treatments-xml/data/03/A2/EB/03A2EB38FF88FF85FC64369BFD59FC6A.xml

<document id="2931766538A89928AEBD248AFA5637D4" ID-DOI="10.26028/cybium/2019-433-004" ID-ISSN="2101-0315" ID-Zenodo-Dep="12711337" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720643445570" checkinUser="felipe" docAuthor="Persat, Henri, Weiss, Steven, Froufe, Elsa, Secci-Petretto, Giulia &amp; Denys, Gaël P. J." docDate="2019" docId="03A2EB38FF88FF85FC64369BFD59FC6A" docLanguage="en" docName="Cybium.43.3.233-238.pdf" docOrigin="Cybium 43 (3)" docSource="http://dx.doi.org/10.26028/cybium/2019-433-004" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Thymallus ligericus Persat, Weiss, Froufe, Secci-Petretto &amp; Denys, 2019, n. sp." docType="treatment" docUuid="8864170D-9718-4FA4-85BD-DA32C60A66A0" docUuidSource="ZooBank" docVersion="2" lastPageNumber="238" masterDocId="FF9B9340FF89FF80FF843202FFE2FFFB" masterDocTitle="A third European species of grayling (Actinopterygii, Salmonidae), endemic to the Loire River basin (France), Thymallus ligericus n. sp." masterLastPageNumber="238" masterPageNumber="233" pageNumber="234" updateTime="1720649658292" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="870D345E52DFD292A52498826D143425">A third European species of grayling (Actinopterygii, Salmonidae), endemic to the Loire River basin (France), Thymallus ligericus n. sp.</mods:title>
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<mods:namePart id="53C005BA8A75865520083BA247491377">Secci-Petretto, Giulia</mods:namePart>
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<taxonomicName id="4C0B21ADFF88FF81FC64369BFB58FB4A" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[992,1210,1177,1201]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="1" pageNumber="234" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF88FF81FC64369BFB58FB4A" bold="true" box="[992,1210,1177,1201]" italics="true" pageId="1" pageNumber="234">Thymallus ligericus</emphasis>
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<paragraph id="8BB45A2EFF88FF81FBBD36B8FB40FB29" blockId="1.[1081,1186,1210,1234]" box="[1081,1186,1210,1234]" pageId="1" pageNumber="234">
(
<figureCitation id="133046ABFF88FF81FBC536B8FB78FB29" box="[1089,1178,1210,1234]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStartId-0="2.[1077,1144,876,898]" captionStartId-1="3.[132,197,996,1018]" captionStartId-2="3.[813,877,680,702]" captionStartId-3="4.[813,878,658,680]" captionStartId-4="4.[813,877,1197,1219]" captionTargetBox-0="[132,1039,206,1228]" captionTargetBox-1="[181,727,215,979]" captionTargetBox-2="[813,1456,215,662]" captionTargetBox-3="[814,1453,218,641]" captionTargetBox-4="[813,1456,833,1181]" captionTargetId-0="figure-423@2.[132,1039,206,1228]" captionTargetId-1="figure-526@3.[181,727,215,979]" captionTargetId-2="figure-603@3.[813,1456,215,662]" captionTargetId-3="figure-775@4.[813,1456,215,641]" captionTargetId-4="figure-734@4.[813,1456,833,1181]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="Figure 2. – Specimens of Thymallus spp. in lateral view. Holotype of Thymallus ligericus n. sp. MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (A) and after preservation (B); Paratype of Thymallus ligericus n. sp. MNHN 2019-0267 (C), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (D), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." captionText-1="Figure 3. – Head profiles of Thymallus spp. A: Paratype of Thymallus ligericus n. sp. MNHN 2018-0728, 300 mm SL J, Dore River at Vertolaye, 29 May 2015, Persat, Juglaret and Buttazzoni coll.; B: Thymallus thymallus UCBLZ – 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." captionText-2="Figure 4. – Mouth in ventral view of Thymallus spp. A: Holotype of Thymallus ligericus n. sp. MNHN 2018-0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll.; B: Thymallus thymallus UCBLZ 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." captionText-3="Figure 5. – Average dot numbers on each scale row on the flank for populations of Thymallus ligericus n. sp. from the Upper Ance (47 specimens), Lower Ance (7 specimens) and Lignon Rivers (16 specimens) in the Loire drainage as well as for Thymallus thymallus populations from the Ain (176 specimens) and Loue rivers (35 specimens) in the Rhône drainage." captionText-4="Figure 6. – Dorsal fin on a live paratype of Thymallus ligericus n. sp. MNHN 2018-0726, 290 mm SL ♀, Lignon River at Boën-sur- Lignon, 29 May 2015, Persat, Juglaret and Buttazzoni coll." figureDoi-0="http://doi.org/10.5281/zenodo.12711341" figureDoi-1="http://doi.org/10.5281/zenodo.12711347" figureDoi-2="http://doi.org/10.5281/zenodo.12711343" figureDoi-3="http://doi.org/10.5281/zenodo.12711349" figureDoi-4="http://doi.org/10.5281/zenodo.12711351" httpUri-0="https://zenodo.org/record/12711341/files/figure.png" httpUri-1="https://zenodo.org/record/12711347/files/figure.png" httpUri-2="https://zenodo.org/record/12711343/files/figure.png" httpUri-3="https://zenodo.org/record/12711349/files/figure.png" httpUri-4="https://zenodo.org/record/12711351/files/figure.png" pageId="1" pageNumber="234">Figs 2-6</figureCitation>
)
</paragraph>
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<emphasis id="B97F863CFF88FF81FCA936F8FBE6FAE9" bold="true" box="[813,1028,1274,1298]" pageId="1" pageNumber="234">Material examined</emphasis>
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<typeStatus id="54B0E48CFF88FF81FCD63718FC5BFAC9" box="[850,953,1306,1330]" pageId="1" pageNumber="234" type="holotype">Holotype</typeStatus>
. –
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<specimenCode id="DBADF255FF88FF81FC5D3718FB49FAC9" box="[985,1195,1306,1330]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0722</specimenCode>
, 
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SL J, 
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: 
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drainage, Cantal Dept., 
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at 
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: 
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, 
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, Persat and FDAAPPMA15 coll.
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</paragraph>
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<typeStatus id="54B0E48CFF88FF81FCD63799FC2BFA49" box="[850,969,1435,1458]" pageId="1" pageNumber="234" type="paratype">Paratypes</typeStatus>
. –
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<specimenCode id="DBADF255FF88FF81FC6F3798FB27FA49" box="[1003,1221,1434,1458]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0723</specimenCode>
, 
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SL 
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, collected together with the 
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; 
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<specimenCode id="DBADF255FF88FF81FB4E37B8FA48FA29" box="[1226,1450,1466,1490]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0724</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FCA937D8FC6FFA0A" box="[813,909,1498,1522]" metricMagnitude="-1" metricUnit="m" metricValue="2.52" pageId="1" pageNumber="234" unit="mm" value="252.0">252 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF88FF81FC4637D8FC34FA0A" box="[962,982,1498,1521]" count="1" pageId="1" pageNumber="234" type="female">♀</specimenCount>
, 
<collectingCountry id="F31C1ABEFF88FF81FC6C37D8FBDCFA0A" box="[1000,1086,1498,1521]" name="France" pageId="1" pageNumber="234">France</collectingCountry>
: 
<collectingRegion id="49CF94CCFF88FF81FBCE37D8FB69FA09" box="[1098,1163,1498,1522]" country="France" name="Pays de la Loire" pageId="1" pageNumber="234">Loire</collectingRegion>
drainage, 
<collectingCounty id="62D522A2FF88FF81FA8837D8FC8EF9EA" pageId="1" pageNumber="234">Haute-Vienne Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF88FF81FCF337F8FBC1F9E9" box="[887,1059,1530,1554]" pageId="1" pageNumber="234">Combade River</collectingMunicipality>
at 
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: 
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, 
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<collectingDate id="EFF18506FF88FF81FCA93418FC29F9C9" box="[813,971,1562,1586]" pageId="1" pageNumber="234" value="2015-05-28">28 May 2015</collectingDate>
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, 
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and FDAAPPMA87 coll
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.; 
<materialsCitation id="3B635073FF88FF81FADF3418FB21F989" collectionCode="MNHN" country="France" location="Masleon" municipality="Combade River" pageId="1" pageNumber="234" specimenCode="MNHN 2018-0725" specimenCount="1" specimenCount-female="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF88FF81FADF3418FC49F9A9" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0725</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FC3C3438FBF7F9AA" box="[952,1045,1594,1618]" metricMagnitude="-1" metricUnit="m" metricValue="2.52" pageId="1" pageNumber="234" unit="mm" value="252.0">252 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF88FF81FBCC3438FBBEF9AA" box="[1096,1116,1594,1617]" count="1" pageId="1" pageNumber="234" type="female">♀</specimenCount>
, 
<collectingMunicipality id="6BD0C054FF88FF81FBE83438FAC3F9A9" box="[1132,1313,1594,1618]" pageId="1" pageNumber="234">Combade River</collectingMunicipality>
at 
<location id="8ED40CF5FF88FF81FAC23438FA49F9A9" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF88FF81FAC23438FA49F9A9" box="[1350,1451,1594,1618]" country="France" municipality="Combade River" name="Masleon" pageId="1" pageNumber="234" stateProvince="Pays de la Loire">Masléon</location>
, same date, location, and collectors
</materialsCitation>
; 
<materialsCitation id="3B635073FF88FF81FB4B3458FBA7F929" collectingDate="2015-05-29" collectionCode="MNHN" collectorName="Persat &amp; Juglaret &amp; Buttazzoni" country="France" county="Dept." latitude="45.751667" location="Boen-sur-Lignon" longLatPrecision="18" longitude="3.9925" municipality="Lignon River" pageId="1" pageNumber="234" specimenCode="MNHN 2018-0726" specimenCount="1" specimenCount-female="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF88FF81FB4B3458FA49F989" box="[1231,1451,1626,1650]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0726</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FCA93478FC69F96A" box="[813,907,1658,1682]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="234" unit="mm" value="290.0">290 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF88FF81FC3A3478FC30F96A" box="[958,978,1658,1681]" count="1" pageId="1" pageNumber="234" type="female">♀</specimenCount>
, 
<collectingRegion id="49CF94CCFF88FF81FC663478FBC2F969" box="[994,1056,1658,1682]" country="France" name="Pays de la Loire" pageId="1" pageNumber="234">Loire</collectingRegion>
<collectingCounty id="62D522A2FF88FF81FBAE3478FB8EF96A" box="[1066,1132,1658,1681]" pageId="1" pageNumber="234">Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF88FF81FBFE3478FAF4F969" box="[1146,1302,1658,1682]" pageId="1" pageNumber="234">Lignon River</collectingMunicipality>
at 
<location id="8ED40CF5FF88FF81FAB83478FC62F949" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF88FF81FAB83478FC62F949" country="France" county="Dept." latitude="45.751667" longLatPrecision="18" longitude="3.9925" municipality="Lignon River" name="Boen-sur-Lignon" pageId="1" pageNumber="234" stateProvince="Pays de la Loire">Boën-sur-Lignon</location>
: 
<geoCoordinate id="EE3F3CE9FF88FF81FC0E3498FBF2F949" box="[906,1040,1690,1714]" degrees="45" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="234" precision="15" seconds="6" value="45.751667">45°45’6”N-</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF88FF81FB943498FB63F94A" box="[1040,1153,1690,1714]" degrees="3" direction="east" minutes="59" orientation="longitude" pageId="1" pageNumber="234" precision="15" seconds="33" value="3.9925">3°59’33”E</geoCoordinate>
, 
<date id="FFB57CEEFF88FF81FB0B3498FAC1F949" box="[1167,1315,1690,1714]" pageId="1" pageNumber="234" value="2015-05-29">
<collectingDate id="EFF18506FF88FF81FB0B3498FAC1F949" box="[1167,1315,1690,1714]" pageId="1" pageNumber="234" value="2015-05-29">29 May 2015</collectingDate>
</date>
, 
<collectorName id="26FE3FF8FF88FF81FAAB3498FA97F94A" box="[1327,1397,1690,1713]" pageId="1" pageNumber="234">Persat</collectorName>
, 
<collectorName id="26FE3FF8FF88FF81FA043498FCBCF929" pageId="1" pageNumber="234">Juglaret</collectorName>
and 
<collectorName id="26FE3FF8FF88FF81FC1734B8FBE8F929" box="[915,1034,1722,1746]" pageId="1" pageNumber="234">Buttazzoni</collectorName>
coll.
</materialsCitation>
; 
<materialsCitation id="3B635073FF88FF81FBD434B8FB8EF8E9" collectionCode="MNHN" country="France" county="Dept." location="Boen-sur-Lignon" municipality="Lignon River" pageId="1" pageNumber="234" specimenCode="MNHN 2018-0727" specimenCount="1" specimenCount-female="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF88FF81FBD434B8FAC6F929" box="[1104,1316,1722,1746]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0727</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FAAB34B8FA6BF92A" box="[1327,1417,1722,1746]" metricMagnitude="-1" metricUnit="m" metricValue="2.78" pageId="1" pageNumber="234" unit="mm" value="278.0">278 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF88FF81FCA934D8FCA3F90A" box="[813,833,1754,1777]" count="1" pageId="1" pageNumber="234" type="female">♀</specimenCount>
, 
<collectingRegion id="49CF94CCFF88FF81FCD534D8FC6DF909" box="[849,911,1754,1778]" country="France" name="Pays de la Loire" pageId="1" pageNumber="234">Loire</collectingRegion>
<collectingCounty id="62D522A2FF88FF81FC1334D8FC3BF90A" box="[919,985,1754,1777]" pageId="1" pageNumber="234">Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF88FF81FC6134D8FB9FF909" box="[997,1149,1754,1778]" pageId="1" pageNumber="234">Lignon River</collectingMunicipality>
at 
<location id="8ED40CF5FF88FF81FB2634D8FA88F909" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF88FF81FB2634D8FA88F909" box="[1186,1386,1754,1778]" country="France" county="Dept." municipality="Lignon River" name="Boen-sur-Lignon" pageId="1" pageNumber="234" stateProvince="Pays de la Loire">Boën-sur-Lignon</location>
: same date, location, and collectors
</materialsCitation>
; 
<materialsCitation id="3B635073FF88FF81FBF334F8FC3AF889" collectingDate="2015-05-29" collectionCode="MNHN" collectorName="Persat &amp; Juglaret &amp; Buttazzoni" country="France" county="Puy-de-Dome Dept." latitude="48.63639" location="Vertolaye" longLatPrecision="18" longitude="3.706389" municipality="Dore River" pageId="1" pageNumber="234" specimenCode="MNHN 2018-0728" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF88FF81FBF334F8FAA8F8E9" box="[1143,1354,1786,1810]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2018-0728</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FAD134F8FA4DF8EA" box="[1365,1455,1786,1810]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="234" unit="mm" value="300.0">300 mm</quantity>
SL J, 
<collectingCounty id="62D522A2FF88FF81FCFB3518FB97F8CA" box="[895,1141,1818,1842]" pageId="1" pageNumber="234">Puy-de-Dôme Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF88FF81FB073518FAE9F8C9" box="[1155,1291,1818,1842]" pageId="1" pageNumber="234">Dore River</collectingMunicipality>
at 
<location id="8ED40CF5FF88FF81FAB03518FA4EF8C9" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF88FF81FAB03518FA4EF8C9" box="[1332,1452,1818,1842]" country="France" county="Puy-de-Dome Dept." latitude="48.63639" longLatPrecision="18" longitude="3.706389" municipality="Dore River" name="Vertolaye" pageId="1" pageNumber="234" stateProvince="Pays de la Loire">Vertolaye</location>
: 
<geoCoordinate id="EE3F3CE9FF88FF81FCA93538FC5FF8A9" box="[813,957,1850,1874]" degrees="48" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="234" precision="15" seconds="11" value="48.63639">48°38’11”N-</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF88FF81FC393538FBCCF8AA" box="[957,1070,1850,1874]" degrees="3" direction="east" minutes="42" orientation="longitude" pageId="1" pageNumber="234" precision="15" seconds="23" value="3.706389">3°42’23”E</geoCoordinate>
, 
<date id="FFB57CEEFF88FF81FBB83538FB32F8A9" box="[1084,1232,1850,1874]" pageId="1" pageNumber="234" value="2015-05-29">
<collectingDate id="EFF18506FF88FF81FBB83538FB32F8A9" box="[1084,1232,1850,1874]" pageId="1" pageNumber="234" value="2015-05-29">29 May 2015</collectingDate>
</date>
, 
<collectorName id="26FE3FF8FF88FF81FB5E3538FAC2F8AA" box="[1242,1312,1850,1873]" pageId="1" pageNumber="234">Persat</collectorName>
, 
<collectorName id="26FE3FF8FF88FF81FAAF3538FA60F8A9" box="[1323,1410,1850,1874]" pageId="1" pageNumber="234">Juglaret</collectorName>
and 
<collectorName id="26FE3FF8FF88FF81FCA93558FC43F889" box="[813,929,1882,1906]" pageId="1" pageNumber="234">Buttazzoni</collectorName>
coll.
</materialsCitation>
; 
<materialsCitation id="3B635073FF88FF82FC663558FEC1FAD7" collectionCode="MNHN" country="France" county="Puy-de-Dome Dept." lastPageId="2" lastPageNumber="235" location="Vertolaye" municipality="Dore River" pageId="1" pageNumber="234" specimenCode="MNHN 2019-0266" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF88FF81FC663558FB52F889" box="[994,1200,1882,1906]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="234">MNHN 2019-0266</specimenCode>
, 
<quantity id="4CF3F7CBFF88FF81FB3D3558FAF2F88A" box="[1209,1296,1882,1906]" metricMagnitude="-1" metricUnit="m" metricValue="2.84" pageId="1" pageNumber="234" unit="mm" value="284.0">284 mm</quantity>
SL J, 
<collectingCounty id="62D522A2FF88FF82FADF3558FEEFFAF0" lastPageId="2" lastPageNumber="235" pageId="1" pageNumber="234">Puy-de-Dôme Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF8BFF82FE9C36F6FE71FAF7" box="[280,403,1268,1292]" pageId="2" pageNumber="235">Dore River</collectingMunicipality>
at 
<location id="8ED40CF5FF8BFF82FE3036F6FDC0FAF7" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FE3036F6FDC0FAF7" box="[436,546,1268,1292]" country="France" county="Puy-de-Dome Dept." municipality="Dore River" name="Vertolaye" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Vertolaye</location>
: same date, location, and collectors
</materialsCitation>
; 
<materialsCitation id="3B635073FF8BFF82FEAB3717FEC4FA96" collectingDate="2016-09-22" collectionCode="MNHN" country="France" county="Haute-Loire Dept." latitude="44.968613" location="Chadron" longLatPrecision="19" longitude="3.9197223" municipality="River" pageId="2" pageNumber="235" specimenCode="MNHN 2019-0267" specimenCount="1" specimenCount-female="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF8BFF82FEAB3717FDE7FAD7" box="[303,517,1300,1324]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="235">MNHN 2019-0267</specimenCode>
, 
<quantity id="4CF3F7CBFF8BFF82FD943716FD89FAD7" box="[528,619,1300,1324]" metricMagnitude="-1" metricUnit="m" metricValue="2.69" pageId="2" pageNumber="235" unit="mm" value="269.0">269 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF8BFF82FD1D3717FD4FFAD7" box="[665,685,1301,1324]" count="1" pageId="2" pageNumber="235" type="female">♀</specimenCount>
, 
<collectingCounty id="62D522A2FF8BFF82FD383717FEE0FAB7" pageId="2" pageNumber="235">Haute-Loire Dept.</collectingCounty>
, 
<collectingRegion id="49CF94CCFF8BFF82FE883737FEA4FAB6" box="[268,326,1333,1357]" country="France" name="Pays de la Loire" pageId="2" pageNumber="235">Loire</collectingRegion>
<collectingMunicipality id="6BD0C054FF8BFF82FEC83737FE65FAB6" box="[332,391,1333,1357]" pageId="2" pageNumber="235">River</collectingMunicipality>
at 
<location id="8ED40CF5FF8BFF82FE233737FDE5FAB6" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FE233737FDE5FAB6" box="[423,519,1333,1357]" country="France" county="Haute-Loire Dept." latitude="44.968613" longLatPrecision="19" longitude="3.9197223" municipality="River" name="Chadron" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Chadron</location>
: 
<geoCoordinate id="EE3F3CE9FF8BFF82FD953737FD71FAB6" box="[529,659,1333,1357]" degrees="44" direction="north" minutes="58" orientation="latitude" pageId="2" pageNumber="235" precision="15" seconds="7" value="44.968613">44°58’7”N-</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF8BFF82FD173737FCE2FAB7" box="[659,768,1333,1357]" degrees="3" direction="east" minutes="55" orientation="longitude" pageId="2" pageNumber="235" precision="15" seconds="11" value="3.9197223">3°55’11”E</geoCoordinate>
, 
<date id="FFB57CEEFF8BFF82FF003757FEC4FA96" box="[132,294,1365,1389]" pageId="2" pageNumber="235" value="2016-09-22">
<collectingDate id="EFF18506FF8BFF82FF003757FEC4FA96" box="[132,294,1365,1389]" pageId="2" pageNumber="235" value="2016-09-22">22 Sep. 2016</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8BFF82FEB03754FE50FA96" box="[308,434,1365,1389]" country="France" location="Persat" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<location id="8ED40CF5FF8BFF82FEB03754FE9CFA96" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FEB03754FE9CFA96" box="[308,382,1366,1389]" country="France" name="Persat" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Persat</location>
and
</materialsCitation>
<materialsCitation id="3B635073FF8BFF82FE3F3757FD9DFA96" box="[443,639,1365,1389]" collectionCode="ONEMA" country="France" pageId="2" pageNumber="235" specimenCode="ONEMA43" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<specimenCode id="DBADF255FF8BFF82FE3F3757FDA3FA96" box="[443,577,1365,1389]" collectionCode="ONEMA" pageId="2" pageNumber="235">ONEMA43</specimenCode>
coll
</materialsCitation>
.; 
<materialsCitation id="3B635073FF8BFF82FD163757FD53FA55" collectingDate="2016-09-22" collectionCode="UCBLZ" country="France" county="Ardeche Dept." latitude="44.631664" location="Laveyrune" municipality="Allier River" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<collectionCode id="ED1AC2EBFF8BFF82FD163757FD12FA96" box="[658,752,1365,1389]" pageId="2" pageNumber="235">UCBLZ</collectionCode>
– 2019.9. 
<quantity id="4CF3F7CBFF8BFF82FF5C3774FE91FA76" box="[216,371,1398,1422]" metricMagnitude="3" metricUnit="m" metricValue="1.0093039999999998" pageId="2" pageNumber="235" unit="mm" value="1009304.0">1009, 304 mm</quantity>
SL J, 
<collectingCounty id="62D522A2FF8BFF82FE3B3777FD83FA76" box="[447,609,1397,1421]" pageId="2" pageNumber="235">Ardèche Dept.</collectingCounty>
, 
<collectingMunicipality id="6BD0C054FF8BFF82FDEE3777FD0EFA76" box="[618,748,1397,1421]" pageId="2" pageNumber="235">Allier River</collectingMunicipality>
at 
<location id="8ED40CF5FF8BFF82FF003794FF1DFA55" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FF003794FF1DFA55" box="[132,255,1430,1454]" country="France" county="Ardeche Dept." latitude="44.631664" municipality="Allier River" name="Laveyrune" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Laveyrune</location>
: 
<geoCoordinate id="EE3F3CE9FF8BFF82FE8E3794FE7CFA55" box="[266,414,1430,1454]" degrees="44" direction="north" minutes="37" orientation="latitude" pageId="2" pageNumber="235" precision="15" seconds="54" value="44.631664">44°37’54”N-</geoCoordinate>
3°53’40’E, 
<date id="FFB57CEEFF8BFF82FD983794FD53FA55" box="[540,689,1430,1454]" pageId="2" pageNumber="235" value="2016-09-22">
<collectingDate id="EFF18506FF8BFF82FD983794FD53FA55" box="[540,689,1430,1454]" pageId="2" pageNumber="235" value="2016-09-22">22 Sep. 2016</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8BFF82FD383794FD1CFA35" country="France" location="Persat" pageId="2" pageNumber="235" specimenCount="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<location id="8ED40CF5FF8BFF82FD383794FCE1FA56" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FD383794FCE1FA56" box="[700,771,1430,1453]" country="France" name="Persat" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Persat</location>
, FDAAPPMA09, FDAAPPMA43 and FDAAPPMA48 coll
</materialsCitation>
.; 
<materialsCitation id="3B635073FF8BFF82FF0037D5FDE8F9D4" collectingDate="2016-06-27" collectionCode="UCBLZ" collectorName="Persat &amp; Lesbre" country="France" latitude="45.17806" location="Alagnon River" longLatPrecision="19" longitude="3.8952777" municipality="Cantal Dept." pageId="2" pageNumber="235" specimenCount="1" specimenCount-female="1" stateProvince="Pays de la Loire" typeStatus="paratype">
<collectionCode id="ED1AC2EBFF8BFF82FF0037D5FF3EFA14" box="[132,220,1495,1519]" pageId="2" pageNumber="235">UCBLZ</collectionCode>
– 2012.9. 
<quantity id="4CF3F7CBFF8BFF82FECF37D5FE04FA15" box="[331,486,1495,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.008405" pageId="2" pageNumber="235" unit="mm" value="1008405.0">1008, 405 mm</quantity>
SL 
<specimenCount id="9D0D91A7FF8BFF82FD9637D5FDC4FA15" box="[530,550,1495,1518]" count="1" pageId="2" pageNumber="235" type="female">♀</specimenCount>
, 
<collectingMunicipality id="6BD0C054FF8BFF82FDB037D5FD22FA15" box="[564,704,1495,1519]" pageId="2" pageNumber="235">Cantal Dept.</collectingMunicipality>
, 
<location id="8ED40CF5FF8BFF82FD4D37D5FF0DF9F4" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8BFF82FD4D37D5FF0DF9F4" country="France" latitude="45.17806" longLatPrecision="19" longitude="3.8952777" municipality="Cantal Dept." name="Alagnon River" pageId="2" pageNumber="235" stateProvince="Pays de la Loire">Alagnon River</location>
at Ferrières-Saint-Mary: 
<geoCoordinate id="EE3F3CE9FF8BFF82FD8437F5FD73F9F4" box="[512,657,1527,1551]" degrees="45" direction="north" minutes="10" orientation="latitude" pageId="2" pageNumber="235" precision="15" seconds="41" value="45.17806">45°10’41”N-</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF8BFF82FD1437F5FCE2F9F4" box="[656,768,1527,1551]" degrees="3" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="235" precision="15" seconds="43" value="3.8952777">3°53’43”E</geoCoordinate>
, 
<date id="FFB57CEEFF8BFF82FF00341AFEEDF9D4" box="[132,271,1559,1584]" pageId="2" pageNumber="235" value="2016-06-27">
<collectingDate id="EFF18506FF8BFF82FF00341AFEEDF9D4" box="[132,271,1559,1584]" pageId="2" pageNumber="235" value="2016-06-27">27 Jun. 2016</collectingDate>
</date>
, 
<collectorName id="26FE3FF8FF8BFF82FE9D341AFEB8F9D4" box="[281,346,1560,1583]" pageId="2" pageNumber="235">Persat</collectorName>
and 
<collectorName id="26FE3FF8FF8BFF82FE083415FE37F9D4" box="[396,469,1559,1583]" pageId="2" pageNumber="235">Lesbre</collectorName>
coll.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF740AA6FF8BFF82FBB1316EFB82FB29" ID-DOI="http://doi.org/10.5281/zenodo.12711341" ID-Zenodo-Dep="12711341" httpUri="https://zenodo.org/record/12711341/files/figure.png" pageId="2" pageNumber="235" startId="2.[1077,1144,876,898]" targetBox="[132,1039,206,1228]" targetPageId="2" targetType="figure">
<paragraph id="8BB45A2EFF8BFF82FBB1316EFB82FB29" blockId="2.[1077,1456,876,1234]" pageId="2" pageNumber="235">
Figure 2. – Specimens of 
<taxonomicName id="4C0B21ADFF8BFF82FAC1316EFBBDFC62" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B97F863CFF8BFF82FAC1316EFA52FC7A" box="[1349,1456,876,897]" italics="true" pageId="2" pageNumber="235">Thymallus</emphasis>
spp.
</taxonomicName>
in lateral view. Holotype of 
<taxonomicName id="4C0B21ADFF8BFF82FA063186FB39FC4A" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8BFF82FA063186FB39FC4A" italics="true" pageId="2" pageNumber="235">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8BFF82FB66319EFAF8FC4A" box="[1250,1306,924,945]" pageId="2" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (
<emphasis id="B97F863CFF8BFF82FBEB31FEFB62FBEA" bold="true" box="[1135,1152,1020,1041]" pageId="2" pageNumber="235">A</emphasis>
) and after preservation (
<emphasis id="B97F863CFF8BFF82FA0A31FEFA7CFBEA" bold="true" box="[1422,1438,1020,1041]" pageId="2" pageNumber="235">B</emphasis>
); Paratype of 
<taxonomicName id="4C0B21ADFF8BFF82FB2A3616FA91FBD2" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[1198,1395,1044,1065]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8BFF82FB2A3616FA91FBD2" box="[1198,1395,1044,1065]" italics="true" pageId="2" pageNumber="235">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8BFF82FAFE3616FA52FBD2" box="[1402,1456,1044,1065]" pageId="2" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2019-0267 (
<emphasis id="B97F863CFF8BFF82FB7F362EFAEEFBB9" bold="true" box="[1275,1292,1068,1090]" pageId="2" pageNumber="235">C</emphasis>
), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (
<emphasis id="B97F863CFF8BFF82FB463676FB31FB72" bold="true" box="[1218,1235,1140,1161]" pageId="2" pageNumber="235">D</emphasis>
), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll.
</paragraph>
</caption>
<subSubSection id="C31109A5FF8BFF82FF00345AFC49FA75" pageId="2" pageNumber="235" type="diagnosis">
<paragraph id="8BB45A2EFF8BFF82FF00345AFF10F98B" blockId="2.[132,775,1624,1907]" box="[132,242,1624,1648]" pageId="2" pageNumber="235">
<emphasis id="B97F863CFF8BFF82FF00345AFF10F98B" bold="true" box="[132,242,1624,1648]" pageId="2" pageNumber="235">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8BFF82FF2E347AFC49FA75" blockId="2.[132,775,1624,1907]" lastBlockId="2.[813,1456,1268,1422]" pageId="2" pageNumber="235">
<taxonomicName id="4C0B21ADFF8BFF82FF2E347AFE64F96B" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[170,390,1656,1680]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8BFF82FF2E347AFE64F96B" box="[170,390,1656,1680]" italics="true" pageId="2" pageNumber="235">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8BFF82FE09347BFE2BF96B" box="[397,457,1657,1680]" pageId="2" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C0B21ADFF8BFF82FDD9347BFD07F96B" authorityName="Linnaeus" authorityYear="1758" box="[605,741,1656,1680]" family="Salmonidae" genus="Salmo" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8BFF82FDD9347BFD07F96B" box="[605,741,1656,1680]" italics="true" pageId="2" pageNumber="235">T. thymallus</emphasis>
</taxonomicName>
by a more elongated body (
<figureCitation id="133046ABFF8BFF82FE17349BFE3BF94A" box="[403,473,1689,1713]" captionStart="Figure 2" captionStartId="2.[1077,1144,876,898]" captionTargetBox="[132,1039,206,1228]" captionTargetId="figure-423@2.[132,1039,206,1228]" captionTargetPageId="2" captionText="Figure 2. – Specimens of Thymallus spp. in lateral view. Holotype of Thymallus ligericus n. sp. MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (A) and after preservation (B); Paratype of Thymallus ligericus n. sp. MNHN 2019-0267 (C), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (D), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711341" httpUri="https://zenodo.org/record/12711341/files/figure.png" pageId="2" pageNumber="235">Fig. 2</figureCitation>
), the presence of a pointed snout revealing a strait or convex snout profile (
<figureCitation id="133046ABFF8BFF82FD0434BBFD30F92A" box="[640,722,1721,1745]" captionStart="Figure 3" captionStartId="3.[132,197,996,1018]" captionTargetBox="[181,727,215,979]" captionTargetId="figure-526@3.[181,727,215,979]" captionTargetPageId="3" captionText="Figure 3. – Head profiles of Thymallus spp. A: Paratype of Thymallus ligericus n. sp. MNHN 2018-0728, 300 mm SL J, Dore River at Vertolaye, 29 May 2015, Persat, Juglaret and Buttazzoni coll.; B: Thymallus thymallus UCBLZ – 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711347" httpUri="https://zenodo.org/record/12711347/files/figure.png" pageId="2" pageNumber="235">Fig. 3A</figureCitation>
) (
<emphasis id="B97F863CFF8BFF82FD6E34B8FCE5F92A" box="[746,775,1722,1745]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
a concave profile for 
<taxonomicName id="4C0B21ADFF8BFF82FEEF34D8FE0DF90A" authorityName="Linnaeus" authorityYear="1758" box="[363,495,1753,1777]" family="Salmonidae" genus="Salmo" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8BFF82FEEF34D8FE0DF90A" box="[363,495,1753,1777]" italics="true" pageId="2" pageNumber="235">T. thymallus</emphasis>
</taxonomicName>
, 
<figureCitation id="133046ABFF8BFF82FE7834DBFDB2F909" box="[508,592,1753,1778]" captionStart="Figure 3" captionStartId="3.[132,197,996,1018]" captionTargetBox="[181,727,215,979]" captionTargetId="figure-526@3.[181,727,215,979]" captionTargetPageId="3" captionText="Figure 3. – Head profiles of Thymallus spp. A: Paratype of Thymallus ligericus n. sp. MNHN 2018-0728, 300 mm SL J, Dore River at Vertolaye, 29 May 2015, Persat, Juglaret and Buttazzoni coll.; B: Thymallus thymallus UCBLZ – 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711347" httpUri="https://zenodo.org/record/12711347/files/figure.png" pageId="2" pageNumber="235">Fig. 3B</figureCitation>
), a more inferior mouth with a transversal aperture and a fleshier upper lip (
<figureCitation id="133046ABFF8BFF82FF093518FF04F8C9" box="[141,230,1818,1842]" captionStart="Figure 4" captionStartId="3.[813,877,680,702]" captionTargetBox="[813,1456,215,662]" captionTargetId="figure-603@3.[813,1456,215,662]" captionTargetPageId="3" captionText="Figure 4. – Mouth in ventral view of Thymallus spp. A: Holotype of Thymallus ligericus n. sp. MNHN 2018-0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll.; B: Thymallus thymallus UCBLZ 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711343" httpUri="https://zenodo.org/record/12711343/files/figure.png" pageId="2" pageNumber="235">Fig. 4A</figureCitation>
), a high number of black dots on the flanks (usually from more than 50 to several hundreds 
<emphasis id="B97F863CFF8BFF82FDE63539FD62F8A9" box="[610,640,1851,1874]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
less than 
<quantity id="4CF3F7CBFF8BFF82FD683538FF78F888" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="235" unit="in" value="50.0">50 in</quantity>
the Rhône populations; 
<figureCitation id="133046ABFF8BFF82FE2C3559FE0DF888" box="[424,495,1883,1907]" captionStart="Figure 2" captionStartId="2.[1077,1144,876,898]" captionTargetBox="[132,1039,206,1228]" captionTargetId="figure-423@2.[132,1039,206,1228]" captionTargetPageId="2" captionText="Figure 2. – Specimens of Thymallus spp. in lateral view. Holotype of Thymallus ligericus n. sp. MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (A) and after preservation (B); Paratype of Thymallus ligericus n. sp. MNHN 2019-0267 (C), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (D), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711341" httpUri="https://zenodo.org/record/12711341/files/figure.png" pageId="2" pageNumber="235">Figs 2</figureCitation>
, 
<figureCitation id="133046ABFF8BFF82FE7F3559FDE8F888" box="[507,522,1883,1907]" captionStart="Figure 5" captionStartId="4.[813,878,658,680]" captionTargetBox="[814,1453,218,641]" captionTargetId="figure-775@4.[813,1456,215,641]" captionTargetPageId="4" captionText="Figure 5. – Average dot numbers on each scale row on the flank for populations of Thymallus ligericus n. sp. from the Upper Ance (47 specimens), Lower Ance (7 specimens) and Lignon Rivers (16 specimens) in the Loire drainage as well as for Thymallus thymallus populations from the Ain (176 specimens) and Loue rivers (35 specimens) in the Rhône drainage." figureDoi="http://doi.org/10.5281/zenodo.12711349" httpUri="https://zenodo.org/record/12711349/files/figure.png" pageId="2" pageNumber="235">5</figureCitation>
), a shorter head length (20.0-23.7% SL 
<emphasis id="B97F863CFF8BFF82FC6036F7FBE0FAF7" box="[996,1026,1269,1292]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
17.5-19.9% SL), smaller occipital and orbital depths of the head (respectively: 54.8-71.7% HL 
<emphasis id="B97F863CFF8BFF82FA163717FA52FAD7" box="[1426,1456,1301,1324]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
73.2-76.1% HL; 43.5-53.4% HL 
<emphasis id="B97F863CFF8BFF82FB123737FB56FAB7" box="[1174,1204,1333,1356]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
51.1-56.4% HL) and a shorter horizontal eye diameter (22.6-26.3% HL 
<emphasis id="B97F863CFF8BFF82FAD43754FA8CFA96" box="[1360,1390,1366,1389]" italics="true" pageId="2" pageNumber="235">vs.</emphasis>
26.1- 31.7% HL).
</paragraph>
</subSubSection>
<subSubSection id="C31109A5FF8BFF83FCA937B4FA92FA12" lastPageId="3" lastPageNumber="236" pageId="2" pageNumber="235" type="description">
<paragraph id="8BB45A2EFF8BFF82FCA937B4FC53FA35" blockId="2.[813,1456,1462,1907]" box="[813,945,1462,1486]" pageId="2" pageNumber="235">
<emphasis id="B97F863CFF8BFF82FCA937B4FC53FA35" bold="true" box="[813,945,1462,1486]" pageId="2" pageNumber="235">Description</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8BFF82FCD637D4FB5CF9F4" blockId="2.[813,1456,1462,1907]" pageId="2" pageNumber="235">
Morphometric measurements are given in table I. The 
<typeStatus id="54B0E48CFF8BFF82FCA937F5FC68F9F4" box="[813,906,1527,1551]" pageId="2" pageNumber="235" type="holotype">holotype</typeStatus>
counts are given in brackets.
</paragraph>
<paragraph id="8BB45A2EFF8BFF83FCD63415FD57FB17" blockId="2.[813,1456,1462,1907]" lastBlockId="3.[132,775,1172,1260]" lastPageId="3" lastPageNumber="236" pageId="2" pageNumber="235">
<taxonomicName id="4C0B21ADFF8BFF82FCD63415FBD3F9D4" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[850,1073,1559,1583]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8BFF82FCD63415FBD3F9D4" box="[850,1073,1559,1583]" italics="true" pageId="2" pageNumber="235">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8BFF82FBBD341AFB94F9D4" box="[1081,1142,1560,1583]" pageId="2" pageNumber="235" rank="species">n. sp.</taxonomicNameLabel>
has an elongated body (
<figureCitation id="133046ABFF8BFF82FA0C3415FCD8F9AB" captionStart="Figure 2" captionStartId="2.[1077,1144,876,898]" captionTargetBox="[132,1039,206,1228]" captionTargetId="figure-423@2.[132,1039,206,1228]" captionTargetPageId="2" captionText="Figure 2. – Specimens of Thymallus spp. in lateral view. Holotype of Thymallus ligericus n. sp. MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (A) and after preservation (B); Paratype of Thymallus ligericus n. sp. MNHN 2019-0267 (C), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (D), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711341" httpUri="https://zenodo.org/record/12711341/files/figure.png" pageId="2" pageNumber="235">Fig 2</figureCitation>
A-C) and a smaller head height (
<figureCitation id="133046ABFF8BFF82FB1F3435FB3CF9AB" box="[1179,1246,1591,1616]" captionStart="Figure 4" captionStartId="3.[813,877,680,702]" captionTargetBox="[813,1456,215,662]" captionTargetId="figure-603@3.[813,1456,215,662]" captionTargetPageId="3" captionText="Figure 4. – Mouth in ventral view of Thymallus spp. A: Holotype of Thymallus ligericus n. sp. MNHN 2018-0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll.; B: Thymallus thymallus UCBLZ 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711343" httpUri="https://zenodo.org/record/12711343/files/figure.png" pageId="2" pageNumber="235">Fig. 4</figureCitation>
). The throat is usually covered by small scales. It has a pointed snout giving a strait or convex snout profile (
<figureCitation id="133046ABFF8BFF82FBF0347AFB29F96B" box="[1140,1227,1656,1680]" captionStart="Figure 3" captionStartId="3.[132,197,996,1018]" captionTargetBox="[181,727,215,979]" captionTargetId="figure-526@3.[181,727,215,979]" captionTargetPageId="3" captionText="Figure 3. – Head profiles of Thymallus spp. A: Paratype of Thymallus ligericus n. sp. MNHN 2018-0728, 300 mm SL J, Dore River at Vertolaye, 29 May 2015, Persat, Juglaret and Buttazzoni coll.; B: Thymallus thymallus UCBLZ – 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711347" httpUri="https://zenodo.org/record/12711347/files/figure.png" pageId="2" pageNumber="235">Fig. 3A</figureCitation>
). Its mouth is clearly inferior with a transversal aperture more resembling the nase 
<taxonomicName id="4C0B21ADFF8BFF82FCA934BBFB2EF92A" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[813,1228,1721,1745]" family="Cyprinidae" genus="Chondrostoma" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="235" phylum="Chordata" rank="species" species="nasus">
<emphasis id="B97F863CFF8BFF82FCA934BBFBECF92A" box="[813,1038,1721,1745]" italics="true" pageId="2" pageNumber="235">Chondrostoma nasus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
(
<figureCitation id="133046ABFF8BFF82FB5E34BBFACEF92A" box="[1242,1324,1721,1745]" captionStart="Figure 4" captionStartId="3.[813,877,680,702]" captionTargetBox="[813,1456,215,662]" captionTargetId="figure-603@3.[813,1456,215,662]" captionTargetPageId="3" captionText="Figure 4. – Mouth in ventral view of Thymallus spp. A: Holotype of Thymallus ligericus n. sp. MNHN 2018-0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll.; B: Thymallus thymallus UCBLZ 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711343" httpUri="https://zenodo.org/record/12711343/files/figure.png" pageId="2" pageNumber="235">Fig. 4A</figureCitation>
). The upper lip is fleshy. The adipose fin is usually rounded. Concern- ing meristic data, this species has VII-VIII 14-15 dorsal fin rays (VII-14), I 14-16 pectoral fin rays (I-15), I 9-11 pelvic fin rays (I-10), IV-V(9-11) anal fin rays (V-11), 73 + 2 to 87 + 4 lateral scales (81 + 3), 8.5-9.5 transverse dorsal scales and 8.5-9.5 transverse pelvic scales. According to Knizhin (unpubl. data), who observed 
<specimenCount id="9D0D91A7FF8AFF83FE5736B7FDBEFB37" box="[467,604,1204,1228]" count="7" pageId="3" pageNumber="236" type="generic">7 specimens</specimenCount>
from the 
<collectingRegion id="49CF94CCFF8AFF83FD4F36B6FCE5FB37" box="[715,775,1204,1228]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
catchment, this species has (15)18(19) pyloric caeca.
</paragraph>
<caption id="DF740AA6FF8AFF83FCA930AAFB07FCCD" ID-DOI="http://doi.org/10.5281/zenodo.12711343" ID-Zenodo-Dep="12711343" httpUri="https://zenodo.org/record/12711343/files/figure.png" pageId="3" pageNumber="236" startId="3.[813,877,680,702]" targetBox="[813,1456,215,662]" targetPageId="3" targetType="figure">
<paragraph id="8BB45A2EFF8AFF83FCA930AAFB07FCCD" blockId="3.[813,1456,679,822]" pageId="3" pageNumber="236">
Figure 4. – Mouth in ventral view of 
<taxonomicName id="4C0B21ADFF8AFF83FB1E30AAFAA4FD47" box="[1178,1350,679,701]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B97F863CFF8AFF83FB1E30AAFAE2FD46" box="[1178,1280,680,701]" italics="true" pageId="3" pageNumber="236">Thymallus</emphasis>
spp. 
<emphasis id="B97F863CFF8AFF83FAB130A5FAA4FD47" bold="true" box="[1333,1350,679,700]" pageId="3" pageNumber="236">A</emphasis>
</taxonomicName>
: Holotype of 
<taxonomicName id="4C0B21ADFF8AFF83FCCF30C2FBF8FD2E" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[843,1050,704,725]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8AFF83FCCF30C2FBF8FD2E" box="[843,1050,704,725]" italics="true" pageId="3" pageNumber="236">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8AFF83FBA630C2FBB9FD2E" box="[1058,1115,704,725]" pageId="3" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2018-0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll.; 
<emphasis id="B97F863CFF8AFF83FBE630F2FB90FCFE" bold="true" box="[1122,1138,752,773]" pageId="3" pageNumber="236">B</emphasis>
: 
<taxonomicName id="4C0B21ADFF8AFF83FB0530F2FA4DFCFE" authority="UCBLZ" authorityName="UCBLZ" box="[1153,1455,752,773]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8AFF83FB0530F2FAB4FCFE" box="[1153,1366,752,773]" italics="true" pageId="3" pageNumber="236">Thymallus thymallus</emphasis>
UCBLZ
</taxonomicName>
2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll.
</paragraph>
</caption>
<caption id="DF740AA6FF8AFF83FF0031E6FEADFB89" ID-DOI="http://doi.org/10.5281/zenodo.12711347" ID-Zenodo-Dep="12711347" httpUri="https://zenodo.org/record/12711347/files/figure.png" pageId="3" pageNumber="236" startId="3.[132,197,996,1018]" targetBox="[181,727,215,979]" targetPageId="3" targetType="figure">
<paragraph id="8BB45A2EFF8AFF83FF0031E6FEADFB89" blockId="3.[132,775,996,1138]" pageId="3" pageNumber="236">
Figure 3. – Head profiles of 
<taxonomicName id="4C0B21ADFF8AFF83FE2731E6FDB0FC02" box="[419,594,996,1017]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="undefined-A">
<emphasis id="B97F863CFF8AFF83FE2731E6FDE9FC02" box="[419,523,996,1017]" italics="true" pageId="3" pageNumber="236">Thymallus</emphasis>
spp. 
<emphasis id="B97F863CFF8AFF83FDC531E6FDB0FC02" bold="true" box="[577,594,996,1017]" pageId="3" pageNumber="236">A</emphasis>
</taxonomicName>
: Paratype of 
<taxonomicName id="4C0B21ADFF8AFF83FD5E31E6FEC5FBEA" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8AFF83FD5E31E6FEC5FBEA" italics="true" pageId="3" pageNumber="236">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8AFF83FEAA31FEFE87FBEA" box="[302,357,1020,1041]" pageId="3" pageNumber="236" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2018-0728, 300 mm SL J, Dore River at Vertolaye, 29 May 2015, Persat, Juglaret and Buttazzoni coll.; 
<emphasis id="B97F863CFF8AFF83FF3F362EFF29FBBA" bold="true" box="[187,203,1068,1089]" pageId="3" pageNumber="236">B</emphasis>
: 
<taxonomicName id="4C0B21ADFF8AFF83FF53362EFE16FBBA" authority="UCBLZ" authorityName="UCBLZ" box="[215,500,1068,1089]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8AFF83FF53362EFE7DFBBA" box="[215,415,1068,1089]" italics="true" pageId="3" pageNumber="236">Thymallus thymallus</emphasis>
UCBLZ
</taxonomicName>
– 2012.9.1010, 376 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll.
</paragraph>
</caption>
<paragraph id="8BB45A2EFF8AFF83FF003716FEB6FAD7" blockId="3.[132,775,1300,1907]" box="[132,340,1300,1324]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FF003716FEB6FAD7" bold="true" box="[132,340,1300,1324]" pageId="3" pageNumber="236">Colouration in life</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FF2E3734FB2BFADF" blockId="3.[132,775,1300,1907]" lastBlockId="3.[813,1456,863,1316]" pageId="3" pageNumber="236">
In adults, overall body colour is usually darker than that of the Rhone or Rhine 
<taxonomicName id="4C0B21ADFF8AFF83FE063754FDEAFA96" box="[386,520,1365,1389]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8AFF83FE063754FDEAFA96" box="[386,520,1365,1389]" italics="true" pageId="3" pageNumber="236">T. thymallus</emphasis>
</taxonomicName>
, but this might depend on the substrate. They can take on a blue metallic sheen in the darkest individuals. The sides are highly spotted by usually more than 50 and up to hundreds of black dots (
<quantity id="4CF3F7CBFF8AFF83FD4437B5FCE5FA35" box="[704,775,1462,1487]" metricMagnitude="0" metricUnit="m" metricValue="6.858" pageId="3" pageNumber="236" unit="in" value="270.0">270 in</quantity>
the 
<typeStatus id="54B0E48CFF8AFF83FF2F37D5FEE8FA14" box="[171,266,1495,1519]" pageId="3" pageNumber="236" type="holotype">holotype</typeStatus>
) (
<figureCitation id="133046ABFF8AFF83FEA437D5FE81FA14" box="[288,355,1495,1519]" captionStart="Figure 2" captionStartId="2.[1077,1144,876,898]" captionTargetBox="[132,1039,206,1228]" captionTargetId="figure-423@2.[132,1039,206,1228]" captionTargetPageId="2" captionText="Figure 2. – Specimens of Thymallus spp. in lateral view. Holotype of Thymallus ligericus n. sp. MNHN 2018- 0722, 337 mm SL J, Alagnon River at La Chapelle d’Alagnon, 9 Sep. 2016, Persat and FDAAPPMA15 coll., in live (A) and after preservation (B); Paratype of Thymallus ligericus n. sp. MNHN 2019-0267 (C), 269 mm SL ♀, Loire River at Chadron, 22 Sep. 2016, Persat and ONEMA43 coll; UCBLZ – 2012.9.1011 (D), 292 mm TL, Rhône drainage, Ain River at Crotenay, 10 Jun. 2014, Persat and FDAAPPMA39 coll." figureDoi="http://doi.org/10.5281/zenodo.12711341" httpUri="https://zenodo.org/record/12711341/files/figure.png" pageId="3" pageNumber="236">Figs 2</figureCitation>
A-C, 5), sometimes extending up to the head, with the exception of the uppermost section of the Alli- er River where the population is less spotted. In some populations, dots might be coalescent forming a kind of short horizontal lines. The lateral large faint wine-red patch in the centre of the side above the pectoral fins, common in many European populations, is often not visible when present because of the dark background colour of the body. Paired and anal fins have a rather dark green-brown colour. The pectoral fin does not develop the patchy colouration pattern usually visible in the largest 
<taxonomicName id="4C0B21ADFF8AFF83FECD3519FE2CF8C9" box="[329,462,1818,1842]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8AFF83FECD3519FE2CF8C9" box="[329,462,1818,1842]" italics="true" pageId="3" pageNumber="236">T. thymallus</emphasis>
</taxonomicName>
of the Rhine and sometimes Rhone catchments. Caudal fin is dark-greenish, sometimes turning to dark purple, with a blue iridescent shine in large individuals. The large dorsal fin of the adults, especially in the males, is a bit less extended and less coloured than in the Rhône or Rhine populations of 
<taxonomicName id="4C0B21ADFF8AFF83FBFF31A0FB1FFC42" box="[1147,1277,929,953]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="3" pageNumber="236" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8AFF83FBFF31A0FB1FFC42" box="[1147,1277,929,953]" italics="true" pageId="3" pageNumber="236">T. thymallus</emphasis>
</taxonomicName>
. The fin rays are contrastingly lighter than the inter ray membrane. The outer margin of the dorsal fin is often underlined by a thin pink line (
<figureCitation id="133046ABFF8AFF83FCE73606FC47FBE7" box="[867,933,1028,1052]" captionStart="Figure 6" captionStartId="4.[813,877,1197,1219]" captionTargetBox="[813,1456,833,1181]" captionTargetId="figure-734@4.[813,1456,833,1181]" captionTargetPageId="4" captionText="Figure 6. – Dorsal fin on a live paratype of Thymallus ligericus n. sp. MNHN 2018-0726, 290 mm SL ♀, Lignon River at Boën-sur- Lignon, 29 May 2015, Persat, Juglaret and Buttazzoni coll." figureDoi="http://doi.org/10.5281/zenodo.12711351" httpUri="https://zenodo.org/record/12711351/files/figure.png" pageId="3" pageNumber="236">Fig. 6</figureCitation>
). Towards the posterior part of the dorsal fin, the edge line takes a brown colour that extends more and deeper between the last dorsal fin rays. A checked pattern of alterna- tively dark grey and brown-reddish blotches covers most of the surface of the fin. This checked pattern is completed on its outer and posterior part by an alignment of increasingly clear blotches showing the same blue iridescent shine as the caudal fin. For large individuals, this blue colour part of the dorsal fin is visible from the riverbank.
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCA9374FFB8AFA9E" blockId="3.[813,1455,1357,1513]" box="[813,1128,1357,1381]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FCA9374FFB8AFA9E" bold="true" box="[813,1128,1357,1381]" pageId="3" pageNumber="236">Colouration in preservation</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCD6376CFA92FA12" blockId="3.[813,1455,1357,1513]" pageId="3" pageNumber="236">Fixed in formalin, then preserved in 70° ethanol, most of the colours, except the black dots, turn to grey. The checked pattern and colours of the dorsal fin, as well as the faint wine-red become faded but persist, for at least several years.</paragraph>
</subSubSection>
<subSubSection id="C31109A5FF8AFF83FCA93410FBEAF934" pageId="3" pageNumber="236" type="etymology">
<paragraph id="8BB45A2EFF8AFF83FCA93410FC4AF9D1" blockId="3.[813,1455,1554,1743]" box="[813,936,1554,1578]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FCA93410FC4AF9D1" bold="true" box="[813,936,1554,1578]" pageId="3" pageNumber="236">Etymology</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCD63436FC7EF997" blockId="3.[813,1455,1554,1743]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FCD63436FC58F9B0" box="[850,954,1588,1611]" italics="true" pageId="3" pageNumber="236">Ligericus</emphasis>
is a Latin adjective meaning “from the 
<collectingRegion id="49CF94CCFF8AFF83FAF73431FA4DF9B0" box="[1395,1455,1587,1611]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
drainage”.
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCA93477FC12F976" blockId="3.[813,1455,1554,1743]" box="[813,1008,1653,1677]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FCA93477FC12F976" bold="true" box="[813,1008,1653,1677]" pageId="3" pageNumber="236">Vernacular name</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCD63494FBEAF934" blockId="3.[813,1455,1554,1743]" pageId="3" pageNumber="236">
<collectingRegion id="49CF94CCFF8AFF83FCD63494FC6CF955" box="[850,910,1686,1710]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
grayling (English), Ombre d’Auvergne,, Ombre de la 
<collectingRegion id="49CF94CCFF8AFF83FCE134B5FC7DF934" box="[869,927,1719,1743]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
(French).
</paragraph>
</subSubSection>
<subSubSection id="C31109A5FF8AFF85FCA934FAFD59FC6A" lastPageId="5" lastPageNumber="238" pageId="3" pageNumber="236" type="distribution">
<paragraph id="8BB45A2EFF8AFF83FCA934FAFBA2F8EB" blockId="3.[813,1455,1784,1907]" box="[813,1088,1784,1808]" pageId="3" pageNumber="236">
<emphasis id="B97F863CFF8AFF83FCA934FAFBA2F8EB" bold="true" box="[813,1088,1784,1808]" pageId="3" pageNumber="236">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8AFF83FCD6351BFA4DF889" blockId="3.[813,1455,1784,1907]" pageId="3" pageNumber="236">
The 
<collectingRegion id="49CF94CCFF8AFF83FC06351BFC5EF8CA" box="[898,956,1817,1841]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
grayling lives in medium to large foothill, canyon and plateau rivers of the mountainous part of the 
<collectingRegion id="49CF94CCFF8AFF83FAF03538FA4DF8A9" box="[1396,1455,1850,1874]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
catchment, i.e. in the Upper Vienne, Upper 
<collectingRegion id="49CF94CCFF8AFF83FB7A3559FADAF888" box="[1278,1336,1883,1907]" country="France" name="Pays de la Loire" pageId="3" pageNumber="236">Loire</collectingRegion>
and Upper
</paragraph>
<caption id="DF740AA6FF8DFF84FF0032DBFD0EFEFC" pageId="4" pageNumber="237" startId="4.[132,189,217,239]" targetType="table">
<paragraph id="8BB45A2EFF8DFF84FF0032DBFD0EFEFC" blockId="4.[132,775,217,263]" pageId="4" pageNumber="237">
Table I. – Morphometric data for the holotype and means and extreme values for the 10 paratypes of 
<taxonomicName id="4C0B21ADFF8DFF84FE7232F3FD51FEFD" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[502,691,241,262]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8DFF84FE7232F3FD51FEFD" box="[502,691,241,262]" italics="true" pageId="4" pageNumber="237">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8DFF84FD3D32F0FD0EFEFC" box="[697,748,242,263]" pageId="4" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB45A2EFF8DFF84FD883314FD1DFA73" pageId="4" pageNumber="237">
<table id="F90BA88EFF8D007FFF093314FD1DFA73" box="[141,767,278,1416]" gridcols="3" gridrows="33" pageId="4" pageNumber="237">
<tr id="353B586CFF8D007FFF093314FD1DFEBD" box="[141,767,278,326]" gridrow="0" pageId="4" pageNumber="237" rowspan-0="1">
<th id="76EA3110FF8D007FFD883314FDA6FEBD" box="[524,580,278,326]" gridcol="1" gridrow="0" pageId="4" pageNumber="237">Holo- type</th>
<th id="76EA3110FF8D007FFDE73314FD1DFEBD" box="[611,767,278,326]" gridcol="2" gridrow="0" pageId="4" pageNumber="237">With paratypes (n = 11)</th>
</tr>
<tr id="353B586CFF8D007FFF093350FD1DFE93" box="[141,767,338,360]" gridrow="1" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093350FE00FE93" box="[141,482,338,360]" gridcol="0" gridrow="1" pageId="4" pageNumber="237">Standard length (SL)</th>
<td id="76EA3110FF8D007FFD883350FDA6FE93" box="[524,580,338,360]" gridcol="1" gridrow="1" pageId="4" pageNumber="237">337</td>
<td id="76EA3110FF8D007FFDE73350FD1DFE93" box="[611,767,338,360]" gridcol="2" gridrow="1" pageId="4" pageNumber="237">300 (252-405)</td>
</tr>
<tr id="353B586CFF8D007FFF093371FD1DFE72" box="[141,767,371,393]" gridrow="2" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093371FD1DFE72" box="[141,767,371,393]" colspan="3" colspanRight="2" gridcol="0" gridrow="2" pageId="4" pageNumber="237">in % SL</th>
</tr>
<tr id="353B586CFF8D007FFF093396FD1DFE51" box="[141,767,404,426]" gridrow="3" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093396FE00FE51" box="[141,482,404,426]" gridcol="0" gridrow="3" pageId="4" pageNumber="237">Head length</th>
<td id="76EA3110FF8D007FFD883396FDA6FE51" box="[524,580,404,426]" gridcol="1" gridrow="3" pageId="4" pageNumber="237">20.7</td>
<td id="76EA3110FF8D007FFDE73396FD1DFE51" box="[611,767,404,426]" gridcol="2" gridrow="3" pageId="4" pageNumber="237">22.0 (20-23.7)</td>
</tr>
<tr id="353B586CFF8D007FFF0933B7FD1DFE30" box="[141,767,437,459]" gridrow="4" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0933B7FE00FE30" box="[141,482,437,459]" gridcol="0" gridrow="4" pageId="4" pageNumber="237">Body depth</th>
<td id="76EA3110FF8D007FFD8833B7FDA6FE30" box="[524,580,437,459]" gridcol="1" gridrow="4" pageId="4" pageNumber="237">20.7</td>
<td id="76EA3110FF8D007FFDE733B7FD1DFE30" box="[611,767,437,459]" gridcol="2" gridrow="4" pageId="4" pageNumber="237">22.1 (20.7-24.1)</td>
</tr>
<tr id="353B586CFF8D007FFF0933D5FD1DFE16" box="[141,767,471,493]" gridrow="5" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0933D5FE00FE16" box="[141,482,471,493]" gridcol="0" gridrow="5" pageId="4" pageNumber="237">Caudal peduncle depth</th>
<td id="76EA3110FF8D007FFD8833D5FDA6FE16" box="[524,580,471,493]" gridcol="1" gridrow="5" pageId="4" pageNumber="237">7.2</td>
<td id="76EA3110FF8D007FFDE733D5FD1DFE16" box="[611,767,471,493]" gridcol="2" gridrow="5" pageId="4" pageNumber="237">7.7 (7.2-8.3)</td>
</tr>
<tr id="353B586CFF8D007FFF0933FAFD1DFDF5" box="[141,767,504,526]" gridrow="6" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0933FAFE00FDF5" box="[141,482,504,526]" gridcol="0" gridrow="6" pageId="4" pageNumber="237">Predorsal length</th>
<td id="76EA3110FF8D007FFD8833FAFDA6FDF5" box="[524,580,504,526]" gridcol="1" gridrow="6" pageId="4" pageNumber="237">35.4</td>
<td id="76EA3110FF8D007FFDE733FAFD1DFDF5" box="[611,767,504,526]" gridcol="2" gridrow="6" pageId="4" pageNumber="237">36.7 (34.4-38.9)</td>
</tr>
<tr id="353B586CFF8D007FFF09301BFD1DFDD4" box="[141,767,537,559]" gridrow="7" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09301BFE00FDD4" box="[141,482,537,559]" gridcol="0" gridrow="7" pageId="4" pageNumber="237">Dorsal adipose distance</th>
<td id="76EA3110FF8D007FFD88301BFDA6FDD4" box="[524,580,537,559]" gridcol="1" gridrow="7" pageId="4" pageNumber="237">23.8</td>
<td id="76EA3110FF8D007FFDE7301BFD1DFDD4" box="[611,767,537,559]" gridcol="2" gridrow="7" pageId="4" pageNumber="237">21.9 (19.0-23.8)</td>
</tr>
<tr id="353B586CFF8D007FFF093038FD1DFDAB" box="[141,767,570,592]" gridrow="8" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093038FE00FDAB" box="[141,482,570,592]" gridcol="0" gridrow="8" pageId="4" pageNumber="237">Post-adipose length</th>
<td id="76EA3110FF8D007FFD883038FDA6FDAB" box="[524,580,570,592]" gridcol="1" gridrow="8" pageId="4" pageNumber="237">16.2</td>
<td id="76EA3110FF8D007FFDE73038FD1DFDAB" box="[611,767,570,592]" gridcol="2" gridrow="8" pageId="4" pageNumber="237">15.4 (14.4-16.6)</td>
</tr>
<tr id="353B586CFF8D007FFF093059FD1DFD8A" box="[141,767,603,625]" gridrow="9" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093059FE00FD8A" box="[141,482,603,625]" gridcol="0" gridrow="9" pageId="4" pageNumber="237">Prepectoral length</th>
<td id="76EA3110FF8D007FFD883059FDA6FD8A" box="[524,580,603,625]" gridcol="1" gridrow="9" pageId="4" pageNumber="237">19.0</td>
<td id="76EA3110FF8D007FFDE73059FD1DFD8A" box="[611,767,603,625]" gridcol="2" gridrow="9" pageId="4" pageNumber="237">20.9 (19.0-22.7)</td>
</tr>
<tr id="353B586CFF8D007FFF09307FFD1DFD68" box="[141,767,637,659]" gridrow="10" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09307FFE00FD68" box="[141,482,637,659]" gridcol="0" gridrow="10" pageId="4" pageNumber="237">Prepelvic length</th>
<td id="76EA3110FF8D007FFD88307FFDA6FD68" box="[524,580,637,659]" gridcol="1" gridrow="10" pageId="4" pageNumber="237">47.7</td>
<td id="76EA3110FF8D007FFDE7307FFD1DFD68" box="[611,767,637,659]" gridcol="2" gridrow="10" pageId="4" pageNumber="237">49.6 (47.7-51.4)</td>
</tr>
<tr id="353B586CFF8D007FFF09309CFD1DFD4F" box="[141,767,670,692]" gridrow="11" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09309CFE00FD4F" box="[141,482,670,692]" gridcol="0" gridrow="11" pageId="4" pageNumber="237">Pre-anal length</th>
<td id="76EA3110FF8D007FFD88309CFDA6FD4F" box="[524,580,670,692]" gridcol="1" gridrow="11" pageId="4" pageNumber="237">74.5</td>
<td id="76EA3110FF8D007FFDE7309CFD1DFD4F" box="[611,767,670,692]" gridcol="2" gridrow="11" pageId="4" pageNumber="237">75.5 (72.8-79.2)</td>
</tr>
<tr id="353B586CFF8D007FFF0930BDFD1DFD2E" box="[141,767,703,725]" gridrow="12" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0930BDFE00FD2E" box="[141,482,703,725]" gridcol="0" gridrow="12" pageId="4" pageNumber="237">Pectoral-pelvic distance</th>
<td id="76EA3110FF8D007FFD8830BDFDA6FD2E" box="[524,580,703,725]" gridcol="1" gridrow="12" pageId="4" pageNumber="237">26.9</td>
<td id="76EA3110FF8D007FFDE730BDFD1DFD2E" box="[611,767,703,725]" gridcol="2" gridrow="12" pageId="4" pageNumber="237">27.4 (26.4-28.2)</td>
</tr>
<tr id="353B586CFF8D007FFF0930E2FD1DFD0D" box="[141,767,736,758]" gridrow="13" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0930E2FE00FD0D" box="[141,482,736,758]" gridcol="0" gridrow="13" pageId="4" pageNumber="237">Pelvic-anal distance</th>
<td id="76EA3110FF8D007FFD8830E2FDA6FD0D" box="[524,580,736,758]" gridcol="1" gridrow="13" pageId="4" pageNumber="237">25.4</td>
<td id="76EA3110FF8D007FFDE730E2FD1DFD0D" box="[611,767,736,758]" gridcol="2" gridrow="13" pageId="4" pageNumber="237">24.3 (22.5-25.9)</td>
</tr>
<tr id="353B586CFF8D007FFF093103FD1DFCEC" box="[141,767,769,791]" gridrow="14" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093103FE00FCEC" box="[141,482,769,791]" gridcol="0" gridrow="14" pageId="4" pageNumber="237">Caudal peduncle length</th>
<td id="76EA3110FF8D007FFD883103FDA6FCEC" box="[524,580,769,791]" gridcol="1" gridrow="14" pageId="4" pageNumber="237">15.6</td>
<td id="76EA3110FF8D007FFDE73103FD1DFCEC" box="[611,767,769,791]" gridcol="2" gridrow="14" pageId="4" pageNumber="237">15.9 (15.1-16.8)</td>
</tr>
<tr id="353B586CFF8D007FFF093121FD1DFCC2" box="[141,767,803,825]" gridrow="15" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093121FE00FCC2" box="[141,482,803,825]" gridcol="0" gridrow="15" pageId="4" pageNumber="237">First dorsal fin branched length</th>
<td id="76EA3110FF8D007FFD883121FDA6FCC2" box="[524,580,803,825]" gridcol="1" gridrow="15" pageId="4" pageNumber="237">13.9</td>
<td id="76EA3110FF8D007FFDE73121FD1DFCC2" box="[611,767,803,825]" gridcol="2" gridrow="15" pageId="4" pageNumber="237">15.3 (13.9-17.5)</td>
</tr>
<tr id="353B586CFF8D007FFF093146FD1DFC8E" box="[141,767,836,885]" gridrow="16" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093146FE00FC8E" box="[141,482,836,885]" gridcol="0" gridrow="16" pageId="4" pageNumber="237">Penultimate dorsal fin branched ray length</th>
<td id="76EA3110FF8D007FFD883146FDA6FC8E" box="[524,580,836,885]" gridcol="1" gridrow="16" pageId="4" pageNumber="237">14.2</td>
<td id="76EA3110FF8D007FFDE73146FD1DFC8E" box="[611,767,836,885]" gridcol="2" gridrow="16" pageId="4" pageNumber="237">13.8 (11.5-16.2)</td>
</tr>
<tr id="353B586CFF8D007FFF093182FD1DFC6D" box="[141,767,896,918]" gridrow="17" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093182FE00FC6D" box="[141,482,896,918]" gridcol="0" gridrow="17" pageId="4" pageNumber="237">Pectoral fin length</th>
<td id="76EA3110FF8D007FFD883182FDA6FC6D" box="[524,580,896,918]" gridcol="1" gridrow="17" pageId="4" pageNumber="237">14.6</td>
<td id="76EA3110FF8D007FFDE73182FD1DFC6D" box="[611,767,896,918]" gridcol="2" gridrow="17" pageId="4" pageNumber="237">15.4 (13.8-16.8)</td>
</tr>
<tr id="353B586CFF8D007FFF0931A3FD1DFC4C" box="[141,767,929,951]" gridrow="18" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0931A3FE00FC4C" box="[141,482,929,951]" gridcol="0" gridrow="18" pageId="4" pageNumber="237">Pelvic fin length</th>
<td id="76EA3110FF8D007FFD8831A3FDA6FC4C" box="[524,580,929,951]" gridcol="1" gridrow="18" pageId="4" pageNumber="237">15.8</td>
<td id="76EA3110FF8D007FFDE731A3FD1DFC4C" box="[611,767,929,951]" gridcol="2" gridrow="18" pageId="4" pageNumber="237">15.9 (14-17.6)</td>
</tr>
<tr id="353B586CFF8D007FFF0931C0FD1DFC23" box="[141,767,962,984]" gridrow="19" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0931C0FE00FC23" box="[141,482,962,984]" gridcol="0" gridrow="19" pageId="4" pageNumber="237">Anal fin length</th>
<td id="76EA3110FF8D007FFD8831C0FDA6FC23" box="[524,580,962,984]" gridcol="1" gridrow="19" pageId="4" pageNumber="237">13.8</td>
<td id="76EA3110FF8D007FFDE731C0FD1DFC23" box="[611,767,962,984]" gridcol="2" gridrow="19" pageId="4" pageNumber="237">14.5 (13.6-15.6)</td>
</tr>
<tr id="353B586CFF8D007FFF0931E1FD1DFC02" box="[141,767,995,1017]" gridrow="20" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0931E1FE00FC02" box="[141,482,995,1017]" gridcol="0" gridrow="20" pageId="4" pageNumber="237">Adipose fin length</th>
<td id="76EA3110FF8D007FFD8831E1FDA6FC02" box="[524,580,995,1017]" gridcol="1" gridrow="20" pageId="4" pageNumber="237">8.8</td>
<td id="76EA3110FF8D007FFDE731E1FD1DFC02" box="[611,767,995,1017]" gridcol="2" gridrow="20" pageId="4" pageNumber="237">8.3 (7.6-9.3)</td>
</tr>
<tr id="353B586CFF8D007FFF093607FD1DFBE0" box="[141,767,1029,1051]" gridrow="21" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093607FD1DFBE0" box="[141,767,1029,1051]" colspan="3" colspanRight="2" gridcol="0" gridrow="21" pageId="4" pageNumber="237">in % HL</th>
</tr>
<tr id="353B586CFF8D007FFF093624FD1DFBC7" box="[141,767,1062,1084]" gridrow="22" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093624FE00FBC7" box="[141,482,1062,1084]" gridcol="0" gridrow="22" pageId="4" pageNumber="237">Occipital depth of head</th>
<td id="76EA3110FF8D007FFD883624FDA6FBC7" box="[524,580,1062,1084]" gridcol="1" gridrow="22" pageId="4" pageNumber="237">64.8</td>
<td id="76EA3110FF8D007FFDE73624FD1DFBC7" box="[611,767,1062,1084]" gridcol="2" gridrow="22" pageId="4" pageNumber="237">66.0 (54.8-71.7)</td>
</tr>
<tr id="353B586CFF8D007FFF093645FD1DFBA6" box="[141,767,1095,1117]" gridrow="23" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093645FE00FBA6" box="[141,482,1095,1117]" gridcol="0" gridrow="23" pageId="4" pageNumber="237">Orbital depth of head</th>
<td id="76EA3110FF8D007FFD883645FDA6FBA6" box="[524,580,1095,1117]" gridcol="1" gridrow="23" pageId="4" pageNumber="237">46.9</td>
<td id="76EA3110FF8D007FFDE73645FD1DFBA6" box="[611,767,1095,1117]" gridcol="2" gridrow="23" pageId="4" pageNumber="237">48.8 (43.5-53.4)</td>
</tr>
<tr id="353B586CFF8D007FFF09366AFD1DFB85" box="[141,767,1128,1150]" gridrow="24" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09366AFE00FB85" box="[141,482,1128,1150]" gridcol="0" gridrow="24" pageId="4" pageNumber="237">Inter-orbital width</th>
<td id="76EA3110FF8D007FFD88366AFDA6FB85" box="[524,580,1128,1150]" gridcol="1" gridrow="24" pageId="4" pageNumber="237">27.0</td>
<td id="76EA3110FF8D007FFDE7366AFD1DFB85" box="[611,767,1128,1150]" gridcol="2" gridrow="24" pageId="4" pageNumber="237">27.2 (24.5-30.0)</td>
</tr>
<tr id="353B586CFF8D007FFF09368BFD1DFB64" box="[141,767,1161,1183]" gridrow="25" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09368BFE00FB64" box="[141,482,1161,1183]" gridcol="0" gridrow="25" pageId="4" pageNumber="237">Horizontal eye diameter</th>
<td id="76EA3110FF8D007FFD88368BFDA6FB64" box="[524,580,1161,1183]" gridcol="1" gridrow="25" pageId="4" pageNumber="237">23.9</td>
<td id="76EA3110FF8D007FFDE7368BFD1DFB64" box="[611,767,1161,1183]" gridcol="2" gridrow="25" pageId="4" pageNumber="237">24.7 (22.6-26.3)</td>
</tr>
<tr id="353B586CFF8D007FFF0936A8FD1DFB3A" box="[141,767,1194,1217]" gridrow="26" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0936A8FE00FB3A" box="[141,482,1194,1217]" gridcol="0" gridrow="26" pageId="4" pageNumber="237">Snout length</th>
<td id="76EA3110FF8D007FFD8836A8FDA6FB3A" box="[524,580,1194,1217]" gridcol="1" gridrow="26" pageId="4" pageNumber="237">30.9</td>
<td id="76EA3110FF8D007FFDE736A8FD1DFB3A" box="[611,767,1194,1217]" gridcol="2" gridrow="26" pageId="4" pageNumber="237">31.3 (28.8-35.0)</td>
</tr>
<tr id="353B586CFF8D007FFF0936CEFD1DFB19" box="[141,767,1228,1250]" gridrow="27" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0936CEFE00FB19" box="[141,482,1228,1250]" gridcol="0" gridrow="27" pageId="4" pageNumber="237">Mandible length</th>
<td id="76EA3110FF8D007FFD8836CEFDA6FB19" box="[524,580,1228,1250]" gridcol="1" gridrow="27" pageId="4" pageNumber="237">43.8</td>
<td id="76EA3110FF8D007FFDE736CEFD1DFB19" box="[611,767,1228,1250]" gridcol="2" gridrow="27" pageId="4" pageNumber="237">42.1 (40.8-44.4)</td>
</tr>
<tr id="353B586CFF8D007FFF0936EFFD1DFAF8" box="[141,767,1261,1283]" gridrow="28" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF0936EFFE00FAF8" box="[141,482,1261,1283]" gridcol="0" gridrow="28" pageId="4" pageNumber="237">Maxillary length</th>
<td id="76EA3110FF8D007FFD8836EFFDA6FAF8" box="[524,580,1261,1283]" gridcol="1" gridrow="28" pageId="4" pageNumber="237">21.0</td>
<td id="76EA3110FF8D007FFDE736EFFD1DFAF8" box="[611,767,1261,1283]" gridcol="2" gridrow="28" pageId="4" pageNumber="237">22.0 (20.5-25.4)</td>
</tr>
<tr id="353B586CFF8D007FFF09370CFD1DFADF" box="[141,767,1294,1316]" gridrow="29" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09370CFE00FADF" box="[141,482,1294,1316]" gridcol="0" gridrow="29" pageId="4" pageNumber="237">Maxillary width</th>
<td id="76EA3110FF8D007FFD88370CFDA6FADF" box="[524,580,1294,1316]" gridcol="1" gridrow="29" pageId="4" pageNumber="237">12.7</td>
<td id="76EA3110FF8D007FFDE7370CFD1DFADF" box="[611,767,1294,1316]" gridcol="2" gridrow="29" pageId="4" pageNumber="237">11.9 (10.6-13.1)</td>
</tr>
<tr id="353B586CFF8D007FFF09372DFD1DFABE" box="[141,767,1327,1349]" gridrow="30" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF09372DFE00FABE" box="[141,482,1327,1349]" gridcol="0" gridrow="30" pageId="4" pageNumber="237">Mandible width</th>
<td id="76EA3110FF8D007FFD88372DFDA6FABE" box="[524,580,1327,1349]" gridcol="1" gridrow="30" pageId="4" pageNumber="237">24.3</td>
<td id="76EA3110FF8D007FFDE7372DFD1DFABE" box="[611,767,1327,1349]" gridcol="2" gridrow="30" pageId="4" pageNumber="237">23.1 (19.9-26.1)</td>
</tr>
<tr id="353B586CFF8D007FFF093752FD1DFA9C" box="[141,767,1360,1383]" gridrow="31" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093752FE00FA9C" box="[141,482,1360,1383]" gridcol="0" gridrow="31" pageId="4" pageNumber="237">Chin length</th>
<td id="76EA3110FF8D007FFD883752FDA6FA9C" box="[524,580,1360,1383]" gridcol="1" gridrow="31" pageId="4" pageNumber="237">27.2</td>
<td id="76EA3110FF8D007FFDE73752FD1DFA9C" box="[611,767,1360,1383]" gridcol="2" gridrow="31" pageId="4" pageNumber="237">29.2 (25.3-32.1)</td>
</tr>
<tr id="353B586CFF8D007FFF093770FD1DFA73" box="[141,767,1394,1416]" gridrow="32" pageId="4" pageNumber="237">
<th id="76EA3110FF8D007FFF093770FE00FA73" box="[141,482,1394,1416]" gridcol="0" gridrow="32" pageId="4" pageNumber="237">Ventral snout length</th>
<td id="76EA3110FF8D007FFD883770FDA6FA73" box="[524,580,1394,1416]" gridcol="1" gridrow="32" pageId="4" pageNumber="237">35.5</td>
<td id="76EA3110FF8D007FFDE73770FD1DFA73" box="[611,767,1394,1416]" gridcol="2" gridrow="32" pageId="4" pageNumber="237">36.9 (32.1-39.6)</td>
</tr>
</table>
</paragraph>
<caption id="DF740AA6FF8DFF84FCA93090FB95FCDB" ID-DOI="http://doi.org/10.5281/zenodo.12711349" ID-Zenodo-Dep="12711349" httpUri="https://zenodo.org/record/12711349/files/figure.png" pageId="4" pageNumber="237" startId="4.[813,878,658,680]" targetBox="[814,1453,218,641]" targetPageId="4" targetType="figure">
<paragraph id="8BB45A2EFF8DFF84FCA93090FB95FCDB" blockId="4.[813,1455,658,800]" pageId="4" pageNumber="237">
Figure 5. – Average dot numbers on each scale row on the flank for populations of 
<taxonomicName id="4C0B21ADFF8DFF84FC6530A8FB40FD44" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[993,1186,682,703]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8DFF84FC6530A8FB40FD44" box="[993,1186,682,703]" italics="true" pageId="4" pageNumber="237">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8DFF84FB2C30A9FB3EFD3B" box="[1192,1244,683,704]" pageId="4" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
from the Upper Ance (47 specimens), Lower Ance (7 specimens) and Lignon Rivers (16 specimens) in the Loire drainage as well as for 
<taxonomicName id="4C0B21ADFF8DFF84FB7D30D8FCABFCFC" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8DFF84FB7D30D8FCABFCFC" italics="true" pageId="4" pageNumber="237">Thymallus thymallus</emphasis>
</taxonomicName>
populations from the Ain (176 specimens) and Loue rivers (35 specimens) in the Rhône drainage.
</paragraph>
</caption>
<caption id="DF740AA6FF8DFF84FCA936AFFABDFB08" ID-DOI="http://doi.org/10.5281/zenodo.12711351" ID-Zenodo-Dep="12711351" httpUri="https://zenodo.org/record/12711351/files/figure.png" pageId="4" pageNumber="237" startId="4.[813,877,1197,1219]" targetBox="[813,1456,833,1181]" targetPageId="4" targetType="figure">
<paragraph id="8BB45A2EFF8DFF84FCA936AFFABDFB08" blockId="4.[813,1456,1197,1267]" pageId="4" pageNumber="237">
Figure 6. – Dorsal fin on a live paratype of 
<taxonomicName id="4C0B21ADFF8DFF84FB5136AFFA75FB39" authorityName="Persat &amp; Weiss &amp; Froufe &amp; Secci-Petretto &amp; Denys" authorityYear="2019" box="[1237,1431,1197,1218]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="ligericus" status="sp. nov.">
<emphasis id="B97F863CFF8DFF84FB5136AFFA75FB39" box="[1237,1431,1197,1218]" italics="true" pageId="4" pageNumber="237">Thymallus ligericus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C3B47FF8DFF84FA1936AFFCABFB21" pageId="4" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2018-0726, 290 mm SL ♀, Lignon River at Boën-surLignon, 29 May 2015, Persat, Juglaret and Buttazzoni coll.
</paragraph>
</caption>
<paragraph id="8BB45A2EFF8DFF84FF0037B8FC5DFA4A" blockId="4.[132,775,1466,1906]" lastBlockId="4.[813,1455,1305,1457]" pageId="4" pageNumber="237">
Allier districts, where it cohabits with brown trout 
<taxonomicName id="4C0B21ADFF8DFF84FD4637B8FE62FA0A" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" family="Salmonidae" genus="Salmo" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="trutta">
<emphasis id="B97F863CFF8DFF84FD4637B8FF26FA0A" italics="true" pageId="4" pageNumber="237">Salmo trutta</emphasis>
Linnaeus, 1758
</taxonomicName>
, and Atlantic salmon 
<taxonomicName id="4C0B21ADFF8DFF84FDFA37D8FED0F9EA" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" family="Salmonidae" genus="Salmo" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="salar">
<emphasis id="B97F863CFF8DFF84FDFA37D8FCE5FA09" box="[638,775,1498,1522]" italics="true" pageId="4" pageNumber="237">Salmo salar</emphasis>
Linnaeus, 1758
</taxonomicName>
, in the Allier branch. It is able to colonise rather small rivers with mean discharges of ca. 
<quantity id="4CF3F7CBFF8DFF84FD053418FD48F9CA" box="[641,682,1562,1586]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="237" unit="m" value="1.0">1 m</quantity>
<superScript id="7C7EF766FF8DFF84FD2E3414FD51F9DC" attach="left" box="[682,691,1558,1575]" fontSize="7" pageId="4" pageNumber="237">3</superScript>
/s when protected from overfishing. It is able to swallow and digest caddisfly larvae with their gravel cases, a common prey in the 
<collectingRegion id="49CF94CCFF8DFF84FF2F3478FF07F969" box="[171,229,1658,1682]" country="France" name="Pays de la Loire" pageId="4" pageNumber="237">Loire</collectingRegion>
catchment (
<bibRefCitation id="EF9A27DFFF8DFF84FEE13478FE0CF96A" author="PERSAT H." box="[357,494,1657,1682]" pageId="4" pageNumber="237" refId="ref4800" refString="PERSAT H., 1976. - Ecologie de l'Ombre commun, Thymallus thymallus (L., 1758) (Poissons Salmonides). PhD Thesis, 69 p., Univ. Lyon 1, France." type="book" year="1976">Persat, 1976</bibRefCitation>
). Only one known population, from the Besbre River, seems to have been driven to extinction, but all populations are potentially threatened by climatic change as droughts result in exceedingly high temperatures and too low water velocities. Up to now, stocking attempts appear to regularly fail in sustaining local native populations (
<bibRefCitation id="EF9A27DFFF8DFF84FE8B3538FE36F8AA" author="PERSAT H. &amp; MATTERSDORFER K. &amp; CHARLAT S. &amp; SCHENEKAR T. &amp; WEISS S." box="[271,468,1849,1874]" pageId="4" pageNumber="237" pagination="7 - 20" refId="ref5193" refString="PERSAT H., MATTERSDORFER K., CHARLAT S., SCHENEKAR T. &amp; WEISS S., 2016. - Genetic integrity of the European grayling (Thymallus thymallus) populations within the Vienne River drainage basin after five decades of stockings. Cybium, 40 (1): 7 - 20." type="journal article" year="2016">
Persat 
<emphasis id="B97F863CFF8DFF84FEDC3538FE6CF8A9" box="[344,398,1850,1874]" italics="true" pageId="4" pageNumber="237">et al.</emphasis>
, 2016
</bibRefCitation>
), suggesting that wild stocks are more competitive than introduced ones. However, some 
<collectingRegion id="49CF94CCFF8DFF84FCA9371BFC88FACA" box="[813,874,1305,1329]" country="France" name="Pays de la Loire" pageId="4" pageNumber="237">Loire</collectingRegion>
grayling populations have been successfully introduced elsewhere, such as in the Corrèze River of the Dordogne catchment, based on stocks reared at the Puy-de-Dôme federal fish farm of Besse-en-Chandesse (M. Mestas, pers. comm.).
</paragraph>
<paragraph id="8BB45A2EFF8DFF84FCA937DAFBCAFA0B" blockId="4.[813,1456,1496,1905]" box="[813,1064,1496,1520]" pageId="4" pageNumber="237">
<emphasis id="B97F863CFF8DFF84FCA937DAFBCAFA0B" bold="true" box="[813,1064,1496,1520]" pageId="4" pageNumber="237">Comparative material</emphasis>
</paragraph>
<paragraph id="8BB45A2EFF8DFF84FCD637FBFA4DF9EB" blockId="4.[813,1456,1496,1905]" box="[850,1455,1528,1553]" pageId="4" pageNumber="237">
<taxonomicName id="4C0B21ADFF8DFF84FCD637FBFAA0F9EA" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[850,1346,1528,1553]" family="Salmonidae" genus="Thymallus" kingdom="Animalia" order="Salmoniformes" pageId="4" pageNumber="237" phylum="Chordata" rank="species" species="thymallus">
<emphasis id="B97F863CFF8DFF84FCD637FBFBB8F9EB" box="[850,1114,1529,1553]" italics="true" pageId="4" pageNumber="237">Thymallus thymallus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
: BMNH
</paragraph>
<paragraph id="8BB45A2EFF8DFF85FCA9341BFE9EFDEA" blockId="4.[813,1456,1496,1905]" lastBlockId="5.[132,775,217,529]" lastPageId="5" lastPageNumber="238" pageId="4" pageNumber="237">
1979.6.22.191-192, 
<materialsCitation id="3B635073FF8DFF84FB86341BFB57F9AA" country="United Kingdom" county="Lambourn" location="Thames" municipality="Lambourn River" pageId="4" pageNumber="237" specimenCount="2">
<specimenCount id="9D0D91A7FF8DFF84FB86341BFB64F9CA" box="[1026,1158,1561,1585]" count="2" pageId="4" pageNumber="237" type="generic">2 specimens</specimenCount>
, 
<collectingCountry id="F31C1ABEFF8DFF84FB0B341BFB0EF9CA" box="[1167,1260,1561,1585]" name="United Kingdom" pageId="4" pageNumber="237">England</collectingCountry>
: 
<location id="8ED40CF5FF8DFF84FB77341BFAA5F9CA" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FB77341BFAA5F9CA" box="[1267,1351,1561,1585]" country="United Kingdom" county="Lambourn" municipality="Lambourn River" name="Thames" pageId="4" pageNumber="237">Thames</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8DFF84FCA9343BFC3DF9AA" box="[813,991,1593,1617]" pageId="4" pageNumber="237">Lambourn River</collectingMunicipality>
at 
<collectingCounty id="62D522A2FF8DFF84FC7B343BFB90F9AA" box="[1023,1138,1593,1617]" pageId="4" pageNumber="237">Lambourn</collectingCounty>
, 1976
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FB44343BFAE4F98A" collectionCode="BMNH" collectorName="Bellamy" country="United Kingdom" location="Kennet River" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FB44343BFAF9F9AA" box="[1216,1307,1593,1617]" pageId="4" pageNumber="237">Bellamy</collectorName>
coll.; 
<collectionCode id="ED1AC2EBFF8DFF84FADB343BFA4DF9AB" box="[1375,1455,1593,1616]" pageId="4" pageNumber="237">BMNH</collectionCode>
1979.6.22.226-238, 4, 
<location id="8ED40CF5FF8DFF84FBAE345BFB23F98A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FBAE345BFB23F98A" box="[1066,1217,1625,1649]" country="United Kingdom" name="Kennet River" pageId="4" pageNumber="237">Kennet River</location>
, 1973
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FA96345BFC16F94A" collectingDate="2017-09-15" collectionCode="BMNH" collectorName="Bellamy" country="United Kingdom" county="Huddersfield" location="River Holme" municipality="Lockwood" pageId="4" pageNumber="237" specimenCode="BMNH 2018.9" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FA96345BFA92F98A" box="[1298,1392,1625,1649]" pageId="4" pageNumber="237">Bellamy</collectorName>
coll.; 
<specimenCode id="DBADF255FF8DFF84FCA9347BFC2DF96A" box="[813,975,1657,1681]" collectionCode="BMNH" pageId="4" pageNumber="237">BMNH 2018.9</specimenCode>
.4.14, 1, 
<location id="8ED40CF5FF8DFF84FBA2347BFB52F96A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FBA2347BFB52F96A" box="[1062,1200,1657,1681]" country="United Kingdom" county="Huddersfield" municipality="Lockwood" name="River Holme" pageId="4" pageNumber="237">River Holme</location>
at 
<collectingMunicipality id="6BD0C054FF8DFF84FB4A347BFAA1F96A" box="[1230,1347,1657,1681]" pageId="4" pageNumber="237">Lockwood</collectingMunicipality>
, 
<collectingCounty id="62D522A2FF8DFF84FAC8347BFCBDF94A" pageId="4" pageNumber="237">Huddersfield</collectingCounty>
, 
<date id="FFB57CEEFF8DFF84FCEC349AFC16F94A" box="[872,1012,1688,1713]" pageId="4" pageNumber="237" value="2017-09-15">
<collectingDate id="EFF18506FF8DFF84FCEC349AFC16F94A" box="[872,1012,1688,1713]" pageId="4" pageNumber="237" value="2017-09-15">15 Sep. 2017</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FC79349BFC15F92B" collectionCode="MNHN" collectorName="Beer" country="France" location="Rhone" pageId="4" pageNumber="237" specimenCode="MNHN 1959-0546, 1" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FC79349BFBCDF94B" box="[1021,1071,1689,1712]" pageId="4" pageNumber="237">Beer</collectorName>
coll.; 
<specimenCode id="DBADF255FF8DFF84FBF4349BFADFF94B" box="[1136,1341,1688,1712]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="237">MNHN 1959-0546</specimenCode>
, 
<specimenCode id="DBADF255FF8DFF84FAC2349BFAB1F94A" box="[1350,1363,1689,1713]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="237">1</specimenCode>
, 
<collectingCountry id="F31C1ABEFF8DFF84FADB349BFA4EF94B" box="[1375,1452,1689,1712]" name="France" pageId="4" pageNumber="237">France</collectingCountry>
: 
<location id="8ED40CF5FF8DFF84FCA934BBFC97F92A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FCA934BBFC97F92A" box="[813,885,1721,1745]" country="France" name="Rhone" pageId="4" pageNumber="237">Rhône</location>
drainage at
</materialsCitation>
<materialsCitation id="3B635073FF8DFF84FC7A34BBFB93F90A" collectionCode="MNHN" collectorName="Nant-de-Sion &amp; Spillmann" country="France" location="Rhone" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FC7A34BBFB75F92A" box="[1022,1175,1721,1745]" pageId="4" pageNumber="237">Nant-de-Sion</collectorName>
, 
<collectorName id="26FE3FF8FF8DFF84FB2634BBFAF6F92A" box="[1186,1300,1721,1745]" pageId="4" pageNumber="237">Spillmann</collectorName>
coll.; 
<collectionCode id="ED1AC2EBFF8DFF84FAD834BBFA4DF92B" box="[1372,1455,1721,1744]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="237">MNHN</collectionCode>
B-2478, 2, 
<location id="8ED40CF5FF8DFF84FC3734DBFC1DF90A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FC3734DBFC1DF90A" box="[947,1023,1753,1777]" country="France" name="Rhone" pageId="4" pageNumber="237">Rhône</location>
drainage
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FBF934DBFBA9F8EA" collectionCode="MNHN" collectorName="de Candolle" country="France" location="Rhone" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FBF934DBFAEEF90A" box="[1149,1292,1753,1777]" pageId="4" pageNumber="237">de Candolle</collectorName>
coll.; 
<collectionCode id="ED1AC2EBFF8DFF84FADE34DBFA4DF90B" box="[1370,1455,1753,1776]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="237">MNHN</collectionCode>
B-2479, 1, 
<location id="8ED40CF5FF8DFF84FC2634FBFC0AF8EA" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FC2634FBFC0AF8EA" box="[930,1000,1785,1809]" country="France" name="Rhone" pageId="4" pageNumber="237">Rhône</location>
drainage
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FBD034FBFA49F8CA" collectionCode="MNHN" collectorName="Blanchard" country="France" location="Rhone" municipality="Dept. Isere" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FBD034FBFB20F8EA" box="[1108,1218,1785,1809]" pageId="4" pageNumber="237">Blanchard</collectorName>
coll.; 
<collectionCode id="ED1AC2EBFF8DFF84FA8634FBFAB1F8EB" box="[1282,1363,1785,1808]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="237">MNHN</collectionCode>
B-2480, 3, 
<location id="8ED40CF5FF8DFF84FCCD351BFC70F8CA" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FCCD351BFC70F8CA" box="[841,914,1817,1841]" country="France" municipality="Dept. Isere" name="Rhone" pageId="4" pageNumber="237">Rhône</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8DFF84FB8E351BFB64F8CA" box="[1034,1158,1817,1841]" pageId="4" pageNumber="237">Dept. Isère</collectingMunicipality>
at Saint-Laurent-du-Pont
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF84FCA9353BFB60F88A" collectingDate="2014-06-11" collectionCode="UCBLZ" collectorName="Galbert" country="France" county="Crotenay" location="Rhone" municipality="Ain River" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FCA9353BFC9FF8AA" box="[813,893,1849,1873]" pageId="4" pageNumber="237">Galbert</collectorName>
coll.; 
<collectionCode id="ED1AC2EBFF8DFF84FC39353BFBF6F8AA" box="[957,1044,1849,1873]" pageId="4" pageNumber="237">UCBLZ</collectionCode>
– 2012.9.1012, 2, 
<location id="8ED40CF5FF8DFF84FB52353BFAFEF8AA" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8DFF84FB52353BFAFEF8AA" box="[1238,1308,1849,1873]" country="France" county="Crotenay" municipality="Ain River" name="Rhone" pageId="4" pageNumber="237">Rhône</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8DFF84FA03353BFC8BF88A" pageId="4" pageNumber="237">Ain River</collectingMunicipality>
at 
<collectingCounty id="62D522A2FF8DFF84FC0D355BFC0FF88A" box="[905,1005,1881,1905]" pageId="4" pageNumber="237">Crotenay</collectingCounty>
, 
<date id="FFB57CEEFF8DFF84FC73355BFB60F88A" box="[1015,1154,1881,1905]" pageId="4" pageNumber="237" value="2014-06-11">
<collectingDate id="EFF18506FF8DFF84FC73355BFB60F88A" box="[1015,1154,1881,1905]" pageId="4" pageNumber="237" value="2014-06-11">11 Jun. 2014</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8DFF85FB09355BFE76FEEA" collectingDate="2016-07-19" collectionCode="UCBLZ" collectorName="Persat" country="France" county="Marigny" lastPageId="5" lastPageNumber="238" location="Rhone" municipality="Ain River" pageId="4" pageNumber="237" specimenCount="1">
<collectorName id="26FE3FF8FF8DFF84FB09355BFB2DF88B" box="[1165,1231,1881,1904]" pageId="4" pageNumber="237">Persat</collectorName>
and FDAAPPMA39 coll.; 
<collectionCode id="ED1AC2EBFF8CFF85FF4632DBFEF8FF0A" box="[194,282,217,241]" pageId="5" pageNumber="238">UCBLZ</collectionCode>
– 2012.9.1013, 1, 
<location id="8ED40CF5FF8CFF85FE6132DBFDCEFF0A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8CFF85FE6132DBFDCEFF0A" box="[485,556,217,241]" country="France" county="Marigny" municipality="Ain River" name="Rhone" pageId="5" pageNumber="238">Rhône</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8CFF85FD1832DBFCE5FF0A" box="[668,775,217,241]" pageId="5" pageNumber="238">Ain River</collectingMunicipality>
at 
<collectingCounty id="62D522A2FF8CFF85FF1B32FBFF1DFEEA" box="[159,255,249,273]" pageId="5" pageNumber="238">Marigny</collectingCounty>
, 
<date id="FFB57CEEFF8CFF85FE8E32FBFE76FEEA" box="[266,404,249,273]" pageId="5" pageNumber="238" value="2016-07-19">
<collectingDate id="EFF18506FF8CFF85FE8E32FBFE76FEEA" box="[266,404,249,273]" pageId="5" pageNumber="238" value="2016-07-19">19 Jul. 2016</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8CFF85FE1B32F8FDDBFEAA" collectingDate="2010-10-19" collectionCode="UCBLZ" collectorName="Persat" country="Virgin Islands" county="Royans" location="Rhone" municipality="Bourne River" pageId="5" pageNumber="238" specimenCount="1">
<collectorName id="26FE3FF8FF8CFF85FE1B32F8FE00FEEA" box="[415,482,250,273]" pageId="5" pageNumber="238">Persat</collectorName>
and FDAAPPMA39 coll.; 
<collectionCode id="ED1AC2EBFF8CFF85FF00331BFF3EFECA" box="[132,220,281,305]" pageId="5" pageNumber="238">UCBLZ</collectionCode>
– 2012.9.1014, 1, 
<location id="8ED40CF5FF8CFF85FE21331BFE09FECA" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8CFF85FE21331BFE09FECA" box="[421,491,281,305]" country="Virgin Islands" county="Royans" municipality="Bourne River" name="Rhone" pageId="5" pageNumber="238">Rhône</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8CFF85FDD83318FD0FFECA" box="[604,749,281,305]" pageId="5" pageNumber="238">Bourne River</collectingMunicipality>
at 
<collectingCountry id="F31C1ABEFF8CFF85FF00333BFEFFFEAA" box="[132,285,313,337]" name="Virgin Islands" pageId="5" pageNumber="238">Saint Thomas</collectingCountry>
en 
<collectingCounty id="62D522A2FF8CFF85FEC13338FE79FEA9" box="[325,411,314,338]" pageId="5" pageNumber="238">Royans</collectingCounty>
, 
<date id="FFB57CEEFF8CFF85FE22333BFDDBFEAA" box="[422,569,313,337]" pageId="5" pageNumber="238" value="2010-10-19">
<collectingDate id="EFF18506FF8CFF85FE22333BFDDBFEAA" box="[422,569,313,337]" pageId="5" pageNumber="238" value="2010-10-19">19 Oct. 2010</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B635073FF8CFF85FDC13338FD5FFE4A" collectingDate="2004-11-10" collectorName="Persat &amp; Mallet" country="Germany" county="Wasserburg" latitude="48.058" location="Danube" longLatPrecision="66" longitude="12.22" municipality="Inn River" pageId="5" pageNumber="238" specimenCount="1">
<collectorName id="26FE3FF8FF8CFF85FDC13338FD6BFEAA" box="[581,649,314,337]" pageId="5" pageNumber="238">Persat</collectorName>
and 
<collectorName id="26FE3FF8FF8CFF85FD3B333BFCE5FEAA" box="[703,775,313,337]" pageId="5" pageNumber="238">Mallet</collectorName>
coll.; ZSM-Pis-032234/537416/389004, 1, 
<collectingCountry id="F31C1ABEFF8CFF85FDD9335BFD25FE8A" box="[605,711,345,369]" name="Germany" pageId="5" pageNumber="238">Germany</collectingCountry>
: 
<location id="8ED40CF5FF8CFF85FD553358FF49FE6A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8CFF85FD553358FF49FE6A" country="Germany" county="Wasserburg" latitude="48.058" longLatPrecision="66" longitude="12.22" municipality="Inn River" name="Danube" pageId="5" pageNumber="238">Danube</location>
drainage, 
<collectingMunicipality id="6BD0C054FF8CFF85FE983378FE60FE6A" box="[284,386,377,401]" pageId="5" pageNumber="238">Inn River</collectingMunicipality>
at 
<collectingCounty id="62D522A2FF8CFF85FE26337BFDC1FE6A" box="[418,547,377,401]" pageId="5" pageNumber="238">Wasserburg</collectingCounty>
, below dam, downriver to “Am Griess”, 
<geoCoordinate id="EE3F3CE9FF8CFF85FEDB339BFE5EFE4A" box="[351,444,409,433]" degrees="48.058" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="48.058">48.058N</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF8CFF85FE46339BFDFEFE4A" box="[450,540,409,433]" degrees="12.220" direction="east" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="12.22">12.220E</geoCoordinate>
, 
<date id="FFB57CEEFF8CFF85FDAD339BFD5FFE4A" box="[553,701,409,433]" pageId="5" pageNumber="238" value="2004-11-10">
<collectingDate id="EFF18506FF8CFF85FDAD339BFD5FFE4A" box="[553,701,409,433]" pageId="5" pageNumber="238" value="2004-11-10">10 Nov. 2004</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B635073FF8CFF85FD43339BFE9AFDEA" collectingDate="2007-09-26" country="Germany" county="East of Munchberg" latitude="50.212" location="Elbe" longLatPrecision="65" longitude="11.917" municipality="North of Forbau" pageId="5" pageNumber="238" specimenCount="1">
ZSM- Pis-036238/541401/392989, 1, 
<location id="8ED40CF5FF8CFF85FE6033BBFDF5FE2A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8CFF85FE6033BBFDF5FE2A" box="[484,535,441,465]" country="Germany" county="East of Munchberg" latitude="50.212" longLatPrecision="65" longitude="11.917" municipality="North of Forbau" name="Elbe" pageId="5" pageNumber="238">Elbe</location>
drainage, 
<location id="8ED40CF5FF8CFF85FD0933BBFF5CFE0A" LSID="urn:lsid:plazi:treatment:03A2EB38FF88FF85FC64369BFD59FC6A:8ED40CF5FF8CFF85FD0933BBFF5CFE0A" country="Germany" county="East of Munchberg" latitude="50.212" longLatPrecision="65" longitude="11.917" municipality="North of Forbau" name="Sachsische Saale" pageId="5" pageNumber="238">Sächsische Saale</location>
stream 
<collectingMunicipality id="6BD0C054FF8CFF85FE9733DBFE26FE0A" box="[275,452,473,497]" pageId="5" pageNumber="238">North of Förbau</collectingMunicipality>
, 
<collectingCounty id="62D522A2FF8CFF85FE4B33D8FD7DFE0A" box="[463,671,473,497]" pageId="5" pageNumber="238">East of Münchberg</collectingCounty>
, 
<geoCoordinate id="EE3F3CE9FF8CFF85FD2D33DBFCE5FE0A" box="[681,775,473,497]" degrees="50.212" direction="north" orientation="latitude" pageId="5" pageNumber="238" precision="55" value="50.212">50.212N</geoCoordinate>
<geoCoordinate id="EE3F3CE9FF8CFF85FF0033FBFF3FFDEA" box="[132,221,505,529]" degrees="11.917" direction="east" orientation="longitude" pageId="5" pageNumber="238" precision="55" value="11.917">11.917E</geoCoordinate>
, 
<date id="FFB57CEEFF8CFF85FF6E33FBFE9AFDEA" box="[234,376,505,529]" pageId="5" pageNumber="238" value="2007-09-26">
<collectingDate id="EFF18506FF8CFF85FF6E33FBFE9AFDEA" box="[234,376,505,529]" pageId="5" pageNumber="238" value="2007-09-26">26 Sep. 2007</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB45A2EFF8CFF85FF003041FD59FC6A" blockId="5.[132,775,579,913]" pageId="5" pageNumber="238">
<emphasis id="B97F863CFF8CFF85FF003041FEA9FDA2" bold="true" box="[132,331,579,601]" pageId="5" pageNumber="238">Acknowledgements</emphasis>
. – This study was supported by the CNRS, the MNHN and the AFB and funded by the partnership MNHN-AFB 2016-2018. We are grateful to the Fishing Authorities, mainly the former CSP-ONEMA, the FDAAPPMA of 
<collectingRegion id="49CF94CCFF8CFF85FDBB3089FD9BFD5A" box="[575,633,651,673]" country="France" name="Pays de la Loire" pageId="5" pageNumber="238">Loire</collectingRegion>
, Haute-Loire, Ardèche, Lozère, Cantal, Haute-Vienne and Puy de Dôme departments, and the Environment Ministry, which physically or finan- cially supported our investigations during so many years. We also thank the fish curators of the NHM, MNHN, UCBLZ and ZSB for providing pictures of specimens of their collections. Our particular thanks go to Pierre Grès, Stéphane Nicolas, Vincent Peyronnet, Valérie Prouha, Romain Max, François Desmolles, Stéphanie Charlat, Lucien Jonard, Marc Mestas, Thierry Millot, Henri Buttazzoni, Grégoire Juglaret, Hervé Brun and Julien Lesbre. Finally, we are very grateful to Agnès Dettai for correcting the manuscript.
</paragraph>
</subSubSection>
</treatment>
</document>