088,1193,1889,1913]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B931F8571F006B02FBD6F8A2FB08566B" box="[1088,1193,1889,1913]" italics="true" pageId="5" pageNumber="524">P. edelmoi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F006B02FB4CF8A2FAC6566B" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[1242,1383,1889,1913]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B931F8571F006B02FB4CF8A2FAC6566B" box="[1242,1383,1889,1913]" italics="true" pageId="5" pageNumber="524">P. brevirostris</emphasis>
</taxonomicName>
. Paratopotypes of 
<taxonomicName id="4C455FC61F006B02FECFF845FDAB568F" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[345,522,1925,1949]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F006B02FECFF845FDAB568F" box="[345,522,1925,1949]" italics="true" pageId="5" pageNumber="524">Phyllodytes iuna</emphasis>
</taxonomicName>
<emphasis id="B931F8571F006B02FD8FF845FDD0568F" bold="true" box="[537,625,1925,1949]" pageId="5" pageNumber="524">
<taxonomicNameLabel id="A202452C1F006B02FD8FF845FDD0568F" box="[537,625,1925,1949]" pageId="5" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MZUESC 18952, A and MZUESC 23001, B, Photos by Rafael O. de Abreu and Marcos Vila Nova, respectively); 
<taxonomicName id="4C455FC61F006B02FDCCF869FC8756D3" box="[602,806,1961,1985]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B931F8571F006B02FDCCF869FC8756D3" box="[602,806,1961,1985]" italics="true" pageId="5" pageNumber="524">Phyllodytes edelmoi</emphasis>
</taxonomicName>
(C, Reserva Biológica de Saltinho, Tamandaré—PE, Photo by Pedro Ivo Simıes) and 
<taxonomicName id="4C455FC61F006B02FE13F80DFDD756F7" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[389,630,1997,2021]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="524" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B931F8571F006B02FE13F80DFDD756F7" box="[389,630,1997,2021]" italics="true" pageId="5" pageNumber="524">Phyllodytes brevirostris</emphasis>
</taxonomicName>
(D, Cruz do Espírito Santo—PB, Photo by Washington Luiz S. Vieira).
</paragraph>
</caption>
<paragraph id="8BFA24451F036B01FF51FF56FC4E500F" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF51FF56FEF951A2" bold="true" box="[199,344,150,176]" pageId="6" pageNumber="525">Color in life</emphasis>
. Based on digital photographs of 
<typeStatus id="54FE9AE71F036B01FD4DFF57FCD951A3" box="[731,888,151,177]" pageId="6" pageNumber="525" type="paratopotype">paratopotypes</typeStatus>
(
<figureCitation id="137E38C01F036B01FC1EFF57FB8C51A3" box="[904,1069,151,177]" captionStart="FIGURE 4" captionStartId="5.[152,255,1889,1913]" captionTargetBox="[170,1417,874,1865]" captionTargetId="figure-110@5.[170,1417,874,1865]" captionTargetPageId="5" captionText="FIGURE 4. Variation pattern of the dorsal coloration of P. iuna sp. nov. and its sister clade P. edelmoi and P. brevirostris. Paratopotypes of Phyllodytes iuna sp. nov. (MZUESC 18952, A and MZUESC 23001, B, Photos by Rafael O. de Abreu and Marcos Vila Nova, respectively); Phyllodytes edelmoi (C, Reserva Biológica de Saltinho, Tamandaré—PE, Photo by Pedro Ivo Simıes) and Phyllodytes brevirostris (D, Cruz do Espírito Santo—PB, Photo by Washington Luiz S. Vieira)." figureDoi="http://doi.org/10.5281/zenodo.10159029" httpUri="https://zenodo.org/record/10159029/files/figure.png" pageId="6" pageNumber="525">Fig. 4A and B</figureCitation>
). In life it is yellow with shades of dark brown in the snout region, but this color smooths throughout the body ending in light yellow in the cloacal region. Dorsum coloration forming a diffuse band-like pattern extending halfway down the body. Dark brown stripe extending from the eyes halfway down the body. Belly uniformly yellowish.
</paragraph>
<paragraph id="8BFA24451F036B01FF51FEE6FF1450C3" blockId="6.[151,1437,150,465]" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF51FEE6FE9E5052" bold="true" box="[199,319,294,320]" pageId="6" pageNumber="525">Variation.</emphasis>
The type series is morphologically consistent with the 
<typeStatus id="54FE9AE71F036B01FC5FFEE7FB8E5053" box="[969,1071,295,321]" pageId="6" pageNumber="525" type="holotype">holotype</typeStatus>
. The measurements of the type series are summarized in 
<tableCitation id="C6C711FE1F036B01FE24FE8BFDA75077" box="[434,518,331,357]" captionStart="TABLE 1" captionStartId="6.[152,244,512,536]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of the type series of Phyllodytes iuna sp. nov." httpUri="http://table.plazi.org/id/DF3A74CD1F036B01FF0EFDC0FC40530B" pageId="6" pageNumber="525" tableUuid="DF3A74CD1F036B01FF0EFDC0FC40530B">Table 1</tableCitation>
. However, 
<materialsCitation id="3B2D2E181F036B01FD10FE8BFCD25077" box="[646,883,330,357]" collectionCode="MZUESC" pageId="6" pageNumber="525" specimenCode="MZUESC 18952" specimenCount="1">
<specimenCode id="DBE38C3E1F036B01FD10FE8BFCE55077" box="[646,836,330,357]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="6" pageNumber="525">MZUESC 18952</specimenCode>
and
</materialsCitation>
MZUESC 
<quantity id="4CBD89A01F036B01FC64FE8BFBF45077" box="[1010,1109,331,357]" metricMagnitude="2" metricUnit="m" metricValue="5.842254" pageId="6" pageNumber="525" unit="in" value="23001.0">23001 in</quantity>
lateral view have a truncated snout; 
<materialsCitation id="3B2D2E181F036B01FF74FEAFFE3E509B" box="[226,415,367,393]" collectionCode="MZUESC" pageId="6" pageNumber="525" specimenCode="MZUESC 23003" specimenCount="1">
<specimenCode id="DBE38C3E1F036B01FF74FEAFFE3E509B" box="[226,415,367,393]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="6" pageNumber="525">MZUESC 23003</specimenCode>
</materialsCitation>
has poorly developed tubercles along the tarsus with a more developed one in the tibiotarsal region; 
<materialsCitation id="3B2D2E181F036B01FF79FE53FE1150BF" box="[239,432,402,429]" collectionCode="MZUESC" pageId="6" pageNumber="525" specimenCode="MZUESC 18952" specimenCount="1">
<specimenCode id="DBE38C3E1F036B01FF79FE53FE1150BF" box="[239,432,402,429]" collectionCode="MZUESC" country="Brazil" httpUri="http://grbio.org/cool/yhj1-k4hf" name="Museu de Zoologia da Universidade Estadual de Santa Cruz" pageId="6" pageNumber="525">MZUESC 18952</specimenCode>
</materialsCitation>
has a dorsal coloration pattern with fewer blotches than the rest of the type-series (
<figureCitation id="137E38C01F036B01FAF9FE53FF0650C3" captionStart="FIGURE 3" captionStartId="5.[152,255,655,679]" captionTargetBox="[151,1435,181,631]" captionTargetId="figure-19@5.[151,1435,181,631]" captionTargetPageId="5" captionText="FIGURE 3. Paratopotypes of Phyllodytes iuna sp. nov. MZUESC 18952 (A and F); MZUESC 23000 (B and G); MZUESC 23001 (C and H); MZUESC 23002 (D and I); MZUESC 23003 (E and J). Scale bar = 10 m" figureDoi="http://doi.org/10.5281/zenodo.10159027" httpUri="https://zenodo.org/record/10159027/files/figure.png" pageId="6" pageNumber="525">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF3A74CD1F036B01FF0EFDC0FC40530B" ID-Table-UUID="DF3A74CD1F036B01FF0EFDC0FC40530B" box="[152,993,512,537]" httpUri="http://table.plazi.org/id/DF3A74CD1F036B01FF0EFDC0FC40530B" pageId="6" pageNumber="525" startId="6.[152,244,512,536]" targetBox="[159,1408,558,1506]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BFA24451F036B01FF0EFDC0FC40530B" blockId="6.[152,993,512,537]" box="[152,993,512,537]" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF0EFDC0FEAE530A" bold="true" box="[152,271,512,536]" pageId="6" pageNumber="525">TABLE 1.</emphasis>
Measurements (mm) of the type series of 
<taxonomicName id="4C455FC61F036B01FD45FDC0FC24530A" authority="Santos &amp; Roseno &amp; Solé &amp; Dias, 2023" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[723,901,512,536]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="525" phylum="Chordata" rank="species" species="iuna" status="sp. nov.">
<emphasis id="B931F8571F036B01FD45FDC0FC24530A" box="[723,901,512,536]" italics="true" pageId="6" pageNumber="525">Phyllodytes iuna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202452C1F036B01FC1AFDC1FC40530B" box="[908,993,513,537]" pageId="6" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BFA24451F036B01FEB5FDEEFA9654F0" pageId="6" pageNumber="525">
<table id="F945D6E51F0394F8FF09FDEEFA2154F0" box="[159,1408,558,1506]" gridcols="9" gridrows="17" pageId="6" pageNumber="525">
<tr id="357526071F0394F8FF09FDEEFA215351" box="[159,1408,558,579]" gridrow="0" pageId="6" pageNumber="525" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="76A44F7B1F0394F8FEB5FDEEFE2B5351" box="[291,394,558,579]" gridcol="1" gridrow="0" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FEB5FDEEFEC15351" bold="true" box="[291,352,558,579]" pageId="6" pageNumber="525">Males</emphasis>
</th>
<th id="76A44F7B1F0394F8FB18FDEEFB545351" box="[1166,1269,558,579]" gridcol="7" gridrow="0" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FB18FDEEFB435351" bold="true" box="[1166,1250,558,579]" pageId="6" pageNumber="525">Females</emphasis>
</th>
</tr>
<tr id="357526071F0394F8FF09FD9AFA2153A5" box="[159,1408,602,695]" gridrow="1" pageId="6" pageNumber="525" rowspan-0="1">
<td id="76A44F7B1F0394F8FEB5FD9AFE2B53A5" box="[291,394,602,695]" gridcol="1" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FEB5FD9AFEFE53A5" bold="true" box="[291,394,602,695]" pageId="6" pageNumber="525">Holotype MZUESC 18950</emphasis>
</td>
<td id="76A44F7B1F0394F8FE21FD9AFDBF53A5" box="[439,542,602,695]" gridcol="2" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FE21FD9AFE5253A5" bold="true" box="[439,542,602,695]" pageId="6" pageNumber="525">Paratype MZUESC 18952</emphasis>
</td>
<td id="76A44F7B1F0394F8FDD0FD9AFD0C53A5" box="[582,685,602,695]" gridcol="3" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FDD0FD9AFD2353A5" bold="true" box="[582,685,602,695]" pageId="6" pageNumber="525">Paratype MZUESC 23000</emphasis>
</td>
<td id="76A44F7B1F0394F8FD40FD9AFC9C53A5" box="[726,829,602,695]" gridcol="4" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FD40FD9AFCB353A5" bold="true" box="[726,829,602,695]" pageId="6" pageNumber="525">Paratype MZUESC 23002</emphasis>
</td>
<td id="76A44F7B1F0394F8FCFCFD9AFC6B53A5" box="[874,970,602,695]" gridcol="5" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FCFCFDBEFC6B5386" bold="true" box="[874,970,638,660]" pageId="6" pageNumber="525">Min/Max</emphasis>
</td>
<td id="76A44F7B1F0394F8FC6FFD9AFBCD53A5" box="[1017,1132,602,695]" gridcol="6" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FC6FFDBEFBCD5386" bold="true" box="[1017,1132,638,660]" pageId="6" pageNumber="525">Mean ± SD</emphasis>
</td>
<td id="76A44F7B1F0394F8FB18FD9AFB5453A5" box="[1166,1269,602,695]" gridcol="7" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FB18FD9AFB6B53A5" bold="true" box="[1166,1269,602,695]" pageId="6" pageNumber="525">Paratype MZUESC 23001</emphasis>
</td>
<td id="76A44F7B1F0394F8FA8FFD9AFA2153A5" box="[1305,1408,602,695]" gridcol="8" gridrow="1" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FA8FFD9AFAF453A5" bold="true" box="[1305,1408,602,695]" pageId="6" pageNumber="525">Paratype MZUESC 23003</emphasis>
</td>
</tr>
<tr id="357526071F0394F8FF09FD11FA2153F5" box="[159,1408,721,743]" gridrow="2" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FD11FF7253F5" box="[159,211,721,743]" gridcol="0" gridrow="2" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FD11FF6F53F5" bold="true" box="[159,206,721,743]" pageId="6" pageNumber="525">SVL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FD11FE2B53F5" box="[291,394,721,743]" gridcol="1" gridrow="2" pageId="6" pageNumber="525">22.2</td>
<td id="76A44F7B1F0394F8FE21FD11FDBF53F5" box="[439,542,721,743]" gridcol="2" gridrow="2" pageId="6" pageNumber="525">23.2</td>
<td id="76A44F7B1F0394F8FDD0FD11FD0C53F5" box="[582,685,721,743]" gridcol="3" gridrow="2" pageId="6" pageNumber="525">25.2</td>
<td id="76A44F7B1F0394F8FD40FD11FC9C53F5" box="[726,829,721,743]" gridcol="4" gridrow="2" pageId="6" pageNumber="525">23.3</td>
<td id="76A44F7B1F0394F8FCFCFD11FC6B53F5" box="[874,970,721,743]" gridcol="5" gridrow="2" pageId="6" pageNumber="525">22.2–25.2</td>
<td id="76A44F7B1F0394F8FC6FFD11FBCD53F5" box="[1017,1132,721,743]" gridcol="6" gridrow="2" pageId="6" pageNumber="525">23.4 ± 1.08</td>
<td id="76A44F7B1F0394F8FB18FD11FB5453F5" box="[1166,1269,721,743]" gridcol="7" gridrow="2" pageId="6" pageNumber="525">24.0</td>
<td id="76A44F7B1F0394F8FA8FFD11FA2153F5" box="[1305,1408,721,743]" gridcol="8" gridrow="2" pageId="6" pageNumber="525">25.9</td>
</tr>
<tr id="357526071F0394F8FF09FCC7FA21520F" box="[159,1408,775,797]" gridrow="3" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FCC7FF72520F" box="[159,211,775,797]" gridcol="0" gridrow="3" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FCC7FF6B520E" bold="true" box="[159,202,775,796]" pageId="6" pageNumber="525">HW</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FCC7FE2B520F" box="[291,394,775,797]" gridcol="1" gridrow="3" pageId="6" pageNumber="525">7.9</td>
<td id="76A44F7B1F0394F8FE21FCC7FDBF520F" box="[439,542,775,797]" gridcol="2" gridrow="3" pageId="6" pageNumber="525">7.7</td>
<td id="76A44F7B1F0394F8FDD0FCC7FD0C520F" box="[582,685,775,797]" gridcol="3" gridrow="3" pageId="6" pageNumber="525">7.9</td>
<td id="76A44F7B1F0394F8FD40FCC7FC9C520F" box="[726,829,775,797]" gridcol="4" gridrow="3" pageId="6" pageNumber="525">7.9</td>
<td id="76A44F7B1F0394F8FCFCFCC7FC6B520F" box="[874,970,775,797]" gridcol="5" gridrow="3" pageId="6" pageNumber="525">7.7–7.9</td>
<td id="76A44F7B1F0394F8FC6FFCC7FBCD520F" box="[1017,1132,775,797]" gridcol="6" gridrow="3" pageId="6" pageNumber="525">7.8 ± 0.09</td>
<td id="76A44F7B1F0394F8FB18FCC7FB54520F" box="[1166,1269,775,797]" gridcol="7" gridrow="3" pageId="6" pageNumber="525">8.1</td>
<td id="76A44F7B1F0394F8FA8FFCC7FA21520F" box="[1305,1408,775,797]" gridcol="8" gridrow="3" pageId="6" pageNumber="525">8.6</td>
</tr>
<tr id="357526071F0394F8FF09FCFEFA215241" box="[159,1408,830,851]" gridrow="4" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FCFEFF725241" box="[159,211,830,851]" gridcol="0" gridrow="4" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FCFEFF635241" bold="true" box="[159,194,830,851]" pageId="6" pageNumber="525">HL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FCFEFE2B5241" box="[291,394,830,851]" gridcol="1" gridrow="4" pageId="6" pageNumber="525">7.7</td>
<td id="76A44F7B1F0394F8FE21FCFEFDBF5241" box="[439,542,830,851]" gridcol="2" gridrow="4" pageId="6" pageNumber="525">8.2</td>
<td id="76A44F7B1F0394F8FDD0FCFEFD0C5241" box="[582,685,830,851]" gridcol="3" gridrow="4" pageId="6" pageNumber="525">8.9</td>
<td id="76A44F7B1F0394F8FD40FCFEFC9C5241" box="[726,829,830,851]" gridcol="4" gridrow="4" pageId="6" pageNumber="525">8.2</td>
<td id="76A44F7B1F0394F8FCFCFCFEFC6B5241" box="[874,970,830,851]" gridcol="5" gridrow="4" pageId="6" pageNumber="525">7.7–8.9</td>
<td id="76A44F7B1F0394F8FC6FFCFEFBCD5241" box="[1017,1132,830,851]" gridcol="6" gridrow="4" pageId="6" pageNumber="525">8.2 ± 0.43</td>
<td id="76A44F7B1F0394F8FB18FCFEFB545241" box="[1166,1269,830,851]" gridcol="7" gridrow="4" pageId="6" pageNumber="525">8.1</td>
<td id="76A44F7B1F0394F8FA8FFCFEFA215241" box="[1305,1408,830,851]" gridcol="8" gridrow="4" pageId="6" pageNumber="525">8.3</td>
</tr>
<tr id="357526071F0394F8FF09FCB4FA215298" box="[159,1408,884,906]" gridrow="5" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FCB4FF725298" box="[159,211,884,906]" gridcol="0" gridrow="5" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FCB4FF6A529B" bold="true" box="[159,203,884,905]" pageId="6" pageNumber="525">IND</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FCB4FE2B5298" box="[291,394,884,906]" gridcol="1" gridrow="5" pageId="6" pageNumber="525">1.2</td>
<td id="76A44F7B1F0394F8FE21FCB4FDBF5298" box="[439,542,884,906]" gridcol="2" gridrow="5" pageId="6" pageNumber="525">1.2</td>
<td id="76A44F7B1F0394F8FDD0FCB4FD0C5298" box="[582,685,884,906]" gridcol="3" gridrow="5" pageId="6" pageNumber="525">1.6</td>
<td id="76A44F7B1F0394F8FD40FCB4FC9C5298" box="[726,829,884,906]" gridcol="4" gridrow="5" pageId="6" pageNumber="525">1.5</td>
<td id="76A44F7B1F0394F8FCFCFCB4FC6B5298" box="[874,970,884,906]" gridcol="5" gridrow="5" pageId="6" pageNumber="525">1.2–1.6</td>
<td id="76A44F7B1F0394F8FC6FFCB4FBCD5298" box="[1017,1132,884,906]" gridcol="6" gridrow="5" pageId="6" pageNumber="525">1.3 ± 0.18</td>
<td id="76A44F7B1F0394F8FB18FCB4FB545298" box="[1166,1269,884,906]" gridcol="7" gridrow="5" pageId="6" pageNumber="525">1.5</td>
<td id="76A44F7B1F0394F8FA8FFCB4FA215298" box="[1305,1408,884,906]" gridcol="8" gridrow="5" pageId="6" pageNumber="525">1.5</td>
</tr>
<tr id="357526071F0394F8FF09FC6AFA2152D3" box="[159,1408,938,961]" gridrow="6" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FC6AFF7252D3" box="[159,211,938,961]" gridcol="0" gridrow="6" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FC6AFF6D52D2" bold="true" box="[159,204,938,960]" pageId="6" pageNumber="525">IOD</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FC6AFE2B52D3" box="[291,394,938,961]" gridcol="1" gridrow="6" pageId="6" pageNumber="525">5.3</td>
<td id="76A44F7B1F0394F8FE21FC6AFDBF52D3" box="[439,542,938,961]" gridcol="2" gridrow="6" pageId="6" pageNumber="525">5.4</td>
<td id="76A44F7B1F0394F8FDD0FC6AFD0C52D3" box="[582,685,938,961]" gridcol="3" gridrow="6" pageId="6" pageNumber="525">5.8</td>
<td id="76A44F7B1F0394F8FD40FC6AFC9C52D3" box="[726,829,938,961]" gridcol="4" gridrow="6" pageId="6" pageNumber="525">5.7</td>
<td id="76A44F7B1F0394F8FCFCFC6AFC6B52D3" box="[874,970,938,961]" gridcol="5" gridrow="6" pageId="6" pageNumber="525">5.3–5.8</td>
<td id="76A44F7B1F0394F8FC6FFC6AFBCD52D3" box="[1017,1132,938,961]" gridcol="6" gridrow="6" pageId="6" pageNumber="525">5.5 ± 0.21</td>
<td id="76A44F7B1F0394F8FB18FC6AFB5452D3" box="[1166,1269,938,961]" gridcol="7" gridrow="6" pageId="6" pageNumber="525">5.3</td>
<td id="76A44F7B1F0394F8FA8FFC6AFA2152D3" box="[1305,1408,938,961]" gridcol="8" gridrow="6" pageId="6" pageNumber="525">5.7</td>
</tr>
<tr id="357526071F0394F8FF09FC21FA2152E5" box="[159,1408,993,1015]" gridrow="7" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FC21FF7252E5" box="[159,211,993,1015]" gridcol="0" gridrow="7" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FC21FF7352E4" bold="true" box="[159,210,993,1014]" pageId="6" pageNumber="525">END</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FC21FE2B52E5" box="[291,394,993,1015]" gridcol="1" gridrow="7" pageId="6" pageNumber="525">2.1</td>
<td id="76A44F7B1F0394F8FE21FC21FDBF52E5" box="[439,542,993,1015]" gridcol="2" gridrow="7" pageId="6" pageNumber="525">2.3</td>
<td id="76A44F7B1F0394F8FDD0FC21FD0C52E5" box="[582,685,993,1015]" gridcol="3" gridrow="7" pageId="6" pageNumber="525">2.6</td>
<td id="76A44F7B1F0394F8FD40FC21FC9C52E5" box="[726,829,993,1015]" gridcol="4" gridrow="7" pageId="6" pageNumber="525">2.9</td>
<td id="76A44F7B1F0394F8FCFCFC21FC6B52E5" box="[874,970,993,1015]" gridcol="5" gridrow="7" pageId="6" pageNumber="525">2.1–2.9</td>
<td id="76A44F7B1F0394F8FC6FFC21FBCD52E5" box="[1017,1132,993,1015]" gridcol="6" gridrow="7" pageId="6" pageNumber="525">2.4 ± 0.3</td>
<td id="76A44F7B1F0394F8FB18FC21FB5452E5" box="[1166,1269,993,1015]" gridcol="7" gridrow="7" pageId="6" pageNumber="525">2.4</td>
<td id="76A44F7B1F0394F8FA8FFC21FA2152E5" box="[1305,1408,993,1015]" gridcol="8" gridrow="7" pageId="6" pageNumber="525">2.9</td>
</tr>
<tr id="357526071F0394F8FF09FBD8FA21553C" box="[159,1408,1048,1070]" gridrow="8" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FBD8FF72553C" box="[159,211,1048,1070]" gridcol="0" gridrow="8" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FBD8FF61553F" bold="true" box="[159,192,1048,1069]" pageId="6" pageNumber="525">ED</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FBD8FE2B553C" box="[291,394,1048,1070]" gridcol="1" gridrow="8" pageId="6" pageNumber="525">2.3</td>
<td id="76A44F7B1F0394F8FE21FBD8FDBF553C" box="[439,542,1048,1070]" gridcol="2" gridrow="8" pageId="6" pageNumber="525">2.5</td>
<td id="76A44F7B1F0394F8FDD0FBD8FD0C553C" box="[582,685,1048,1070]" gridcol="3" gridrow="8" pageId="6" pageNumber="525">2.6</td>
<td id="76A44F7B1F0394F8FD40FBD8FC9C553C" box="[726,829,1048,1070]" gridcol="4" gridrow="8" pageId="6" pageNumber="525">2.6</td>
<td id="76A44F7B1F0394F8FCFCFBD8FC6B553C" box="[874,970,1048,1070]" gridcol="5" gridrow="8" pageId="6" pageNumber="525">2.3–2.6</td>
<td id="76A44F7B1F0394F8FC6FFBD8FBCD553C" box="[1017,1132,1048,1070]" gridcol="6" gridrow="8" pageId="6" pageNumber="525">2.5 ± 0.12</td>
<td id="76A44F7B1F0394F8FB18FBD8FB54553C" box="[1166,1269,1048,1070]" gridcol="7" gridrow="8" pageId="6" pageNumber="525">2.4</td>
<td id="76A44F7B1F0394F8FA8FFBD8FA21553C" box="[1305,1408,1048,1070]" gridcol="8" gridrow="8" pageId="6" pageNumber="525">2.6</td>
</tr>
<tr id="357526071F0394F8FF09FB8EFA215576" box="[159,1408,1102,1124]" gridrow="9" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FB8EFF725576" box="[159,211,1102,1124]" gridcol="0" gridrow="9" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FB8EFF615571" bold="true" box="[159,192,1102,1123]" pageId="6" pageNumber="525">TD</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FB8EFE2B5576" box="[291,394,1102,1124]" gridcol="1" gridrow="9" pageId="6" pageNumber="525">1.5</td>
<td id="76A44F7B1F0394F8FE21FB8EFDBF5576" box="[439,542,1102,1124]" gridcol="2" gridrow="9" pageId="6" pageNumber="525">1.7</td>
<td id="76A44F7B1F0394F8FDD0FB8EFD0C5576" box="[582,685,1102,1124]" gridcol="3" gridrow="9" pageId="6" pageNumber="525">1.6</td>
<td id="76A44F7B1F0394F8FD40FB8EFC9C5576" box="[726,829,1102,1124]" gridcol="4" gridrow="9" pageId="6" pageNumber="525">1.5</td>
<td id="76A44F7B1F0394F8FCFCFB8EFC6B5576" box="[874,970,1102,1124]" gridcol="5" gridrow="9" pageId="6" pageNumber="525">1.5–1.7</td>
<td id="76A44F7B1F0394F8FC6FFB8EFBCD5576" box="[1017,1132,1102,1124]" gridcol="6" gridrow="9" pageId="6" pageNumber="525">1.5 ± 0.08</td>
<td id="76A44F7B1F0394F8FB18FB8EFB545576" box="[1166,1269,1102,1124]" gridcol="7" gridrow="9" pageId="6" pageNumber="525">1.4</td>
<td id="76A44F7B1F0394F8FA8FFB8EFA215576" box="[1305,1408,1102,1124]" gridcol="8" gridrow="9" pageId="6" pageNumber="525">1.5</td>
</tr>
<tr id="357526071F0394F8FF09FB44FA215588" box="[159,1408,1156,1178]" gridrow="10" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FB44FF725588" box="[159,211,1156,1178]" gridcol="0" gridrow="10" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FB44FF73558B" bold="true" box="[159,210,1156,1177]" pageId="6" pageNumber="525">THL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FB44FE2B5588" box="[291,394,1156,1178]" gridcol="1" gridrow="10" pageId="6" pageNumber="525">10.9</td>
<td id="76A44F7B1F0394F8FE21FB44FDBF5588" box="[439,542,1156,1178]" gridcol="2" gridrow="10" pageId="6" pageNumber="525">11.1</td>
<td id="76A44F7B1F0394F8FDD0FB44FD0C5588" box="[582,685,1156,1178]" gridcol="3" gridrow="10" pageId="6" pageNumber="525">11.0</td>
<td id="76A44F7B1F0394F8FD40FB44FC9C5588" box="[726,829,1156,1178]" gridcol="4" gridrow="10" pageId="6" pageNumber="525">10.5</td>
<td id="76A44F7B1F0394F8FCFCFB44FC6B5588" box="[874,970,1156,1178]" gridcol="5" gridrow="10" pageId="6" pageNumber="525">10.5–11.1</td>
<td id="76A44F7B1F0394F8FC6FFB44FBCD5588" box="[1017,1132,1156,1178]" gridcol="6" gridrow="10" pageId="6" pageNumber="525">10.8 ± 0.23</td>
<td id="76A44F7B1F0394F8FB18FB44FB545588" box="[1166,1269,1156,1178]" gridcol="7" gridrow="10" pageId="6" pageNumber="525">10.0</td>
<td id="76A44F7B1F0394F8FA8FFB44FA215588" box="[1305,1408,1156,1178]" gridcol="8" gridrow="10" pageId="6" pageNumber="525">11.5</td>
</tr>
<tr id="357526071F0394F8FF09FB7BFA2155C3" box="[159,1408,1211,1233]" gridrow="11" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FB7BFF7255C3" box="[159,211,1211,1233]" gridcol="0" gridrow="11" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FB7BFF6E55C2" bold="true" box="[159,207,1211,1232]" pageId="6" pageNumber="525">TBL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FB7BFE2B55C3" box="[291,394,1211,1233]" gridcol="1" gridrow="11" pageId="6" pageNumber="525">12.4</td>
<td id="76A44F7B1F0394F8FE21FB7BFDBF55C3" box="[439,542,1211,1233]" gridcol="2" gridrow="11" pageId="6" pageNumber="525">12.8</td>
<td id="76A44F7B1F0394F8FDD0FB7BFD0C55C3" box="[582,685,1211,1233]" gridcol="3" gridrow="11" pageId="6" pageNumber="525">13.1</td>
<td id="76A44F7B1F0394F8FD40FB7BFC9C55C3" box="[726,829,1211,1233]" gridcol="4" gridrow="11" pageId="6" pageNumber="525">11.7</td>
<td id="76A44F7B1F0394F8FCFCFB7BFC6B55C3" box="[874,970,1211,1233]" gridcol="5" gridrow="11" pageId="6" pageNumber="525">11.7–13.1</td>
<td id="76A44F7B1F0394F8FC6FFB7BFBCD55C3" box="[1017,1132,1211,1233]" gridcol="6" gridrow="11" pageId="6" pageNumber="525">12.5 ± 0.52</td>
<td id="76A44F7B1F0394F8FB18FB7BFB5455C3" box="[1166,1269,1211,1233]" gridcol="7" gridrow="11" pageId="6" pageNumber="525">12.5</td>
<td id="76A44F7B1F0394F8FA8FFB7BFA2155C3" box="[1305,1408,1211,1233]" gridcol="8" gridrow="11" pageId="6" pageNumber="525">13.4</td>
</tr>
<tr id="357526071F0394F8FF09FB31FA215415" box="[159,1408,1265,1287]" gridrow="12" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FB31FF725415" box="[159,211,1265,1287]" gridcol="0" gridrow="12" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FB31FF615414" bold="true" box="[159,192,1265,1286]" pageId="6" pageNumber="525">AL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FB31FE2B5415" box="[291,394,1265,1287]" gridcol="1" gridrow="12" pageId="6" pageNumber="525">6.6</td>
<td id="76A44F7B1F0394F8FE21FB31FDBF5415" box="[439,542,1265,1287]" gridcol="2" gridrow="12" pageId="6" pageNumber="525">6.4</td>
<td id="76A44F7B1F0394F8FDD0FB31FD0C5415" box="[582,685,1265,1287]" gridcol="3" gridrow="12" pageId="6" pageNumber="525">7.1</td>
<td id="76A44F7B1F0394F8FD40FB31FC9C5415" box="[726,829,1265,1287]" gridcol="4" gridrow="12" pageId="6" pageNumber="525">6.7</td>
<td id="76A44F7B1F0394F8FCFCFB31FC6B5415" box="[874,970,1265,1287]" gridcol="5" gridrow="12" pageId="6" pageNumber="525">6.4–7.1</td>
<td id="76A44F7B1F0394F8FC6FFB31FBCD5415" box="[1017,1132,1265,1287]" gridcol="6" gridrow="12" pageId="6" pageNumber="525">6.7 ± 0.25</td>
<td id="76A44F7B1F0394F8FB18FB31FB545415" box="[1166,1269,1265,1287]" gridcol="7" gridrow="12" pageId="6" pageNumber="525">6.9</td>
<td id="76A44F7B1F0394F8FA8FFB31FA215415" box="[1305,1408,1265,1287]" gridcol="8" gridrow="12" pageId="6" pageNumber="525">7.3</td>
</tr>
<tr id="357526071F0394F8FF09FAE8FA21542F" box="[159,1408,1320,1341]" gridrow="13" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FAE8FF72542F" box="[159,211,1320,1341]" gridcol="0" gridrow="13" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FAE8FF1F542F" bold="true" box="[159,190,1320,1341]" pageId="6" pageNumber="525">FL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FAE8FE2B542F" box="[291,394,1320,1341]" gridcol="1" gridrow="13" pageId="6" pageNumber="525">10.1</td>
<td id="76A44F7B1F0394F8FE21FAE8FDBF542F" box="[439,542,1320,1341]" gridcol="2" gridrow="13" pageId="6" pageNumber="525">9.8</td>
<td id="76A44F7B1F0394F8FDD0FAE8FD0C542F" box="[582,685,1320,1341]" gridcol="3" gridrow="13" pageId="6" pageNumber="525">10.0</td>
<td id="76A44F7B1F0394F8FD40FAE8FC9C542F" box="[726,829,1320,1341]" gridcol="4" gridrow="13" pageId="6" pageNumber="525">9.9</td>
<td id="76A44F7B1F0394F8FCFCFAE8FC6B542F" box="[874,970,1320,1341]" gridcol="5" gridrow="13" pageId="6" pageNumber="525">9.8–10.1</td>
<td id="76A44F7B1F0394F8FC6FFAE8FBCD542F" box="[1017,1132,1320,1341]" gridcol="6" gridrow="13" pageId="6" pageNumber="525">9.9 ± 0.11</td>
<td id="76A44F7B1F0394F8FB18FAE8FB54542F" box="[1166,1269,1320,1341]" gridcol="7" gridrow="13" pageId="6" pageNumber="525">9.8</td>
<td id="76A44F7B1F0394F8FA8FFAE8FA21542F" box="[1305,1408,1320,1341]" gridcol="8" gridrow="13" pageId="6" pageNumber="525">11.0</td>
</tr>
<tr id="357526071F0394F8FF09FA9EFA215466" box="[159,1408,1374,1396]" gridrow="14" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FA9EFF725466" box="[159,211,1374,1396]" gridcol="0" gridrow="14" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FA9EFF725461" bold="true" box="[159,211,1374,1395]" pageId="6" pageNumber="525">HAL</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FA9EFE2B5466" box="[291,394,1374,1396]" gridcol="1" gridrow="14" pageId="6" pageNumber="525">7.1</td>
<td id="76A44F7B1F0394F8FE21FA9EFDBF5466" box="[439,542,1374,1396]" gridcol="2" gridrow="14" pageId="6" pageNumber="525">6.8</td>
<td id="76A44F7B1F0394F8FDD0FA9EFD0C5466" box="[582,685,1374,1396]" gridcol="3" gridrow="14" pageId="6" pageNumber="525">7.7</td>
<td id="76A44F7B1F0394F8FD40FA9EFC9C5466" box="[726,829,1374,1396]" gridcol="4" gridrow="14" pageId="6" pageNumber="525">7.0</td>
<td id="76A44F7B1F0394F8FCFCFA9EFC6B5466" box="[874,970,1374,1396]" gridcol="5" gridrow="14" pageId="6" pageNumber="525">6.8–7.7</td>
<td id="76A44F7B1F0394F8FC6FFA9EFBCD5466" box="[1017,1132,1374,1396]" gridcol="6" gridrow="14" pageId="6" pageNumber="525">7.1 ± 0.34</td>
<td id="76A44F7B1F0394F8FB18FA9EFB545466" box="[1166,1269,1374,1396]" gridcol="7" gridrow="14" pageId="6" pageNumber="525">7.0</td>
<td id="76A44F7B1F0394F8FA8FFA9EFA215466" box="[1305,1408,1374,1396]" gridcol="8" gridrow="14" pageId="6" pageNumber="525">8.1</td>
</tr>
<tr id="357526071F0394F8FF09FA54FA2154B8" box="[159,1408,1428,1450]" gridrow="15" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FA54FF7254B8" box="[159,211,1428,1450]" gridcol="0" gridrow="15" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FA54FF6B54BB" bold="true" box="[159,202,1428,1449]" pageId="6" pageNumber="525">DF3</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FA54FE2B54B8" box="[291,394,1428,1450]" gridcol="1" gridrow="15" pageId="6" pageNumber="525">1.4</td>
<td id="76A44F7B1F0394F8FE21FA54FDBF54B8" box="[439,542,1428,1450]" gridcol="2" gridrow="15" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FDD0FA54FD0C54B8" box="[582,685,1428,1450]" gridcol="3" gridrow="15" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FD40FA54FC9C54B8" box="[726,829,1428,1450]" gridcol="4" gridrow="15" pageId="6" pageNumber="525">1.1</td>
<td id="76A44F7B1F0394F8FCFCFA54FC6B54B8" box="[874,970,1428,1450]" gridcol="5" gridrow="15" pageId="6" pageNumber="525">1.1–1.4</td>
<td id="76A44F7B1F0394F8FC6FFA54FBCD54B8" box="[1017,1132,1428,1450]" gridcol="6" gridrow="15" pageId="6" pageNumber="525">1.2 ± 0.11</td>
<td id="76A44F7B1F0394F8FB18FA54FB5454B8" box="[1166,1269,1428,1450]" gridcol="7" gridrow="15" pageId="6" pageNumber="525">1.4</td>
<td id="76A44F7B1F0394F8FA8FFA54FA2154B8" box="[1305,1408,1428,1450]" gridcol="8" gridrow="15" pageId="6" pageNumber="525">1.3</td>
</tr>
<tr id="357526071F0394F8FF09FA0CFA2154F0" box="[159,1408,1484,1506]" gridrow="16" pageId="6" pageNumber="525">
<th id="76A44F7B1F0394F8FF09FA0CFF7254F0" box="[159,211,1484,1506]" gridcol="0" gridrow="16" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF09FA0CFF6D54F3" bold="true" box="[159,204,1484,1505]" pageId="6" pageNumber="525">4TD</emphasis>
</th>
<td id="76A44F7B1F0394F8FEB5FA0CFE2B54F0" box="[291,394,1484,1506]" gridcol="1" gridrow="16" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FE21FA0CFDBF54F0" box="[439,542,1484,1506]" gridcol="2" gridrow="16" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FDD0FA0CFD0C54F0" box="[582,685,1484,1506]" gridcol="3" gridrow="16" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FD40FA0CFC9C54F0" box="[726,829,1484,1506]" gridcol="4" gridrow="16" pageId="6" pageNumber="525">1.2</td>
<td id="76A44F7B1F0394F8FCFCFA0CFC6B54F0" box="[874,970,1484,1506]" gridcol="5" gridrow="16" pageId="6" pageNumber="525">1.2–1.3</td>
<td id="76A44F7B1F0394F8FC6FFA0CFBCD54F0" box="[1017,1132,1484,1506]" gridcol="6" gridrow="16" pageId="6" pageNumber="525">1.2 ± 0.04</td>
<td id="76A44F7B1F0394F8FB18FA0CFB5454F0" box="[1166,1269,1484,1506]" gridcol="7" gridrow="16" pageId="6" pageNumber="525">1.3</td>
<td id="76A44F7B1F0394F8FA8FFA0CFA2154F0" box="[1305,1408,1484,1506]" gridcol="8" gridrow="16" pageId="6" pageNumber="525">1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFA24451F036B01FF01FA39FC025703" blockId="6.[151,931,1529,1553]" box="[151,931,1529,1553]" pageId="6" pageNumber="525">
<tableNote id="76A325CB1F036B01FF01FA39FC025703" box="[151,931,1529,1553]" pageId="6" pageNumber="525" targetBox="[159,1408,558,1506]" targetPageId="6">Characters presented as informed in M&amp;M section. SD = standard deviation.</tableNote>
</paragraph>
<paragraph id="8BFA24451F036B01FF51F9A3FBC35623" blockId="6.[151,1436,1635,1842]" pageId="6" pageNumber="525">
<emphasis id="B931F8571F036B01FF51F9A3FD50576F" bold="true" box="[199,753,1635,1661]" pageId="6" pageNumber="525">Natural history and geographic distribution.</emphasis>
Specimens were sighted at night and males vocalized in terrestrial bromeliads within well-preserved forests between 650 and 
<quantity id="4CBD89A01F036B01FC2AF947FBA657B0" box="[956,1031,1671,1698]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="525" unit="m" value="800.0">800 m</quantity>
a.s.l. 
<taxonomicName id="4C455FC61F036B01FBC6F947FAAE57B3" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1104,1295,1671,1697]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="6" pageNumber="525" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F036B01FBC6F947FAAE57B3" box="[1104,1295,1671,1697]" italics="true" pageId="6" pageNumber="525">Phyllodytes iuna</emphasis>
</taxonomicName>
is currently known only from Estaç ã o Ecológica de Wenceslau Guimar ã es (EEWG), a conservation unit managed by the state of 
<collectingRegion id="4981EAA71F036B01FF21F90FFF5A57FB" box="[183,251,1743,1769]" country="Brazil" name="Bahia" pageId="6" pageNumber="525">Bahia</collectingRegion>
, located in the north of the Central Corridor of the Atlantic Forest with a total area of ~ 
<quantity id="4CBD89A01F036B01FB71F90FFA9157FE" box="[1255,1328,1743,1772]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="525" unit="km" value="24.0">24 km</quantity>
² (
<figureCitation id="137E38C01F036B01FADEF90FFA2F57F8" box="[1352,1422,1743,1770]" captionStart="FIGURE 5" captionStartId="7.[152,255,1131,1155]" captionTargetBox="[151,1436,181,1090]" captionTargetId="figure-19@7.[151,1436,181,1090]" captionTargetPageId="7" captionText="FIGURE 5. Geographic distribution of Phyllodytes iuna sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10159031" httpUri="https://zenodo.org/record/10159031/files/figure.png" pageId="6" pageNumber="525">Fig. 5</figureCitation>
). This area was previously exploited for logging and cacao (
<taxonomicName id="4C455FC61F036B01FC8AF933FC44561F" box="[796,997,1779,1805]" class="Magnoliopsida" family="Malvaceae" genus="Theobroma" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="525" phylum="Tracheophyta" rank="species" species="cacao">
<emphasis id="B931F8571F036B01FC8AF933FC44561F" box="[796,997,1779,1805]" italics="true" pageId="6" pageNumber="525">Theobroma cacao</emphasis>
</taxonomicName>
) cultivation; however, many areas have remained untouched, especially near streams and upland areas (Rodrigues 
<emphasis id="B931F8571F036B01FC4CF8D8FBAA5623" box="[986,1035,1815,1841]" italics="true" pageId="6" pageNumber="525">et al</emphasis>
. 2013).
</paragraph>
<caption id="DF3A74CD1F026B00FF0EFBABFC905591" ID-DOI="http://doi.org/10.5281/zenodo.10159031" ID-Zenodo-Dep="10159031" box="[152,817,1131,1155]" httpUri="https://zenodo.org/record/10159031/files/figure.png" pageId="7" pageNumber="526" startId="7.[152,255,1131,1155]" targetBox="[151,1436,181,1090]" targetPageId="7" targetType="figure">
<paragraph id="8BFA24451F026B00FF0EFBABFC905591" blockId="7.[152,817,1131,1155]" box="[152,817,1131,1155]" pageId="7" pageNumber="526">
<emphasis id="B931F8571F026B00FF0EFBABFEB95591" bold="true" box="[152,280,1131,1155]" pageId="7" pageNumber="526">FIGURE 5.</emphasis>
Geographic distribution of 
<emphasis id="B931F8571F026B00FDA4FBABFD4D5591" box="[562,748,1131,1155]" italics="true" pageId="7" pageNumber="526">
<taxonomicName id="4C455FC61F026B00FDA4FBABFD7A5591" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[562,731,1131,1155]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="526" phylum="Chordata" rank="species" species="iuna">Phyllodytes iuna</taxonomicName>
s
</emphasis>
p. nov.
</paragraph>
</caption>
<paragraph id="8BFA24451F026B00FF51FB71FCED5406" blockId="7.[151,1436,1201,1300]" pageId="7" pageNumber="526">
<emphasis id="B931F8571F026B00FF51FB71FE2F55D9" bold="true" box="[199,398,1201,1227]" pageId="7" pageNumber="526">Genetic distance</emphasis>
. Genetic divergence with uncorrected p-distance between 
<taxonomicName id="4C455FC61F026B00FBBFFB71FB4655D9" authorityName="Santos &amp; Roseno &amp; Solé &amp; Dias" authorityYear="2023" box="[1065,1255,1201,1227]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="526" phylum="Chordata" rank="species" species="iuna">
<emphasis id="B931F8571F026B00FBBFFB71FB4655D9" box="[1065,1255,1201,1227]" italics="true" pageId="7" pageNumber="526">Phyllodytes iuna</emphasis>
</taxonomicName>
and the species that form its sister clade (
<taxonomicName id="4C455FC61F026B00FE2EFB16FDF755FD" authorityName="Peixoto &amp; Cruz" authorityYear="1988" box="[440,598,1237,1263]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="526" phylum="Chordata" rank="species" species="brevirostris">
<emphasis id="B931F8571F026B00FE2EFB16FDF755FD" box="[440,598,1237,1263]" italics="true" pageId="7" pageNumber="526">P. brevirostris</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C455FC61F026B00FD1BFB16FCA355FD" box="[653,770,1237,1263]" class="Amphibia" family="Hylidae" genus="Phyllodytes" kingdom="Animalia" order="Anura" pageId="7" pageNumber="526" phylum="Chordata" rank="species" species="edelmoi">
<emphasis id="B931F8571F026B00FD1BFB16FCA355FD" box="[653,770,1237,1263]" italics="true" pageId="7" pageNumber="526">P. edelmoi</emphasis>
</taxonomicName>
) were 4.4–5.0% for the partial 16S rDNA, whereas for the other species of the genus varied among 5.8–14.2% (
<tableCitation id="C6C711FE1F026B00FD71FB39FC9E5406" box="[743,831,1273,1300]" captionStart="TABLE 2" captionStartId="8.[151,243,651,677]" captionTargetPageId="8" captionText="TABLE 2. Uncorrected pairwise genetic distances of partial 16S rDNA (520 bp) between species of Phyllodytes." httpUri="http://table.plazi.org/id/DF3A74CD1F0D6B0FFF01FD4BFAF053B7" pageId="7" pageNumber="526" tableUuid="DF3A74CD1F0D6B0FFF01FD4BFAF053B7">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>