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<document id="9289CDFD7AFF6803119882471A5FDBDD" ID-CLB-Dataset="55245" ID-DOI="10.11646/zootaxa.5309.1.1" ID-GBIF-Dataset="048abd7b-8c78-494e-b466-b614aeef7f6b" ID-ISSN="1175-5326" ID-Zenodo-Dep="8090046" ID-ZooBank="C930242A-041C-4CEC-8B2F-2DB671A2395D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1687949280324" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="03DA1A1EFFACFF98FF053900E4AF3C46" docLanguage="en" docName="zootaxa.5309.1.1.pdf" docOrigin="Zootaxa 5309 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5309.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lagynochthonius zhijinensis Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="FFE36266FF9AFFA4FF923A59E133393B" masterDocTitle="New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China" masterLastPageNumber="64" masterPageNumber="1" pageNumber="55" updateTime="1699537060991" updateUser="ExternalLinkService">
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<mods:title id="4256D7CC3BA9CBA83D12748A479EBDFD">New cave-dwelling pseudoscorpions of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae) from Guizhou in China</mods:title>
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<mods:namePart id="2130E0E07516BE35A5CFF54757345F88">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="C66CB24CAAC4FF63482C2760A0514B1C">Feng, Zegang</mods:namePart>
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<mods:namePart id="5DF5ECA6D47C26B981165A53293DE31E">Zhang, Feng</mods:namePart>
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<treatment id="03DA1A1EFFACFF98FF053900E4AF3C46" ID-DOI="http://doi.org/10.5281/zenodo.8091594" ID-GBIF-Taxon="210479507" ID-Zenodo-Dep="8091594" LSID="urn:lsid:plazi:treatment:03DA1A1EFFACFF98FF053900E4AF3C46" httpUri="http://treatment.plazi.org/id/03DA1A1EFFACFF98FF053900E4AF3C46" lastPageId="60" lastPageNumber="61" pageId="54" pageNumber="55">
<subSubSection id="C369F883FFACFF92FF053900E2233A4A" box="[151,784,857,884]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="8BCCAB08FFACFF92FF053900E2233A4A" blockId="54.[151,784,857,919]" box="[151,784,857,884]" pageId="54" pageNumber="55">
<heading id="D0841C64FFACFF92FF053900E2233A4A" bold="true" box="[151,784,857,884]" fontSize="11" level="2" pageId="54" pageNumber="55" reason="4">
<emphasis id="B907771AFFACFF92FF053900E2233A4A" bold="true" box="[151,784,857,884]" pageId="54" pageNumber="55">
<taxonomicName id="4C73D08BFFACFF92FF053900E0C03A4F" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,499,857,884]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFACFF92FF053900E0C03A4F" bold="true" box="[151,499,857,884]" italics="true" pageId="54" pageNumber="55">Lagynochthonius zhijinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234CA61FFACFF92FE693900E36B3A48" box="[507,600,857,883]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
(ṂĤṄṁae)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369F883FFACFF92FF053924E0273AAC" box="[151,276,893,919]" pageId="54" pageNumber="55" type="description">
<paragraph id="8BCCAB08FFACFF92FF053924E0273AAC" blockId="54.[151,784,857,919]" box="[151,276,893,919]" pageId="54" pageNumber="55">
<figureCitation id="1348B78DFFACFF92FF053924E0273AAC" box="[151,276,893,919]" captionStart-0="FIGURE 44" captionStart-1="FIGURE 45" captionStart-2="FIGURE 46" captionStart-3="FIGURE 47" captionStartId-0="55.[151,250,1764,1788]" captionStartId-1="56.[151,250,1411,1435]" captionStartId-2="57.[151,250,1623,1647]" captionStartId-3="58.[151,250,1906,1930]" captionTargetBox-0="[151,1436,777,1741]" captionTargetBox-1="[311,1277,181,1387]" captionTargetBox-2="[178,1436,181,1600]" captionTargetBox-3="[184,1403,181,1880]" captionTargetId-0="figure-308@55.[151,1436,777,1741]" captionTargetId-1="figure-20@56.[311,1277,181,1387]" captionTargetId-2="figure-18@57.[151,1436,181,1600]" captionTargetId-3="figure-20@58.[184,1403,181,1882]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionTargetPageId-2="57" captionTargetPageId-3="58" captionText-0="FIGURE 44. Long Cave, type locality of Lagynochthonius zhijinensis sp. nov., A. Entrance; B. Inside the cave entrance; C. A narrow tunnel to the interior of the cave; D. Area where L. zhijinensis sp. nov. specimens were collected." captionText-1="FIGURE 45. Lagynochthonius zhijinensis sp. nov., holotype female, habitus (dorsal view)." captionText-2="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." captionText-3="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.8090206" figureDoi-1="http://doi.org/10.5281/zenodo.8090208" figureDoi-2="http://doi.org/10.5281/zenodo.8090210" figureDoi-3="http://doi.org/10.5281/zenodo.8090212" httpUri-0="https://zenodo.org/record/8090206/files/figure.png" httpUri-1="https://zenodo.org/record/8090208/files/figure.png" httpUri-2="https://zenodo.org/record/8090210/files/figure.png" httpUri-3="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">Figs 4447</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C369F883FFACFF92FF05399CE04D3D70" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="8BCCAB08FFACFF92FF05399CE04D3D70" blockId="54.[151,1437,965,2036]" pageId="54" pageNumber="55">
<emphasis id="B907771AFFACFF92FF05399CE0743AE4" bold="true" box="[151,327,965,991]" pageId="54" pageNumber="55">Type material.</emphasis>
<materialsCitation id="3B1BA155FFACFF92FEC2399CE3C23D1C" ID-GBIF-Occurrence="4138993330" collectingDate="2019-10-02" collectorName="Zegang Feng &amp; Lingchen Zhao" country="China" county="Zhijin County" elevation="1345" latitude="26.67934" location="Yangliu Village" longLatPrecision="1" longitude="105.78786" municipality="Chengguan Town" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="54C815AAFFACFF92FEC2399CE08B3AE4" box="[336,440,965,991]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
<specimenCount id="9D756081FFACFF92FE50399FE0E43ADB" box="[450,471,966,992]" pageId="54" pageNumber="55" type="female"></specimenCount>
(Ps.-MHBU-GZZJ-19-04-01):
<collectingCountry id="F364EB98FFACFF92FCAF399CE2B03AE4" box="[829,899,965,991]" name="China" pageId="54" pageNumber="55">China</collectingCountry>
,
<collectingRegion id="49B765EAFFACFF92FC02399CE5533AE4" box="[912,1120,965,991]" country="China" name="Guizhou" pageId="54" pageNumber="55">Guizhou Province</collectingRegion>
,
<collectingCounty id="62ADD384FFACFF92FBFC399CE4233AE4" box="[1134,1296,965,991]" pageId="54" pageNumber="55">Zhijin County</collectingCounty>
,
<collectingMunicipality id="6BA83172FFACFF92FA8F399CE1EA3D3F" pageId="54" pageNumber="55">Chengguan Town</collectingMunicipality>
,
<location id="8EACFDD3FFACFF92FF7639B0E0A63D38" LSID="urn:lsid:plazi:treatment:03DA1A1EFFACFF98FF053900E4AF3C46:8EACFDD3FFACFF92FF7639B0E0A63D38" box="[228,405,1001,1027]" country="China" county="Zhijin County" latitude="26.67934" longLatPrecision="1" longitude="105.78786" municipality="Chengguan Town" name="Yangliu Village" pageId="54" pageNumber="55" stateProvince="Guizhou">Yangliu Village</location>
,
<location id="8EACFDD3FFACFF92FE3239B3E32D3D38" LSID="urn:lsid:plazi:treatment:03DA1A1EFFACFF98FF053900E4AF3C46:8EACFDD3FFACFF92FE3239B3E32D3D38" box="[416,542,1001,1028]" country="China" county="Zhijin County" latitude="26.67934" longLatPrecision="1" longitude="105.78786" municipality="Chengguan Town" name="Long Cave" pageId="54" pageNumber="55" stateProvince="Guizhou">Long Cave</location>
, under stones in the deep zone [
<geoCoordinate id="EE47CDCFFFACFF92FC1039B2E51C3D3D" box="[898,1071,1003,1030]" degrees="26" direction="north" minutes="40" orientation="latitude" pageId="54" pageNumber="55" precision="1" seconds="45.63" value="26.67934">26°4045.63″N</geoCoordinate>
,
<geoCoordinate id="EE47CDCFFFACFF92FBA939B2E5C03D3D" box="[1083,1267,1003,1030]" degrees="105" direction="east" minutes="47" orientation="longitude" pageId="54" pageNumber="55" precision="1" seconds="16.30" value="105.78786">105°4716.30″E</geoCoordinate>
],
<elevation id="005E4C3BFFACFF92FA9739B0E4A43D38" box="[1285,1431,1001,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="54" pageNumber="55" unit="m" value="1345.0">
<quantity id="4C8B06EDFFACFF92FA9739B0E4693D3F" box="[1285,1370,1001,1028]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="54" pageNumber="55" unit="m" value="1345.0">1345 m</quantity>
a.s.l.
</elevation>
,
<date id="FFCD8DC8FFACFF92FF053E54E0783D1C" box="[151,331,1037,1063]" pageId="54" pageNumber="55" value="2019-10-02">
<collectingDate id="EF897420FFACFF92FF053E54E0783D1C" box="[151,331,1037,1063]" pageId="54" pageNumber="55" value="2019-10-02">2 October 2019</collectingDate>
</date>
,
<collectorName id="2686CEDEFFACFF92FECA3E57E0DF3D13" box="[344,492,1038,1064]" pageId="54" pageNumber="55">Zegang Feng</collectorName>
&amp;
<collectorName id="2686CEDEFFACFF92FD813E54E38C3D1C" box="[531,703,1037,1063]" pageId="54" pageNumber="55">Lingchen Zhao</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B1BA155FFACFF92FD6B3E57E0493D70" ID-GBIF-Occurrence="4138993322" collectingDate="2019-10-02" collectorName="Zegang Feng &amp; Lingchen Zhao" country="China" county="Zhijin County" elevation="1345" latitude="26.67934" location="Yangliu Village" longLatPrecision="1" longitude="105.78786" municipality="Chengguan Town" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="54C815AAFFACFF92FD6B3E57E26C3D13" box="[761,863,1038,1064]" pageId="54" pageNumber="55" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D756081FFACFF92FCF93E54E2A43D13" box="[875,919,1037,1064]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
(Ps.-MSWU-GZZJ-19-04-02), with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FFACFF92FF553E0CE29F3D54" box="[199,940,1109,1136]" pageId="54" pageNumber="55" type="etymology">
<paragraph id="8BCCAB08FFACFF92FF553E0CE29F3D54" blockId="54.[151,1437,965,2036]" box="[199,940,1109,1136]" pageId="54" pageNumber="55">
<emphasis id="B907771AFFACFF92FF553E0CE07D3D54" bold="true" box="[199,334,1109,1135]" pageId="54" pageNumber="55">Etymology.</emphasis>
Named after the Zhijin County, near the
<typeStatus id="54C815AAFFACFF92FC893E0FE2783D4B" box="[795,843,1110,1136]" pageId="54" pageNumber="55">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FFACFF92FF553E20E2433C7C" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="8BCCAB08FFACFF92FF553E20E2433C7C" blockId="54.[151,1437,965,2036]" pageId="54" pageNumber="55">
<emphasis id="B907771AFFACFF92FF553E20E0433DAC" bold="true" box="[199,368,1145,1175]" pageId="54" pageNumber="55">
Diagnosis (
<specimenCount id="9D756081FFACFF92FEDE3E22E0523DAC" box="[332,353,1147,1175]" pageId="54" pageNumber="55" type="female"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites IIIII each with two setae, tergites IVVII each with four setae. Pedipalps slender, femur 8.478.53 times longer than broad; chela 7.467.71 times longer than broad; only fixed chelal finger with intercalary teeth and a modified accessory tooth (
<emphasis id="B907771AFFACFF92FD623F50E2353C18" box="[752,774,1289,1315]" italics="true" pageId="54" pageNumber="55">td</emphasis>
) on retrolateral face; chelal fingers slightly curved in dorsal view; chemosensory setae (
<emphasis id="B907771AFFACFF92FE5E3F77E0D03C7C" box="[460,483,1326,1351]" italics="true" pageId="54" pageNumber="55">sc</emphasis>
) present on dorsum of chelal hand.
</paragraph>
</subSubSection>
<subSubSection id="C369F883FFACFF9CFF553F08E5853E83" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="description">
<paragraph id="8BCCAB08FFACFF92FF553F08E5123C57" blockId="54.[151,1437,965,2036]" box="[199,1057,1361,1388]" pageId="54" pageNumber="55">
<emphasis id="B907771AFFACFF92FF553F08E06A3C50" bold="true" box="[199,345,1361,1387]" pageId="54" pageNumber="55">Description.</emphasis>
Females (
<typeStatus id="54C815AAFFACFF92FE593F08E31C3C50" box="[459,559,1361,1387]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
and
<typeStatus id="54C815AAFFACFF92FDF43F08E3F83C50" box="[614,715,1361,1387]" pageId="54" pageNumber="55" type="paratype">paratype</typeStatus>
), male unknown (
<figureCitation id="1348B78DFFACFF92FC003F08E5253C57" box="[914,1046,1361,1388]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStart-2="FIGURE 47" captionStartId-0="56.[151,250,1411,1435]" captionStartId-1="57.[151,250,1623,1647]" captionStartId-2="58.[151,250,1906,1930]" captionTargetBox-0="[311,1277,181,1387]" captionTargetBox-1="[178,1436,181,1600]" captionTargetBox-2="[184,1403,181,1880]" captionTargetId-0="figure-20@56.[311,1277,181,1387]" captionTargetId-1="figure-18@57.[151,1436,181,1600]" captionTargetId-2="figure-20@58.[184,1403,181,1882]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionTargetPageId-2="58" captionText-0="FIGURE 45. Lagynochthonius zhijinensis sp. nov., holotype female, habitus (dorsal view)." captionText-1="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." captionText-2="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.8090208" figureDoi-1="http://doi.org/10.5281/zenodo.8090210" figureDoi-2="http://doi.org/10.5281/zenodo.8090212" httpUri-0="https://zenodo.org/record/8090208/files/figure.png" httpUri-1="https://zenodo.org/record/8090210/files/figure.png" httpUri-2="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">Figs 4547</figureCitation>
).
</paragraph>
<paragraph id="8BCCAB08FFACFF92FF553F2CE5DC3CB4" blockId="54.[151,1437,965,2036]" box="[199,1263,1397,1424]" pageId="54" pageNumber="55">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="8BCCAB08FFACFF92FF553FC0E5513FEF" blockId="54.[151,1437,965,2036]" pageId="54" pageNumber="55">
Cephalothorax (
<figureCitation id="1348B78DFFACFF92FEED3FC0E0DF3C88" box="[383,492,1433,1459]" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="54" pageNumber="55">Figs 46C</figureCitation>
,
<figureCitation id="1348B78DFFACFF92FE6E3FC0E3023C88" box="[508,561,1433,1459]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">47A</figureCitation>
): carapace 0.930.98 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, 3, II 4, III 5, IV 56; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa with small, rounded anteromedial process; coxae II with 1112 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="1348B78DFFACFF92FE543CE0E31E3FE8" box="[454,557,1721,1747]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">Fig. 47C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="8BCCAB08FFACFF92FF553C84E3CE3EC8" blockId="54.[151,1437,965,2036]" pageId="54" pageNumber="55">
Chelicera (
<figureCitation id="1348B78DFFACFF92FED73C84E0803FCC" box="[325,435,1757,1783]" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="54" pageNumber="55">Figs 46D</figureCitation>
,
<figureCitation id="1348B78DFFACFF92FE503C84E0C53FCC" box="[450,502,1757,1783]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">47B</figureCitation>
): large, about as long as carapace, 2.292.31 times longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 2024 teeth, distal one largest; movable finger with 1819 retrorse contiguous small teeth; galea absent (
<figureCitation id="1348B78DFFACFF92FB583D34E4073EBC" box="[1226,1332,1901,1927]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">Fig. 47B</figureCitation>
). Serrula exterior with 2021 blades and serrula interior with 1315 blades. Rallum with seven blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="1348B78DFFACFF92FD1A3D80E3C23EC8" box="[648,753,2009,2035]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="54" pageNumber="55">Fig. 47D</figureCitation>
).
</paragraph>
<paragraph id="8BCCAB08FFADFF93FF553ACEE0263BCA" blockId="55.[151,1437,150,753]" pageId="55" pageNumber="56">
Pedipalp (
<figureCitation id="1348B78DFFADFF93FEAA3ACEE0EC398A" box="[312,479,151,177]" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="55" pageNumber="56">Figs 46AB, F</figureCitation>
,
<figureCitation id="1348B78DFFADFF93FE783ACEE30C398A" box="[490,575,151,177]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="55" pageNumber="56">47EG</figureCitation>
): long and slender, trochanter 1.251.32, femur 8.478.53, patella 2.212.37, chela 7.467.71, hand 3.043.17 times longer than broad; femur 2.843.02 times longer than patella; movable chelal finger 1.421.45 times longer than hand and 0.580.59 times longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (
<figureCitation id="1348B78DFFADFF93FDC23B5AE3873826" box="[592,692,259,285]" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="55" pageNumber="56">Figs 46F</figureCitation>
,
<figureCitation id="1348B78DFFADFF93FD2C3B5AE3DC3826" box="[702,751,259,285]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="55" pageNumber="56">47E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
<emphasis id="B907771AFFADFF93FABB3B12E40C385E" box="[1321,1343,331,357]" italics="true" pageId="55" pageNumber="56">ib</emphasis>
and
<emphasis id="B907771AFFADFF93FAEE3B12E4AE385E" box="[1404,1437,331,357]" italics="true" pageId="55" pageNumber="56">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="B907771AFFADFF93FCC13B36E25E38B2" box="[851,877,367,393]" italics="true" pageId="55" pageNumber="56">eb</emphasis>
,
<emphasis id="B907771AFFADFF93FCE93B36E29338B2" box="[891,928,367,393]" italics="true" pageId="55" pageNumber="56">esb</emphasis>
and
<emphasis id="B907771AFFADFF93FC443B36E2C338B3" box="[982,1008,367,392]" italics="true" pageId="55" pageNumber="56">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis id="B907771AFFADFF93FE063BCAE0973897" box="[404,420,403,428]" italics="true" pageId="55" pageNumber="56">it</emphasis>
slightly distal to
<emphasis id="B907771AFFADFF93FDFB3BCAE3BB3897" box="[617,648,403,428]" italics="true" pageId="55" pageNumber="56">est</emphasis>
, situated subdistally;
<emphasis id="B907771AFFADFF93FCEC3BCAE2A13897" box="[894,914,403,428]" italics="true" pageId="55" pageNumber="56">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="B907771AFFADFF93FE153BEEE09238EA" box="[391,417,439,465]" italics="true" pageId="55" pageNumber="56">dx</emphasis>
situated distal to
<emphasis id="B907771AFFADFF93FDF83BEEE34D38EB" box="[618,638,439,464]" italics="true" pageId="55" pageNumber="56">et</emphasis>
;
<emphasis id="B907771AFFADFF93FD1C3BEEE39438EA" box="[654,679,439,465]" italics="true" pageId="55" pageNumber="56">sb</emphasis>
situated midway between
<emphasis id="B907771AFFADFF93FC473BEEE2D038EA" box="[981,995,439,465]" italics="true" pageId="55" pageNumber="56">b</emphasis>
and
<emphasis id="B907771AFFADFF93FB893BEEE51D38EB" box="[1051,1070,439,464]" italics="true" pageId="55" pageNumber="56">st</emphasis>
;
<emphasis id="B907771AFFADFF93FBAC3BEEE57F38EA" box="[1086,1100,439,465]" italics="true" pageId="55" pageNumber="56">b</emphasis>
and
<emphasis id="B907771AFFADFF93FB163BEEE5BF38EB" box="[1156,1164,439,464]" italics="true" pageId="55" pageNumber="56">t</emphasis>
situated subdistally and situated at the same level as
<emphasis id="B907771AFFADFF93FE473B82E0C738CF" box="[469,500,475,500]" italics="true" pageId="55" pageNumber="56">est</emphasis>
and
<emphasis id="B907771AFFADFF93FDB83B82E30938CF" box="[554,570,475,500]" italics="true" pageId="55" pageNumber="56">it</emphasis>
, respectively (
<figureCitation id="1348B78DFFADFF93FD4C3B82E27138CE" box="[734,834,475,501]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="55" pageNumber="56">Fig. 47F</figureCitation>
). A tiny retrolateral lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="B907771AFFADFF93FE683BA6E3273B23" box="[506,532,511,536]" italics="true" pageId="55" pageNumber="56">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 23 macrodenticles, slightly retrorse and pointed, plus 11
<specimenCount id="9D756081FFADFF93FC77381EE5B33B5A" box="[997,1152,583,609]" pageId="55" pageNumber="56" type="intercalary">12 intercalary</specimenCount>
microdenticles,
<quantity id="4C8B06EDFFADFF93FAA8381FE4AF3B5A" box="[1338,1436,582,609]" metricMagnitude="-1" metricUnit="m" metricValue="8.763" metricValueMax="8.89" metricValueMin="8.636" pageId="55" pageNumber="56" unit="in" value="34.5" valueMax="35.0" valueMin="34.0">3435 in</quantity>
total; movable chelal finger with 2122 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 78 vestigial, rounded and contiguous basal teeth,
<quantity id="4C8B06EDFFADFF93FB8F38D6E5B73B92" box="[1053,1156,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="7.62" metricValueMin="7.112" pageId="55" pageNumber="56" unit="in" value="29.0" valueMax="30.0" valueMin="28.0">2830 in</quantity>
total (
<figureCitation id="1348B78DFFADFF93FB4638D6E40F3B92" box="[1236,1340,655,681]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="55" pageNumber="56">Fig. 47F</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="1348B78DFFADFF93FAF938EAE1FB3BCA" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="55" pageNumber="56">Figs 46B</figureCitation>
,
<figureCitation id="1348B78DFFADFF93FF46388EE03B3BCA" box="[212,264,727,753]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="55" pageNumber="56">47G</figureCitation>
).
</paragraph>
<caption id="DF0CFB80FFADFF93FF053CBDE58C3E1B" ID-DOI="http://doi.org/10.5281/zenodo.8090206" ID-Zenodo-Dep="8090206" httpUri="https://zenodo.org/record/8090206/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1764,1788]" targetBox="[151,1436,777,1741]" targetPageId="55" targetType="figure">
<paragraph id="8BCCAB08FFADFF93FF053CBDE58C3E1B" blockId="55.[151,1436,1764,1824]" pageId="55" pageNumber="56">
<emphasis id="B907771AFFADFF93FF053CBDE0133FC7" bold="true" box="[151,288,1764,1788]" pageId="55" pageNumber="56">FIGURE 44.</emphasis>
Long Cave, type locality of
<taxonomicName id="4C73D08BFFADFF93FDD63CBDE2573FC7" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[580,868,1764,1788]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFADFF93FDD63CBDE2573FC7" box="[580,868,1764,1788]" italics="true" pageId="55" pageNumber="56">Lagynochthonius zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFADFF93FCF93CBDE28F3FC7" bold="true" box="[875,956,1764,1788]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A234CA61FFADFF93FCF93CBDE28F3FC7" box="[875,956,1764,1788]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. A narrow tunnel to the interior of the cave; D. Area where
<taxonomicName id="4C73D08BFFADFF93FD433D50E2643E1B" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[721,855,1800,1824]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFADFF93FD433D50E2643E1B" box="[721,855,1800,1824]" italics="true" pageId="55" pageNumber="56">L. zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFADFF93FCCF3D51E29D3E1B" bold="true" box="[861,942,1800,1824]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A234CA61FFADFF93FCCF3D51E29D3E1B" box="[861,942,1800,1824]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected.
</paragraph>
</caption>
<paragraph id="8BCCAB08FFADFF93FF553D17E55B3EEF" blockId="55.[151,1437,1870,2005]" pageId="55" pageNumber="56">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy XII: 23: 2: 2: 4: 4: 4: 4: 5: 5: 2: T2T: 0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IVXII: 13: 9: 910: 89: 9: 9: 78: 0: 2. Anterior genital operculum with 910 setae, posterior margin with 1214 marginal setae,
<quantity id="4C8B06EDFFADFF93FCD93DE2E29D3EEF" box="[843,942,1978,2005]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.096" metricValueMin="5.334" pageId="55" pageNumber="56" unit="in" value="22.5" valueMax="24.0" valueMin="21.0">2124 in</quantity>
total (
<figureCitation id="1348B78DFFADFF93FC643DE3E56F3EEF" box="[1014,1116,1978,2004]" captionStart="FIGURE 46" captionStartId="57.[151,250,1623,1647]" captionTargetBox="[178,1436,181,1600]" captionTargetId="figure-18@57.[151,1436,181,1600]" captionTargetPageId="57" captionText="FIGURE 46. Lagynochthonius zhijinensis sp. nov., holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="55" pageNumber="56">Fig. 46E</figureCitation>
).
</paragraph>
<caption id="DF0CFB80FFA2FF9CFF053FDAE5043CA0" ID-DOI="http://doi.org/10.5281/zenodo.8090208" ID-Zenodo-Dep="8090208" box="[151,1079,1411,1436]" httpUri="https://zenodo.org/record/8090208/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1411,1435]" targetBox="[311,1277,181,1387]" targetPageId="56" targetType="figure">
<paragraph id="8BCCAB08FFA2FF9CFF053FDAE5043CA0" blockId="56.[151,1079,1411,1436]" box="[151,1079,1411,1436]" pageId="56" pageNumber="57">
<emphasis id="B907771AFFA2FF9CFF053FDAE0133CA0" bold="true" box="[151,288,1411,1435]" pageId="56" pageNumber="57">FIGURE 45.</emphasis>
<taxonomicName id="4C73D08BFFA2FF9CFEB43FDAE3753CA0" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[294,582,1411,1435]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFA2FF9CFEB43FDAE3753CA0" box="[294,582,1411,1435]" italics="true" pageId="56" pageNumber="57">Lagynochthonius zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA2FF9CFDDE3FDDE3AE3CA7" bold="true" box="[588,669,1412,1436]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A234CA61FFA2FF9CFDDE3FDDE3AE3CA7" box="[588,669,1412,1436]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="8BCCAB08FFA2FF9CFF553F93E5343FA3" blockId="56.[151,1437,1482,2012]" pageId="56" pageNumber="57">
Legs (
<figureCitation id="1348B78DFFA2FF9CFE9F3F93E0BD3CDF" box="[269,398,1482,1508]" captionStart="FIGURE 47" captionStartId="58.[151,250,1906,1930]" captionTargetBox="[184,1403,181,1880]" captionTargetId="figure-20@58.[184,1403,181,1882]" captionTargetPageId="58" captionText="FIGURE 47. Lagynochthonius zhijinensis sp. nov., holotype female:A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="56" pageNumber="57">Fig. 47HI</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg 1.761.79 times longer than patella and with one lyrifissure at the base of femur; tarsus 2.482.56 times longer than tibia. Femoropatella of leg IV 4.124.13 times longer than deep; tibia 6.276.36 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.56 times longer than deep (TS = 0.280.34), tarsus 14.1714.33 times longer than deep and 2.662.69 times longer than metatarsus (TS = 0.310.34). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="8BCCAB08FFA2FF9CFF553CFBE5853E83" blockId="56.[151,1437,1482,2012]" pageId="56" pageNumber="57">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Females: body length 2.172.36. Pedipalps: trochanter 0.25/0.190.20 (1.251.32), femur 1.271.28/0.15 (8.478.53), patella 0.420.45/0.19 (2.212.37), chela 1.791.85/0.24 (7.467.71), hand 0.730.76/0.24 (3.043.17), movable chelal finger length 1.061.08. Chelicera 0.710.74/0.310.32 (2.292.31), movable finger length 0.370.38. Carapace 0.620.65/0.660.67 (0.930.98). Leg : trochanter 0.190.21/0.16 (1.191.31), femur 0.700.72/0.09 (7.788.00), patella 0.390.41/0.080.09 (4.564.88), tibia 0.310.32/0.06 (5.175.33), tarsus 0.770.82/0.060.07 (11.71 12.83). Leg IV: trochanter 0.280.29/0.160.18 (1.611.75), femoropatella 0.991.03/0.240.25 (4.124.13), tibia 0.690.70/0.11 (6.276.36), metatarsus 0.32/0.09 (3.56), tarsus 0.850.86/0.06 (14.1714.33).</paragraph>
</subSubSection>
<subSubSection id="C369F883FFA2FF9CFF553D98E3C23EE7" box="[199,753,1985,2012]" pageId="56" pageNumber="57" type="distribution">
<paragraph id="8BCCAB08FFA2FF9CFF553D98E3C23EE7" blockId="56.[151,1437,1482,2012]" box="[199,753,1985,2012]" pageId="56" pageNumber="57">
<emphasis id="B907771AFFA2FF9CFF553D98E0523EE0" bold="true" box="[199,353,1985,2011]" pageId="56" pageNumber="57">Distribution.</emphasis>
Known only from the
<typeStatus id="54C815AAFFA2FF9CFDF23D9BE3A33EE7" box="[608,656,1986,2012]" pageId="56" pageNumber="57">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF0CFB80FFA3FF9DFF053C0EE02F3F83" ID-DOI="http://doi.org/10.5281/zenodo.8090210" ID-Zenodo-Dep="8090210" httpUri="https://zenodo.org/record/8090210/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1623,1647]" targetBox="[178,1436,181,1600]" targetPageId="57" targetType="figure">
<paragraph id="8BCCAB08FFA3FF9DFF053C0EE02F3F83" blockId="57.[151,1436,1623,1720]" pageId="57" pageNumber="58">
<emphasis id="B907771AFFA3FF9DFF053C0EE0123F4B" bold="true" box="[151,289,1623,1648]" pageId="57" pageNumber="58">FIGURE 46.</emphasis>
<taxonomicName id="4C73D08BFFA3FF9DFEB53C01E37B3F4B" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[295,584,1624,1648]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFA3FF9DFEB53C01E37B3F4B" box="[295,584,1624,1648]" italics="true" pageId="57" pageNumber="58">Lagynochthonius zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA3FF9DFDDD3C01E3933F4B" bold="true" box="[591,672,1624,1648]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A234CA61FFA3FF9DFDDD3C01E3933F4B" box="[591,672,1624,1648]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Female genital area (ventral view); F. Left pedipalp (minus chela, dorsal view).
</paragraph>
</caption>
<caption id="DF0CFB80FFA0FF9EFF053D2BE27A3ECD" ID-DOI="http://doi.org/10.5281/zenodo.8090212" ID-Zenodo-Dep="8090212" httpUri="https://zenodo.org/record/8090212/files/figure.png" pageId="58" pageNumber="59" startId="58.[151,250,1906,1930]" targetBox="[184,1403,181,1880]" targetPageId="58" targetType="figure">
<paragraph id="8BCCAB08FFA0FF9EFF053D2BE27A3ECD" blockId="58.[151,1436,1906,2038]" pageId="58" pageNumber="59">
<emphasis id="B907771AFFA0FF9EFF053D2BE02C3EB1" bold="true" box="[151,287,1906,1930]" pageId="58" pageNumber="59">FIGURE 47.</emphasis>
<taxonomicName id="4C73D08BFFA0FF9EFEB63D2BE3703EB1" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[292,579,1906,1930]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFA0FF9EFEB63D2BE3703EB1" box="[292,579,1906,1930]" italics="true" pageId="58" pageNumber="59">Lagynochthonius zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA0FF9EFDDA3D2BE3A43EB1" bold="true" box="[584,663,1906,1930]" pageId="58" pageNumber="59">
<taxonomicNameLabel id="A234CA61FFA0FF9EFDDA3D2BE3A43EB1" box="[584,663,1906,1930]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype female: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and trichobothrial pattern; G. Left chela (dorsal view); H. Leg (lateral view); I. Leg IV (lateral view). Scale bars: 0.20 mm (AB, EI); 0.10 mm (CD).
</paragraph>
</caption>
<subSubSection id="C369F883FFA1FF98FF053ACEE4AF3C46" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="key">
<paragraph id="8BCCAB08FFA1FF9FFF053ACEE239398A" blockId="59.[151,778,151,178]" box="[151,778,151,178]" pageId="59" pageNumber="60">
<emphasis id="B907771AFFA1FF9FFF053ACEE239398A" bold="true" box="[151,778,151,178]" pageId="59" pageNumber="60">
Key to the species of
<taxonomicName id="4C73D08BFFA1FF9FFE0F3ACEE3423989" ID-CoL="62VPZ" authorityName="Beier" authorityYear="1951" box="[413,625,151,178]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B907771AFFA1FF9FFE0F3ACEE3423989" bold="true" box="[413,625,151,178]" italics="true" pageId="59" pageNumber="60">Lagynochthonius</emphasis>
</taxonomicName>
from
<collectingCountry id="F364EB98FFA1FF9FFD2F3ACEE239398A" box="[701,778,151,177]" name="China" pageId="59" pageNumber="60">China</collectingCountry>
</emphasis>
</paragraph>
<key id="FFE2FC39FFA1FF98FF053A85E4AF3C46" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<keyStep id="3087B3ADFFA1FF9FFF053A85E4AF3834" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053A85E4AF39CA" blockId="59.[151,1437,220,2031]" box="[151,1436,220,242]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053A85E4AF39CA" box="[151,1436,220,242]" pageId="59" pageNumber="60">
1 Carapace without antero-median setae; trichobothrium
<emphasis id="B907771AFFA1FF9FFD663A85E23A39CA" box="[756,777,220,241]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
closer to
<emphasis id="B907771AFFA1FF9FFCF53A85E24039CA" box="[871,883,220,241]" italics="true" pageId="59" pageNumber="60">b</emphasis>
than to
<emphasis id="B907771AFFA1FF9FFC533A84E2E239C9" box="[961,977,221,242]" italics="true" pageId="59" pageNumber="60">st</emphasis>
..................................... 2
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053AA3E4AF3834" blockId="59.[151,1437,220,2031]" box="[151,1436,249,271]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053AA3E4AF3834" box="[151,1436,249,271]" pageId="59" pageNumber="60">
- Carapace with antero-median setae; trichobothrium
<emphasis id="B907771AFFA1FF9FFD473AA0E3D93835" box="[725,746,249,270]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
closer to
<emphasis id="B907771AFFA1FF9FFCDA3AA3E26B3834" box="[840,856,250,271]" italics="true" pageId="59" pageNumber="60">st</emphasis>
than to
<emphasis id="B907771AFFA1FF9FFC343AA0E2813835" box="[934,946,249,270]" italics="true" pageId="59" pageNumber="60">b</emphasis>
or midway between
<emphasis id="B907771AFFA1FF9FFBEE3AA3E5BF3834" box="[1148,1164,250,271]" italics="true" pageId="59" pageNumber="60">st</emphasis>
and
<emphasis id="B907771AFFA1FF9FFB283AA0E5F53835" box="[1210,1222,249,270]" italics="true" pageId="59" pageNumber="60">b</emphasis>
................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053B4EE4AF38BF" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053B4EE4AF3872" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053B4EE4AF3872" pageId="59" pageNumber="60">
2 Chela 7.03 (♂) times as long as broad (length
<quantity id="4C8B06EDFFA1FF9FFD0F3B4EE3C53817" box="[669,758,279,300]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" pageId="59" pageNumber="60" unit="mm" value="2.04">2.04 mm</quantity>
); epistome pointed and small, triangular............................................................................................
<taxonomicName id="4C73D08BFFA1FF9FFB873B6CE4AF3872" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1045,1436,308,330]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="xinjiaoensis">
<emphasis id="B907771AFFA1FF9FFB873B6CE5923871" box="[1045,1185,309,330]" italics="true" pageId="59" pageNumber="60">L. xinjiaoensis</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053B0BE4AF38BF" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053B0BE4AF38BF" pageId="59" pageNumber="60">
- Chela 6.446.46 (♂), 5.876.04 (
<specimenCount id="9D756081FFA1FF9FFDB83B08E309385D" box="[554,570,337,358]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.551.61 (♂), 1.691.76 (
<specimenCount id="9D756081FFA1FF9FFB113B08E5A0385D" box="[1155,1171,337,358]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); epistome small and obtuse, rounded.....................................................
<taxonomicName id="4C73D08BFFA1FF9FFC683B36E4AF38BF" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1018,1436,367,389]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="magnidentatus">
<emphasis id="B907771AFFA1FF9FFC683B36E59238BF" box="[1018,1185,367,388]" italics="true" pageId="59" pageNumber="60">L. magnidentatus</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053BD5E4AF3884" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053BD5E4AF389A" blockId="59.[151,1437,220,2031]" box="[151,1436,396,418]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053BD5E4AF389A" box="[151,1436,396,418]" pageId="59" pageNumber="60">3 Carapace without eyes or eyespots........................................................................ 4</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053BF3E4AF3884" blockId="59.[151,1437,220,2031]" box="[151,1436,425,447]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053BF3E4AF3884" box="[151,1436,425,447]" pageId="59" pageNumber="60">- Carapace with eyes or eyespots......................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053B9EE4AF38C2" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053B9EE4AF38E6" blockId="59.[151,1437,220,2031]" box="[151,1436,455,477]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053B9EE4AF38E6" box="[151,1436,455,477]" pageId="59" pageNumber="60">4 At least one chela finger with intercalary teeth.............................................................. 5</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053BBDE4AF38C2" blockId="59.[151,1437,220,2031]" box="[151,1436,484,506]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053BBDE4AF38C2" box="[151,1436,484,506]" pageId="59" pageNumber="60">- Both chelal fingers without intercalary teeth............................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053858E4AF3B0F" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053858E4AF3B2C" blockId="59.[151,1437,220,2031]" box="[151,1436,513,535]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053858E4AF3B2C" box="[151,1436,513,535]" pageId="59" pageNumber="60">5 Only fixed chelal finger with intercalary teeth............................................................... 6</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053846E4AF3B0F" blockId="59.[151,1437,220,2031]" box="[151,1436,543,565]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053846E4AF3B0F" box="[151,1436,543,565]" pageId="59" pageNumber="60">- Both chelal fingers with intercalary teeth................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053865E4AF3B54" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053865E4AF3B69" blockId="59.[151,1437,220,2031]" box="[151,1436,572,594]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053865E4AF3B69" box="[151,1436,572,594]" pageId="59" pageNumber="60">
6 Trichobothrium
<emphasis id="B907771AFFA1FF9FFEED3865E0A73B6A" box="[383,404,572,593]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
closer to
<emphasis id="B907771AFFA1FF9FFE603864E3313B69" box="[498,514,573,594]" italics="true" pageId="59" pageNumber="60">st</emphasis>
than to
<emphasis id="B907771AFFA1FF9FFDC23865E36F3B6A" box="[592,604,572,593]" italics="true" pageId="59" pageNumber="60">b</emphasis>
; tergite with four setae............................................... 7
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053803E4AF3B54" blockId="59.[151,1437,220,2031]" box="[151,1436,601,623]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053803E4AF3B54" box="[151,1436,601,623]" pageId="59" pageNumber="60">
- Trichobothrium
<emphasis id="B907771AFFA1FF9FFEED3800E0A73B55" box="[383,404,601,622]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA1FF9FFD063803E3973B54" box="[660,676,602,623]" italics="true" pageId="59" pageNumber="60">st</emphasis>
and
<emphasis id="B907771AFFA1FF9FFD413800E3EC3B55" box="[723,735,601,622]" italics="true" pageId="59" pageNumber="60">b</emphasis>
; tergite with two setae.....................
<taxonomicName id="4C73D08BFFA1FF9FFB593803E47A3B55" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1227,1353,601,623]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="zhijinensis" status="sp. nov.">
<emphasis id="B907771AFFA1FF9FFB593803E47A3B55" box="[1227,1353,601,623]" italics="true" pageId="59" pageNumber="60">L. zhijinensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA1FF9FFAC23803E4AF3B54" bold="true" box="[1360,1436,602,623]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="A234CA61FFA1FF9FFAC23803E4AF3B54" box="[1360,1436,602,623]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF05382EE4AF3A3A" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF05382EE4AF3B92" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF05382EE4AF3B92" pageId="59" pageNumber="60">
7 The intercalary teeth round and obtuse; the teeth on movable chelal fingers retrorse and contiguous; chela length 1.571.61 (♂), 1.641.69 (
<specimenCount id="9D756081FFA1FF9FFEC338CDE0523B92" box="[337,353,660,681]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; movable chelal finger length 0.930.96 (♂), 0.991.01 (
<specimenCount id="9D756081FFA1FF9FFC2C38CDE2FD3B92" box="[958,974,660,681]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm....
<taxonomicName id="4C73D08BFFA1FF9FFBA338CCE4AF3B92" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1073,1436,660,682]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="retrorsus">
<emphasis id="B907771AFFA1FF9FFBA338CCE5923B91" box="[1073,1185,661,682]" italics="true" pageId="59" pageNumber="60">L. retrorsus</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF0538EBE4AF3A3A" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF0538EBE4AF3A3A" pageId="59" pageNumber="60">
- The intercalary teeth small and pointed; the teeth on movable chelal fingers pointed and well-spaced; chela length 2.072.16 (♂), 2.062.19 (
<specimenCount id="9D756081FFA1FF9FFEED3896E0BC3BDF" box="[383,399,719,740]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; movable chelal finger length 1.241.25 (♂), 1.201.26 (
<specimenCount id="9D756081FFA1FF9FFC7E3896E2CF3BDF" box="[1004,1020,719,740]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm..................................................................................................
<taxonomicName id="4C73D08BFFA1FF9FFB8838B4E4AF3A3A" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1050,1436,748,770]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="spinulentus">
<emphasis id="B907771AFFA1FF9FFB8838B4E5923A3A" box="[1050,1185,748,770]" italics="true" pageId="59" pageNumber="60">L. spinulentus</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053950E4AF3A07" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053950E4AF3A24" blockId="59.[151,1437,220,2031]" box="[151,1436,777,799]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053950E4AF3A24" box="[151,1436,777,799]" pageId="59" pageNumber="60">8 Tergites IIIII and each with four setae.................................................................. 9</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF05397EE4AF3A07" blockId="59.[151,1437,220,2031]" box="[151,1436,807,829]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF05397EE4AF3A07" box="[151,1436,807,829]" pageId="59" pageNumber="60">- Tergites IIIII and each with two setae.................................................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF05391DE4AF3D32" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF05391DE4AF3AAF" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF05391DE4AF3AAF" pageId="59" pageNumber="60">
9 Trichobothrium
<emphasis id="B907771AFFA1FF9FFE12391DE0A63A62" box="[384,405,836,857]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
closer to
<emphasis id="B907771AFFA1FF9FFE67391CE3363A61" box="[501,517,837,858]" italics="true" pageId="59" pageNumber="60">st</emphasis>
than to
<emphasis id="B907771AFFA1FF9FFDC7391DE3523A62" box="[597,609,836,857]" italics="true" pageId="59" pageNumber="60">b</emphasis>
; body length 1.42 (♂), 1.59 (
<specimenCount id="9D756081FFA1FF9FFC13391DE2A23A62" box="[897,913,836,857]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; chela 6.65 (♂), 6.63 (
<specimenCount id="9D756081FFA1FF9FFB37391DE5863A62" box="[1189,1205,836,857]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.33 (♂), 1.26 (
<specimenCount id="9D756081FFA1FF9FFE51393BE0E03A4C" box="[451,467,866,887]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); palpal femur 7.15 (♂), 7.42 (
<specimenCount id="9D756081FFA1FF9FFCA0393BE2713A4C" box="[818,834,866,887]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 0.93 (♂), 0.89 (
<specimenCount id="9D756081FFA1FF9FFA9E393BE42F3A4C" box="[1292,1308,866,887]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); coxae II with seven terminally indented coxal spines on each side.................................
<taxonomicName id="4C73D08BFFA1FF9FFB383926E4793AAF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1194,1354,895,916]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="longedentatus" status="sp. nov.">
<emphasis id="B907771AFFA1FF9FFB383926E4793AAF" box="[1194,1354,895,916]" italics="true" pageId="59" pageNumber="60">L. longedentatus</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA1FF9FFAC23926E4AF3AAF" bold="true" box="[1360,1436,895,916]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="A234CA61FFA1FF9FFAC23926E4AF3AAF" box="[1360,1436,895,916]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF0539C5E4AF3D32" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF0539C5E4AF3D32" pageId="59" pageNumber="60">
- Trichobothrium
<emphasis id="B907771AFFA1FF9FFEED39C5E0A73A8A" box="[383,404,924,945]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA1FF9FFD0139C4E3903A89" box="[659,675,925,946]" italics="true" pageId="59" pageNumber="60">st</emphasis>
and
<emphasis id="B907771AFFA1FF9FFD4339C5E3EE3A8A" box="[721,733,924,945]" italics="true" pageId="59" pageNumber="60">b</emphasis>
; body length 1.63 (♂), 1.812.20 (
<specimenCount id="9D756081FFA1FF9FFBBC39C5E50D3A8A" box="[1070,1086,924,945]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; chela 8.76 (♂), 8.308.32 (
<specimenCount id="9D756081FFA1FF9FFA1139C5E4A03A8A" box="[1411,1427,924,945]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.84 (♂), 1.831.91 (
<specimenCount id="9D756081FFA1FF9FFD4B39E3E3DA3AF4" box="[729,745,954,975]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); palpal femur 10.46 (♂), 10.0710.38 (
<specimenCount id="9D756081FFA1FF9FFB3439E3E5853AF4" box="[1190,1206,954,975]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.36 (♂), 1.351.41 (
<specimenCount id="9D756081FFA1FF9FFE6B398EE33A3AD7" box="[505,521,983,1004]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); coxae II with 911 terminally indented coxal spines on each side..................................................................................
<taxonomicName id="4C73D08BFFA1FF9FFC7B39ACE4AF3D32" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1001,1436,1012,1034]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="yaowangguensis">
<emphasis id="B907771AFFA1FF9FFC7B39ACE5923D31" box="[1001,1185,1013,1034]" italics="true" pageId="59" pageNumber="60">L. yaowangguensis</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053E48E4AF3D44" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053E48E4AF3D7F" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053E48E4AF3D7F" pageId="59" pageNumber="60">10 Chelal fingers straight or slightly curved in dorsal view; fixed chelal finger with max. 47 teeth, movable chelal finger with max. 42 teeth............................................................................................ 11</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053E15E4AF3D44" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053E15E4AF3D44" pageId="59" pageNumber="60">
- Chelal fingers markedly curved in dorsal view; fixed chelal finger with 5860 teeth, movable chelal finger with 6061 teeth..................................................................................
<taxonomicName id="4C73D08BFFA1FF9FFB233E33E4793D44" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1201,1354,1130,1151]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="guanniuensis" status="sp. nov.">
<emphasis id="B907771AFFA1FF9FFB233E33E4793D44" box="[1201,1354,1130,1151]" italics="true" pageId="59" pageNumber="60">L. guanniuensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA1FF9FFAC23E33E4AF3D44" bold="true" box="[1360,1436,1130,1151]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="A234CA61FFA1FF9FFAC23E33E4AF3D44" box="[1360,1436,1130,1151]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053EDEE4AF3DCF" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053EDEE4AF3D82" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053EDEE4AF3D82" pageId="59" pageNumber="60">
11 Palpal femur 8.078.93 (♂) times as long as broad (length
<quantity id="4C8B06EDFFA1FF9FFC853EDEE2953DA7" box="[791,934,1159,1181]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.25" metricValueMin="1.21" pageId="59" pageNumber="60" unit="mm" value="1.23" valueMax="1.25" valueMin="1.21">1.211.25 mm</quantity>
), chela length 1.681.76 (♂) mm; trichobothrium
<emphasis id="B907771AFFA1FF9FFA153EDEE4AF3DA7" box="[1415,1436,1159,1180]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
closer to
<emphasis id="B907771AFFA1FF9FFEA83EFCE0793D81" box="[314,330,1189,1210]" italics="true" pageId="59" pageNumber="60">st</emphasis>
than to
<emphasis id="B907771AFFA1FF9FFE0A3EFDE0973D82" box="[408,420,1188,1209]" italics="true" pageId="59" pageNumber="60">b</emphasis>
.................................................................................. 12
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053E9BE4AF3DCF" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053E9BE4AF3DCF" pageId="59" pageNumber="60">
- Palpal femur 7.577.64 (♂) times as long as broad (length
<quantity id="4C8B06EDFFA1FF9FFC853E98E2953DED" box="[791,934,1217,1239]" metricMagnitude="-3" metricUnit="m" metricValue="1.065" metricValueMax="1.07" metricValueMin="1.06" pageId="59" pageNumber="60" unit="mm" value="1.065" valueMax="1.07" valueMin="1.06">1.061.07 mm</quantity>
), chela length 1.531.54 (♂) mm; trichobothrium
<emphasis id="B907771AFFA1FF9FFA153E98E4AF3DED" box="[1415,1436,1217,1238]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA1FF9FFE4E3E86E0DF3DCF" box="[476,492,1247,1268]" italics="true" pageId="59" pageNumber="60">st</emphasis>
and
<emphasis id="B907771AFFA1FF9FFD883E86E3153DCF" box="[538,550,1247,1268]" italics="true" pageId="59" pageNumber="60">b</emphasis>
..............................................
<taxonomicName id="4C73D08BFFA1FF9FFBCA3E86E4AF3DCF" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1112,1436,1247,1269]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="fengi">
<emphasis id="B907771AFFA1FF9FFBCA3E86E5923DCF" box="[1112,1185,1247,1268]" italics="true" pageId="59" pageNumber="60">L. fengi</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053EA5E4AF3C9F" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053EA5E4AF3C77" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053EA5E4AF3C77" pageId="59" pageNumber="60">
12 Carapace markedly narrowed posteriorly; chela 8.388.84 (♂) times as long as broad, hand 3.383.58 (♂) times longer than broad; proximal half of movable chelal finger without vestigial teeth; trichobothrium
<emphasis id="B907771AFFA1FF9FFBBA3F40E50E3C15" box="[1064,1085,1305,1326]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
of movable chelal finger 1.221.24 times as far from
<emphasis id="B907771AFFA1FF9FFE193F6EE0A83C77" box="[395,411,1335,1356]" italics="true" pageId="59" pageNumber="60">st</emphasis>
as from
<emphasis id="B907771AFFA1FF9FFE623F6EE0CF3C77" box="[496,508,1335,1356]" italics="true" pageId="59" pageNumber="60">b</emphasis>
...............................................
<taxonomicName id="4C73D08BFFA1FF9FFBAB3F6EE4AF3C77" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1081,1436,1335,1357]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="B907771AFFA1FF9FFBAB3F6EE5923C77" box="[1081,1185,1335,1356]" italics="true" pageId="59" pageNumber="60">L. serratus</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053F0DE4AF3C9F" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053F0DE4AF3C9F" pageId="59" pageNumber="60">
- Carapace gently narrowed posteriorly; chela 7.00 (♂) times as long as broad, hand 2.92 (♂) times longer than broad; proximal half of movable chelal finger with six vestigial teeth; trichobothrium
<emphasis id="B907771AFFA1FF9FFC113F28E2AB3CBD" box="[899,920,1393,1414]" italics="true" pageId="59" pageNumber="60">sb</emphasis>
of movable chelal finger 1.67 times as far from
<emphasis id="B907771AFFA1FF9FFAE33F2BE4B23CBC" box="[1393,1409,1394,1415]" italics="true" pageId="59" pageNumber="60">st</emphasis>
as from
<emphasis id="B907771AFFA1FF9FFE853FD6E0103C9F" box="[279,291,1423,1444]" italics="true" pageId="59" pageNumber="60">b</emphasis>
...............................................................
<taxonomicName id="4C73D08BFFA1FF9FFB873FD6E4AF3C9F" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1045,1436,1423,1445]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="xiaolinensis">
<emphasis id="B907771AFFA1FF9FFB873FD6E5923C9F" box="[1045,1185,1423,1444]" italics="true" pageId="59" pageNumber="60">L. xiaolinensis</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053FF5E4AF3F21" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053FF5E4AF3CE4" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053FF5E4AF3CE4" pageId="59" pageNumber="60">
13 Troglomorphic species with a larger body size and elongate appendages: body length not less than 1.71 (♂,
<specimenCount id="9D756081FFA1FF9FFB483FF5E5D93CFA" box="[1242,1258,1452,1473]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm, chela length not less than 1.32 (♂,
<specimenCount id="9D756081FFA1FF9FFE243F93E0F53CE4" box="[438,454,1482,1503]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; without corneate eyes, only anterior pair of eyespots present.......................... 14
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053FBEE4AF3F21" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053FBEE4AF3F21" pageId="59" pageNumber="60">
- Epigean species with smaller body size and without elongate appendages: body length not more than 1.60 (♂,
<specimenCount id="9D756081FFA1FF9FFA843FBEE4153CC7" box="[1302,1318,1511,1532]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm, chela length not more than 1.00 (♂,
<specimenCount id="9D756081FFA1FF9FFD973C5DE3263F22" box="[517,533,1540,1561]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm; with corneate eyes, anterior pair of eyes well developed....................... 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053C78E4AF3FF2" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053C78E4AF3F4A" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053C78E4AF3F4A" pageId="59" pageNumber="60">
14 Chela 7.007.14 (♂), 6.406.46 (
<specimenCount id="9D756081FFA1FF9FFDB33C78E3023F0D" box="[545,561,1569,1590]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.541.57 (♂), 1.551.60 (
<specimenCount id="9D756081FFA1FF9FFBF73C78E5463F0D" box="[1125,1141,1569,1590]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); palpal femur 8.218.50 (♂), 7.507.87 (
<specimenCount id="9D756081FFA1FF9FFE133C66E0A23F6F" box="[385,401,1599,1620]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.151.19 (♂), 1.181.20 (
<specimenCount id="9D756081FFA1FF9FFC413C66E2D03F6F" box="[979,995,1599,1620]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); coxae II with 89 terminally indented coxal spines on each side...................................................
<taxonomicName id="4C73D08BFFA1FF9FFBD23C04E4AF3F4A" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1088,1436,1628,1650]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B907771AFFA1FF9FFBD23C04E5923F49" box="[1088,1185,1629,1650]" italics="true" pageId="59" pageNumber="60">L. crassus</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053C23E4AF3FF2" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053C23E4AF3FF2" pageId="59" pageNumber="60">
- Chela 7.337.67 (♂), 6.716.82 (
<specimenCount id="9D756081FFA1FF9FFDB33C20E3023FB5" box="[545,561,1657,1678]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 1.321.38 (♂), 1.411.50 (
<specimenCount id="9D756081FFA1FF9FFBF73C20E5463FB5" box="[1125,1141,1657,1678]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); palpal femur 6.927.23 (♂), 6.807.29 (
<specimenCount id="9D756081FFA1FF9FFE123CCEE0A33F97" box="[384,400,1687,1708]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times as long as broad (length 0.900.94 (♂), 1.02 (
<specimenCount id="9D756081FFA1FF9FFC013CCEE2903F97" box="[915,931,1687,1708]" pageId="59" pageNumber="60" type="female"></specimenCount>
) mm); coxae II with 911 terminally indented coxal spines on each side....................................................................
<taxonomicName id="4C73D08BFFA1FF9FFB413CECE47A3FF2" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1235,1353,1716,1738]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="latipectus" status="sp. nov.">
<emphasis id="B907771AFFA1FF9FFB413CECE47A3FF2" box="[1235,1353,1716,1738]" italics="true" pageId="59" pageNumber="60">L. latipectus</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA1FF9FFAC23CEDE4AF3FF2" bold="true" box="[1360,1436,1716,1737]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="A234CA61FFA1FF9FFAC23CEDE4AF3FF2" box="[1360,1436,1716,1737]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053C88E4AF3E1A" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053C88E4AF3FDC" blockId="59.[151,1437,220,2031]" box="[151,1436,1745,1767]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053C88E4AF3FDC" box="[151,1436,1745,1767]" pageId="59" pageNumber="60">15 Two corneate eyes, posterior pair of eyes reduced to eyespots; tergites and II each with four setae................... 16</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053CB6E4AF3E1A" blockId="59.[151,1437,220,2031]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053CB6E4AF3E1A" pageId="59" pageNumber="60">
- Four corneate eyes, posterior pair of eyes well developed; tergites and II each with two setae...................................................................................................
<taxonomicName id="4C73D08BFFA1FF9FFBFA3D54E4AF3E1A" ID-CoL="3RY2R" authority="Zhang &amp; Zhang, 2014" authorityName="Zhang &amp; Zhang" authorityYear="2014" box="[1128,1436,1804,1826]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="medog">
<emphasis id="B907771AFFA1FF9FFBFA3D54E5F23E1A" box="[1128,1217,1804,1826]" italics="true" pageId="59" pageNumber="60">L. medog</emphasis>
Zhang &amp; Zhang, 2014
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053D70E4AF3E67" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053D70E4AF3E04" blockId="59.[151,1437,220,2031]" box="[151,1436,1833,1855]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053D70E4AF3E04" box="[151,1436,1833,1855]" pageId="59" pageNumber="60">16 Carapace without epistome or with extremely indistinct epistome.............................................. 17</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053D1EE4AF3E67" blockId="59.[151,1437,220,2031]" box="[151,1436,1863,1885]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053D1EE4AF3E67" box="[151,1436,1863,1885]" pageId="59" pageNumber="60">
- Carapace with a distinct triangular epistome...........................................
<taxonomicName id="4C73D08BFFA1FF9FFB073D1EE4AF3E67" ID-CoL="3RY2X" authority="Hu &amp; Zhang, 2012" authorityName="Hu &amp; Zhang" authorityYear="2012" box="[1173,1436,1863,1885]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B907771AFFA1FF9FFB073D1EE5D23E67" box="[1173,1249,1863,1884]" italics="true" pageId="59" pageNumber="60">L. niger</emphasis>
Hu &amp; Zhang, 2012
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053D3DE4AF3EAC" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053D3DE4AF3E42" blockId="59.[151,1437,220,2031]" box="[151,1436,1892,1914]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053D3DE4AF3E42" box="[151,1436,1892,1914]" pageId="59" pageNumber="60">17 At least one finger of chela with intercalary teeth........................................................... 18</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053DDBE4AF3EAC" blockId="59.[151,1437,220,2031]" box="[151,1436,1921,1943]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053DDBE4AF3EAC" box="[151,1436,1921,1943]" pageId="59" pageNumber="60">- Both chelal fingers without intercalary teeth............................................................... 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF9FFF053DC6E4AF3EE9" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF9FFF053DC6E4AF3E8F" blockId="59.[151,1437,220,2031]" box="[151,1436,1951,1973]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053DC6E4AF3E8F" box="[151,1436,1951,1973]" pageId="59" pageNumber="60">
18 Only fixed chelal finger with intercalary teeth........................................
<taxonomicName id="4C73D08BFFA1FF9FFB123DC6E4AF3E8F" ID-CoL="3RY2C" authority="Hu &amp; Zhang, 2014" authorityName="Hu &amp; Zhang" authorityYear="2014" box="[1152,1436,1951,1973]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="B907771AFFA1FF9FFB123DC6E5D23E8F" box="[1152,1249,1951,1972]" italics="true" pageId="59" pageNumber="60">L. harveyi</emphasis>
Hu &amp; Zhang, 2014
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA1FF9FFF053DE5E4AF3EE9" blockId="59.[151,1437,220,2031]" box="[151,1436,1980,2002]" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF9FFF053DE5E4AF3EE9" box="[151,1436,1980,2002]" pageId="59" pageNumber="60">
- Both chelal fingers with intercalary teeth..............................................
<taxonomicName id="4C73D08BFFA1FF9FFB003DE4E4AF3EE9" ID-CoL="3RY3D" authority="(Beier, 1951)" baseAuthorityName="Beier" baseAuthorityYear="1951" box="[1170,1436,1980,2002]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="tonkinensis">
<emphasis id="B907771AFFA1FF9FFB003DE4E4243EEA" box="[1170,1303,1980,2002]" italics="true" pageId="59" pageNumber="60">L. tonkinensis</emphasis>
(
<bibRefCitation id="EFE2D6F9FFA1FF9FFAB63DE5E4A63EE9" author="Beier, M." box="[1316,1429,1980,2002]" pageId="59" pageNumber="60" pagination="47 - 123" refId="ref34847" refString="Beier, M. (1951) Die Pseudoscorpione Indochinas. Memoires du Museum National d'Histoire Naturelle, Paris, Nouvelle Serie, 1, 47 - 123." type="journal article" year="1951">Beier, 1951</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA1FF98FF053D80E4AF39DC" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<paragraph id="8BCCAB08FFA1FF98FF053D80E4AF3996" blockId="59.[151,1437,220,2031]" lastBlockId="60.[151,1437,151,1405]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<keyLead id="3082083DFFA1FF98FF053D80E4AF3996" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
19 Movable chelal finger more than 1.20 (♂,
<specimenCount id="9D756081FFA1FF9FFDEF3D80E3BE3ED5" box="[637,653,2009,2030]" pageId="59" pageNumber="60" type="female"></specimenCount>
) times longer than hand; proximal half of movable finger with rounded vestigial teeth.............................................................................................. 20
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053AECE4AF39DC" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053AECE4AF39DC" pageId="60" pageNumber="61">
- Movable chelal finger 1.03 (♂) times, 0.971.00 (
<specimenCount id="9D756081FFA6FF98FD5B3AECE3EA39F1" box="[713,729,181,202]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than hand; proximal half of movable chelal finger without vestigial teeth........................................................
<taxonomicName id="4C73D08BFFA6FF98FB833A8AE4AF39DC" ID-CoL="3RXZW" authority="Zhang &amp; Zhang, 2014" authorityName="Zhang &amp; Zhang" authorityYear="2014" box="[1041,1436,210,232]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="brachydigitatus">
<emphasis id="B907771AFFA6FF98FB833A8AE5F239DC" box="[1041,1217,210,232]" italics="true" pageId="60" pageNumber="61">L. brachydigitatus</emphasis>
Zhang &amp; Zhang, 2014
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053AA9E4AF3866" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053AA9E4AF3819" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053AA9E4AF3819" pageId="60" pageNumber="61">
20 Chelal hand 2.502.87 (♂) times, 2.392.75 (
<specimenCount id="9D756081FFA6FF98FD0B3AA9E39A383E" box="[665,681,240,261]" pageId="60" pageNumber="61" type="female"></specimenCount>
) as long as broad; trichobothrium
<emphasis id="B907771AFFA6FF98FC643AB6E538383F" box="[1014,1035,239,260]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
of movable chelal finger two times as far from
<emphasis id="B907771AFFA6FF98FE853B54E0143819" box="[279,295,269,290]" italics="true" pageId="60" pageNumber="61">st</emphasis>
as from
<emphasis id="B907771AFFA6FF98FEE93B54E0B43819" box="[379,391,269,290]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; tergites VII with four setae...................................
<taxonomicName id="4C73D08BFFA6FF98FBCA3B54E4AF3819" ID-CoL="3RY2P" authority="Hu &amp; Zhang, 2012" authorityName="Hu &amp; Zhang" authorityYear="2012" box="[1112,1436,269,291]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="leptopalpus">
<emphasis id="B907771AFFA6FF98FBCA3B54E5D23819" box="[1112,1249,269,290]" italics="true" pageId="60" pageNumber="61">L. leptopalpus</emphasis>
Hu &amp; Zhang, 2012
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053B72E4AF3866" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053B72E4AF3866" pageId="60" pageNumber="61">
- Chelal hand 2.00 (♂) times as long as broad; trichobothrium
<emphasis id="B907771AFFA6FF98FCB63B73E20A3804" box="[804,825,298,319]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
of movable chelal finger 1.5 times as far from
<emphasis id="B907771AFFA6FF98FB653B72E434387B" box="[1271,1287,299,320]" italics="true" pageId="60" pageNumber="61">st</emphasis>
as from
<emphasis id="B907771AFFA6FF98FAC83B73E4553804" box="[1370,1382,298,319]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; only the tergites II with four setae.........................................................
<taxonomicName id="4C73D08BFFA6FF98FB203B11E4AF3866" ID-CoL="3RY39" authority="(Beier, 1967)" baseAuthorityName="Beier" baseAuthorityYear="1967" box="[1202,1436,327,349]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B907771AFFA6FF98FB203B11E4243866" box="[1202,1303,328,349]" italics="true" pageId="60" pageNumber="61">L. sinensis</emphasis>
(Beier, 1967)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053B3CE4AF38A3" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053B3CE4AF3841" blockId="60.[151,1437,151,1405]" box="[151,1436,357,379]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053B3CE4AF3841" box="[151,1436,357,379]" pageId="60" pageNumber="61">21 Tergites II each with two setae........................................................................ 22</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053BDAE4AF38A3" blockId="60.[151,1437,151,1405]" box="[151,1436,386,408]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053BDAE4AF38A3" box="[151,1436,386,408]" pageId="60" pageNumber="61">- Tergites II each with three or four setae................................................................. 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053BF9E4AF38E9" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053BF9E4AF388E" blockId="60.[151,1437,151,1405]" box="[151,1436,415,437]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053BF9E4AF388E" box="[151,1436,415,437]" pageId="60" pageNumber="61">
22 Carapace with a round and obtuse epistome; body length max.
<quantity id="4C8B06EDFFA6FF98FCAC3BF9E2A5388E" box="[830,918,416,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="60" pageNumber="61" unit="mm" value="1.83">1.83 mm</quantity>
, chelal length max.
<quantity id="4C8B06EDFFA6FF98FBC43BF9E599388E" box="[1110,1194,416,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="60" pageNumber="61" unit="mm" value="1.11">1.11 mm</quantity>
.................. 23
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053BE4E4AF38E9" blockId="60.[151,1437,151,1405]" box="[151,1436,445,467]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053BE4E4AF38E9" box="[151,1436,445,467]" pageId="60" pageNumber="61">
- Carapace with a distinct triangular epistome; body length min.
<quantity id="4C8B06EDFFA6FF98FCAD3BE4E2A438E9" box="[831,919,445,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="60" pageNumber="61" unit="mm" value="1.85">1.85 mm</quantity>
, chelal length min.
<quantity id="4C8B06EDFFA6FF98FBC13BE4E59B38E9" box="[1107,1192,445,467]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" pageId="60" pageNumber="61" unit="mm" value="1.51">1.51 mm</quantity>
.................. 24
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053B83E4AF3B7C" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053B83E4AF3B36" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053B83E4AF3B36" pageId="60" pageNumber="61">
23 Palpal femur 6.456.80 (
<specimenCount id="9D756081FFA6FF98FE5D3B83E0EC38D4" box="[463,479,474,495]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times as long as broad; chelal finger 1.481.50 (
<specimenCount id="9D756081FFA6FF98FC263B83E2F738D4" box="[948,964,474,495]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than hand; tergite with 56 setae; anterior genital operculum with 12 setae; trichobothrium
<emphasis id="B907771AFFA6FF98FC9A3BA1E22C3B36" box="[776,799,504,525]" italics="true" pageId="60" pageNumber="61">ist</emphasis>
situated distal to
<emphasis id="B907771AFFA6FF98FC5A3BA1E2EB3B36" box="[968,984,504,525]" italics="true" pageId="60" pageNumber="61">st</emphasis>
...................
<taxonomicName id="4C73D08BFFA6FF98FB513BA1E47A3B37" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1219,1353,503,525]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tuoluoensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB513BA1E47A3B37" box="[1219,1353,503,525]" italics="true" pageId="60" pageNumber="61">L. tuoluoensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23BA1E4AF3B36" bold="true" box="[1360,1436,504,525]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23BA1E4AF3B36" box="[1360,1436,504,525]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF05384CE4AF3B7C" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF05384CE4AF3B7C" pageId="60" pageNumber="61">
-
<materialsCitation id="3B1BA155FFA6FF98FF70384CE4AB3B7C" ID-GBIF-Occurrence="4138993314" collectionCode="V" pageId="60" pageNumber="61" specimenCount="2" specimenCount-female="2">
Palpal femur 5.435.75 (
<specimenCount id="9D756081FFA6FF98FE45384CE0D43B11" box="[471,487,533,554]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times as long as broad; chelal finger 1.551.56 (
<specimenCount id="9D756081FFA6FF98FC47384CE2D63B11" box="[981,997,533,554]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than hand; tergite
<collectionCode id="ED6233CDFFA6FF98FABB384CE4093B11" box="[1321,1338,533,554]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="60" pageNumber="61" type="Museum"></collectionCode>
with four setae; anterior genital operculum with ten setae; trichobothrium
<emphasis id="B907771AFFA6FF98FCD8386AE2523B73" box="[842,865,563,584]" italics="true" pageId="60" pageNumber="61">ist</emphasis>
situated proximal
<emphasis id="B907771AFFA6FF98FB86386AE5173B73" box="[1044,1060,563,584]" italics="true" pageId="60" pageNumber="61">st</emphasis>
..............
<taxonomicName id="4C73D08BFFA6FF98FB59386AE47A3B73" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1227,1353,563,584]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="maanensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB59386AE47A3B73" box="[1227,1353,563,584]" italics="true" pageId="60" pageNumber="61">L. maanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234CA61FFA6FF98FAC2386BE4AB3B7C" box="[1360,1432,562,583]" pageId="60" pageNumber="61" rank="species">
<emphasis id="B907771AFFA6FF98FAC2386BE4AB3B7C" bold="true" box="[1360,1432,562,583]" pageId="60" pageNumber="61">sp. nov</emphasis>
</taxonomicNameLabel>
</materialsCitation>
<emphasis id="B907771AFFA6FF98FA0A386BE4AF3B7C" bold="true" box="[1432,1436,562,583]" pageId="60" pageNumber="61">.</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053809E4AF3BE1" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053809E4AF3BB9" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053809E4AF3BB9" pageId="60" pageNumber="61">
24 Tergite with two setae; body length min. 2.00 (♂), 2.17 (
<specimenCount id="9D756081FFA6FF98FC833809E2123B5E" box="[785,801,592,613]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, chelal length min. 1.79 (♂), 1.84 (
<specimenCount id="9D756081FFA6FF98FB363809E5873B5E" box="[1188,1204,592,613]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, palpal femur min. 1.25 (♂), 1.30 (
<specimenCount id="9D756081FFA6FF98FEEB3834E0BA3BB9" box="[377,393,621,642]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; coxae II with 1315 terminally indented coxal spines on each side...........
<taxonomicName id="4C73D08BFFA6FF98FB5D3834E47A3BB9" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1231,1353,621,642]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nigriculus" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB5D3834E47A3BB9" box="[1231,1353,621,642]" italics="true" pageId="60" pageNumber="61">L. nigriculus</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23834E4AF3BB9" bold="true" box="[1360,1436,621,642]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23834E4AF3BB9" box="[1360,1436,621,642]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF0538D2E4AF3BE1" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF0538D2E4AF3BE1" pageId="60" pageNumber="61">
- Tergite with four setae; body length max. 1.78 (♂), 2.05 (
<specimenCount id="9D756081FFA6FF98FCB938D3E2083BA4" box="[811,827,650,671]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, chelal length max. 1.62 (♂), 1.66 (
<specimenCount id="9D756081FFA6FF98FB4238D3E5D33BA4" box="[1232,1248,650,671]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, palpal femur max. 1.11 (♂), 1.17 (
<specimenCount id="9D756081FFA6FF98FE3F38F1E08E3B86" box="[429,445,680,701]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; coxae II with nine terminally indented coxal spines on each side..........................................................................................
<taxonomicName id="4C73D08BFFA6FF98FC67389CE4AF3BE1" authority="Hou, Gao &amp; Zhang, 2022" authorityName="Hou, Gao &amp; Zhang" authorityYear="2022" box="[1013,1436,709,731]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="laoxueyanensis">
<emphasis id="B907771AFFA6FF98FC67389CE5923BE1" box="[1013,1185,709,730]" italics="true" pageId="60" pageNumber="61">L. laoxueyanensis</emphasis>
Hou, Gao &amp; Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF0538BBE4AF3A2E" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF0538BBE4AF3BC3" blockId="60.[151,1437,151,1405]" box="[151,1436,738,760]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF0538BBE4AF3BC3" box="[151,1436,738,760]" pageId="60" pageNumber="61">25 Carapace with a distinct triangular epistome............................................................... 26</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053959E4AF3A2E" blockId="60.[151,1437,151,1405]" box="[151,1436,767,789]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053959E4AF3A2E" box="[151,1436,767,789]" pageId="60" pageNumber="61">- Carapace with a round and obtuse epistome............................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053944E4AF3A9C" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053944E4AF3A74" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053944E4AF3A74" pageId="60" pageNumber="61">
26 Chelal finger 1.51 (
<specimenCount id="9D756081FFA6FF98FE0E3944E09F3A09" box="[412,428,797,818]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than hand; trichobothrium
<emphasis id="B907771AFFA6FF98FCA03944E2743A09" box="[818,839,797,818]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA6FF98FBD03944E5613A09" box="[1090,1106,797,818]" italics="true" pageId="60" pageNumber="61">st</emphasis>
and
<emphasis id="B907771AFFA6FF98FBED3944E5B83A09" box="[1151,1163,797,818]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; coxae II with ten terminally indented coxal spines on each side; rallum with nine blades; movable chelal finger with 2729 teeth...
<taxonomicName id="4C73D08BFFA6FF98FB593962E47A3A74" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1227,1353,826,848]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="biyunensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB593962E47A3A74" box="[1227,1353,826,848]" italics="true" pageId="60" pageNumber="61">L. biyunensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23963E4AF3A74" bold="true" box="[1360,1436,826,847]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23963E4AF3A74" box="[1360,1436,826,847]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053901E4AF3A9C" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053901E4AF3A9C" pageId="60" pageNumber="61">
- Chelal finger 1.24 (
<specimenCount id="9D756081FFA6FF98FE313901E0803A56" box="[419,435,856,877]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than hand; trichobothrium
<emphasis id="B907771AFFA6FF98FCD4390EE2683A57" box="[838,859,855,876]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
closer to
<emphasis id="B907771AFFA6FF98FC2F3901E2FE3A56" box="[957,973,856,877]" italics="true" pageId="60" pageNumber="61">st</emphasis>
than to
<emphasis id="B907771AFFA6FF98FB8D390EE5183A57" box="[1055,1067,855,876]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; coxae II with 12 terminally indented coxal spines on each side; rallum with seven blades; movable chelal finger with 42 teeth..................................................................................................
<taxonomicName id="4C73D08BFFA6FF98FB8A39CAE4AF3A9C" authority="Hou, Gao and Zhang, 2022" authorityName="Hou, Gao and Zhang" authorityYear="2022" box="[1048,1436,914,936]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="xibaiensis">
<emphasis id="B907771AFFA6FF98FB8A39CAE5A23A9C" box="[1048,1169,914,936]" italics="true" pageId="60" pageNumber="61">L. xibaiensis</emphasis>
Hou, Gao and Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF0539E9E4AF3D26" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF0539E9E4AF3AD9" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF0539E9E4AF3AD9" pageId="60" pageNumber="61">
27 Smaller: body length max. 1.23 (♂), 1.47 (
<specimenCount id="9D756081FFA6FF98FD1739E9E3A63AFE" box="[645,661,944,965]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; chela length max. 0.85 (♂), 0.97 (
<specimenCount id="9D756081FFA6FF98FBB639E9E5073AFE" box="[1060,1076,944,965]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; palpal femur length max. 0.60 (♂), 0.67 (
<specimenCount id="9D756081FFA6FF98FEDB3994E06A3AD9" box="[329,345,973,994]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm.................................................................................... 28
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF0539B2E4AF3D26" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF0539B2E4AF3D26" pageId="60" pageNumber="61">
- Larger: body length min. 1.59 (♂), 1.64 (
<specimenCount id="9D756081FFA6FF98FDFC39B3E34D3AC4" box="[622,638,1002,1023]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; chela length min. 1.03 (♂), 1.12 (
<specimenCount id="9D756081FFA6FF98FB9339B3E5223AC4" box="[1025,1041,1002,1023]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; palpal femur length min. 0.74 (♂), 0.80 (
<specimenCount id="9D756081FFA6FF98FE893E51E0183D26" box="[283,299,1032,1053]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm........................................................................................ 29
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053E7CE4AF3DA9" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053E7CE4AF3D6C" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053E7CE4AF3D6C" pageId="60" pageNumber="61">
28 Fixed chelal finger with 2425 teeth; trichobothrium
<emphasis id="B907771AFFA6FF98FD733E7CE3C53D01" box="[737,758,1061,1082]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA6FF98FC643E7CE5353D01" box="[1014,1030,1061,1082]" italics="true" pageId="60" pageNumber="61">st</emphasis>
and
<emphasis id="B907771AFFA6FF98FBA73E7CE5723D01" box="[1077,1089,1061,1082]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; rallum with nine blades............................................................................................
<taxonomicName id="4C73D08BFFA6FF98FB453E1AE47A3D63" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1239,1353,1091,1112]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="xiaoensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB453E1AE47A3D63" box="[1239,1353,1091,1112]" italics="true" pageId="60" pageNumber="61">L. xiaoensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23E1BE4AF3D6C" bold="true" box="[1360,1436,1090,1111]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23E1BE4AF3D6C" box="[1360,1436,1090,1111]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053E39E4AF3DA9" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053E39E4AF3DA9" pageId="60" pageNumber="61">
- Fixed chelal finger with 3537 teeth; trichobothrium
<emphasis id="B907771AFFA6FF98FD733E06E3C53D4F" box="[737,758,1119,1140]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
closer to
<emphasis id="B907771AFFA6FF98FCC63E39E2573D4E" box="[852,868,1120,1141]" italics="true" pageId="60" pageNumber="61">st</emphasis>
than to
<emphasis id="B907771AFFA6FF98FC203E06E28D3D4F" box="[946,958,1119,1140]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; rallum with eight blades........................................................................................
<taxonomicName id="4C73D08BFFA6FF98FBB43E24E4AF3DA9" authority="Hou, Gao and Zhang, 2022" authorityName="Hou, Gao and Zhang" authorityYear="2022" box="[1062,1436,1149,1171]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B907771AFFA6FF98FBB43E24E5A23DA9" box="[1062,1169,1149,1170]" italics="true" pageId="60" pageNumber="61">L. minimus</emphasis>
Hou, Gao and Zhang, 2022
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053EC3E4AF3C3C" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053EC3E4AF3DF6" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053EC3E4AF3DF6" pageId="60" pageNumber="61">
29 Larger: body length min. 1.76 (♂), 1.93 (
<specimenCount id="9D756081FFA6FF98FDFC3EC3E34D3D94" box="[622,638,1178,1199]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; chela length min. 1.26 (♂), 1.38 (
<specimenCount id="9D756081FFA6FF98FB933EC3E5223D94" box="[1025,1041,1178,1199]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; palpal femur length min. 0.90 (♂), 0.99 (
<specimenCount id="9D756081FFA6FF98FE893EE1E0183DF6" box="[283,299,1208,1229]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm........................................................................................ 30
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053E8CE4AF3C3C" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053E8CE4AF3C3C" pageId="60" pageNumber="61">
- Smaller: body length max. 1.66 (♂), 1.73 (
<specimenCount id="9D756081FFA6FF98FDE53E8CE3B43DD1" box="[631,647,1237,1258]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; chela length max. 1.11(♂), 1.14 (
<specimenCount id="9D756081FFA6FF98FB903E8CE5213DD1" box="[1026,1042,1237,1258]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm; palpal femur length max. 0.79 (♂), 0.80 (
<specimenCount id="9D756081FFA6FF98FE893EABE0183C3C" box="[283,299,1266,1287]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm......................................................................
<taxonomicName id="4C73D08BFFA6FF98FB3D3EAAE47A3C33" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1199,1353,1267,1288]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="mawangensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB3D3EAAE47A3C33" box="[1199,1353,1267,1288]" italics="true" pageId="60" pageNumber="61">L. mawangensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23EABE4AF3C3C" bold="true" box="[1360,1436,1266,1287]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23EABE4AF3C3C" box="[1360,1436,1266,1287]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3087B3ADFFA6FF98FF053F49E4AF3C46" pageId="60" pageNumber="61">
<paragraph id="8BCCAB08FFA6FF98FF053F49E4AF3C79" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053F49E4AF3C79" pageId="60" pageNumber="61">
30 Trichobothrium
<emphasis id="B907771AFFA6FF98FEED3F56E0A73C1F" box="[383,404,1295,1316]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
situated midway between
<emphasis id="B907771AFFA6FF98FD013F49E3903C1E" box="[659,675,1296,1317]" italics="true" pageId="60" pageNumber="61">st</emphasis>
and
<emphasis id="B907771AFFA6FF98FD403F56E3ED3C1F" box="[722,734,1295,1316]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; body length 1.761.95 (♂), 1.93 (
<specimenCount id="9D756081FFA6FF98FBBD3F49E50C3C1E" box="[1071,1087,1296,1317]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, chela 6.636.95 (♂), 5.75 (
<specimenCount id="9D756081FFA6FF98FA113F49E4A03C1E" box="[1411,1427,1296,1317]" pageId="60" pageNumber="61" type="female"></specimenCount>
) times longer than broad (length 1.261.32 (♂), 1.38 (
<specimenCount id="9D756081FFA6FF98FD703F74E3C13C79" box="[738,754,1325,1346]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm)................................
<taxonomicName id="4C73D08BFFA6FF98FB213F74E47A3C79" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1203,1353,1325,1346]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="longyanensis" status="sp. nov.">
<emphasis id="B907771AFFA6FF98FB213F74E47A3C79" box="[1203,1353,1325,1346]" italics="true" pageId="60" pageNumber="61">L. longyanensis</emphasis>
</taxonomicName>
<emphasis id="B907771AFFA6FF98FAC23F74E4AF3C79" bold="true" box="[1360,1436,1325,1346]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="A234CA61FFA6FF98FAC23F74E4AF3C79" box="[1360,1436,1325,1346]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCCAB08FFA6FF98FF053F12E4AF3C46" blockId="60.[151,1437,151,1405]" pageId="60" pageNumber="61">
<keyLead id="3082083DFFA6FF98FF053F12E4AF3C46" pageId="60" pageNumber="61">
- Trichobothrium
<emphasis id="B907771AFFA6FF98FEEC3F13E0A03C64" box="[382,403,1354,1375]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
closer to
<emphasis id="B907771AFFA6FF98FE7D3F12E0CC3C5B" box="[495,511,1355,1376]" italics="true" pageId="60" pageNumber="61">st</emphasis>
than to
<emphasis id="B907771AFFA6FF98FDD83F13E3653C64" box="[586,598,1354,1375]" italics="true" pageId="60" pageNumber="61">b</emphasis>
; body length 2.552.92 (♂), 2.722.95 (
<specimenCount id="9D756081FFA6FF98FC4B3F13E2DA3C64" box="[985,1001,1354,1375]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm, chela 8.5010.20 (♂) times longer than broad (length 1.852.22 (♂), 1.942.03 (
<specimenCount id="9D756081FFA6FF98FDFE3F31E34F3C46" box="[620,636,1384,1405]" pageId="60" pageNumber="61" type="female"></specimenCount>
) mm)..............................
<taxonomicName id="4C73D08BFFA6FF98FBBA3F31E4AF3C46" ID-CoL="6NTXL" authority="Li, Liu &amp; Shi, 2019" authorityName="Li, Liu &amp; Shi" authorityYear="2019" box="[1064,1436,1383,1405]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bailongtanensis">
<emphasis id="B907771AFFA6FF98FBBA3F31E5EA3C47" box="[1064,1241,1383,1405]" italics="true" pageId="60" pageNumber="61">L. bailongtanensis</emphasis>
<bibRefCitation id="EFE2D6F9FFA6FF98FB4D3F3EE4AF3C46" author="Li, Y. C. &amp; Liu, H. &amp; Shi, A. M." box="[1247,1436,1383,1405]" pageId="60" pageNumber="61" pagination="28 - 34" refId="ref35901" refString="Li, Y. C., Liu, H. &amp; Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">Li, Liu &amp; Shi, 2019</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>