260 lines
39 KiB
XML
260 lines
39 KiB
XML
<document id="BA13D3BF4CE1664BE6B0EC514DC024C7" ID-DOI="10.37828/em.2024.78.8" ID-ISSN="2336-9744" ID-Zenodo-Dep="14654239" ID-ZooBank="CC0FAE19-9610-4EBA-8EE9-2353CE7C48DD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736952339350" checkinUser="felipe" docAuthor="Volynkin, Anton V." docDate="2024" docId="03BF816AFFC0AA62FF126AD903F86AFC" docLanguage="en" docName="EcolMontenegrina.78.72-78.pdf" docOrigin="Ecologica Montenegrina 78" docSource="https://doi.org/10.37828/em.2024.78.8" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Biampullisna Volynkin, 2024, gen. n." docType="treatment" docVersion="2" lastPageNumber="76" masterDocId="FF86F912FFC1AA66FFAF6906006C6E73" masterDocTitle="Biampullisna, a new genus for “ Teulisna ” uniplaga Hampson (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="78" masterPageNumber="72" pageNumber="73" updateTime="1736954568525" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7A88F30E0ACA832EA5B89620BBB2AE42">Biampullisna, a new genus for “ Teulisna ” uniplaga Hampson (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="B854EBB2AE7770E7E142FDA8D3E96895">Volynkin, Anton V.</mods:namePart>
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<mods:title id="60527517B2021710652B79F8F348B4FD">Ecologica Montenegrina</mods:title>
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<mods:part id="4A09AAEAF456FC5ABE313DB908B3B0A0">
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<mods:date id="369C6E49CAE1F5CD66A3652D06123522">2024</mods:date>
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<mods:number id="7AF04BABC423ECE5E4AFC1B7A8FDC2AD">2024-09-23</mods:number>
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<mods:url id="27B8BB52D7CF08C2AA78D2039E30FAA9">https://doi.org/10.37828/em.2024.78.8</mods:url>
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<treatment id="03BF816AFFC0AA62FF126AD903F86AFC" ID-DOI="http://doi.org/10.5281/zenodo.14654241" ID-Zenodo-Dep="14654241" LSID="urn:lsid:plazi:treatment:03BF816AFFC0AA62FF126AD903F86AFC" httpUri="http://treatment.plazi.org/id/03BF816AFFC0AA62FF126AD903F86AFC" lastPageId="4" lastPageNumber="76" pageId="1" pageNumber="73">
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<subSubSection id="C30C63F7FFC0AA67FF126AD904DC6A6B" pageId="1" pageNumber="73" type="nomenclature">
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<paragraph id="8BA9307CFFC0AA67FF126AD901D66D8A" blockId="1.[189,1200,991,1082]" box="[189,442,991,1017]" pageId="1" pageNumber="73">
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<heading id="D0E18710FFC0AA67FF126AD901D66D8A" box="[189,442,991,1017]" fontSize="11" level="2" pageId="1" pageNumber="73" reason="2">
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<emphasis id="B962EC6EFFC0AA67FF126AD901D66D8A" bold="true" box="[189,442,991,1017]" pageId="1" pageNumber="73">
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<taxonomicName id="4C164BFFFFC0AA67FF126AD9010C6D8A" ID-CoL="CL3YN" authorityName="Volynkin" authorityYear="2024" box="[189,352,991,1017]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B962EC6EFFC0AA67FF126AD9010C6D8A" bold="true" box="[189,352,991,1017]" italics="true" pageId="1" pageNumber="73">Biampullisna</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2515115FFC0AA67FEC76AD901D66D8A" box="[360,442,991,1017]" pageId="1" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA9307CFFC0AA67FF126D0604DC6A6B" blockId="1.[189,1200,991,1082]" box="[189,1200,1024,1048]" pageId="1" pageNumber="73">
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="C30C63F7FFC0AA67FF126D2604756A49" box="[189,1049,1055,1082]" pageId="1" pageNumber="73" type="materials_examined">
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<paragraph id="8BA9307CFFC0AA67FF126D2604756A49" blockId="1.[189,1200,991,1082]" box="[189,1049,1055,1082]" pageId="1" pageNumber="73">
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<typeStatus id="54AD8EDEFFC0AA67FF126D2600956A49" box="[189,249,1056,1082]" pageId="1" pageNumber="73">Type</typeStatus>
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species:
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<taxonomicName id="4C164BFFFFC0AA67FECA6D1902916A49" authority="Hampson, 1894" authorityName="Hampson" authorityYear="1894" box="[357,765,1055,1082]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="species" species="uniplaga">
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<emphasis id="B962EC6EFFC0AA67FECA6D1902566A49" box="[357,570,1055,1082]" italics="true" pageId="1" pageNumber="73">Teulisna uniplaga</emphasis>
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<bibRefCitation id="EF874D8DFFC0AA67FDEE6D2602916A49" author="Hampson, G. F." box="[577,765,1055,1082]" pageId="1" pageNumber="73" pagination="1 - 609" refId="ref2739" refString="Hampson, G. F. (1894) The Fauna of British India, including Ceylon and Burma. Moths. Vol. 2. London, Calcutta, Bombay & Berlin, pp. v - xxii, 1 - 609." type="book chapter" year="1894">Hampson, 1894</bibRefCitation>
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</taxonomicName>
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, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C63F7FFC0AA67FF126D6504316910" pageId="1" pageNumber="73" type="diagnosis">
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<paragraph id="8BA9307CFFC0AA67FF126D6504316910" blockId="1.[189,1400,1122,2026]" pageId="1" pageNumber="73">
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<emphasis id="B962EC6EFFC0AA67FF126D65015A6A0E" bold="true" box="[189,310,1123,1149]" pageId="1" pageNumber="73">Diagnosis</emphasis>
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. The
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<typeStatus id="54AD8EDEFFC0AA67FED66D6501C06A0E" box="[377,428,1123,1149]" pageId="1" pageNumber="73">type</typeStatus>
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species of the new genus (
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<figureCitation id="132D2CF9FFC0AA67FD4C6D6403536A0E" box="[739,831,1122,1149]" captionStart="Figures 1–8" captionStartId="2.[189,275,1815,1839]" captionTargetBox="[197,1392,223,1778]" captionTargetId="figure-151@2.[189,1392,223,1808]" captionTargetPageId="2" captionText="Figures 1–8. Adults of Biampullisna uniplaga (type species of Biampullisna gen. n.) (1, 2), Teulisna plagiata (type species of Teulisna) (3, 4), Teulisna quadratella Holloway, 2001 (5), Witteulisna paraxena Volynkin, 2020 (type species of Witteulisna) (6), and Teulisna basigera (Walker, [1865]) (type species of Cossa Walker, [1865], a junior synonym of Teulisna) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="1" pageNumber="73">Fig 1, 2</figureCitation>
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) is externally reminiscent of certain members of the genus
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<taxonomicName id="4C164BFFFFC0AA67FE966D8201F26AEC" authorityName="Walker" authorityYear="1862" box="[313,414,1156,1183]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EC6EFFC0AA67FE966D8201F26AEC" box="[313,414,1156,1183]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
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</taxonomicName>
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(including the
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<typeStatus id="54AD8EDEFFC0AA67FDF96D8302E46AEC" box="[598,648,1157,1183]" pageId="1" pageNumber="73">type</typeStatus>
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one) (
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<figureCitation id="132D2CF9FFC0AA67FD776D82032E6AEC" box="[728,834,1156,1183]" captionStart="Figures 1–8" captionStartId="2.[189,275,1815,1839]" captionTargetBox="[197,1392,223,1778]" captionTargetId="figure-151@2.[189,1392,223,1808]" captionTargetPageId="2" captionText="Figures 1–8. Adults of Biampullisna uniplaga (type species of Biampullisna gen. n.) (1, 2), Teulisna plagiata (type species of Teulisna) (3, 4), Teulisna quadratella Holloway, 2001 (5), Witteulisna paraxena Volynkin, 2020 (type species of Witteulisna) (6), and Teulisna basigera (Walker, [1865]) (type species of Cossa Walker, [1865], a junior synonym of Teulisna) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="1" pageNumber="73">Figs 3–5</figureCitation>
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) as well as the male of the sexually dimorphic genus
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<taxonomicName id="4C164BFFFFC0AA67FEA36DA0023B6AB2" authority="Volynkin, 2020" authorityName="Volynkin" authorityYear="2020" box="[268,599,1190,1217]" class="Insecta" family="Erebidae" genus="Witteulisna" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EC6EFFC0AA67FEA36DA001FE6AB2" box="[268,402,1190,1217]" italics="true" pageId="1" pageNumber="73">Witteulisna</emphasis>
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Volynkin, 2020
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</taxonomicName>
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(
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<figureCitation id="132D2CF9FFC0AA67FDC66DA002D86AB2" box="[617,692,1190,1217]" captionStart="Figures 1–8" captionStartId="2.[189,275,1815,1839]" captionTargetBox="[197,1392,223,1778]" captionTargetId="figure-151@2.[189,1392,223,1808]" captionTargetPageId="2" captionText="Figures 1–8. Adults of Biampullisna uniplaga (type species of Biampullisna gen. n.) (1, 2), Teulisna plagiata (type species of Teulisna) (3, 4), Teulisna quadratella Holloway, 2001 (5), Witteulisna paraxena Volynkin, 2020 (type species of Witteulisna) (6), and Teulisna basigera (Walker, [1865]) (type species of Cossa Walker, [1865], a junior synonym of Teulisna) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="1" pageNumber="73">Fig. 6</figureCitation>
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), which has, however, fundamentally different genitalia of both sexes (
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<figureCitation id="132D2CF9FFC0AA67FEE86DC101A16A91" box="[327,461,1223,1250]" captionStart="Figures 13–17" captionStartId="5.[189,275,1879,1903]" captionTargetBox="[189,1380,241,1843]" captionTargetId="figure-114@5.[189,1398,223,1859]" captionTargetPageId="5" captionText="Figures 13–17. Female genitalia of Biampullisna uniplaga (13), Teulisna spp. (14, 15) and Witteulisna paraxena (17), and male genitalia of Witteulisna paraxena (16). Depositories of the specimens dissected: 13, 14, 16 and 17 in MWM/ZSM; 15 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654249" httpUri="https://zenodo.org/record/14654249/files/figure.png" pageId="1" pageNumber="73">Figs 16, 17</figureCitation>
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), but differs in the forewing with a nearly straight costal margin and unmodified venation, and without patches of androconial scales. The male genital capsule of
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<taxonomicName id="4C164BFFFFC0AA67FBD06DEF05776B77" authorityName="Volynkin" authorityYear="2024" box="[1151,1307,1257,1284]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B962EC6EFFC0AA67FBD06DEF05776B77" box="[1151,1307,1257,1284]" italics="true" pageId="1" pageNumber="73">Biampullisna</emphasis>
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</taxonomicName>
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<emphasis id="B962EC6EFFC0AA67FA8B6DEF051B6B70" bold="true" box="[1316,1399,1257,1283]" pageId="1" pageNumber="73">
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<taxonomicNameLabel id="A2515115FFC0AA67FA8B6DEF051B6B70" box="[1316,1399,1257,1283]" pageId="1" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="132D2CF9FFC0AA67FF696C0D01626B55" box="[198,270,1291,1318]" captionStart="Figures 9–12" captionStartId="3.[189,275,1912,1936]" captionTargetBox="[199,1398,263,1876]" captionTargetId="figure-103@3.[189,1398,257,1893]" captionTargetPageId="3" captionText="Figures 9–12. Male genitalia of Biampullisna uniplaga (9) and Teulisna spp. (10–12). Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 in NHMUK; 12 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654247" httpUri="https://zenodo.org/record/14654247/files/figure.png" pageId="1" pageNumber="73">Fig. 9</figureCitation>
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) is characterised by the combination of the following three features. (1) The uncus is weakly sclerotised, dorso-ventrally flattened, relatively short and fused with the dorsal wall of the anal cone whereas in
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<taxonomicName id="4C164BFFFFC0AA67FEEA6C4801C66B1A" authorityName="Walker" authorityYear="1862" box="[325,426,1358,1385]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EC6EFFC0AA67FEEA6C4801C66B1A" box="[325,426,1358,1385]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
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</taxonomicName>
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(
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<figureCitation id="132D2CF9FFC0AA67FE156C48022C6B1A" box="[442,576,1358,1385]" captionStart="Figures 9–12" captionStartId="3.[189,275,1912,1936]" captionTargetBox="[199,1398,263,1876]" captionTargetId="figure-103@3.[189,1398,257,1893]" captionTargetPageId="3" captionText="Figures 9–12. Male genitalia of Biampullisna uniplaga (9) and Teulisna spp. (10–12). Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 in NHMUK; 12 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654247" httpUri="https://zenodo.org/record/14654247/files/figure.png" pageId="1" pageNumber="73">Figs 10–12</figureCitation>
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) it is longer, heavily sclerotised, laterally flattened and separate from the anal cone, which, in turn, has a thin scaphium. (2) The tegumen has narrow and weakly sclerotised arms with only dorsal ends being fused whereas that of
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<taxonomicName id="4C164BFFFFC0AA67FCF66C9403D26BDE" authorityName="Walker" authorityYear="1862" box="[857,958,1426,1453]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EC6EFFC0AA67FCF66C9403D26BDE" box="[857,958,1426,1453]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
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</taxonomicName>
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is bell-shaped with strongly dorsally dilated arms, which are fused in their dorsal three-quarters. The vinculum is short, more or less Ushaped, lacking the intravincular structures whereas in
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<taxonomicName id="4C164BFFFFC0AA67FCCC6CD303A46B83" authorityName="Walker" authorityYear="1862" box="[867,968,1493,1520]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EC6EFFC0AA67FCCC6CD303A46B83" box="[867,968,1493,1520]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
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</taxonomicName>
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it is longer, frame-shaped and has intravincular sclerotised structures (discussed by
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<bibRefCitation id="EF874D8DFFC0AA67FCA46CF103B26861" author="Volynkin, A. V." box="[779,990,1527,1554]" pageId="1" pageNumber="73" pagination="176 - 207" refId="ref2981" refString="Volynkin, A. V. (2024) On the terminology of the genitalia structures of lichen moths (Lepidoptera: Erebidae: Arctiinae: Lithosiini) with some references to Noctuidae. Ecologica Montenegrina, 73, 176 - 207. https: // dx. doi. org / 10.37828 / em. 2024.73.18" type="journal article" year="2024">Volynkin (2024))</bibRefCitation>
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, which may be reduced in certain groups (
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<figureCitation id="132D2CF9FFC0AA67FE8E6F1F01FF6847" box="[289,403,1561,1588]" captionStart="Figures 1–8" captionStartId="2.[189,275,1815,1839]" captionTargetBox="[197,1392,223,1778]" captionTargetId="figure-151@2.[189,1392,223,1808]" captionTargetPageId="2" captionText="Figures 1–8. Adults of Biampullisna uniplaga (type species of Biampullisna gen. n.) (1, 2), Teulisna plagiata (type species of Teulisna) (3, 4), Teulisna quadratella Holloway, 2001 (5), Witteulisna paraxena Volynkin, 2020 (type species of Witteulisna) (6), and Teulisna basigera (Walker, [1865]) (type species of Cossa Walker, [1865], a junior synonym of Teulisna) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="1" pageNumber="73">Figs 7, 8</figureCitation>
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,
|
||
<figureCitation id="132D2CF9FFC0AA67FE0B6F1F01A86847" box="[420,452,1561,1588]" captionStart="Figures 9–12" captionStartId="3.[189,275,1912,1936]" captionTargetBox="[199,1398,263,1876]" captionTargetId="figure-103@3.[189,1398,257,1893]" captionTargetPageId="3" captionText="Figures 9–12. Male genitalia of Biampullisna uniplaga (9) and Teulisna spp. (10–12). Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 in NHMUK; 12 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654247" httpUri="https://zenodo.org/record/14654247/files/figure.png" pageId="1" pageNumber="73">12</figureCitation>
|
||
,
|
||
<figureCitation id="132D2CF9FFC0AA67FE7A6F1F019F6847" box="[469,499,1561,1588]" captionStart="Figures 13–17" captionStartId="5.[189,275,1879,1903]" captionTargetBox="[189,1380,241,1843]" captionTargetId="figure-114@5.[189,1398,223,1859]" captionTargetPageId="5" captionText="Figures 13–17. Female genitalia of Biampullisna uniplaga (13), Teulisna spp. (14, 15) and Witteulisna paraxena (17), and male genitalia of Witteulisna paraxena (16). Depositories of the specimens dissected: 13, 14, 16 and 17 in MWM/ZSM; 15 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654249" httpUri="https://zenodo.org/record/14654249/files/figure.png" pageId="1" pageNumber="73">15</figureCitation>
|
||
) but their remnants are still recognisable. (3) The editum is elongate, its proximal part occupies the base of the dorsal margin of the valva while the distal section is reaching the base of the distal saccular process; the editum bears two ampullae: a stake-shaped proximal and a lobe-like, spinulose distal ones. Unlike in
|
||
<taxonomicName id="4C164BFFFFC0AA67FDD46F78037B68EA" authorityName="Volynkin" authorityYear="2024" box="[635,791,1662,1689]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B962EC6EFFC0AA67FDD46F78037B68EA" box="[635,791,1662,1689]" italics="true" pageId="1" pageNumber="73">Biampullisna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EC6EFFC0AA67FC8F6F78031968EB" bold="true" box="[800,885,1662,1688]" pageId="1" pageNumber="73">
|
||
<taxonomicNameLabel id="A2515115FFC0AA67FC8F6F78031968EB" box="[800,885,1662,1688]" pageId="1" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the editum of
|
||
<taxonomicName id="4C164BFFFFC0AA67FB806F7804F868EA" authorityName="Walker" authorityYear="1862" box="[1071,1172,1662,1689]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC0AA67FB806F7804F868EA" box="[1071,1172,1662,1689]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
is shorter, situated more ventrally, and bears a medial ampulla or lacks it, while the costa occupies the whole proximal section of the dorsal margin of the valva, and additionally bears a proximal process in a number of species. The phallus of
|
||
<taxonomicName id="4C164BFFFFC0AA67FE7B6FE5021C688D" authorityName="Volynkin" authorityYear="2024" box="[468,624,1763,1790]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B962EC6EFFC0AA67FE7B6FE5021C688D" box="[468,624,1763,1790]" italics="true" pageId="1" pageNumber="73">Biampullisna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EC6EFFC0AA67FDD76FE202A7688D" bold="true" box="[632,715,1764,1790]" pageId="1" pageNumber="73">
|
||
<taxonomicNameLabel id="A2515115FFC0AA67FDD76FE202A7688D" box="[632,715,1764,1790]" pageId="1" pageNumber="73" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is markedly smaller than in
|
||
<taxonomicName id="4C164BFFFFC0AA67FBB46FE504EC688D" authorityName="Walker" authorityYear="1862" box="[1051,1152,1763,1790]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC0AA67FBB46FE504EC688D" box="[1051,1152,1763,1790]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
(in proportion to the genital capsule), but the vesicae of the two genera display no fundamental differences. In the female genitalia, the membranous corpus bursae is characteristic of the new genus (
|
||
<figureCitation id="132D2CF9FFC0AA67FB836E2004136932" box="[1068,1151,1830,1857]" captionStart="Figures 13–17" captionStartId="5.[189,275,1879,1903]" captionTargetBox="[189,1380,241,1843]" captionTargetId="figure-114@5.[189,1398,223,1859]" captionTargetPageId="5" captionText="Figures 13–17. Female genitalia of Biampullisna uniplaga (13), Teulisna spp. (14, 15) and Witteulisna paraxena (17), and male genitalia of Witteulisna paraxena (16). Depositories of the specimens dissected: 13, 14, 16 and 17 in MWM/ZSM; 15 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654249" httpUri="https://zenodo.org/record/14654249/files/figure.png" pageId="1" pageNumber="73">Fig. 13</figureCitation>
|
||
) whereas in
|
||
<taxonomicName id="4C164BFFFFC0AA67FABD6E20051B6932" authorityName="Walker" authorityYear="1862" box="[1298,1399,1830,1857]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC0AA67FABD6E20051B6932" box="[1298,1399,1830,1857]" italics="true" pageId="1" pageNumber="73">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132D2CF9FFC0AA67FF6A6E4E01276910" box="[197,331,1864,1891]" captionStart="Figures 13–17" captionStartId="5.[189,275,1879,1903]" captionTargetBox="[189,1380,241,1843]" captionTargetId="figure-114@5.[189,1398,223,1859]" captionTargetPageId="5" captionText="Figures 13–17. Female genitalia of Biampullisna uniplaga (13), Teulisna spp. (14, 15) and Witteulisna paraxena (17), and male genitalia of Witteulisna paraxena (16). Depositories of the specimens dissected: 13, 14, 16 and 17 in MWM/ZSM; 15 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654249" httpUri="https://zenodo.org/record/14654249/files/figure.png" pageId="1" pageNumber="73">Figs 14, 15</figureCitation>
|
||
) the corpus bursae has broad gelatinous areas or sclerotised plates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C63F7FFC0AA62FEB36E6C047E6A59" lastPageId="4" lastPageNumber="76" pageId="1" pageNumber="73" type="description">
|
||
<paragraph id="8BA9307CFFC0AA62FEB36E6C047E6A59" blockId="1.[189,1400,1122,2026]" lastBlockId="4.[189,1400,196,1167]" lastPageId="4" lastPageNumber="76" pageId="1" pageNumber="73">
|
||
<emphasis id="B962EC6EFFC0AA67FEB36E6C01C169F7" bold="true" box="[284,429,1898,1924]" pageId="1" pageNumber="73">Description</emphasis>
|
||
.
|
||
<emphasis id="B962EC6EFFC0AA67FE6D6E6C027969F7" bold="true" box="[450,533,1898,1924]" pageId="1" pageNumber="73">Adults</emphasis>
|
||
(
|
||
<figureCitation id="132D2CF9FFC0AA67FD846E6C02CD69F6" box="[555,673,1898,1925]" captionStart="Figures 1–8" captionStartId="2.[189,275,1815,1839]" captionTargetBox="[197,1392,223,1778]" captionTargetId="figure-151@2.[189,1392,223,1808]" captionTargetPageId="2" captionText="Figures 1–8. Adults of Biampullisna uniplaga (type species of Biampullisna gen. n.) (1, 2), Teulisna plagiata (type species of Teulisna) (3, 4), Teulisna quadratella Holloway, 2001 (5), Witteulisna paraxena Volynkin, 2020 (type species of Witteulisna) (6), and Teulisna basigera (Walker, [1865]) (type species of Cossa Walker, [1865], a junior synonym of Teulisna) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK." figureDoi="http://doi.org/10.5281/zenodo.14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="1" pageNumber="73">Figs 1, 2</figureCitation>
|
||
). Sexual dimorphism limited: female larger than male, its forewing somewhat broader and with more elongate apex. Antenna weakly ciliate in both sexes, ochreous proximally and blackish medially and distally. Head pale ochreous yellow. Thorax black with deep ochreous yellow tegula and patagia. Forewing elongate and narrow, with tapered apex and convex termen and anal margin. Forewing ground colour pale ochreous yellow with intense dark grey suffusion. Forewing pattern black, consisting of diffuse subbasal dash, large pentagonal medial patch touching the anal margin but not reaching the costal one, and small trapezoidal costal postmedial spot. Cilia ochreous yellow. Hindwing ground colour and cilia uniformly brownish-grey with ochreous irroration. Abdomen brownish-grey, with ochreous yellow hair-shaped scales distally.
|
||
<emphasis id="B962EC6EFFC5AA62FC1A684D040B6F16" bold="true" box="[949,1127,331,357]" pageId="4" pageNumber="76">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="132D2CF9FFC5AA62FBDA684D04D06F15" box="[1141,1212,331,358]" captionStart="Figures 9–12" captionStartId="3.[189,275,1912,1936]" captionTargetBox="[199,1398,263,1876]" captionTargetId="figure-103@3.[189,1398,257,1893]" captionTargetPageId="3" captionText="Figures 9–12. Male genitalia of Biampullisna uniplaga (9) and Teulisna spp. (10–12). Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 in NHMUK; 12 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654247" httpUri="https://zenodo.org/record/14654247/files/figure.png" pageId="4" pageNumber="76">Fig. 9</figureCitation>
|
||
). Uncus weakly sclerotised, dorso-ventrally flattened, relatively short, distally tapered and apically rounded, fused with dorsal wall of anal cone. Anal cone with very broad basal section occupying most of intrategumenal area. Tegumen with narrow and weakly sclerotised arms fused only apically (dorsally). Vinculum as long as tegumen but more heavily sclerotised, U-shaped. Valva lobular, gradually tapered distally, with apically rounded valvula. Tendon basally triangular, distally (entally) thin and ribbon-like, fused with its counterpart. Editum large, stretching distad along the costa and reaching the base of distal saccular process, bearing two ampullae. Proximal ampulla smoothly upcurved stake-shaped, apically pointed. Distal ampulla lobe-like flattened, with densely spinulose ventral margin. Sacculus moderately broad, its dorsal margin connected to juxta by membrane of annelifer while ental end firmly articulated with its counterpart. Distal saccular process thin but heavily sclerotised, upcurved, distally rugose and serrulate, and apically pointed. Juxta trapezoidal and weakly sclerotised, with trapezoidal caulis reaching the base of anellus. Phallus cylindrical, S-like curved, with short and rounded coecum. Proximal section of vesica tubular, somewhat tapered distally, bearing cluster of numerous denticles of various sizes dorsally and laterally. Medial section of vesica covered with spinulose scobination, with large clavate ventral diverticulum also densely covered with minute spinules and bearing cluster of four or five spikelike cornuti of various sizes. Vesica ejaculatorius originating distally.
|
||
<emphasis id="B962EC6EFFC5AA62FC4D6A6104C36DF2" bold="true" box="[994,1199,871,897]" pageId="4" pageNumber="76">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="132D2CF9FFC5AA62FB126A61057E6DF1" box="[1213,1298,871,898]" captionStart="Figures 13–17" captionStartId="5.[189,275,1879,1903]" captionTargetBox="[189,1380,241,1843]" captionTargetId="figure-114@5.[189,1398,223,1859]" captionTargetPageId="5" captionText="Figures 13–17. Female genitalia of Biampullisna uniplaga (13), Teulisna spp. (14, 15) and Witteulisna paraxena (17), and male genitalia of Witteulisna paraxena (16). Depositories of the specimens dissected: 13, 14, 16 and 17 in MWM/ZSM; 15 in OUMNH." figureDoi="http://doi.org/10.5281/zenodo.14654249" httpUri="https://zenodo.org/record/14654249/files/figure.png" pageId="4" pageNumber="76">Fig. 13</figureCitation>
|
||
). Papilla analis trapezoidal with rounded corners, weakly setose. Apophyses elongate and thin, anterior one somewhat longer and thicker than posterior one. 8
|
||
<superScript id="7C639D34FFC5AA62FC956AA103246DCB" attach="left" box="[826,840,935,952]" fontSize="7" pageId="4" pageNumber="76">th</superScript>
|
||
segment narrow and moderately sclerotised. Postvaginal area weakly sclerotised and rugose. Ventral margin of ostium bursae with deep medial Vshaped incision. Ductus bursae goblet-shaped, gelatinous. Corpus bursae pyriform, membranous. Appendix bursae short, conical, membranous, situated postero-ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6960F4FFC3AA64FF126E11014569DA" ID-DOI="http://doi.org/10.5281/zenodo.14654243" ID-Zenodo-Dep="14654243" httpUri="https://zenodo.org/record/14654243/files/figure.png" pageId="2" pageNumber="74" startId="2.[189,275,1815,1839]" targetBox="[197,1392,223,1778]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BA9307CFFC3AA64FF126E11014569DA" blockId="2.[189,1400,1815,1961]" pageId="2" pageNumber="74">
|
||
<emphasis id="B962EC6EFFC3AA64FF126E110128695C" bold="true" box="[189,324,1815,1839]" pageId="2" pageNumber="74">Figures 1–8</emphasis>
|
||
. Adults of
|
||
<taxonomicName id="4C164BFFFFC3AA64FE6C6E1102D5695C" authorityName="Volynkin" authorityYear="2024" baseAuthorityName="Hampson" baseAuthorityYear="1894" box="[451,697,1815,1839]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="species" species="uniplaga">
|
||
<emphasis id="B962EC6EFFC3AA64FE6C6E1102D5695C" box="[451,697,1815,1839]" italics="true" pageId="2" pageNumber="74">Biampullisna uniplaga</emphasis>
|
||
</taxonomicName>
|
||
(type species of
|
||
<taxonomicName id="4C164BFFFFC3AA64FCD86E110469695C" authorityName="Volynkin" authorityYear="2024" box="[887,1029,1815,1839]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B962EC6EFFC3AA64FCD86E110469695C" box="[887,1029,1815,1839]" italics="true" pageId="2" pageNumber="74">Biampullisna</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B962EC6EFFC3AA64FBA16E110437695C" bold="true" box="[1038,1115,1815,1839]" pageId="2" pageNumber="74">
|
||
<taxonomicNameLabel id="A2515115FFC3AA64FBA16E110437695C" box="[1038,1115,1815,1839]" pageId="2" pageNumber="74" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) (1, 2),
|
||
<taxonomicName id="4C164BFFFFC3AA64FB166E11051B695C" authorityName="Walker" authorityYear="1862" box="[1209,1399,1815,1839]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="species" species="plagiata">
|
||
<emphasis id="B962EC6EFFC3AA64FB166E11051B695C" box="[1209,1399,1815,1839]" italics="true" pageId="2" pageNumber="74">Teulisna plagiata</emphasis>
|
||
</taxonomicName>
|
||
(type species of
|
||
<taxonomicName id="4C164BFFFFC3AA64FEC36E3001A4693D" authorityName="Walker" authorityYear="1862" box="[364,456,1846,1870]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC3AA64FEC36E3001A4693D" box="[364,456,1846,1870]" italics="true" pageId="2" pageNumber="74">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
) (3, 4),
|
||
<taxonomicName id="4C164BFFFFC3AA64FDB06E3003DC693D" authority="Holloway, 2001" authorityName="Holloway" authorityYear="2001" box="[543,944,1846,1870]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="species" species="quadratella">
|
||
<emphasis id="B962EC6EFFC3AA64FDB06E300291693D" box="[543,765,1846,1870]" italics="true" pageId="2" pageNumber="74">Teulisna quadratella</emphasis>
|
||
<bibRefCitation id="EF874D8DFFC3AA64FCAB6E3003DC693D" author="Holloway, J. D." box="[772,944,1846,1870]" pageId="2" pageNumber="74" pagination="279 - 486" refId="ref2822" refString="Holloway, J. D. (2001) The Moths of Borneo, part 7. Family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 55, 279 - 486." type="journal article" year="2001">Holloway, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
(5),
|
||
<taxonomicName id="4C164BFFFFC3AA64FC4B6E30051A693D" authority="Volynkin, 2020" authorityName="Volynkin" authorityYear="2020" box="[996,1398,1846,1870]" class="Insecta" family="Erebidae" genus="Witteulisna" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="species" species="paraxena">
|
||
<emphasis id="B962EC6EFFC3AA64FC4B6E3004A5693D" box="[996,1225,1846,1870]" italics="true" pageId="2" pageNumber="74">Witteulisna paraxena</emphasis>
|
||
Volynkin, 2020
|
||
</taxonomicName>
|
||
(type species of
|
||
<taxonomicName id="4C164BFFFFC3AA64FEC26E52018A691F" authorityName="Volynkin" authorityYear="2020" box="[365,486,1876,1900]" class="Insecta" family="Erebidae" genus="Witteulisna" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC3AA64FEC26E52018A691F" box="[365,486,1876,1900]" italics="true" pageId="2" pageNumber="74">Witteulisna</emphasis>
|
||
</taxonomicName>
|
||
) (6), and
|
||
<taxonomicName id="4C164BFFFFC3AA64FDFE6E5203A1691F" authority="(Walker, [1865])" baseAuthorityName="Walker" baseAuthorityYear="1865" box="[593,973,1876,1900]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="species" species="basigera">
|
||
<emphasis id="B962EC6EFFC3AA64FDFE6E52037D691F" box="[593,785,1876,1900]" italics="true" pageId="2" pageNumber="74">Teulisna basigera</emphasis>
|
||
(Walker, [1865])
|
||
</taxonomicName>
|
||
(type species of
|
||
<taxonomicName id="4C164BFFFFC3AA64FB2B6E53051D691F" authority="Walker, [1865]" authorityName="Walker" authorityYear="1865" box="[1156,1393,1876,1900]" class="Insecta" family="Erebidae" genus="Cossa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC3AA64FB2B6E5304A9691F" box="[1156,1221,1877,1900]" italics="true" pageId="2" pageNumber="74">Cossa</emphasis>
|
||
Walker, [1865]
|
||
</taxonomicName>
|
||
, a junior synonym of
|
||
<taxonomicName id="4C164BFFFFC3AA64FE0E6E75019169F8" authorityName="Walker" authorityYear="1862" box="[417,509,1907,1931]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="74" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC3AA64FE0E6E75019169F8" box="[417,509,1907,1931]" italics="true" pageId="2" pageNumber="74">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
) (7, 8). Depositories of the specimens: 1, 3, 4 and 6–8 in MWM/ZSM; 2 and 5 in NHMUK.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6960F4FFC2AA65FF126E7E046169DC" ID-DOI="http://doi.org/10.5281/zenodo.14654247" ID-Zenodo-Dep="14654247" httpUri="https://zenodo.org/record/14654247/files/figure.png" pageId="3" pageNumber="75" startId="3.[189,275,1912,1936]" targetBox="[199,1398,263,1876]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BA9307CFFC2AA65FF126E7E046169DC" blockId="3.[189,1398,1912,1967]" pageId="3" pageNumber="75">
|
||
<emphasis id="B962EC6EFFC2AA65FF126E7E013969E3" bold="true" box="[189,341,1912,1936]" pageId="3" pageNumber="75">Figures 9–12</emphasis>
|
||
. Male genitalia of
|
||
<taxonomicName id="4C164BFFFFC2AA65FD9A6E7E034369E3" authorityName="Volynkin" authorityYear="2024" baseAuthorityName="Hampson" baseAuthorityYear="1894" box="[565,815,1912,1936]" class="Insecta" family="Erebidae" genus="Biampullisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="75" phylum="Arthropoda" rank="species" species="uniplaga">
|
||
<emphasis id="B962EC6EFFC2AA65FD9A6E7E034369E3" box="[565,815,1912,1936]" italics="true" pageId="3" pageNumber="75">Biampullisna uniplaga</emphasis>
|
||
</taxonomicName>
|
||
(9) and
|
||
<taxonomicName id="4C164BFFFFC2AA65FC366E7E044269E3" box="[921,1070,1912,1936]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="75" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B962EC6EFFC2AA65FC366E7E039969E3" box="[921,1013,1912,1936]" italics="true" pageId="3" pageNumber="75">Teulisna</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(10–12). Depositories of the specimens dissected: 9 and 10 in MWM/ZSM; 11 in NHMUK; 12 in OUMNH.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C63F7FFC5AA62FEB36D3703F86AFC" pageId="4" pageNumber="76" type="etymology">
|
||
<paragraph id="8BA9307CFFC5AA62FEB36D3703F86AFC" blockId="4.[189,1400,196,1167]" pageId="4" pageNumber="76">
|
||
<emphasis id="B962EC6EFFC5AA62FEB36D3701C86A38" bold="true" box="[284,420,1073,1099]" pageId="4" pageNumber="76">Etymology</emphasis>
|
||
. The new genus name is an aggregate of the Latin adjective ‘biampullalis’ meaning ‘double- ampullar’ and the genus- group name
|
||
<taxonomicName id="4C164BFFFFC5AA62FD4D6D55032B6A1D" authorityName="Walker" authorityYear="1862" box="[738,839,1107,1134]" class="Insecta" family="Erebidae" genus="Teulisna" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="76" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EC6EFFC5AA62FD4D6D55032B6A1D" box="[738,839,1107,1134]" italics="true" pageId="4" pageNumber="76">Teulisna</emphasis>
|
||
</taxonomicName>
|
||
, and refers to the presence of two ampullae in the male genitalia of the
|
||
<typeStatus id="54AD8EDEFFC5AA62FE4F6D73027E6AFC" box="[480,530,1141,1167]" pageId="4" pageNumber="76">type</typeStatus>
|
||
species. The gender is feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |