752 lines
90 KiB
XML
752 lines
90 KiB
XML
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<mods:title id="87A0502299913478ADE8B34636E21E8F">Integrative revision of the Lygodactylus madagascariensis group reveals an unexpected diversity of little brown geckos in Madagascar’s rainforest</mods:title>
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sp.
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18—
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Gippner
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(2021)
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.
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<quantity id="C05A22ED1302BF13FC4AFC4BFBC141D4" box="[1013,1106,973,1000]" metricMagnitude="4" metricUnit="m" metricValue="1.5699999999999998" pageId="40" pageNumber="41" unit="km" value="15.7">15.7 km</quantity>
|
||
NNW of Bealanana
|
||
</locationDeviation>
|
||
(
|
||
<locationDeviation id="16AF962A1302BF13FAF4FC4BFE3F4638" location="Beandrarezona" pageId="40" pageNumber="41">
|
||
<quantity id="C05A22ED1302BF13FAF4FC4BFA0F41D4" box="[1355,1436,973,999]" metricMagnitude="3" metricUnit="m" metricValue="8.7" pageId="40" pageNumber="41" unit="km" value="8.7">8.7 km</quantity>
|
||
NNW of Beandrarezona
|
||
</locationDeviation>
|
||
), northern
|
||
<collectingCountry id="7FB5CF981302BF13FD9AFC77FD3D4638" box="[549,686,1009,1035]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||
, at geographical coordinates
|
||
<geoCoordinate id="6296E9CF1302BF13FC4AFC77FBE24638" box="[1013,1137,1009,1035]" degrees="14.41900" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-14.419">S14.41900</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6296E9CF1302BF13FBC3FC77FB6A4638" box="[1148,1273,1009,1035]" degrees="48.71883" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="48.71883">E48.71883</geoCoordinate>
|
||
,
|
||
<elevation id="8C8F683B1302BF13FABAFC77FA044638" box="[1285,1431,1009,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="40" pageNumber="41" unit="m" value="1364.0">
|
||
<quantity id="C05A22ED1302BF13FABAFC77FAC9463F" box="[1285,1370,1009,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="40" pageNumber="41" unit="m" value="1364.0">1364 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81302BF13FF05FB93FE1C4603" box="[186,399,1045,1072]" pageId="40" pageNumber="41" value="2015-12-22">
|
||
<collectingDate id="635850201302BF13FF05FB93FE1C4603" box="[186,399,1045,1072]" pageId="40" pageNumber="41" value="2015-12-22">22 December 2015</collectingDate>
|
||
</date>
|
||
(
|
||
<figureCitation id="9F99938D1302BF13FE20FB93FE614603" box="[415,498,1045,1072]" captionStart="FIGURE 15" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[244,1351,181,1906]" captionTargetId="figure-18@25.[236,1352,181,1919]" captionTargetPageId="25" captionText="FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study." figureDoi="http://doi.org/10.5281/zenodo.7040784" httpUri="https://zenodo.org/record/7040784/files/figure.png" pageId="40" pageNumber="41">Fig. 15</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="071D8F081302BF13FF78FBBFFEC447A4" blockId="40.[151,1437,937,2044]" pageId="40" pageNumber="41">
|
||
<materialsCitation id="B7CA85551302BF13FF78FBBFFB25464B" ID-GBIF-Occurrence="3897988310" collectingDate="2015-12-29" collectionCode="ZSM, MSZC" collectorName="M. D. Scherz & Scherz, J & Borrell, L & Starnes, E & Razafimandimby, D. H & J. Rabearivony" country="Madagascar" elevation="1315" latitude="-14.42843" location="Ampotsidy" longLatPrecision="1" longitude="48.72285" pageId="40" pageNumber="41" specimenCode="ZSM 48, MSZC 0110" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="35D6531A1302BF13FF78FBBFFED54660" bold="true" box="[199,326,1081,1107]" pageId="40" pageNumber="41">
|
||
<typeStatus id="D81931AA1302BF13FF78FBBFFED24660" box="[199,321,1081,1107]" pageId="40" pageNumber="41" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="570427731302BF13FEF2FBBFFE3B4660" box="[333,424,1081,1107]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="40" pageNumber="41" type="Museum">ZSM 48</specimenCode>
|
||
/2016 (
|
||
<specimenCode id="570427731302BF13FE48FBBFFD144660" box="[503,647,1081,1107]" collectionCode="MSZC" pageId="40" pageNumber="41">MSZC 0110</specimenCode>
|
||
), adult male, collected by same collectors and at same locality as holotype, at geographical coordinates
|
||
<geoCoordinate id="6296E9CF1302BF13FDAFFBDBFD184644" box="[528,651,1117,1143]" degrees="14.42843" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-14.42843">S14.42843</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6296E9CF1302BF13FD29FBDBFC80464B" box="[662,787,1117,1144]" degrees="48.72285" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="48.72285">E48.72285</geoCoordinate>
|
||
,
|
||
<elevation id="8C8F683B1302BF13FCA1FBDBFC3C4644" box="[798,943,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.315" pageId="40" pageNumber="41" unit="m" value="1315.0">
|
||
<quantity id="C05A22ED1302BF13FCA1FBDBFCE0464B" box="[798,883,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.315" pageId="40" pageNumber="41" unit="m" value="1315.0">1315 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81302BF13FC61FBDBFB25464B" box="[990,1206,1117,1144]" pageId="40" pageNumber="41" value="2015-12-29">
|
||
<collectingDate id="635850201302BF13FC61FBDBFB25464B" box="[990,1206,1117,1144]" pageId="40" pageNumber="41" value="2015-12-29">29 December 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7CA85551302BF13FB7DFBD8FE6F46D7" ID-GBIF-Occurrence="3897988329" collectingDate="2015-12-18" collectingDateMax="2015-12-22" collectingDateMin="2015-12-18" collectionCode="MSZC" collectorName="M. D. Scherz & Scherz, J & Borrell, L & Starnes, E & Razafimandimby, D. H & J. Rabearivony" country="Madagascar" elevation="1362" latitude="48.71149" location="Ampotsidy" longLatPrecision="1" longitude="-48.71916" pageId="40" pageNumber="41" specimenCode="UADBA-R 70856- 70859, MSZC 0011, 0019, 0023, 0077" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCode id="570427731302BF13FB7DFBD8FF4E46AF" collectionCode="UADBA-R" pageId="40" pageNumber="41">UADBA-R 70856– 70859</specimenCode>
|
||
(
|
||
<specimenCode id="570427731302BF13FF50FB07FE1246A8" box="[239,385,1153,1179]" collectionCode="MSZC" pageId="40" pageNumber="41">MSZC 0011</specimenCode>
|
||
,
|
||
<specimenCode id="570427731302BF13FE30FB07FE5946A8" box="[399,458,1153,1179]" collectionCode="MSZC" pageId="40" pageNumber="41">0019</specimenCode>
|
||
,
|
||
<specimenCode id="570427731302BF13FE67FB07FD8046A8" box="[472,531,1153,1179]" collectionCode="MSZC" pageId="40" pageNumber="41">0023</specimenCode>
|
||
,
|
||
<specimenCode id="570427731302BF13FD9EFB07FDCE46A8" box="[545,605,1153,1179]" collectionCode="MSZC" pageId="40" pageNumber="41">0077</specimenCode>
|
||
),
|
||
<specimenCount id="11A444811302BF13FDCCFB04FD7B46A8" box="[627,744,1153,1180]" pageId="40" pageNumber="41" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="11A444811302BF13FC9BFB04FC2146A8" box="[804,946,1153,1180]" pageId="40" pageNumber="41" type="female">two females</specimenCount>
|
||
, respectively, collected by same collectors and at same locality as holotype, between coordinates
|
||
<geoCoordinate id="6296E9CF1302BF13FCBDFB23FCE946F3" box="[770,890,1189,1216]" degrees="14.41455" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-14.41455">S14.41455</geoCoordinate>
|
||
–14.42317,
|
||
<geoCoordinate id="6296E9CF1302BF13FBBEFB23FBEF468C" box="[1025,1148,1189,1215]" degrees="48.71149" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="48.71149">E48.71149</geoCoordinate>
|
||
–48.71916,
|
||
<elevation id="8C8F683B1302BF13FABFFB23FF5E46D0" metricMagnitude="3" metricUnit="m" metricValue="1.3619999999999999" metricValueMax="1.4040000000000001" metricValueMin="1.3199999999999998" pageId="40" pageNumber="41" unit="m" value="1362.0" valueMax="1404.0" valueMin="1320.0">
|
||
<quantity id="C05A22ED1302BF13FABFFB23FA0F46F3" box="[1280,1436,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.3619999999999999" metricValueMax="1.4040000000000001" metricValueMin="1.3199999999999998" pageId="40" pageNumber="41" unit="m" value="1362.0" valueMax="1404.0" valueMin="1320.0">1320–1404 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81302BF13FF44FB4FFE6F46D7" box="[251,508,1225,1252]" pageId="40" pageNumber="41" value="2015-12-18" valueMax="2015-12-22" valueMin="2015-12-18">
|
||
<collectingDate id="635850201302BF13FF44FB4FFE6F46D7" box="[251,508,1225,1252]" pageId="40" pageNumber="41" value="2015-12-18" valueMax="2015-12-22" valueMin="2015-12-18">18–22 December 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7CA85551302BF13FDB8FB4FFB0F4718" ID-GBIF-Occurrence="3897988309" collectingDate="2010-06-27" collectionCode="ZSM, ZCMV" collectorName="Vieites" country="Madagascar" elevation="1466" latitude="-14.43061" longLatPrecision="1" longitude="48.60179" pageId="40" pageNumber="41" specimenCode="ZSM 1763, ZCMV 12502" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="570427731302BF13FDB8FB4FFDEE46D0" box="[519,637,1225,1251]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="40" pageNumber="41" type="Museum">ZSM 1763</specimenCode>
|
||
/2010 (
|
||
<specimenCode id="570427731302BF13FD72FB4FFCFC46D7" box="[717,879,1225,1252]" collectionCode="ZCMV" pageId="40" pageNumber="41">ZCMV 12502</specimenCode>
|
||
), by M. Vences, D.
|
||
<collectionCode id="61B317CD1302BF13FBFDFB4CFBCB46D7" box="[1090,1112,1226,1252]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="40" pageNumber="41">R</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FBDDFB4FFB2046D0" box="[1122,1203,1225,1251]" pageId="40" pageNumber="41">Vieites</collectorName>
|
||
,
|
||
<collectionCode id="61B317CD1302BF13FB02FB4CFB4046D7" box="[1213,1235,1226,1252]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="40" pageNumber="41">R</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FB68FB4CFA0B46D0" box="[1239,1432,1225,1252]" pageId="40" pageNumber="41">D. Randrianiaina</collectorName>
|
||
,
|
||
<collectorName id="AA57EADE1302BF13FF28FB68FEC24734" box="[151,337,1261,1288]" pageId="40" pageNumber="41">F.M. Ratsoavina</collectorName>
|
||
,
|
||
<collectorName id="AA57EADE1302BF13FEE3FB6BFE644734" box="[348,503,1261,1287]" pageId="40" pageNumber="41">S. Rasamison</collectorName>
|
||
, A.
|
||
<collectorName id="AA57EADE1302BF13FD9EFB6BFD2A4734" box="[545,697,1261,1287]" pageId="40" pageNumber="41">Rakotoarison</collectorName>
|
||
, E.
|
||
<collectorName id="AA57EADE1302BF13FD5CFB6BFCFC4734" box="[739,879,1261,1287]" pageId="40" pageNumber="41">Rajeriarison</collectorName>
|
||
, and
|
||
<collectionCode id="61B317CD1302BF13FC17FB68FC29473B" box="[936,954,1262,1288]" name="Tavera, Department of Geology and Geophysics" pageId="40" pageNumber="41">T</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FC7BFB6BFBCD4734" box="[964,1118,1261,1287]" pageId="40" pageNumber="41">Rajoafiarison</collectorName>
|
||
, at
|
||
<collectorName id="AA57EADE1302BF13FB3CFB6BFA814734" box="[1155,1298,1261,1287]" pageId="40" pageNumber="41">Bemanevika</collectorName>
|
||
,
|
||
<collectorName id="AA57EADE1302BF13FAA4FB6BFA0F4734" box="[1307,1436,1261,1287]" pageId="40" pageNumber="41">Antsirakala</collectorName>
|
||
campsite, geographical coordinates
|
||
<geoCoordinate id="6296E9CF1302BF13FD94FA97FD354718" box="[555,678,1297,1323]" degrees="14.43061" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="1" value="-14.43061">S14.43061</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6296E9CF1302BF13FD0CFA97FCA34718" box="[691,816,1297,1323]" degrees="48.60179" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="1" value="48.60179">E48.60179</geoCoordinate>
|
||
,
|
||
<elevation id="8C8F683B1302BF13FC83FA97FC5C4718" box="[828,975,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.466" pageId="40" pageNumber="41" unit="m" value="1466.0">
|
||
<quantity id="C05A22ED1302BF13FC83FA97FC01471F" box="[828,914,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.466" pageId="40" pageNumber="41" unit="m" value="1466.0">1466 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81302BF13FBBFFA97FB0F4718" box="[1024,1180,1297,1324]" pageId="40" pageNumber="41" value="2010-06-27">
|
||
<collectingDate id="635850201302BF13FBBFFA97FB0F4718" box="[1024,1180,1297,1324]" pageId="40" pageNumber="41" value="2010-06-27">27 June 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7CA85551302BF13FB18FA97FEC747A4" ID-GBIF-Occurrence="3897988304" collectingDate="2010-06-21" collectionCode="ZSM, DRV" collectorName="F. M. Ratsoavina" country="Madagascar" elevation="1717" latitude="-14.3464" longLatPrecision="7" longitude="49.1028" pageId="40" pageNumber="41" specimenCode="ZSM 555, DRV 6288" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="570427731302BF13FB18FA97FA81471F" box="[1191,1298,1297,1324]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="40" pageNumber="41" type="Museum">ZSM 555</specimenCode>
|
||
/2014 (
|
||
<specimenCode id="570427731302BF13FADDFA97FF40477C" collectionCode="DRV" pageId="40" pageNumber="41">DRV 6288</specimenCode>
|
||
), collected by
|
||
<collectorName id="AA57EADE1302BF13FECAFAB0FDBC477C" box="[373,559,1333,1360]" pageId="40" pageNumber="41">F.M. Ratsoavina</collectorName>
|
||
, D.
|
||
<collectionCode id="61B317CD1302BF13FDEAFAB0FDF84763" box="[597,619,1334,1360]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="40" pageNumber="41">R</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FDCAFAB3FD7F4763" box="[629,748,1333,1360]" pageId="40" pageNumber="41">Vieites, M</collectorName>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FD49FAB0FCDB4763" box="[758,840,1334,1360]" pageId="40" pageNumber="41">Vences</collectorName>
|
||
,
|
||
<collectionCode id="61B317CD1302BF13FCECFAB0FCFA4763" box="[851,873,1334,1360]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="40" pageNumber="41">R</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FCD2FAB0FBBC477C" box="[877,1071,1333,1360]" pageId="40" pageNumber="41">D. Randrianiaina</collectorName>
|
||
,
|
||
<collectorName id="AA57EADE1302BF13FB85FAB3FB46477C" box="[1082,1237,1333,1359]" pageId="40" pageNumber="41">S. Rasamison</collectorName>
|
||
,
|
||
<collectorName id="AA57EADE1302BF13FB61FAB0FA0B477C" box="[1246,1432,1333,1360]" pageId="40" pageNumber="41">A. Rakotoarison</collectorName>
|
||
, E.
|
||
<collectorName id="AA57EADE1302BF13FF09FADFFED14740" box="[182,322,1369,1395]" pageId="40" pageNumber="41">Rajeriarison</collectorName>
|
||
, and
|
||
<collectionCode id="61B317CD1302BF13FEC3FADCFE1D4747" box="[380,398,1370,1396]" name="Tavera, Department of Geology and Geophysics" pageId="40" pageNumber="41">T</collectionCode>
|
||
.
|
||
<collectorName id="AA57EADE1302BF13FE26FADFFDA34740" box="[409,560,1369,1395]" pageId="40" pageNumber="41">Rajoafiarison</collectorName>
|
||
at
|
||
<collectorName id="AA57EADE1302BF13FDEEFADFFD7B4740" box="[593,744,1369,1395]" pageId="40" pageNumber="41">Andrevorevo</collectorName>
|
||
, geographical coordinates
|
||
<geoCoordinate id="6296E9CF1302BF13FBADFADFFB134740" box="[1042,1152,1369,1395]" degrees="14.3464" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="5" value="-14.3464">S14.3464</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6296E9CF1302BF13FB34FADFFB694740" box="[1163,1274,1369,1395]" degrees="49.1028" direction="east" orientation="longitude" pageId="40" pageNumber="41" precision="5" value="49.1028">E49.1028</geoCoordinate>
|
||
,
|
||
<elevation id="8C8F683B1302BF13FABAFADFFA044740" box="[1285,1431,1369,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="40" pageNumber="41" unit="m" value="1717.0">
|
||
<quantity id="C05A22ED1302BF13FABAFADFFAC94747" box="[1285,1370,1369,1396]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="40" pageNumber="41" unit="m" value="1717.0">1717 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, on
|
||
<date id="731CA9C81302BF13FF05FAFBFEC747A4" box="[186,340,1405,1432]" pageId="40" pageNumber="41" value="2010-06-21">
|
||
<collectingDate id="635850201302BF13FF05FAFBFEC747A4" box="[186,340,1405,1432]" pageId="40" pageNumber="41" value="2010-06-21">21 June 2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831302BF12FF78FA27FF5446CD" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
|
||
<paragraph id="071D8F081302BF13FF78FA27FD6945E4" blockId="40.[151,1437,937,2044]" pageId="40" pageNumber="41">
|
||
<emphasis id="35D6531A1302BF13FF78FA27FED14788" bold="true" box="[199,322,1441,1467]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
|
||
<taxonomicName id="C0A2F48B1302BF13FEF5FA27FDB74788" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[330,548,1441,1467]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="winki" status="sp. nov.">
|
||
<emphasis id="35D6531A1302BF13FEF5FA27FDB74788" box="[330,548,1441,1467]" italics="true" pageId="40" pageNumber="41">Lygodactylus winki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1302BF13FD92FA27FD1B4788" bold="true" box="[557,648,1441,1467]" pageId="40" pageNumber="41">
|
||
<taxonomicNameLabel id="2EE5EE611302BF13FD92FA27FD1B4788" box="[557,648,1441,1467]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
corresponds to a lineage forming part of the subclade A5 of
|
||
<taxonomicName id="C0A2F48B1302BF13FAF9FA24FF4E47EC" authorityName="Pasteur" authorityYear="1964" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1302BF13FAF9FA24FF4E47EC" italics="true" pageId="40" pageNumber="41">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
, and is one of only two representatives of this subclade known to reach northern
|
||
<collectingCountry id="7FB5CF981302BF13FB36FA43FA8147EC" box="[1161,1298,1477,1503]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||
. The lowest genetic divergences of the lineages are 8.7% uncorrected 16S distance to
|
||
<taxonomicName id="C0A2F48B1302BF13FC56FA6CFBDC4430" authorityName=": Rosler" authorityYear="2000" box="[1001,1103,1513,1539]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1302BF13FC56FA6CFBDC4430" box="[1001,1103,1513,1539]" italics="true" pageId="40" pageNumber="41">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
and 10.3% to
|
||
<emphasis id="35D6531A1302BF13FB44FA6CFAF54430" box="[1275,1382,1514,1539]" italics="true" pageId="40" pageNumber="41">
|
||
<taxonomicName id="C0A2F48B1302BF13FB44FA6CFAF14430" authorityName="Gunther" authorityYear="1891" box="[1275,1378,1514,1539]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="miops">L. miops</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It is characterized by the presence of very distinct lateral spine-like scales at the base of the tail in males, as are found in several representatives of subclade A5 but not in other
|
||
<taxonomicName id="C0A2F48B1302BF13FD49F9B7FC1A4478" authorityName="Pasteur" authorityYear="1964" box="[758,905,1585,1611]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1302BF13FD49F9B7FC1A4478" box="[758,905,1585,1611]" italics="true" pageId="40" pageNumber="41">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
. It can also be assigned to the subgenus
|
||
<taxonomicName id="C0A2F48B1302BF13FAF9F9B4FF4E445C" authorityName="Pasteur" authorityYear="1964" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1302BF13FAF9F9B4FF4E445C" italics="true" pageId="40" pageNumber="41">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
by an undivided mental scale with two postmentals, absence of a claw on the first finger, and 7 preanal pores in males. Within
|
||
<taxonomicName id="C0A2F48B1302BF13FEE2F9FFFE6344A0" authorityName="Pasteur" authorityYear="1964" box="[349,496,1657,1683]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1302BF13FEE2F9FFFE6344A0" box="[349,496,1657,1683]" italics="true" pageId="40" pageNumber="41">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
, the new species is one of only two species of subclade A5 known from northern
|
||
<collectingCountry id="7FB5CF981302BF13FF28F91BFEB34484" box="[151,288,1693,1719]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
|
||
. It differs from the other nominal species of
|
||
<taxonomicName id="C0A2F48B1302BF13FCB6F91BFC0F4484" authorityName="Pasteur" authorityYear="1964" box="[777,924,1693,1719]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
|
||
<emphasis id="35D6531A1302BF13FCB6F91BFC0F4484" box="[777,924,1693,1719]" italics="true" pageId="40" pageNumber="41">Domerguella</emphasis>
|
||
</taxonomicName>
|
||
occurring in the same general area as follows: from
|
||
<taxonomicName id="C0A2F48B1302BF13FF6BF944FEFB44E8" authorityName=": Rosler" authorityYear="2000" box="[212,360,1730,1755]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="expectatus">
|
||
<emphasis id="35D6531A1302BF13FF6BF944FEFB44E8" box="[212,360,1730,1755]" italics="true" pageId="40" pageNumber="41">L. expectatus</emphasis>
|
||
</taxonomicName>
|
||
by non-enlarged dorsolateral scales (longitudinal count of dorsal scales>185 vs. <170); from
|
||
<taxonomicName id="C0A2F48B1302BF13FA39F944FF4744CC" authorityName=": Rosler" authorityYear="2000" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="rarus">
|
||
<emphasis id="35D6531A1302BF13FA39F944FF4744CC" italics="true" pageId="40" pageNumber="41">L. rarus</emphasis>
|
||
</taxonomicName>
|
||
by lack of regular crossbands on tail (vs. presence) and different body shape with less elongated limbs (relative hindlimb length 0.49–0.54 vs.>0.55); from
|
||
<emphasis id="35D6531A1302BF13FD11F88CFB9C4510" box="[686,1039,1801,1827]" italics="true" pageId="40" pageNumber="41">
|
||
<taxonomicName id="C0A2F48B1302BF13FD11F88CFC0B4510" authorityName=": Pasteur" authorityYear="1965" box="[686,920,1801,1827]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="madagascariensis">L. madagascariensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="C0A2F48B1302BF13FC1DF88CFB984510" authorityName="Pasteur & Blanc" authorityYear="1967" box="[930,1035,1802,1827]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="petteri">L. petteri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="C0A2F48B1302BF13FBFCF88CFB064510" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1091,1173,1801,1827]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="salvi">
|
||
<emphasis id="35D6531A1302BF13FBFCF88CFB064510" box="[1091,1173,1801,1827]" italics="true" pageId="40" pageNumber="41">L. salvi</emphasis>
|
||
</taxonomicName>
|
||
by a lower longitudinal count of ventral scales (83–98 vs.>100); from
|
||
<taxonomicName id="C0A2F48B1302BF13FD05F8A8FCBB4574" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[698,808,1837,1863]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="roellae">
|
||
<emphasis id="35D6531A1302BF13FD05F8A8FCBB4574" box="[698,808,1837,1863]" italics="true" pageId="40" pageNumber="41">L. roellae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="35D6531A1302BF13FCDCF8A8FC5B4574" box="[867,968,1837,1863]" italics="true" pageId="40" pageNumber="41">
|
||
<taxonomicName id="C0A2F48B1302BF13FCDCF8A8FC574574" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[867,964,1837,1863]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="hapei">L. hapei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
by a higher longitudinal count of dorsal scales (187–222 vs. 159–179); and from
|
||
<taxonomicName id="C0A2F48B1302BF13FDD3F8D4FD7F4558" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[620,748,1873,1899]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="tantsaha">
|
||
<emphasis id="35D6531A1302BF13FDD3F8D4FD7F4558" box="[620,748,1873,1899]" italics="true" pageId="40" pageNumber="41">L. tantsaha</emphasis>
|
||
</taxonomicName>
|
||
by a lower longitudinal count of dorsal scales (187–222 vs. 239–240). Furthermore, the new species differs from all of these species of the subclades A1–A4 by the presence of a distinct lateral spine at the base of the tail, especially large in males but also clearly recognizable in females (vs. more weakly expressed or absent in the other species).
|
||
</paragraph>
|
||
<paragraph id="071D8F081302BF12FF78F867FAB043FF" blockId="40.[151,1437,937,2044]" lastBlockId="41.[151,1437,151,461]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
|
||
From the other two nominal species in subclade A5 (
|
||
<taxonomicName id="C0A2F48B1302BF13FC99F864FC1845C8" authorityName=": Rosler" authorityYear="2000" box="[806,907,2017,2043]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1302BF13FC99F864FC1845C8" box="[806,907,2017,2043]" italics="true" pageId="40" pageNumber="41">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0A2F48B1302BF13FC7AF864FBB445C8" authorityName="Gunther" authorityYear="1891" box="[965,1063,2018,2043]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="miops">
|
||
<emphasis id="35D6531A1302BF13FC7AF864FBB445C8" box="[965,1063,2018,2043]" italics="true" pageId="40" pageNumber="41">L. miops</emphasis>
|
||
</taxonomicName>
|
||
, according to current taxonomy; see above) the new species differs as follows: from
|
||
<taxonomicName id="C0A2F48B1303BF12FD51FF11FCC34283" authorityName="Gunther" authorityYear="1891" box="[750,848,151,176]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="miops">
|
||
<emphasis id="35D6531A1303BF12FD51FF11FCC34283" box="[750,848,151,176]" italics="true" pageId="41" pageNumber="42">L. miops</emphasis>
|
||
</taxonomicName>
|
||
by a lower longitudinal count of ventral scales despite minimal overlap (83–98 vs. 98–113); and from
|
||
<taxonomicName id="C0A2F48B1303BF12FD57FF3DFCD842E7" authorityName=": Rosler" authorityYear="2000" box="[744,843,186,212]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1303BF12FD57FF3DFCD842E7" box="[744,843,186,212]" italics="true" pageId="41" pageNumber="42">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
by apparently relatively longer hindlimbs (HIL/SVL 0.49–0.54 vs. 0.42–0.49). Further comparative examination of specimens also revealed a different head shape in
|
||
<taxonomicName id="C0A2F48B1303BF12FF28FE84FF674328" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[151,244,257,283]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1303BF12FF28FE84FF674328" box="[151,244,257,283]" italics="true" pageId="41" pageNumber="42">L. winki</emphasis>
|
||
</taxonomicName>
|
||
compared to
|
||
<taxonomicName id="C0A2F48B1303BF12FE2DFE84FE644328" authorityName=": Rosler" authorityYear="2000" box="[402,503,257,283]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1303BF12FE2DFE84FE644328" box="[402,503,257,283]" italics="true" pageId="41" pageNumber="42">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
, with apparently more expressed supraocular bulges (also visible in specimens in life;
|
||
<figureCitation id="9F99938D1303BF12FF73FEA2FE8F430C" box="[204,284,292,319]" captionStart="FIGURE 14" captionStartId="23.[151,250,1683,1707]" captionTargetBox="[151,1437,181,1659]" captionTargetId="figure-18@23.[151,1437,181,1659]" captionTargetPageId="23" captionText="FIGURE 14. Photos of specimens of L. guibei in life. A, female specimen ZSM 525/2009 (ZCMV 11210) from the western slope of the Makira Massif, photographed on 23 June 2009. B–C, male and D–E, female, from Andasibe, in dorsolateral and ventral views, photographed 2003, not attributable to a voucher specimen and not genotyped, and thus only tentatively attributable to the species." figureDoi="http://doi.org/10.5281/zenodo.7040777" httpUri="https://zenodo.org/record/7040777/files/figure.png" pageId="41" pageNumber="42">Fig. 14</figureCitation>
|
||
vs.
|
||
<figureCitation id="9F99938D1303BF12FEF4FEA2FE0C430C" box="[331,415,292,319]" captionStart="FIGURE 20" captionStartId="42.[151,250,1872,1896]" captionTargetBox="[228,1359,181,1848]" captionTargetId="figure-18@42.[228,1359,181,1848]" captionTargetPageId="42" captionText="FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika." figureDoi="http://doi.org/10.5281/zenodo.7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="41" pageNumber="42">Fig. 20</figureCitation>
|
||
). Additional measurements taken on selected specimens in good, fully comparable state of preservation (
|
||
<tableCitation id="4A20BAB31303BF12FE90FECEFE114351" box="[303,386,328,354]" captionStart="TABLE 3" captionStartId="41.[151,239,508,532]" captionTargetPageId="41" captionText="TABLE 3. Measurements of head proportions (in mm) in selected specimens of Lygodactylus guibei and L. winki sp. nov. to illustrate relatively shallower and shorter heads in L. guibei. Only specimens in good state of preservation were included. STD, snout-tympanum distance; HH, head height; relSTD, relHH are relative values (in percent) given as ratio to SVL. For additional measurements and metadata of specimens, see Table 1." httpUri="http://table.plazi.org/id/53DDDF801303BF12FF28FE7AFC4940B3" pageId="41" pageNumber="42" tableUuid="53DDDF801303BF12FF28FE7AFC4940B3">Table 3</tableCitation>
|
||
) revealed that
|
||
<taxonomicName id="C0A2F48B1303BF12FD99FECFFD134351" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[550,640,328,354]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1303BF12FD99FECFFD134351" box="[550,640,328,354]" italics="true" pageId="41" pageNumber="42">L. winki</emphasis>
|
||
</taxonomicName>
|
||
individuals have proportionally longer and higher heads than
|
||
<emphasis id="35D6531A1303BF12FA8CFECFFA0F4351" box="[1331,1436,328,354]" italics="true" pageId="41" pageNumber="42">
|
||
<taxonomicName id="C0A2F48B1303BF12FA8CFECFFA0B4351" authorityName=": Rosler" authorityYear="2000" box="[1331,1432,328,354]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">L. guibei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with non-overlapping values (relative snout tip to tympanum distance in percent, 24.7–28.8 vs. 23.0–24.6; relative head height in percent, 12.5–14.2 vs. 11.1–12.4; see
|
||
<tableCitation id="4A20BAB31303BF12FD5FFE09FCA6439A" box="[736,821,399,425]" captionStart="TABLE 3" captionStartId="41.[151,239,508,532]" captionTargetPageId="41" captionText="TABLE 3. Measurements of head proportions (in mm) in selected specimens of Lygodactylus guibei and L. winki sp. nov. to illustrate relatively shallower and shorter heads in L. guibei. Only specimens in good state of preservation were included. STD, snout-tympanum distance; HH, head height; relSTD, relHH are relative values (in percent) given as ratio to SVL. For additional measurements and metadata of specimens, see Table 1." httpUri="http://table.plazi.org/id/53DDDF801303BF12FF28FE7AFC4940B3" pageId="41" pageNumber="42" tableUuid="53DDDF801303BF12FF28FE7AFC4940B3">Table 3</tableCitation>
|
||
).
|
||
<taxonomicName id="C0A2F48B1303BF12FCF5FE16FC37439A" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[842,932,399,425]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="winki" status="sp. nov.">
|
||
<emphasis id="35D6531A1303BF12FCF5FE16FC37439A" box="[842,932,399,425]" italics="true" pageId="41" pageNumber="42">L. winki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1303BF12FC15FE09FB97439A" bold="true" box="[938,1028,399,425]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="2EE5EE611303BF12FC15FE09FB97439A" box="[938,1028,399,425]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
does not share haplotypes in CMOS or RAG1 with
|
||
<taxonomicName id="C0A2F48B1303BF12FE81FE35FE0C43FF" authorityName="Gunther" authorityYear="1891" box="[318,415,435,460]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="miops">
|
||
<emphasis id="35D6531A1303BF12FE81FE35FE0C43FF" box="[318,415,435,460]" italics="true" pageId="41" pageNumber="42">L. miops</emphasis>
|
||
</taxonomicName>
|
||
, and only one instance of haplotype sharing in RAG1 is detected with
|
||
<emphasis id="35D6531A1303BF12FB07FE35FAB043FF" box="[1208,1315,434,460]" italics="true" pageId="41" pageNumber="42">
|
||
<taxonomicName id="C0A2F48B1303BF12FB07FE35FAB343FF" authorityName=": Rosler" authorityYear="2000" box="[1208,1312,434,460]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">L. guibei</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="53DDDF801303BF12FF28FE7AFC4940B3" ID-Table-UUID="53DDDF801303BF12FF28FE7AFC4940B3" httpUri="http://table.plazi.org/id/53DDDF801303BF12FF28FE7AFC4940B3" pageId="41" pageNumber="42" startId="41.[151,239,508,532]" targetBox="[160,1404,653,1170]" targetIsTable="true" targetPageId="41">
|
||
<paragraph id="071D8F081303BF12FF28FE7AFC4940B3" blockId="41.[151,1436,508,640]" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FF28FE7AFE964027" bold="true" box="[151,261,508,532]" pageId="41" pageNumber="42">TABLE 3</emphasis>
|
||
. Measurements of head proportions (in mm) in selected specimens of
|
||
<taxonomicName id="C0A2F48B1303BF12FBB4FE7AFB774027" authorityName=": Rosler" authorityYear="2000" box="[1035,1252,508,532]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1303BF12FBB4FE7AFB774027" box="[1035,1252,508,532]" italics="true" pageId="41" pageNumber="42">Lygodactylus guibei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C0A2F48B1303BF12FAA3FE7BFAE74027" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[1308,1396,508,532]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="winki" status="sp. nov.">
|
||
<emphasis id="35D6531A1303BF12FAA3FE7BFAE74027" box="[1308,1396,508,532]" italics="true" pageId="41" pageNumber="42">L. winki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1303BF12FAC3FE7AFF55400B" bold="true" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="2EE5EE611303BF12FAC3FE7AFF55400B" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
to illustrate relatively shallower and shorter heads in
|
||
<taxonomicName id="C0A2F48B1303BF12FCB4FDA7FCF8400B" authorityName=": Rosler" authorityYear="2000" box="[779,875,544,568]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="guibei">
|
||
<emphasis id="35D6531A1303BF12FCB4FDA7FCF8400B" box="[779,875,544,568]" italics="true" pageId="41" pageNumber="42">L. guibei</emphasis>
|
||
</taxonomicName>
|
||
. Only specimens in good state of preservation were included. STD, snout-tympanum distance; HH, head height; relSTD, relHH are relative values (in percent) given as ratio to SVL. For additional measurements and metadata of specimens, see Table 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="071D8F081303BF12FF1FFD08FAA146A1" pageId="41" pageNumber="42">
|
||
<table id="75A27DA8130340C4FF1FFD0BFAEF46A1" box="[160,1404,653,1170]" gridcols="8" gridrows="14" pageId="41" pageNumber="42">
|
||
<tr id="B9928D4A130340C4FF1FFD0BFAEF4095" box="[160,1404,653,678]" gridrow="0" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFD0BFEEF4095" box="[160,380,653,678]" gridcol="0" gridrow="0" pageId="41" pageNumber="42">Specimen voucher</th>
|
||
<th id="FA43E436130340C4FE6AFD0BFDF04095" box="[469,611,653,678]" gridcol="1" gridrow="0" pageId="41" pageNumber="42">Field number</th>
|
||
<th id="FA43E436130340C4FD05FD0BFD6C4095" box="[698,767,653,678]" gridcol="2" gridrow="0" pageId="41" pageNumber="42">Sex</th>
|
||
<th id="FA43E436130340C4FC85FD0BFCF94095" box="[826,874,653,678]" gridcol="3" gridrow="0" pageId="41" pageNumber="42">SVL</th>
|
||
<th id="FA43E436130340C4FC1FFD0BFC434095" box="[928,976,653,678]" gridcol="4" gridrow="0" pageId="41" pageNumber="42">STD</th>
|
||
<th id="FA43E436130340C4FBB8FD0BFBBF4095" box="[1031,1068,653,678]" gridcol="5" gridrow="0" pageId="41" pageNumber="42">HH</th>
|
||
<th id="FA43E436130340C4FBDEFD0BFB734095" box="[1121,1248,653,678]" gridcol="6" gridrow="0" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FBDEFD0BFB734096" bold="true" box="[1121,1248,653,677]" pageId="41" pageNumber="42">relSTD (%)</emphasis>
|
||
</th>
|
||
<th id="FA43E436130340C4FAB9FD0BFAEF4095" box="[1286,1404,653,678]" gridcol="7" gridrow="0" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FAB9FD08FAEF4096" bold="true" box="[1286,1404,653,678]" pageId="41" pageNumber="42">relHH (%)</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFD32FAEF40FF" box="[160,1404,692,716]" gridrow="1" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFD32FAEF40FF" box="[160,1404,692,716]" colspan="8" colspanRight="7" gridcol="0" gridrow="1" pageId="41" pageNumber="42">
|
||
<taxonomicName id="C0A2F48B1303BF12FF1FFD33FF6140FF" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[160,242,692,716]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="winki" status="sp. nov.">
|
||
<emphasis id="35D6531A1303BF12FF1FFD33FF6140FF" box="[160,242,692,716]" italics="true" pageId="41" pageNumber="42">L. winki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1303BF12FF47FD32FEDA40FF" bold="true" box="[248,329,692,716]" pageId="41" pageNumber="42">
|
||
<taxonomicNameLabel id="2EE5EE611303BF12FF47FD32FEDA40FF" box="[248,329,692,716]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(sp. 18)
|
||
</th>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFD5CFAEF40C1" box="[160,1404,730,754]" gridrow="2" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFD5CFEEF40C1" box="[160,380,730,754]" gridcol="0" gridrow="2" pageId="41" pageNumber="42">ZSM 47/2016 (HT)</th>
|
||
<td id="FA43E436130340C4FE6AFD5CFDF040C1" box="[469,611,730,754]" gridcol="1" gridrow="2" pageId="41" pageNumber="42">MSZC 0075</td>
|
||
<td id="FA43E436130340C4FD05FD5CFD6C40C1" box="[698,767,730,754]" gridcol="2" gridrow="2" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FD5CFCF940C1" box="[826,874,730,754]" gridcol="3" gridrow="2" pageId="41" pageNumber="42">29.5</td>
|
||
<td id="FA43E436130340C4FC1FFD5CFC4340C1" box="[928,976,730,754]" gridcol="4" gridrow="2" pageId="41" pageNumber="42">7.3</td>
|
||
<td id="FA43E436130340C4FBB8FD5CFBBF40C1" box="[1031,1068,730,754]" gridcol="5" gridrow="2" pageId="41" pageNumber="42">3.7</td>
|
||
<td id="FA43E436130340C4FBDEFD5CFB7340C1" box="[1121,1248,730,754]" gridcol="6" gridrow="2" pageId="41" pageNumber="42">24.7</td>
|
||
<td id="FA43E436130340C4FAB9FD5CFAEF40C1" box="[1286,1404,730,754]" gridcol="7" gridrow="2" pageId="41" pageNumber="42">12.5</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFD79FAEF412B" box="[160,1404,767,792]" gridrow="3" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFD79FEEF412B" box="[160,380,767,792]" gridcol="0" gridrow="3" pageId="41" pageNumber="42">ZSM 48/2016 (PT)</th>
|
||
<td id="FA43E436130340C4FE6AFD79FDF0412B" box="[469,611,767,792]" gridcol="1" gridrow="3" pageId="41" pageNumber="42">MSZC 0110</td>
|
||
<td id="FA43E436130340C4FD05FD79FD6C412B" box="[698,767,767,792]" gridcol="2" gridrow="3" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FD79FCF9412B" box="[826,874,767,792]" gridcol="3" gridrow="3" pageId="41" pageNumber="42">29.8</td>
|
||
<td id="FA43E436130340C4FC1FFD79FC43412B" box="[928,976,767,792]" gridcol="4" gridrow="3" pageId="41" pageNumber="42">7.6</td>
|
||
<td id="FA43E436130340C4FBB8FD79FBBF412B" box="[1031,1068,767,792]" gridcol="5" gridrow="3" pageId="41" pageNumber="42">4.0</td>
|
||
<td id="FA43E436130340C4FBDEFD79FB73412B" box="[1121,1248,767,792]" gridcol="6" gridrow="3" pageId="41" pageNumber="42">25.5</td>
|
||
<td id="FA43E436130340C4FAB9FD79FAEF412B" box="[1286,1404,767,792]" gridcol="7" gridrow="3" pageId="41" pageNumber="42">13.4</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFCA3FAEF410E" box="[160,1404,805,829]" gridrow="4" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFCA3FEEF410E" box="[160,380,805,829]" gridcol="0" gridrow="4" pageId="41" pageNumber="42">ZSM 555/2014 (PT)</th>
|
||
<td id="FA43E436130340C4FE6AFCA3FDF0410E" box="[469,611,805,829]" gridcol="1" gridrow="4" pageId="41" pageNumber="42">DRV 6288</td>
|
||
<td id="FA43E436130340C4FD05FCA3FD6C410E" box="[698,767,805,829]" gridcol="2" gridrow="4" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FCA3FCF9410E" box="[826,874,805,829]" gridcol="3" gridrow="4" pageId="41" pageNumber="42">33.0</td>
|
||
<td id="FA43E436130340C4FC1FFCA3FC43410E" box="[928,976,805,829]" gridcol="4" gridrow="4" pageId="41" pageNumber="42">9.5</td>
|
||
<td id="FA43E436130340C4FBB8FCA3FBBF410E" box="[1031,1068,805,829]" gridcol="5" gridrow="4" pageId="41" pageNumber="42">4.7</td>
|
||
<td id="FA43E436130340C4FBDEFCA3FB73410E" box="[1121,1248,805,829]" gridcol="6" gridrow="4" pageId="41" pageNumber="42">28.8</td>
|
||
<td id="FA43E436130340C4FAB9FCA3FAEF410E" box="[1286,1404,805,829]" gridcol="7" gridrow="4" pageId="41" pageNumber="42">14.2</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFCCDFAEF4150" box="[160,1404,843,867]" gridrow="5" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFCCDFEEF4150" box="[160,380,843,867]" gridcol="0" gridrow="5" pageId="41" pageNumber="42">ZSM 1763/2010 (PT)</th>
|
||
<td id="FA43E436130340C4FE6AFCCDFDF04150" box="[469,611,843,867]" gridcol="1" gridrow="5" pageId="41" pageNumber="42">ZCMV 12502</td>
|
||
<td id="FA43E436130340C4FD05FCCDFD6C4150" box="[698,767,843,867]" gridcol="2" gridrow="5" pageId="41" pageNumber="42">female</td>
|
||
<td id="FA43E436130340C4FC85FCCDFCF94150" box="[826,874,843,867]" gridcol="3" gridrow="5" pageId="41" pageNumber="42">33.4</td>
|
||
<td id="FA43E436130340C4FC1FFCCDFC434150" box="[928,976,843,867]" gridcol="4" gridrow="5" pageId="41" pageNumber="42">9.3</td>
|
||
<td id="FA43E436130340C4FBB8FCCDFBBF4150" box="[1031,1068,843,867]" gridcol="5" gridrow="5" pageId="41" pageNumber="42">4.5</td>
|
||
<td id="FA43E436130340C4FBDEFCCDFB734150" box="[1121,1248,843,867]" gridcol="6" gridrow="5" pageId="41" pageNumber="42">27.8</td>
|
||
<td id="FA43E436130340C4FAB9FCCDFAEF4150" box="[1286,1404,843,867]" gridcol="7" gridrow="5" pageId="41" pageNumber="42">13.5</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFCF7FAEF41BA" box="[160,1404,881,905]" gridrow="6" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFCF7FAEF41BA" box="[160,1404,881,905]" colspan="8" colspanRight="7" gridcol="0" gridrow="6" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FF1FFCF4FF6941BA" box="[160,250,881,905]" italics="true" pageId="41" pageNumber="42">L. guibei</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFC11FAEF419C" box="[160,1404,919,943]" gridrow="7" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFC11FEEF419C" box="[160,380,919,943]" gridcol="0" gridrow="7" pageId="41" pageNumber="42">ZSM 320/2004</th>
|
||
<td id="FA43E436130340C4FE6AFC11FDF0419C" box="[469,611,919,943]" gridcol="1" gridrow="7" pageId="41" pageNumber="42">FGZC 616</td>
|
||
<td id="FA43E436130340C4FD05FC11FD6C419C" box="[698,767,919,943]" gridcol="2" gridrow="7" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FC11FCF9419C" box="[826,874,919,943]" gridcol="3" gridrow="7" pageId="41" pageNumber="42">34.9</td>
|
||
<td id="FA43E436130340C4FC1FFC11FC43419C" box="[928,976,919,943]" gridcol="4" gridrow="7" pageId="41" pageNumber="42">8.6</td>
|
||
<td id="FA43E436130340C4FBB8FC11FBBF419C" box="[1031,1068,919,943]" gridcol="5" gridrow="7" pageId="41" pageNumber="42">4</td>
|
||
<td id="FA43E436130340C4FBDEFC11FB73419C" box="[1121,1248,919,943]" gridcol="6" gridrow="7" pageId="41" pageNumber="42">24.6</td>
|
||
<td id="FA43E436130340C4FAB9FC11FAEF419C" box="[1286,1404,919,943]" gridcol="7" gridrow="7" pageId="41" pageNumber="42">11.5</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFC3AFAEF41E6" box="[160,1404,956,981]" gridrow="8" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFC3AFEEF41E6" box="[160,380,956,981]" gridcol="0" gridrow="8" pageId="41" pageNumber="42">ZSM 321/2004</th>
|
||
<td id="FA43E436130340C4FE6AFC3AFDF041E6" box="[469,611,956,981]" gridcol="1" gridrow="8" pageId="41" pageNumber="42">FGZC 617</td>
|
||
<td id="FA43E436130340C4FD05FC3AFD6C41E6" box="[698,767,956,981]" gridcol="2" gridrow="8" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FC3AFCF941E6" box="[826,874,956,981]" gridcol="3" gridrow="8" pageId="41" pageNumber="42">33.2</td>
|
||
<td id="FA43E436130340C4FC1FFC3AFC4341E6" box="[928,976,956,981]" gridcol="4" gridrow="8" pageId="41" pageNumber="42">8.1</td>
|
||
<td id="FA43E436130340C4FBB8FC3AFBBF41E6" box="[1031,1068,956,981]" gridcol="5" gridrow="8" pageId="41" pageNumber="42">3.7</td>
|
||
<td id="FA43E436130340C4FBDEFC3AFB7341E6" box="[1121,1248,956,981]" gridcol="6" gridrow="8" pageId="41" pageNumber="42">24.4</td>
|
||
<td id="FA43E436130340C4FAB9FC3AFAEF41E6" box="[1286,1404,956,981]" gridcol="7" gridrow="8" pageId="41" pageNumber="42">11.1</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFC64FAEF41C9" box="[160,1404,994,1018]" gridrow="9" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFC64FEEF41C9" box="[160,380,994,1018]" gridcol="0" gridrow="9" pageId="41" pageNumber="42">ZSM 349/2010</th>
|
||
<td id="FA43E436130340C4FE6AFC64FDF041C9" box="[469,611,994,1018]" gridcol="1" gridrow="9" pageId="41" pageNumber="42">FGZC 4362</td>
|
||
<td id="FA43E436130340C4FD05FC64FD6C41C9" box="[698,767,994,1018]" gridcol="2" gridrow="9" pageId="41" pageNumber="42">male</td>
|
||
<td id="FA43E436130340C4FC85FC64FCF941C9" box="[826,874,994,1018]" gridcol="3" gridrow="9" pageId="41" pageNumber="42">37.6</td>
|
||
<td id="FA43E436130340C4FC1FFC64FC4341C9" box="[928,976,994,1018]" gridcol="4" gridrow="9" pageId="41" pageNumber="42">9.0</td>
|
||
<td id="FA43E436130340C4FBB8FC64FBBF41C9" box="[1031,1068,994,1018]" gridcol="5" gridrow="9" pageId="41" pageNumber="42">4.5</td>
|
||
<td id="FA43E436130340C4FBDEFC64FB7341C9" box="[1121,1248,994,1018]" gridcol="6" gridrow="9" pageId="41" pageNumber="42">23.9</td>
|
||
<td id="FA43E436130340C4FAB9FC64FAEF41C9" box="[1286,1404,994,1018]" gridcol="7" gridrow="9" pageId="41" pageNumber="42">12.0</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFB8EFAEF4613" box="[160,1404,1032,1056]" gridrow="10" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFB8EFEEF4613" box="[160,380,1032,1056]" gridcol="0" gridrow="10" pageId="41" pageNumber="42">ZSM 319/2004</th>
|
||
<td id="FA43E436130340C4FE6AFB8EFDF04613" box="[469,611,1032,1056]" gridcol="1" gridrow="10" pageId="41" pageNumber="42">FGZC 613</td>
|
||
<td id="FA43E436130340C4FD05FB8EFD6C4613" box="[698,767,1032,1056]" gridcol="2" gridrow="10" pageId="41" pageNumber="42">female</td>
|
||
<td id="FA43E436130340C4FC85FB8EFCF94613" box="[826,874,1032,1056]" gridcol="3" gridrow="10" pageId="41" pageNumber="42">35.8</td>
|
||
<td id="FA43E436130340C4FC1FFB8EFC434613" box="[928,976,1032,1056]" gridcol="4" gridrow="10" pageId="41" pageNumber="42">8.9</td>
|
||
<td id="FA43E436130340C4FBB8FB8EFBBF4613" box="[1031,1068,1032,1056]" gridcol="5" gridrow="10" pageId="41" pageNumber="42">4.3</td>
|
||
<td id="FA43E436130340C4FBDEFB8EFB734613" box="[1121,1248,1032,1056]" gridcol="6" gridrow="10" pageId="41" pageNumber="42">24.9</td>
|
||
<td id="FA43E436130340C4FAB9FB8EFAEF4613" box="[1286,1404,1032,1056]" gridcol="7" gridrow="10" pageId="41" pageNumber="42">12.0</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFBA8FAEF4675" box="[160,1404,1070,1094]" gridrow="11" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFBA8FEEF4675" box="[160,380,1070,1094]" gridcol="0" gridrow="11" pageId="41" pageNumber="42">ZSM 330/2004</th>
|
||
<td id="FA43E436130340C4FE6AFBA8FDF04675" box="[469,611,1070,1094]" gridcol="1" gridrow="11" pageId="41" pageNumber="42">FGZC 627</td>
|
||
<td id="FA43E436130340C4FD05FBA8FD6C4675" box="[698,767,1070,1094]" gridcol="2" gridrow="11" pageId="41" pageNumber="42">female</td>
|
||
<td id="FA43E436130340C4FC85FBA8FCF94675" box="[826,874,1070,1094]" gridcol="3" gridrow="11" pageId="41" pageNumber="42">39.5</td>
|
||
<td id="FA43E436130340C4FC1FFBA8FC434675" box="[928,976,1070,1094]" gridcol="4" gridrow="11" pageId="41" pageNumber="42">9.1</td>
|
||
<td id="FA43E436130340C4FBB8FBA8FBBF4675" box="[1031,1068,1070,1094]" gridcol="5" gridrow="11" pageId="41" pageNumber="42">4.6</td>
|
||
<td id="FA43E436130340C4FBDEFBA8FB734675" box="[1121,1248,1070,1094]" gridcol="6" gridrow="11" pageId="41" pageNumber="42">23.0</td>
|
||
<td id="FA43E436130340C4FAB9FBA8FAEF4675" box="[1286,1404,1070,1094]" gridcol="7" gridrow="11" pageId="41" pageNumber="42">11.6</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFBD2FAEF465F" box="[160,1404,1108,1132]" gridrow="12" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFBD2FEEF465F" box="[160,380,1108,1132]" gridcol="0" gridrow="12" pageId="41" pageNumber="42">ZSM 5117/2005</th>
|
||
<td id="FA43E436130340C4FE6AFBD2FDF0465F" box="[469,611,1108,1132]" gridcol="1" gridrow="12" pageId="41" pageNumber="42">FGZC 2689</td>
|
||
<td id="FA43E436130340C4FD05FBD2FD6C465F" box="[698,767,1108,1132]" gridcol="2" gridrow="12" pageId="41" pageNumber="42">female</td>
|
||
<td id="FA43E436130340C4FC85FBD2FCF9465F" box="[826,874,1108,1132]" gridcol="3" gridrow="12" pageId="41" pageNumber="42">34.0</td>
|
||
<td id="FA43E436130340C4FC1FFBD2FC43465F" box="[928,976,1108,1132]" gridcol="4" gridrow="12" pageId="41" pageNumber="42">8.0</td>
|
||
<td id="FA43E436130340C4FBB8FBD2FBBF465F" box="[1031,1068,1108,1132]" gridcol="5" gridrow="12" pageId="41" pageNumber="42">4.2</td>
|
||
<td id="FA43E436130340C4FBDEFBD2FB73465F" box="[1121,1248,1108,1132]" gridcol="6" gridrow="12" pageId="41" pageNumber="42">23.5</td>
|
||
<td id="FA43E436130340C4FAB9FBD2FAEF465F" box="[1286,1404,1108,1132]" gridcol="7" gridrow="12" pageId="41" pageNumber="42">12.4</td>
|
||
</tr>
|
||
<tr id="B9928D4A130340C4FF1FFBFFFAEF46A1" box="[160,1404,1145,1170]" gridrow="13" pageId="41" pageNumber="42">
|
||
<th id="FA43E436130340C4FF1FFBFFFEEF46A1" box="[160,380,1145,1170]" gridcol="0" gridrow="13" pageId="41" pageNumber="42">ZSM 525/2009</th>
|
||
<td id="FA43E436130340C4FE6AFBFFFDF046A1" box="[469,611,1145,1170]" gridcol="1" gridrow="13" pageId="41" pageNumber="42">ZCMV 11210</td>
|
||
<td id="FA43E436130340C4FD05FBFFFD6C46A1" box="[698,767,1145,1170]" gridcol="2" gridrow="13" pageId="41" pageNumber="42">female</td>
|
||
<td id="FA43E436130340C4FC85FBFFFCF946A1" box="[826,874,1145,1170]" gridcol="3" gridrow="13" pageId="41" pageNumber="42">31.3</td>
|
||
<td id="FA43E436130340C4FC1FFBFFFC4346A1" box="[928,976,1145,1170]" gridcol="4" gridrow="13" pageId="41" pageNumber="42">7.6</td>
|
||
<td id="FA43E436130340C4FBB8FBFFFBBF46A1" box="[1031,1068,1145,1170]" gridcol="5" gridrow="13" pageId="41" pageNumber="42">3.8</td>
|
||
<td id="FA43E436130340C4FBDEFBFFFB7346A1" box="[1121,1248,1145,1170]" gridcol="6" gridrow="13" pageId="41" pageNumber="42">24.3</td>
|
||
<td id="FA43E436130340C4FAB9FBFFFAEF46A1" box="[1286,1404,1145,1170]" gridcol="7" gridrow="13" pageId="41" pageNumber="42">12.1</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="071D8F081303BF12FF78FB47FF5446CD" blockId="41.[151,1437,1216,2024]" pageId="41" pageNumber="42">For a distinction from additional species newly named and described herein, see the respective diagnoses below.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831303BF12FF78FA81FD7D475B" pageId="41" pageNumber="42" type="etymology">
|
||
<paragraph id="071D8F081303BF12FF78FA81FD7D475B" blockId="41.[151,1437,1216,2024]" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FF78FA81FEDD4712" bold="true" box="[199,334,1287,1313]" pageId="41" pageNumber="42">Etymology.</emphasis>
|
||
This species is dedicated to Michael Wink, pharmacologist, herpetologist, ornithologist and professor emeritus of the University of Heidelberg, in recognition for his support of research in squamate systematics. The name is a patronym (i.e., a noun in the genitive case).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831303BF10FF78FAF7FD934052" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42" type="description">
|
||
<paragraph id="071D8F081303BF12FF78FAF7FDB2456A" blockId="41.[151,1437,1216,2024]" pageId="41" pageNumber="42">
|
||
<emphasis id="35D6531A1303BF12FF78FAF7FD8047BF" bold="true" box="[199,531,1393,1420]" pageId="41" pageNumber="42">
|
||
Description of the
|
||
<typeStatus id="D81931AA1303BF12FE1AFAF4FD9D47BF" box="[421,526,1394,1420]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male, hemipenes everted, in good state of preservation, tail is broken and missing, second toe on the left forelimb is removed as source of tissue for molecular analysis (
|
||
<figureCitation id="9F99938D1303BF12FB79FA13FA894783" box="[1222,1306,1429,1456]" captionStart="FIGURE 15" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[244,1351,181,1906]" captionTargetId="figure-18@25.[236,1352,181,1919]" captionTargetPageId="25" captionText="FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study." figureDoi="http://doi.org/10.5281/zenodo.7040784" httpUri="https://zenodo.org/record/7040784/files/figure.png" pageId="41" pageNumber="42">Fig. 15</figureCitation>
|
||
). SVL
|
||
<quantity id="C05A22ED1303BF12FAD4FA13FF5647E0" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="41" pageNumber="42" unit="mm" value="29.5">29.5 mm</quantity>
|
||
, TAL
|
||
<quantity id="C05A22ED1303BF12FEB6FA3EFEFC47E0" box="[265,367,1464,1491]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="41" pageNumber="42" unit="mm" value="15.6">15.6 mm</quantity>
|
||
; for other measurements see
|
||
<tableCitation id="4A20BAB31303BF12FD0FFA3FFC9047E0" box="[688,771,1465,1491]" captionStart="TABLE 1" captionStartId="31.[151,239,459,485]" captionTargetPageId="31" captionText="TABLE 1. Selected morphometric and meristic data for voucher specimens of the Lygodactylus madagascariensis group examined for this study. All morphometric measurements (SVL, TAL, HIL, HW, ED, SED) in mm. All specimens in the table were genotyped by 16S sequences, except for historical types from BMNH, MNHN, and SMF, and two further individuals of L. expectatus. Abbreviations are those described in Materials and Methods section; additional abbreviations and symbols as follows: NA, not assessed or not applicable; * counts and measurements partly or fully taken by M. Puente (Puente et al. 2009); § specimen not genotyped; # tail broken or incomplete; ## tail regenerated. HT (sep) and HT (spi) mark the holotypes of Lygodactylus septemtuberculatus Angel, 1942 and Microscalabotes spinulifer Boettger, 1913, here considered to be junior synonyms of L. miops. First LCDS value for L. expectatus refers to the enlarged dorsolateral scales, the second value to the non-enlarged vertebral scales. See species account of L. fritzi sp. nov for the identity of the historical locality “Senbendrana”." httpUri="http://table.plazi.org/id/53DDDF801335BF24FF28FE4DFD0E408E" pageId="41" pageNumber="42" tableUuid="53DDDF801335BF24FF28FE4DFD0E408E">Table 1</tableCitation>
|
||
. Head and neck short, head broader than body. The distance from the tip of the snout to the anterior border of the eye (
|
||
<quantity id="C05A22ED1303BF12FC91FA5AFC1447C5" box="[814,903,1500,1526]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="41" pageNumber="42" unit="mm" value="3.9">3.9 mm</quantity>
|
||
) is less than the interorbital distance anteriorly (4.0 mm), and greater than the distance between the eye and ear opening. Snout covered with granular scales larger than those on the rest of the dorsum. Nostril surrounded by five scales: rostral, first supralabial, and three supranasals. Mental scale undivided; no contact between posterior projection of mental scale and first infralabial; two symmetrical postmental scales with four postpostmental scales; six infralabial scales; seven supralabial scales; two internasal scales; granular dorsal scales; dorsum with small, homogeneous, granular, and unkeeled scales of similar size to those on trunk, smaller than on head and tail, the scales on limbs can be slightly larger; 222 dorsal scales longitudinally along the body; 93 ventral scales between mental and cloaca; venter with large homogeneous smooth scales; first finger present but very small, not bearing a claw; three pairs of subdigital lamellae on the fourth toe; eight not very distinct dorsolateral tubercles, consisting of one scale; seven preanal pores; tail without whorls; large lateral spines at the base of the tail.
|
||
</paragraph>
|
||
<paragraph id="071D8F081303BF10FF78F8E4FD72432E" blockId="41.[151,1437,1216,2024]" lastBlockId="43.[151,1437,151,789]" lastPageId="43" lastPageNumber="44" pageId="41" pageNumber="42">
|
||
Based on available photographs (
|
||
<figureCitation id="9F99938D1303BF12FDECF8E4FD3F454F" box="[595,684,1890,1916]" captionStart="FIGURE 20" captionStartId="42.[151,250,1872,1896]" captionTargetBox="[228,1359,181,1848]" captionTargetId="figure-18@42.[228,1359,181,1848]" captionTargetPageId="42" captionText="FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika." figureDoi="http://doi.org/10.5281/zenodo.7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="41" pageNumber="42">Fig. 20</figureCitation>
|
||
), the
|
||
<typeStatus id="D81931AA1303BF12FD49F8E4FCCB454F" box="[758,856,1890,1916]" pageId="41" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
displayed a light marbled yellow grayish dorsal coloration with more brownish flanks before preservation. Distinct yellow spots are present on the head, the flank, the limbs, and the tail. Two black markings are present on the shoulder on either side of the body, reminiscent of double scapular semi-ocelli. Along the spine a narrow brown line reaches from the neck to the base of the tail. Adjacent to this line two pairs of symmetrical dark spots are present (forming two disrupted chevrons), one pair on forelimb level and one pair
|
||
<quantity id="C05A22ED1301BF10FE69FF3DFDB542E6" box="[470,550,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="43" pageNumber="44" unit="mm" value="10.0">10 mm</quantity>
|
||
posterior (
|
||
<figureCitation id="9F99938D1301BF10FD1CFF3DFC9D42E6" box="[675,782,187,213]" captionStart="FIGURE 20" captionStartId="42.[151,250,1872,1896]" captionTargetBox="[228,1359,181,1848]" captionTargetId="figure-18@42.[228,1359,181,1848]" captionTargetPageId="42" captionText="FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika." figureDoi="http://doi.org/10.5281/zenodo.7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="43" pageNumber="44">Fig. 20D</figureCitation>
|
||
). After six years of preservation in ethanol, the preserved specimen is more uniformly brownish and most of the marbled pattern is faded. The ventral side is fawn with few small brown spots, most of them in the gular region.
|
||
</paragraph>
|
||
<caption id="53DDDF801300BF11FF28F8D6FE6945CB" ID-DOI="http://doi.org/10.5281/zenodo.7040803" ID-Zenodo-Dep="7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1872,1896]" targetBox="[228,1359,181,1848]" targetPageId="42">
|
||
<paragraph id="071D8F081300BF11FF28F8D6FE6945CB" blockId="42.[151,1437,1872,2040]" pageId="42" pageNumber="43">
|
||
<emphasis id="35D6531A1300BF11FF28F8D6FEB2455B" bold="true" box="[151,289,1872,1896]" pageId="42" pageNumber="43">FIGURE 20.</emphasis>
|
||
Photos of
|
||
<taxonomicName id="C0A2F48B1300BF11FE2DF8D6FDC4455B" authority="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz, 2022" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[402,599,1872,1896]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="winki" status="sp. nov.">
|
||
<emphasis id="35D6531A1300BF11FE2DF8D6FDC4455B" box="[402,599,1872,1896]" italics="true" pageId="42" pageNumber="43">Lygodactylus winki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="35D6531A1300BF11FDE1F8D6FD23455B" bold="true" box="[606,688,1872,1896]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="2EE5EE611300BF11FDE1F8D6FD23455B" box="[606,688,1872,1896]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="071D8F081301BF10FF78FEA0FD2D43BA" blockId="43.[151,1437,151,789]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78FEA0FEAC4373" bold="true" box="[199,319,294,320]" pageId="43" pageNumber="44">Variation.</emphasis>
|
||
Males display a light marbled yellow grayish dorsal coloration with more brownish flanks and yellow spots. The dorsal and ventral coloration on females is darker without a pattern except for a few dark dorsal spots (
|
||
<figureCitation id="9F99938D1301BF10FF20FEE9FECE43BA" box="[159,349,367,393]" captionStart="FIGURE 20" captionStartId="42.[151,250,1872,1896]" captionTargetBox="[228,1359,181,1848]" captionTargetId="figure-18@42.[228,1359,181,1848]" captionTargetPageId="42" captionText="FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika." figureDoi="http://doi.org/10.5281/zenodo.7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="43" pageNumber="44">Fig. 20E and 20I</figureCitation>
|
||
in comparison to
|
||
<figureCitation id="9F99938D1301BF10FD98FEE9FD2143BA" box="[551,690,367,393]" captionStart="FIGURE 20" captionStartId="42.[151,250,1872,1896]" captionTargetBox="[228,1359,181,1848]" captionTargetId="figure-18@42.[228,1359,181,1848]" captionTargetPageId="42" captionText="FIGURE 20. Photos of Lygodactylus winki sp. nov. in life. A–C, adult male paratype UADBA-R 70857 (MSZC 0019) from Ampotsidy in dorsolateral and ventral view, and ventral view of throat. D, male holotype ZSM 47/2016 (MSZC 0075) from Ampotsidy. E, female paratype ZSM 555/2014 (DRV 6288) from Andrevorevo. F–G, female paratype UADBA-R 70859 (MSZC 0077) from Ampotsidy. H, male paratype ZSM 48/2016 (MSZC 0110) from Ampotsidy. I, female paratype ZSM 1763/2010 (ZCMV 12502) from Bemanevika." figureDoi="http://doi.org/10.5281/zenodo.7040803" httpUri="https://zenodo.org/record/7040803/files/figure.png" pageId="43" pageNumber="44">Fig. 20A–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="071D8F081301BF10FF78FE15FD934052" blockId="43.[151,1437,151,789]" pageId="43" pageNumber="44">
|
||
<materialsCitation id="B7CA85551301BF10FF78FE15FE6F4052" ID-GBIF-Occurrence="3897988315" collectionCode="SVL" pageId="43" pageNumber="44" specimenCode="ZSM 555, ZSM 48, ZSM 1763" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
|
||
Three additional specimens (
|
||
<specimenCount id="11A444811301BF10FDB2FE15FD11439E" box="[525,642,403,429]" pageId="43" pageNumber="44" type="male">two males</specimenCount>
|
||
[
|
||
<specimenCode id="570427731301BF10FD2FFE15FD6C439E" box="[656,767,402,429]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="43" pageNumber="44" type="Museum">ZSM 555</specimenCode>
|
||
/2014,
|
||
<specimenCode id="570427731301BF10FCF1FE15FC3E439E" box="[846,941,403,429]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="43" pageNumber="44" type="Museum">ZSM 48</specimenCode>
|
||
/2016],
|
||
<specimenCount id="11A444811301BF10FBBBFE15FB13439E" box="[1028,1152,403,429]" pageId="43" pageNumber="44" type="female">one female</specimenCount>
|
||
[
|
||
<specimenCode id="570427731301BF10FB30FE15FA98439E" box="[1167,1291,403,429]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="43" pageNumber="44" type="Museum">ZSM 1763</specimenCode>
|
||
/2010]) were examined. The
|
||
<collectionCode id="61B317CD1301BF10FEFDFE31FEE543E2" box="[322,374,439,465]" pageId="43" pageNumber="44">SVL</collectionCode>
|
||
ranges between 29.8 and
|
||
<quantity id="C05A22ED1301BF10FD2BFE31FD6B43E2" box="[660,760,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="43" pageNumber="44" unit="mm" value="33.4">33.4 mm</quantity>
|
||
with the female being the largest. The
|
||
<specimenCount id="11A444811301BF10FB1AFE31FA8543E2" box="[1189,1302,439,465]" pageId="43" pageNumber="44" type="male">two males</specimenCount>
|
||
have a TAL of 37.8 and 45.0 mm. The relative hindlimb length is 0.49 to 0.53 with the female having the smallest. The female also has the smallest eyes relative to the size of the body and with a medium size smaller tubercles at the tail base than the males, which have large tubercles. The number of dorsal scales ranges between 187 and 208. The ventral scales range between 83 and 98
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831301BF10FF78FDEDFC5840FE" pageId="43" pageNumber="44" type="biology_ecology">
|
||
<paragraph id="071D8F081301BF10FF78FDEDFC5840FE" blockId="43.[151,1437,151,789]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78FDEDFE1540B7" bold="true" box="[199,390,618,645]" pageId="43" pageNumber="44">Natural history.</emphasis>
|
||
<materialsCitation id="B7CA85551301BF10FE32FDEDFC5440FE" ID-GBIF-Occurrence="3897988301" accessionNumber="R 70856" collectionCode="R" elevation="2" pageId="43" pageNumber="44" specimenCount="2" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="D81931AA1301BF10FE7FFDEDFDB140B6" box="[448,546,619,645]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
was collected in primary rainforest, on the trunk of a big tree,
|
||
<quantity id="C05A22ED1301BF10FB57FDEDFABB40B6" box="[1256,1320,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="0.2">
|
||
<elevation id="8C8F683B1301BF10FB57FDEDFABB40B6" box="[1256,1320,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="m" value="0.2">0.2 m</elevation>
|
||
</quantity>
|
||
above the ground. The
|
||
<typeStatus id="D81931AA1301BF10FE9AFD09FE03409A" box="[293,400,655,681]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
|
||
from Ampotsidy were mostly collected at night sleeping on leaves, twigs, or vines.
|
||
<collectionCode id="61B317CD1301BF10FA90FD09FF3940FE" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="44">UADBA-R</collectionCode>
|
||
70856 was collected in the afternoon, on the ground during heavy rain
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FB8DC831301BF10FF78FD50FC794126" pageId="43" pageNumber="44" type="distribution">
|
||
<paragraph id="071D8F081301BF10FF78FD50FC794126" blockId="43.[151,1437,151,789]" pageId="43" pageNumber="44">
|
||
<emphasis id="35D6531A1301BF10FF78FD50FEF240C3" bold="true" box="[199,353,726,752]" pageId="43" pageNumber="44">Distribution.</emphasis>
|
||
<taxonomicName id="C0A2F48B1301BF10FED8FD51FE5240C2" authorityName="Vences & Multzsch & Gippner & Miralles & Crottini & Gehring & Rakotoarison & Ratsoavina & Glaw & Scherz" authorityYear="2022" box="[359,449,727,753]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="winki">
|
||
<emphasis id="35D6531A1301BF10FED8FD51FE5240C2" box="[359,449,727,753]" italics="true" pageId="43" pageNumber="44">L. winki</emphasis>
|
||
</taxonomicName>
|
||
is known from three localities in the North and Sambirano regions in northern
|
||
<collectingCountry id="7FB5CF981301BF10FA91FD51FF2E4126" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
|
||
: (1) the
|
||
<typeStatus id="D81931AA1301BF10FEA6FD7DFEDA4126" box="[281,329,763,789]" pageId="43" pageNumber="44">type</typeStatus>
|
||
locality, Ampotsidy, (2) Andrevorevo, and (3) Bemanevika.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |