treatments-xml/data/03/88/9E/03889E18FF1355B9FF5C40D5FA1AFE92.xml
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<document id="E03FC9183C304B6318E00743A7F2B3E8" ID="10.11646/zootaxa.4246.1.1SLASH10606" ID-CLB-Dataset="36212" ID-DOI="10.5281/zenodo.437611" ID-GBIF-Dataset="3ff74501-6241-43b2-a52f-0e7580bb2f45" ID-ISSN="1175-5326" ID-Zenodo-Dep="437611" ID-ZooBank="D8594E96-D561-4A37-9073-B138EC7E95A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490260213340" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2017" docId="03889E18FF1355B9FF5C40D5FA1AFE92" docLanguage="en" docName="zootaxa.4246.1.1SLASH10606.pdf" docOrigin="Zootaxa 4246 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Occidenchthonius mateui Zaragoza, 2017, n. sp." docType="treatment" docUuid="B1EE35F4-529E-4DED-8305-A30EEDDDA81F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="176" masterDocId="FFB1E660FFBF5516FFCB4161FF86FFD2" masterDocTitle="Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="221" masterPageNumber="1" pageNumber="173" updateTime="1698438337209" updateUser="ExternalLinkService">
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<mods:title id="25E623F0919B9AC47049A054C1935E11">Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="372FC8DD8B93D54808D2E70086DBF3AF">Zaragoza, Juan A.</mods:namePart>
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<mods:title id="E2C9C25DC3F0893DD776EAB0BFA84480">Zootaxa</mods:title>
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<mods:date id="3717CC1DF014407504DEB900288F7AFB">2017</mods:date>
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<mods:number id="CB9D6B7C6749AE21501DC64C016C6752">4246</mods:number>
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<mods:identifier id="6AF45F18580B43C6A5AF3B0BEC5F92C2" type="DOI">10.5281/zenodo.437611</mods:identifier>
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</mods:mods>
<treatment id="03889E18FF1355B9FF5C40D5FA1AFE92" ID-DOI="http://doi.org/10.5281/zenodo.5687173" ID-GBIF-Taxon="127852112" ID-Zenodo-Dep="5687173" LSID="urn:lsid:zoobank.org:act:B1EE35F4-529E-4DED-8305-A30EEDDDA81F" httpUri="http://treatment.plazi.org/id/03889E18FF1355B9FF5C40D5FA1AFE92" lastPageId="175" lastPageNumber="176" pageId="172" pageNumber="173">
<subSubSection id="C33B7C85FF1355BAFF5C40D5FEC5FE22" pageId="172" pageNumber="173" type="nomenclature">
<paragraph id="8B9E2F0EFF1355BAFF5C40D5FEC5FE22" blockId="172.[151,531,436,496]" pageId="172" pageNumber="173">
<taxonomicName id="4C21548DFF1355BAFF5C40D5FE4BFE1C" ID-CoL="48BQY" box="[151,461,436,462]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="mateui" status="sp. nov.">
<emphasis id="B955F31CFF1355BAFF5C40D5FE4BFE1C" bold="true" box="[151,461,436,462]" italics="true" pageId="172" pageNumber="173">Occidenchthonius mateui</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1355BAFE1F40D4FD95FE1C" bold="true" box="[468,531,437,462]" pageId="172" pageNumber="173">
<taxonomicNameLabel id="A2664E67FF1355BAFE1F40D4FD95FE1C" box="[468,531,437,462]" pageId="172" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131A338BFF1355BAFF5440B6FEBDFE22" box="[159,315,471,496]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="172" pageNumber="173">Figs 272276</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF5C437EFE70FD43" pageId="172" pageNumber="173" type="description">
<paragraph id="8B9E2F0EFF1355BAFF5C437EFD04FD86" blockId="172.[151,1436,543,657]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="AB310820FF1355BAFF5C437EFD04FD86" pageId="172" pageNumber="173">
<taxonomicName id="4C21548DFF1355BAFF5C437EFDD6FDE7" ID-CoL="5Z5FF" box="[151,592,543,565]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="cazorlensis" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFF1355BAFF5C437EFF79FDE7" box="[151,255,543,565]" italics="true" pageId="172" pageNumber="173">Chthonius</emphasis>
(
<emphasis id="B955F31CFF1355BAFEC4437EFE48FDE7" box="[271,462,543,565]" italics="true" pageId="172" pageNumber="173">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFF1355BAFE15437EFDD6FDE7" box="[478,592,543,565]" italics="true" pageId="172" pageNumber="173">cazorlensis</emphasis>
</taxonomicName>
(not Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández 2001):
<treatmentCitation id="0A80091FFF1355BAFAF54341FE03FD86" author="Zaragoza" page="204" pageId="172" pageNumber="173" year="2013">
<bibRefCitation id="EFB052FFFF1355BAFAF54341FED3FD86" author="Zaragoza" pageId="172" pageNumber="173" refString="Zaragoza, J. A. &amp; Perez, T. (2013) Hypogean pseudoscorpions (Arachnida) from Jaen province (Andalusia, Spain), with descriptions of four new species and a new synonymy. Zootaxa, 3700 (2), 201 - 225." type="journal article" year="2013">Zaragoza &amp; Pérez 2013</bibRefCitation>
: 204
</treatmentCitation>
205 (misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9E2F0EFF1355BAFF5C433DFE70FD43" blockId="172.[151,1436,543,657]" pageId="172" pageNumber="173">
<treatmentCitationGroup id="AB310820FF1355BAFF5C433DFE70FD43" pageId="172" pageNumber="173">
<taxonomicName id="4C21548DFF1355BAFF5C433DFDD6FDA0" ID-CoL="5Z5FF" box="[151,592,604,626]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="cazorlensis" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFF1355BAFF5C433DFF79FDA0" box="[151,255,604,626]" italics="true" pageId="172" pageNumber="173">Chthonius</emphasis>
(
<emphasis id="B955F31CFF1355BAFEC4433DFE48FDA0" box="[271,462,604,626]" italics="true" pageId="172" pageNumber="173">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFF1355BAFE15433DFDD6FDA0" box="[478,592,604,626]" italics="true" pageId="172" pageNumber="173">cazorlensis</emphasis>
</taxonomicName>
(not Carabajal Márquez, García Carrillo &amp; Rodríguez Fernández 2001):
<treatmentCitation id="0A80091FFF1355BAFAF5433CFEADFD43" author="Zaragoza" page="66" pageId="172" pageNumber="173" year="2013">
<bibRefCitation id="EFB052FFFF1355BAFAF5433CFE8FFD43" author="Zaragoza" pageId="172" pageNumber="173" refString="Zaragoza, J. A. (2013 a) Los pseudoscorpiones (Arachnida) subterraneos de Jaen. In: Grupo de Espeleologia de Villacarrillo (G. E. V.) (Ed.), Los invertebrados de habitats subterraneos de Jaen, 2013, pp. 65 - 71. [Perez Fernandez, T. y Perez Ruiz, A. (Coord.)]" type="journal article" year="2013" yearSuffix="a">Zaragoza 2013a</bibRefCitation>
: 66
</treatmentCitation>
(misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF5C43D9FE48FD27" pageId="172" pageNumber="173" type="materials_examined">
<paragraph id="8B9E2F0EFF1355BAFF5C43D9FE48FD27" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
<materialsCitation id="3B492553FF1355BAFF5C43D9FE48FD27" ID-GBIF-Occurrence="1449740709" collectorName="Sima de la Fractura II" country="Spain" elevation="1016" latitude="38.18667" location="Jaen province" longLatPrecision="19" longitude="-2.7883334" pageId="172" pageNumber="173" specimenCount="1" stateProvince="Andalusia" typeStatus="holotype">
<emphasis id="B955F31CFF1355BAFF5C43D9FEB8FD03" bold="true" box="[151,318,696,721]" pageId="172" pageNumber="173">
<typeStatus id="549A91ACFF1355BAFF5C43D9FF54FD03" box="[151,210,696,721]" pageId="172" pageNumber="173">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3366F9EFF1355BAFE8543D9FE17FD03" box="[334,401,696,721]" name="Spain" pageId="172" pageNumber="173">Spain</collectingCountry>
,
<collectingRegion id="49E5E1ECFF1355BAFE6F43D9FD9FFD03" box="[420,537,696,721]" country="Spain" name="Andalucia" pageId="172" pageNumber="173">Andalusia</collectingRegion>
,
<location id="8EFE79D5FF1355BAFDE643D8FD52FD03" LSID="urn:lsid:plazi:treatment:03889E18FF1355B9FF5C40D5FA1AFE92:8EFE79D5FF1355BAFDE643D8FD52FD03" box="[557,724,696,721]" country="Spain" latitude="38.18667" longLatPrecision="19" longitude="-2.7883334" name="Jaen province" pageId="172" pageNumber="173" stateProvince="Andalusia">Jaén province</location>
, Hornos del Segura,
<collectorName id="26D44AD8FF1355BAFC2A43D9FB7CFD02" box="[993,1274,696,721]" pageId="172" pageNumber="173">Sima de la Fractura II</collectorName>
(
<geoCoordinate id="EE1549C9FF1355BAFAD943D9FA11FD03" box="[1298,1431,696,721]" direction="north" orientation="latitude" pageId="172" pageNumber="173" precision="15" value="38.18667">38°1112″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FF1355BAFF5C43BDFEA5FD27" box="[151,291,732,757]" direction="west" orientation="longitude" pageId="172" pageNumber="173" precision="15" value="-2.7883334">02°4718″W</geoCoordinate>
;
<elevation id="000CC83DFF1355BAFEE643BDFE39FD27" box="[301,447,732,757]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="172" pageNumber="173" unit="m" value="1016.0">
<quantity id="4CD982EBFF1355BAFEE643BDFE05FD26" box="[301,387,732,757]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="172" pageNumber="173" unit="m" value="1016.0">1016 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF0C439EFDA9FCCA" box="[199,559,767,792]" pageId="172" pageNumber="173" type="distribution">
<paragraph id="8B9E2F0EFF1355BAFF0C439EFDA9FCCA" blockId="172.[151,1437,696,1930]" box="[199,559,767,792]" pageId="172" pageNumber="173">
<emphasis id="B955F31CFF1355BAFF0C439EFEE6FCCA" bold="true" box="[199,352,767,792]" pageId="172" pageNumber="173">Distribution.</emphasis>
<collectingCountry id="F3366F9EFF1355BAFEAC439EFE2DFCCA" box="[359,427,767,792]" name="Spain" pageId="172" pageNumber="173">Spain</collectingCountry>
:
<collectingRegion id="49E5E1ECFF1355BAFE7E439EFDACFCCA" box="[437,554,767,792]" country="Spain" name="Andalucia" pageId="172" pageNumber="173">Andalusia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF0C4242FD17FC74" pageId="172" pageNumber="173" type="etymology">
<paragraph id="8B9E2F0EFF1355BAFF0C4242FD17FC74" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
<emphasis id="B955F31CFF1355BAFF0C4242FEC8FCEE" bold="true" box="[199,334,803,828]" pageId="172" pageNumber="173">Etymology.</emphasis>
At the suggestion of Toni Pérez (G.E.V.), I am pleased to dedicate this species to the memory of the late Dr Joaquim Mateu (
<date id="FF9F09CEFF1355BAFE1F4227FDD1FC8D" box="[468,599,838,863]" pageId="172" pageNumber="173" value="1921" valueMax="2015">19212015</date>
), one of the worlds greatest specialists of
<taxonomicName id="4C21548DFF1355BAFBF34227FB2AFC8D" box="[1080,1196,838,863]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
, in recognition of his great contribution to improving knowledge of the faunas of
<collectingCountry id="F3366F9EFF1355BAFC8D420BFC00FC51" box="[838,902,874,899]" name="Spain" pageId="172" pageNumber="173">Spain</collectingCountry>
and the Saharan region. His highly interesting life and work are described by
<bibRefCitation id="EFB052FFFF1355BAFE3942ECFD0BFC74" author="Belles" box="[498,653,909,934]" pageId="172" pageNumber="173" refString="Belles, X. (2015). Joaquim Mateu (1921 - 2015), tota una vida dedicada a l'estudi dels insects. Museu de Ciencies Naturals de Barcelona. El Blog del Museu. Available from: http: // www. blogmuseuciencies. org / 2015 / 04 / joaquim-mateu- 1921 - 2015 - tota-una-vida-dedicada-a-lestudi-dels-insectes / (accessed 30 December 2015)" type="book chapter" year="2015">Bellés (2015)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF0C42D0FDEAFAFD" pageId="172" pageNumber="173" type="diagnosis">
<paragraph id="8B9E2F0EFF1355BAFF0C42D0FDEAFAFD" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
<emphasis id="B955F31CFF1355BAFF0C42D0FECBFC18" bold="true" box="[199,333,945,970]" pageId="172" pageNumber="173">Diagnosis (</emphasis>
<specimenCount id="9D27E487FF1355BAFE8642D3FEE0FC1B" box="[333,358,946,969]" pageId="172" pageNumber="173" type="male"></specimenCount>
<specimenCount id="9D27E487FF1355BAFEAE42D3FEF0FC1B" box="[357,374,946,969]" pageId="172" pageNumber="173" type="female"></specimenCount>
<emphasis id="B955F31CFF1355BAFEBC42D0FE01FC18" bold="true" box="[375,391,945,970]" pageId="172" pageNumber="173">).</emphasis>
A medium-sized, weakly troglomorphic, hypogean
<taxonomicName id="4C21548DFF1355BAFC1542D0FB2EFC18" box="[990,1192,945,970]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="172" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B955F31CFF1355BAFC1542D0FB2EFC18" box="[990,1192,945,970]" italics="true" pageId="172" pageNumber="173">Occidenchthonius</emphasis>
</taxonomicName>
species of the
<taxonomicName id="4C21548DFF1355BAFA9242D3FA15FC1B" box="[1369,1427,946,969]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="verai">
<emphasis id="B955F31CFF1355BAFA9242D3FA15FC1B" box="[1369,1427,946,969]" italics="true" pageId="172" pageNumber="173">verai</emphasis>
</taxonomicName>
- group. Movable cheliceral finger without isolated subapical tooth (
<emphasis id="B955F31CFF1355BAFC6C42B5FC3BFC3F" box="[935,957,980,1005]" italics="true" pageId="172" pageNumber="173">di</emphasis>
) and spinneret small in female, strongly reduced in male, lyrifissure
<emphasis id="B955F31CFF1355BAFE134296FE7AFBC2" box="[472,508,1015,1040]" italics="true" pageId="172" pageNumber="173">ldb</emphasis>
absent; without eyes or eye-spots, anterior margin of carapace without preocular microsetae, posterior margin with 2 macrosetae; chelal hand strongly depressed at level of
<emphasis id="B955F31CFF1355BAFB4C457AFB1BFBE6" box="[1159,1181,1051,1076]" italics="true" pageId="172" pageNumber="173">ib</emphasis>
/
<emphasis id="B955F31CFF1355BAFB6E457AFB40FBE6" box="[1189,1222,1051,1076]" italics="true" pageId="172" pageNumber="173">isb</emphasis>
, with a short hump distad of
<emphasis id="B955F31CFF1355BAFF34455FFE93FB85" box="[255,277,1086,1111]" italics="true" pageId="172" pageNumber="173">ib</emphasis>
/
<emphasis id="B955F31CFF1355BAFED6455FFEB8FB85" box="[285,318,1086,1111]" italics="true" pageId="172" pageNumber="173">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFF1355BAFCEA455FFCB1FB85" box="[801,823,1086,1111]" italics="true" pageId="172" pageNumber="173">ib</emphasis>
/
<emphasis id="B955F31CFF1355BAFCF4455FFCE6FB85" box="[831,864,1086,1111]" italics="true" pageId="172" pageNumber="173">isb</emphasis>
and
<emphasis id="B955F31CFF1355BAFC5D455FFC36FB85" box="[918,944,1086,1111]" italics="true" pageId="172" pageNumber="173">eb</emphasis>
; fixed chelal finger with 2022 teeth; distal two-thirds of movable chelal finger with 1617 pointed teeth with dental canals, basal third of movable chelal finger with 47 rounded, partially fused, vestigial teeth, without canals, on raised and short lamina; pedipalpal femur (
<specimenCount id="9D27E487FF1355BAFF2045CBFE87FB10" box="[235,257,1194,1218]" pageId="172" pageNumber="173" type="male"></specimenCount>
) 6.87.1, (
<specimenCount id="9D27E487FF1355BAFE4B45CBFE14FB10" box="[384,402,1194,1218]" pageId="172" pageNumber="173" type="female"></specimenCount>
) 6.97.1 times longer than broad, length (
<specimenCount id="9D27E487FF1355BAFCBF45CBFC0CFB10" box="[884,906,1194,1218]" pageId="172" pageNumber="173" type="male"></specimenCount>
)
<quantity id="4CD982EBFF1355BAFC5045C8FBC7FB13" box="[923,1089,1193,1218]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="172" pageNumber="173" unit="mm" value="0.73" valueMax="0.78" valueMin="0.68">0.680.78 mm</quantity>
, (
<specimenCount id="9D27E487FF1355BAFB9345CBFBECFB10" box="[1112,1130,1194,1218]" pageId="172" pageNumber="173" type="female"></specimenCount>
)
<quantity id="4CD982EBFF1355BAFBB545CBFAA2FB13" box="[1150,1316,1194,1218]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.0" metricValueMin="7.8" pageId="172" pageNumber="173" unit="mm" value="0.79" valueMax="0.8" valueMin="0.78">0.780.80 mm</quantity>
; chela (
<specimenCount id="9D27E487FF1355BAFAB645CBFA15FB10" box="[1405,1427,1194,1218]" pageId="172" pageNumber="173" type="male"></specimenCount>
) 6.26.6, (
<specimenCount id="9D27E487FF1355BAFECE45ACFE91FB37" box="[261,279,1229,1253]" pageId="172" pageNumber="173" type="female"></specimenCount>
) 5.96.1 times longer than deep, length (
<specimenCount id="9D27E487FF1355BAFD3845ACFC8FFB37" box="[755,777,1229,1253]" pageId="172" pageNumber="173" type="male"></specimenCount>
)
<quantity id="4CD982EBFF1355BAFCD145ACFC47FB36" box="[794,961,1229,1253]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" metricValueMax="10.3" metricValueMin="9.3" pageId="172" pageNumber="173" unit="mm" value="0.98" valueMax="1.03" valueMin="0.93">0.931.03 mm</quantity>
, (
<specimenCount id="9D27E487FF1355BAFC1245ACFC6DFB37" box="[985,1003,1229,1253]" pageId="172" pageNumber="173" type="female"></specimenCount>
)
<quantity id="4CD982EBFF1355BAFC3445ACFB21FB36" box="[1023,1191,1229,1253]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.08" metricValueMin="1.07" pageId="172" pageNumber="173" unit="mm" value="1.075" valueMax="1.08" valueMin="1.07">1.071.08 mm</quantity>
; ratio movable chelal finger/chelal hand (
<specimenCount id="9D27E487FF1355BAFEBD4591FE0AFADA" box="[374,396,1264,1288]" pageId="172" pageNumber="173" type="male"></specimenCount>
) 1.41.5, (
<specimenCount id="9D27E487FF1355BAFDC64591FD99FADA" box="[525,543,1264,1288]" pageId="172" pageNumber="173" type="female"></specimenCount>
) 1.5; lacking lyrifissures
<emphasis id="B955F31CFF1355BAFC824590FCF5FADF" box="[841,883,1265,1293]" italics="true" pageId="172" pageNumber="173">ma1</emphasis>
,
<emphasis id="B955F31CFF1355BAFC484590FC2BFADF" box="[899,941,1265,1293]" italics="true" pageId="172" pageNumber="173">ma2</emphasis>
and
<emphasis id="B955F31CFF1355BAFC234591FB82FADB" box="[1000,1028,1264,1289]" italics="true" pageId="172" pageNumber="173">hp</emphasis>
, all other chelal lyrifissure patterns present, with their standard complements.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355BAFF0C445BFDB1FA4B" pageId="172" pageNumber="173" type="materials_examined">
<paragraph id="8B9E2F0EFF1355BAFF0C445BFDB1FA4B" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
<emphasis id="B955F31CFF1355BAFF0C445BFE77FA81" bold="true" box="[199,497,1338,1363]" pageId="172" pageNumber="173">Type material examined.</emphasis>
♂ holotype (DEUA), 3 ♂ (DEUA, MCNB, MNCN),
<materialsCitation id="3B492553FF1355BAFB90445BFDB1FA4B" ID-GBIF-Occurrence="1449740718" collectingDate="2009-05-31" collectionCode="DEUA" collectorName="Sima de la Fractura II" country="Spain" elevation="1016" latitude="38.18667" location="Jaen province" longLatPrecision="19" longitude="-2.7883334" pageId="172" pageNumber="173" specimenCount="2" specimenCount-female="2" stateProvince="Andalusia" typeStatus="paratype">
<specimenCount id="9D27E487FF1355BAFB90445BFB00FA80" box="[1115,1158,1338,1362]" pageId="172" pageNumber="173" type="female">2 ♀</specimenCount>
, 1 tritonymph
<typeStatus id="549A91ACFF1355BAFAFA445AFA1AFA81" box="[1329,1436,1339,1363]" pageId="172" pageNumber="173" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED30B7CBFF1355BAFF54443FFF74FAA4" box="[159,242,1374,1398]" pageId="172" pageNumber="173">DEUA</collectionCode>
),
<collectingCountry id="F3366F9EFF1355BAFECC443CFECCFAA4" box="[263,330,1373,1398]" name="Spain" pageId="172" pageNumber="173">Spain</collectingCountry>
,
<collectingRegion id="49E5E1ECFF1355BAFE9C443CFE4AFAA4" box="[343,460,1373,1398]" country="Spain" name="Andalucia" pageId="172" pageNumber="173">Andalusia</collectingRegion>
,
<location id="8EFE79D5FF1355BAFE12443FFDFFFAA4" LSID="urn:lsid:plazi:treatment:03889E18FF1355B9FF5C40D5FA1AFE92:8EFE79D5FF1355BAFE12443FFDFFFAA4" box="[473,633,1373,1398]" country="Spain" latitude="38.18667" longLatPrecision="19" longitude="-2.7883334" name="Jaen province" pageId="172" pageNumber="173" stateProvince="Andalusia">Jaén province</location>
, Hornos del Segura,
<collectorName id="26D44AD8FF1355BAFCA0443CFBEEFAA7" box="[875,1128,1373,1398]" pageId="172" pageNumber="173">Sima de la Fractura II</collectorName>
(
<geoCoordinate id="EE1549C9FF1355BAFBB2443CFB78FAA4" box="[1145,1278,1373,1398]" direction="north" orientation="latitude" pageId="172" pageNumber="173" precision="15" value="38.18667">38°1112″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FF1355BAFAC7443CFA1EFAA4" box="[1292,1432,1373,1398]" direction="west" orientation="longitude" pageId="172" pageNumber="173" precision="15" value="-2.7883334">02°4718″W</geoCoordinate>
;
<elevation id="000CC83DFF1355BAFF5C44E1FEA1FA4B" box="[151,295,1408,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="172" pageNumber="173" unit="m" value="1016.0">
<quantity id="4CD982EBFF1355BAFF5C44E1FF6AFA4A" box="[151,236,1408,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.016" pageId="172" pageNumber="173" unit="m" value="1016.0">1016 m</quantity>
a.s.l.
</elevation>
),
<date id="FF9F09CEFF1355BAFEF544E0FE35FA4B" box="[318,435,1409,1433]" pageId="172" pageNumber="173" value="2009-05-31">
<collectingDate id="EFDBF026FF1355BAFEF544E0FE35FA4B" box="[318,435,1409,1433]" pageId="172" pageNumber="173" value="2009-05-31">31.V.2009</collectingDate>
</date>
, leg. G.E.V.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1355B8FF0C44C5FE8DF837" lastPageId="174" lastPageNumber="175" pageId="172" pageNumber="173" type="description">
<paragraph id="8B9E2F0EFF1355BAFF0C44C5FF6AF9D6" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
<emphasis id="B955F31CFF1355BAFF0C44C5FEF7FA6F" bold="true" box="[199,369,1444,1469]" pageId="172" pageNumber="173">Description (</emphasis>
<specimenCount id="9D27E487FF1355BAFEB944C4FE0DFA6E" box="[370,395,1445,1468]" pageId="172" pageNumber="173" type="male"></specimenCount>
<specimenCount id="9D27E487FF1355BAFE4144C4FE1DFA6E" box="[394,411,1445,1468]" pageId="172" pageNumber="173" type="female"></specimenCount>
<emphasis id="B955F31CFF1355BAFE5744C5FE2BFA6F" bold="true" box="[412,429,1444,1469]" pageId="172" pageNumber="173">).</emphasis>
Medium-sized, hypogean species of weakly troglomorphic facies. Integument depigmented; weak hispid granulation on lateral surfaces of carapace, on cheliceral hand and on bases of chelal fingers.
</paragraph>
<paragraph id="8B9E2F0EFF1355BAFF0C476EFB79F940" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
Carapace (
<figureCitation id="131A338BFF1355BAFEF5476FFE19F9F5" box="[318,415,1550,1575]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="172" pageNumber="173">Fig. 273</figureCitation>
) slightly longer than broad, constricted posteriorly; medial part of anterior margin strongly prominent, without a well-defined epistome, and strongly dentate (
<figureCitation id="131A338BFF1355BAFC054753FBBFF999" box="[974,1081,1586,1611]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="172" pageNumber="173">Fig. 272</figureCitation>
). Without eyes or eye-spots. Chaetotaxy: 18 setae, without preocular microsetae, 2 setae in posterior row, formula 4:6:4:2:2, anteromedial setae
<quantity id="4CD982EBFF1355BAFF5C4718FEBFF943" box="[151,313,1657,1681]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="172" pageNumber="173" unit="mm" value="0.12" valueMax="0.13" valueMin="0.11">0.110.13 mm</quantity>
long, sublateral ocular setae
<quantity id="4CD982EBFF1355BAFDB44718FCA2F943" box="[639,804,1657,1681]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="172" pageNumber="173" unit="mm" value="0.11499999999999999" valueMax="0.12" valueMin="0.11">0.110.12 mm</quantity>
; 4 lyrifissures anteriorly and 2 posteriorly.
</paragraph>
<paragraph id="8B9E2F0EFF1355BAFF0C47FDFEE5F858" blockId="172.[151,1437,696,1930]" pageId="172" pageNumber="173">
Chelicera (
<figureCitation id="131A338BFF1355BAFE8D47FDFE6BF967" box="[326,493,1692,1717]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="172" pageNumber="173">Figs 274, 275</figureCitation>
) with 6 setae and 1 lateral microseta on hand, seta
<emphasis id="B955F31CFF1355BAFBA947FDFBFBF967" box="[1122,1149,1692,1717]" italics="true" pageId="172" pageNumber="173">vb</emphasis>
short (
<quantity id="4CD982EBFF1355BAFB1F47FDFAC8F966" box="[1236,1358,1692,1717]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" pageId="172" pageNumber="173" unit="mm" value="0.045">0.045 mm</quantity>
long), microseta
<quantity id="4CD982EBFF1355BAFEC047A1FE06F90A" box="[267,384,1728,1753]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="172" pageNumber="173" unit="mm" value="0.035">0.035 mm</quantity>
; hand with 4 dorsal and 1 ventral lyrifissure, lyrifissure
<emphasis id="B955F31CFF1355BAFC3C47A1FB9DF90B" box="[1015,1051,1728,1753]" italics="true" pageId="172" pageNumber="173">ldb</emphasis>
absent. Fixed finger with 11 teeth decreasing in size proximally, two distal teeth distinctly larger than others, 3 proximal microtubercles. Movable finger without an isolated subapical tooth (
<emphasis id="B955F31CFF1355BAFD534666FD2BF8F2" box="[664,685,1799,1824]" italics="true" pageId="172" pageNumber="173">di</emphasis>
), with 78 teeth decreasing in size proximally and 2 proximal microtubercles, the distal tooth larger than others; spinneret strongly reduced in male (
<figureCitation id="131A338BFF1355BAFBA6464BFB49F891" box="[1133,1231,1834,1859]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="172" pageNumber="173">Fig. 275</figureCitation>
), small in female; seta
<emphasis id="B955F31CFF1355BAFF07462CFF64F8B4" box="[204,226,1869,1894]" italics="true" pageId="172" pageNumber="173">gl</emphasis>
0.530.56 from base of movable finger. Rallum with 11 blades. Serrula exterior with 15 blades, serrula interior 13 blades.
</paragraph>
<caption id="DF5E7F86FF1255BBFF5C46C1FB4AF804" httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="173" pageNumber="174" targetBox="[228,1376,224,1902]" targetPageId="173">
<paragraph id="8B9E2F0EFF1255BBFF5C46C1FB4AF804" blockId="173.[151,1436,1952,2006]" pageId="173" pageNumber="174">
<emphasis id="B955F31CFF1255BBFF5C46C1FEF6F864" bold="true" box="[151,368,1952,1975]" pageId="173" pageNumber="174">FIGURES 272276.</emphasis>
<taxonomicName id="4C21548DFF1255BBFEB146C0FD06F865" box="[378,640,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="173" pageNumber="174" phylum="Arthropoda" rank="species" species="mateui" status="sp. nov.">
<emphasis id="B955F31CFF1255BBFEB146C0FD06F865" box="[378,640,1953,1975]" italics="true" pageId="173" pageNumber="174">Occidenchthonius mateui</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1255BBFD4146C0FD41F864" bold="true" box="[650,711,1953,1974]" pageId="173" pageNumber="174">
<taxonomicNameLabel id="A2664E67FF1255BBFD4146C0FD41F864" box="[650,711,1953,1974]" pageId="173" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view.
</paragraph>
</caption>
<paragraph id="8B9E2F0EFF1155B8FF0C41F6FC57FECE" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
Chaetotaxy of tergites 4:4:4:4:6:6:6:6:1T2T1:4:1T2T1:0, tergites IX and XI each with 2 sublateral tactile setae (0.24 and
<quantity id="4CD982EBFF1155B8FEC141DDFEF6FF06" box="[266,368,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="174" pageNumber="175" unit="mm" value="0.33">0.33 mm</quantity>
long, respectively). Chaetotaxy of sternites 910:(3)7 9(3):(2)67(2):78:6:6:6:6:2T1T2:0:2, lateral setae on sternite III macrosetal in size, sternite X with 2 submedial tactile setae; in addition, genital notch of male flanked by 610 setae on each side and 4+4 internal glandular setae.
</paragraph>
<paragraph id="8B9E2F0EFF1155B8FF0C4046FC47FE5A" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
Pedipalpal coxa with 5 setae (including 2 on manducatory process), distal marginal seta of disk
<quantity id="4CD982EBFF1155B8FB314049FA1BFEED" box="[1274,1437,296,320]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="174" pageNumber="175" unit="mm" value="0.09" valueMax="0.1" valueMin="0.08">0.080.10 mm</quantity>
long; coxa
<date id="FF9F09CEFF1155B8FED9402CFEB6FEB6" box="[274,304,332,356]" pageId="174" pageNumber="175">I 3</date>
+ 3 marginal microsetae, distal marginal seta
<quantity id="4CD982EBFF1155B8FCF9402DFC52FEB6" box="[818,980,332,357]" metricMagnitude="-5" metricUnit="m" metricValue="7" metricValueMax="8.0" metricValueMin="6.0" pageId="174" pageNumber="175" unit="mm" value="0.07" valueMax="0.08" valueMin="0.06">0.060.08 mm</quantity>
long;
<date id="FF9F09CEFF1155B8FBD0402CFBC7FEB6" box="[1051,1089,332,356]" pageId="174" pageNumber="175">II 4</date>
+ 712 bipinnate coxal spines,
<date id="FF9F09CEFF1155B8FF5C4011FF4EFE5A" box="[151,200,367,392]" pageId="174" pageNumber="175">III 5</date>
+ 36 bipinnate coxal spines and
<date id="FF9F09CEFF1155B8FD8D4011FDFCFE5A" box="[582,634,367,392]" pageId="174" pageNumber="175">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="8B9E2F0EFF1155B8FF0C40F5FDE3FB50" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
Pedipalp with femoral chaetotaxy 3:67:3:5:1. Chela (
<figureCitation id="131A338BFF1155B8FCEE40F5FC03FE7F" box="[805,901,404,429]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="174" pageNumber="175">Fig. 276</figureCitation>
) with hand strongly depressed at level of
<emphasis id="B955F31CFF1155B8FA9C40F5FAEBFE7F" box="[1367,1389,404,429]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FABF40F5FA13FE7F" box="[1396,1429,404,429]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
, with a short hump distad of
<emphasis id="B955F31CFF1155B8FE0640D6FE65FE02" box="[461,483,439,464]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FE2040D6FD8AFE02" box="[491,524,439,464]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFF1155B8FC2040D6FB87FE02" box="[1003,1025,439,464]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FBC340D6FBAFFE02" box="[1032,1065,439,464]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
and
<emphasis id="B955F31CFF1155B8FB9540D6FBFFFE02" box="[1118,1145,439,464]" italics="true" pageId="174" pageNumber="175">eb</emphasis>
; weak hollow before base of movable finger with thicker cuticle; width slightly shorter than depth, maximum width proximad of
<emphasis id="B955F31CFF1155B8FA9D40BDFAEAFE27" box="[1366,1388,476,501]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FABF40BDFA13FE27" box="[1396,1429,476,501]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
; chaetotaxy 4:5:4, seta
<emphasis id="B955F31CFF1155B8FE5B409EFE32FDCE" box="[400,436,511,540]" italics="true" pageId="174" pageNumber="175">ph3</emphasis>
present; distal end of hand and bases of chelal fingers with sclerotized condylar complex. Fixed finger with 2022 pointed teeth and with dental canals, first two distal teeth small, third tooth of normal row (
<emphasis id="B955F31CFF1155B8FF6B432DFF3AFDB1" box="[160,188,588,611]" italics="true" pageId="174" pageNumber="175">mt</emphasis>
) of fixed finger distinctly modified in shape and deviated in orientation with respect to the others, dental row reaching up to halfway between sensilla
<emphasis id="B955F31CFF1155B8FDAE4311FDF9FD55" box="[613,639,624,647]" italics="true" pageId="174" pageNumber="175">pc</emphasis>
and trichobothrium
<emphasis id="B955F31CFF1155B8FCA3430FFC07FD55" box="[872,897,622,647]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
, about 10 proximal microtubercles; tip of fixed finger with an accessory tooth (
<emphasis id="B955F31CFF1155B8FDC243F3FD99FD79" box="[521,543,658,683]" italics="true" pageId="174" pageNumber="175">td</emphasis>
) on antiaxial face; tip of fixed chelal finger of male with a weak hollow on paraxial face, without subdistal protuberance (
<emphasis id="B955F31CFF1155B8FD5043D9FD32FD1D" box="[667,692,696,719]" italics="true" pageId="174" pageNumber="175">sp</emphasis>
); one pair of long antiaxial sensory setae (
<emphasis id="B955F31CFF1155B8FB4743D9FB23FD1D" box="[1164,1189,696,719]" italics="true" pageId="174" pageNumber="175">as</emphasis>
) at the base, one level with and other markedly distad of lyrifissure
<emphasis id="B955F31CFF1155B8FD5043BBFD37FD21" box="[667,689,730,755]" italics="true" pageId="174" pageNumber="175">fb</emphasis>
,
<quantity id="4CD982EBFF1155B8FD0B43BBFC07FD20" box="[704,897,730,755]" metricMagnitude="-5" metricUnit="m" metricValue="4.249999999999999" metricValueMax="4.5" metricValueMin="4.0" pageId="174" pageNumber="175" unit="mm" value="0.042499999999999996" valueMax="0.045" valueMin="0.04">0.0400.045 mm</quantity>
long, distance between them
<quantity id="4CD982EBFF1155B8FB1E43BBFA1EFD20" box="[1237,1432,730,755]" metricMagnitude="-5" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="5.0" metricValueMin="4.299999999999999" pageId="174" pageNumber="175" unit="mm" value="0.0465" valueMax="0.05" valueMin="0.043">0.0430.050 mm</quantity>
, fixed finger depth at the base
<quantity id="4CD982EBFF1155B8FE30439FFD46FCC5" box="[507,704,766,791]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="174" pageNumber="175" unit="mm" value="0.06" valueMax="0.065" valueMin="0.055">0.0550.065 mm</quantity>
; 5 teeth at level of
<emphasis id="B955F31CFF1155B8FC674261FC4DFCC5" box="[940,971,768,791]" italics="true" pageId="174" pageNumber="175">est</emphasis>
/
<emphasis id="B955F31CFF1155B8FC184261FC65FCC5" box="[979,995,768,791]" italics="true" pageId="174" pageNumber="175">it</emphasis>
occupying
<quantity id="4CD982EBFF1155B8FBA6439EFB4FFCC5" box="[1133,1225,767,791]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="174" pageNumber="175" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<quantity id="4CD982EBFF1155B8FEA14243FDB7FCE8" box="[362,561,802,827]" metricMagnitude="-5" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="174" pageNumber="175" unit="mm" value="0.0225" valueMax="0.025" valueMin="0.02">0.0200.025 mm</quantity>
. Distal two-thirds of movable finger with 1617 pointed teeth with dental canals that reach up to slightly proximad of halfway between trichobothria
<emphasis id="B955F31CFF1155B8FC174229FC69FC8D" box="[988,1007,840,863]" italics="true" pageId="174" pageNumber="175">st</emphasis>
and
<emphasis id="B955F31CFF1155B8FBEF4227FBBBFC8D" box="[1060,1085,838,863]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
, 23 proximal teeth decreasing in size, the distal tooth tiny, the subdistal tooth small; basal third of movable chelal finger with only 47 rounded, partially fused, vestigial teeth, without canals, on weakly raised and short lamina; dental row reaching up to level of trichobothrium
<emphasis id="B955F31CFF1155B8FEAF42D3FEFBFC19" box="[356,381,946,971]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
or sensilla
<emphasis id="B955F31CFF1155B8FE3542D5FD9FFC19" box="[510,537,948,971]" italics="true" pageId="174" pageNumber="175">pc</emphasis>
, 5 proximal microtubercles; basal apodeme long and apically indented; coupled sensilla
<emphasis id="B955F31CFF1155B8FF3D42B9FE97FC3D" box="[246,273,984,1007]" italics="true" pageId="174" pageNumber="175">pc</emphasis>
halfway between
<emphasis id="B955F31CFF1155B8FE2342B7FD87FC3D" box="[488,513,982,1007]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
and
<emphasis id="B955F31CFF1155B8FD8B42B7FDC8FC3D" box="[576,590,982,1007]" italics="true" pageId="174" pageNumber="175">b</emphasis>
or slightly closer to
<emphasis id="B955F31CFF1155B8FC8242B7FCE4FC3D" box="[841,866,982,1007]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
. Trichobothria as in
<figureCitation id="131A338BFF1155B8FB9042B7FB46FC3D" box="[1115,1216,982,1007]" captionStart="FIGURES 272 276" captionStartId="173.[151,264,1952,1975]" captionTargetBox="[228,1376,224,1902]" captionTargetId="figure@173.[151,1436,193,1932]" captionTargetPageId="173" captionText="FIGURES 272 276. Occidenchthonius mateui n. sp., male holotype. 272, anterior margin of carapace, partial view; 273, carapace; 274, right chelicera; 275, fingers of right chelicera, partial view; 276, right chela, antiaxial view." httpUri="https://zenodo.org/record/437670/files/figure.png" pageId="174" pageNumber="175">Fig. 276</figureCitation>
; trichobothrium
<emphasis id="B955F31CFF1155B8FA4A42B9FA1AFC3D" box="[1409,1436,984,1007]" italics="true" pageId="174" pageNumber="175">ist</emphasis>
distinctly distad of
<emphasis id="B955F31CFF1155B8FEA4429BFE12FBC1" box="[367,404,1018,1043]" italics="true" pageId="174" pageNumber="175">esb</emphasis>
, trichobothria
<emphasis id="B955F31CFF1155B8FDF1429BFD20FBC1" box="[570,678,1018,1043]" italics="true" pageId="174" pageNumber="175">eb -esb -ist</emphasis>
in a straight line, and distinctly proximad of lyrifissure
<emphasis id="B955F31CFF1155B8FAD0429BFAB7FBC1" box="[1307,1329,1018,1043]" italics="true" pageId="174" pageNumber="175">fb</emphasis>
; distance between
<emphasis id="B955F31CFF1155B8FF36457FFE95FBE5" box="[253,275,1054,1079]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FED0457FFEBAFBE5" box="[283,316,1054,1079]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
and base of hand 1.1 longer than that between
<emphasis id="B955F31CFF1155B8FC94457FFCF3FBE5" box="[863,885,1054,1079]" italics="true" pageId="174" pageNumber="175">ib</emphasis>
/
<emphasis id="B955F31CFF1155B8FCB6457FFC18FBE5" box="[893,926,1054,1079]" italics="true" pageId="174" pageNumber="175">isb</emphasis>
and
<emphasis id="B955F31CFF1155B8FC13457FFC7BFBE5" box="[984,1021,1054,1079]" italics="true" pageId="174" pageNumber="175">esb</emphasis>
; distance between
<emphasis id="B955F31CFF1155B8FB124541FB6AFBE5" box="[1241,1260,1056,1079]" italics="true" pageId="174" pageNumber="175">st</emphasis>
and
<emphasis id="B955F31CFF1155B8FAED457FFAB9FBE5" box="[1318,1343,1054,1079]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
1.92.2 times longer than that between
<emphasis id="B955F31CFF1155B8FE364523FD90FB89" box="[509,534,1090,1115]" italics="true" pageId="174" pageNumber="175">sb</emphasis>
and
<emphasis id="B955F31CFF1155B8FD9B4523FDD8FB89" box="[592,606,1090,1115]" italics="true" pageId="174" pageNumber="175">b</emphasis>
; lacking lyrifissures
<emphasis id="B955F31CFF1155B8FC854525FCFFFB8D" box="[846,889,1092,1119]" italics="true" pageId="174" pageNumber="175">ma1</emphasis>
,
<emphasis id="B955F31CFF1155B8FC434525FC34FB8D" box="[904,946,1092,1119]" italics="true" pageId="174" pageNumber="175">ma2</emphasis>
and
<emphasis id="B955F31CFF1155B8FC204523FB81FB89" box="[1003,1031,1090,1115]" italics="true" pageId="174" pageNumber="175">hp</emphasis>
, all other chelal lyrifissure patterns present with their standard complements.
</paragraph>
<paragraph id="8B9E2F0EFF1155B8FF0C45EFFD70FA14" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
<emphasis id="B955F31CFF1155B8FF0C45EFFE62FB77" box="[199,484,1165,1190]" italics="true" pageId="174" pageNumber="175">Measurements and ratios</emphasis>
. Male
<typeStatus id="549A91ACFF1155B8FDFE45ECFD1CFB74" box="[565,666,1165,1190]" pageId="174" pageNumber="175" type="holotype">holotype</typeStatus>
, followed, when different, by one male
<typeStatus id="549A91ACFF1155B8FBAB45EFFB46FB74" box="[1120,1216,1166,1190]" pageId="174" pageNumber="175" type="paratype">paratype</typeStatus>
in square brackets: Body 1.52 [1.69]. Carapace 0.43/0.41 (1.0) [0.49/0.48]. Chelicera 0.440.20 (2.2), movable finger 0.23. Pedipalp: femur 0.68/0.10 (7.1) [0.78/0.12 (6.8)], patella 0.30/0.12 (2.4) [0.31/0.14 (2.2)], chela 0.93/0.14 (6.6) [1.03/0.17 (6.2)], hand 0.37 (2.6) [0.41 (2.5)], movable finger 0.56 [0.60]; ratio movable finger/hand 1.5 [1.4], femur/movable finger 1.2 [1.3], femur/carapace 1.6, chela/carapace 2.2 [2.1], chela/femur 1.4 [1.3]. Female
<typeStatus id="549A91ACFF1155B8FB6B447FFA96FAE4" box="[1184,1296,1310,1334]" pageId="174" pageNumber="175" type="paratype">paratypes</typeStatus>
: Body 1.58 1.78. Carapace 0.480.5/0.470.48 (1.01.1). Chelicera 0.450.47/0.210.22 (2.12.2), movable finger 0.230.24. Pedipalp: femur 0.780.80/0.110.12 (6.97.1), patella 0.330.34/0.15 (2.2), chela 1.071.08/0.18 (5.96.1), hand 0.430.44 (2.4), movable finger 0.640.65; ratio movable finger/hand 1.5, femur/movable finger 1.2, femur/ carapace 1.6, chela/carapace 2.12.2, chela/femur 1.4.
</paragraph>
<paragraph id="8B9E2F0EFF1155B8FF0C44B0FAF9F883" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
<emphasis id="B955F31CFF1155B8FF0C44B0FED8FA38" bold="true" box="[199,350,1489,1514]" pageId="174" pageNumber="175">Tritonymph.</emphasis>
Carapace as long as broad; medial part of anterior margin straight and strongly dentate; without eyes; chaetotaxy as in adult. Cheliceral hand with 5 setae and 1 lateral microseta; fixed finger with 9 teeth, two distal teeth larger than others; movable finger without an isolated subapical tooth (
<emphasis id="B955F31CFF1155B8FB8B4778FBD3F9E0" box="[1088,1109,1561,1586]" italics="true" pageId="174" pageNumber="175">di</emphasis>
), with 6 teeth, the distal one larger than others; spinneret prominent as in adult female; seta
<emphasis id="B955F31CFF1155B8FC97475CFCF4F984" box="[860,882,1597,1622]" italics="true" pageId="174" pageNumber="175">gl</emphasis>
0.56 from base of movable finger. Chaetotaxy of tergites as in adults, except tergite IX with 5 setae: 1T3; sternites 6:(2)6(2):(1)6(1):7:6:6:6:6:2T1T2:0:2. Pedipalpal coxa 5 setae (including 2 on manducatory process), distal marginal seta of disk
<quantity id="4CD982EBFF1155B8FB8847E4FB3EF94F" box="[1091,1208,1669,1694]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="174" pageNumber="175" unit="mm" value="0.06">0.060 mm</quantity>
long; coxa
<date id="FF9F09CEFF1155B8FA8C47E7FAEFF94C" box="[1351,1385,1670,1694]" pageId="174" pageNumber="175">I 3</date>
+ 2 marginal microsetae, distal marginal seta
<quantity id="4CD982EBFF1155B8FDA547CBFD67F913" box="[622,737,1706,1730]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="174" pageNumber="175" unit="mm" value="0.04">0.040 mm</quantity>
long;
<date id="FF9F09CEFF1155B8FCE147CBFCD5F910" box="[810,851,1706,1730]" pageId="174" pageNumber="175">II 4</date>
+ 79 bipinnate coxal spines,
<date id="FF9F09CEFF1155B8FB7B47CBFB64F910" box="[1200,1250,1705,1730]" pageId="174" pageNumber="175">III 5</date>
+ 45 bipinnate coxal spines and
<date id="FF9F09CEFF1155B8FE9547AFFE12F934" box="[350,404,1741,1766]" pageId="174" pageNumber="175">IV 5</date>
; intercoxal tubercle bisetose. Pedipalp: chelal hand chaetotaxy 4:5:4; fixed finger with 16 pointed teeth with dental canals, first two distal teeth small, third tooth of normal row modified (
<emphasis id="B955F31CFF1155B8FB124793FB73F8DB" box="[1241,1269,1778,1801]" italics="true" pageId="174" pageNumber="175">mt</emphasis>
); distal half of movable finger with 11 pointed teeth with dental canals; proximal half of finger with slightly raised lamina with 6 rounded, vestigial teeth; coupled sensilla
<emphasis id="B955F31CFF1155B8FDAD465BFD07F883" box="[614,641,1850,1873]" italics="true" pageId="174" pageNumber="175">pc</emphasis>
distad of trichobothrium
<emphasis id="B955F31CFF1155B8FC544658FC2BF880" box="[927,941,1849,1874]" italics="true" pageId="174" pageNumber="175">b</emphasis>
; lacking lyrifissures
<emphasis id="B955F31CFF1155B8FB534658FB30F884" box="[1176,1206,1849,1878]" italics="true" pageId="174" pageNumber="175">fd3</emphasis>
,
<emphasis id="B955F31CFF1155B8FB0F4658FB66F880" box="[1220,1248,1849,1874]" italics="true" pageId="174" pageNumber="175">hp</emphasis>
,
<emphasis id="B955F31CFF1155B8FB25465BFA9EF884" box="[1262,1304,1850,1878]" italics="true" pageId="174" pageNumber="175">ma1</emphasis>
and
<emphasis id="B955F31CFF1155B8FA85465BFAFEF884" box="[1358,1400,1850,1878]" italics="true" pageId="174" pageNumber="175">ma2</emphasis>
.
</paragraph>
<paragraph id="8B9E2F0EFF1155B8FF0C4600FE8DF837" blockId="174.[151,1437,151,2021]" pageId="174" pageNumber="175">
<emphasis id="B955F31CFF1155B8FF0C4600FE6DF8AA" box="[199,491,1888,1913]" italics="true" pageId="174" pageNumber="175">Measurements and ratios</emphasis>
. Body 1.18. Carapace 0.33/0.33 (1.0). Chelicera 0.30/0.15 (2.1), movable finger 0.16. Pedipalp: femur 0.46/0.08 (5.8), patella 0.21/0.10 (2.1), chela 0.65/0.12 (5.7), hand 0.26 (2.3), movable finger 0.39; ratio movable finger/hand 1.5, femur/movable finger 1.2, femur/carapace 1.4, chela/carapace 2.0, chela/ femur 1.4.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF1055B9FF0C41F6FA1AFE92" pageId="175" pageNumber="176" type="discussion">
<paragraph id="8B9E2F0EFF1055B9FF0C41F6FA1AFE92" blockId="175.[151,1436,151,320]" pageId="175" pageNumber="176">
<emphasis id="B955F31CFF1055B9FF0C41F6FEBDFF62" bold="true" box="[199,315,151,176]" pageId="175" pageNumber="176">Remarks.</emphasis>
Within the
<taxonomicName id="4C21548DFF1055B9FE0241F8FD85FF62" box="[457,515,153,176]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="verai">
<emphasis id="B955F31CFF1055B9FE0241F8FD85FF62" box="[457,515,153,176]" italics="true" pageId="175" pageNumber="176">verai</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C21548DFF1055B9FD9541F9FD56FF62" box="[606,720,152,176]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="mateui" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FD9541F9FD56FF62" box="[606,720,152,176]" italics="true" pageId="175" pageNumber="176">O. mateui</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FD1041F6FC9AFF62" bold="true" box="[731,796,151,176]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FD1041F6FC9AFF62" box="[731,796,151,176]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shares with
<taxonomicName id="4C21548DFF1055B9FC7F41F9FBD1FF62" box="[948,1111,151,176]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="cazorlensis">
<emphasis id="B955F31CFF1055B9FC7F41F9FBD1FF62" box="[948,1111,151,176]" italics="true" pageId="175" pageNumber="176">O. cazorlensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21548DFF1055B9FBA241F9FB70FF62" box="[1129,1270,151,176]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="hoerwegi" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FBA241F9FB70FF62" box="[1129,1270,151,176]" italics="true" pageId="175" pageNumber="176">O. hoerwegi</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FACA41F6FAC5FF62" bold="true" box="[1281,1347,151,176]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FACA41F6FAC5FF62" box="[1281,1347,151,176]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C21548DFF1055B9FA4A41F9FF59FF06" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="riopar" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FA4A41F9FF59FF06" italics="true" pageId="175" pageNumber="176">O. riopar</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FF2E41DDFEA3FF07" bold="true" box="[229,293,188,213]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FF2E41DDFEA3FF07" box="[229,293,188,213]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
a short row of only 38 low, rounded, vestigial teeth in the basal third of the movable chelal finger. It differs from
<taxonomicName id="4C21548DFF1055B9FEEF4181FE43FF2A" box="[292,453,223,248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="cazorlensis">
<emphasis id="B955F31CFF1055B9FEEF4181FE43FF2A" box="[292,453,223,248]" italics="true" pageId="175" pageNumber="176">O. cazorlensis</emphasis>
</taxonomicName>
by the absence of cheliceral lyrifissure
<emphasis id="B955F31CFF1055B9FC4E41BEFC2FFF2A" box="[901,937,223,248]" italics="true" pageId="175" pageNumber="176">ldb</emphasis>
and from
<taxonomicName id="4C21548DFF1055B9FBD54181FB0EFF2A" box="[1054,1160,224,248]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="riopar" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FBD54181FB0EFF2A" box="[1054,1160,224,248]" italics="true" pageId="175" pageNumber="176">O. riopar</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FB4441BEFB48FF2A" bold="true" box="[1167,1230,223,248]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FB4441BEFB48FF2A" box="[1167,1230,223,248]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the position of trichobothrium
<emphasis id="B955F31CFF1055B9FE8D4064FEE6FECE" box="[326,352,261,284]" italics="true" pageId="175" pageNumber="176">ist</emphasis>
, distinctly proximad of lyrifissure
<emphasis id="B955F31CFF1055B9FD234065FD78FECF" box="[744,766,260,285]" italics="true" pageId="175" pageNumber="176">fb</emphasis>
, as opposed to level with it in
<taxonomicName id="4C21548DFF1055B9FB9C4064FB47FECE" box="[1111,1217,261,284]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="riopar" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FB9C4064FB47FECE" box="[1111,1217,261,284]" italics="true" pageId="175" pageNumber="176">O. riopar</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FB024065FA8EFECF" bold="true" box="[1225,1288,260,285]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FB024065FA8EFECF" box="[1225,1288,260,285]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C21548DFF1055B9FAC44064FAF8FECE" box="[1295,1406,261,284]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="mateui" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FAC44064FAF8FECE" box="[1295,1406,261,284]" italics="true" pageId="175" pageNumber="176">O. mateui</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FA4E4065FF3EFE92" bold="true" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FA4E4065FF3EFE92" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a more slender pedipalp than
<taxonomicName id="4C21548DFF1055B9FDF94049FD3AFE92" box="[562,700,295,320]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="175" pageNumber="176" phylum="Arthropoda" rank="species" species="hoerwegi" status="sp. nov.">
<emphasis id="B955F31CFF1055B9FDF94049FD3AFE92" box="[562,700,295,320]" italics="true" pageId="175" pageNumber="176">O. hoerwegi</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF1055B9FD094046FC86FE92" bold="true" box="[706,768,295,320]" pageId="175" pageNumber="176">
<taxonomicNameLabel id="A2664E67FF1055B9FD094046FC86FE92" box="[706,768,295,320]" pageId="175" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, e.g. femur (
<specimenCount id="9D27E487FF1055B9FC5A4049FC25FE92" box="[913,931,296,320]" pageId="175" pageNumber="176" type="female"></specimenCount>
) 6.97.1 times longer than broad, versus 6.1.
</paragraph>
</subSubSection>
</treatment>
</document>