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<mods:title id="1833512A0C7A6C83E82EE7A590921D0C">New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda)</mods:title>
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<emphasis id="D9CBEAEEFF98FFA4E2C27E9AF261FD1D" bold="true" box="[151,374,693,720]" italics="true" pageId="3" pageNumber="254">Alpheus edwardsii</emphasis>
(
<bibRefCitation id="8F2E4B0DFF98FFA4E3D07E9AF128FD02" author="Audouin, V." box="[389,575,693,719]" pageId="3" pageNumber="254" pagination="77 - 98" refId="ref11726" refString="Audouin, V. (1826) Explication sommaire des planches de crustaces de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, Membre de l'Institut; offrant un expos des characteres naturels des genres avec la distinction des especes. Animaux invertebres. In: Description de l'Egypte ou receuil des observations et des recherches qui ont et faites en Egypte pendant l'expedition de l'arme francaise, publi par les ordres de sa Majest l'Empereur Napoleon le Grand. Imprimerie Imperiale, Paris, pp. 77 - 98." type="book chapter" year="1826">Audouin, 1826</bibRefCitation>
)
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<taxonomicNameLabel id="C2F85795FF98FFA4E01B7E9AF1DEFD02" box="[590,713,693,719]" pageId="3" pageNumber="254" sensu="lato">sensu lato</taxonomicNameLabel>
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(
<figureCitation id="73842A79FF98FFA4E2F57EF6F3F4FD3E" box="[160,227,729,755]" captionStart="FIGURE 2" captionStartId="4.[151,250,1702,1727]" captionTargetBox="[151,1435,181,1677]" captionTargetId="figure@4.[151,1435,181,1677]" captionTargetPageId="4" captionText="FIGURE 2. Alpheusedwardsii (Audouin, 1826) [sensu lato]:A—male from Kubbar Island, Kuwait (OUMNH.ZC.2020.01.018), dorsal view; B—same, major chela with dactylus opened. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909081" httpUri="https://zenodo.org/record/3909081/files/figure.png" pageId="3" pageNumber="254">Fig. 2</figureCitation>
)
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<treatmentCitation id="6A1E10EDFF98FFA4E2C27F32F11EFCF8" author="Audouin, V." box="[151,521,796,821]" page="91" pageId="3" pageNumber="254" year="1826">
<taxonomicName id="2CBF4D7FFF98FFA4E2C27F32F11EFCF8" ID-CoL="84LMQ" authority="Edwarsii Audouin 1826: 91" authorityName="Edwarsii Audouin" authorityPageNumber="91" authorityYear="1826" box="[151,521,796,821]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="D9CBEAEEFF98FFA4E2C27F32F25AFCF8" box="[151,333,797,821]" italics="true" pageId="3" pageNumber="254">Athanas Edwarsii</emphasis>
<bibRefCitation id="8F2E4B0DFF98FFA4E3047F33F11EFCF8" author="Audouin, V." box="[337,521,796,821]" pageId="3" pageNumber="254" pagination="77 - 98" refId="ref11726" refString="Audouin, V. (1826) Explication sommaire des planches de crustaces de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, Membre de l'Institut; offrant un expos des characteres naturels des genres avec la distinction des especes. Animaux invertebres. In: Description de l'Egypte ou receuil des observations et des recherches qui ont et faites en Egypte pendant l'expedition de l'arme francaise, publi par les ordres de sa Majest l'Empereur Napoleon le Grand. Imprimerie Imperiale, Paris, pp. 77 - 98." type="book chapter" year="1826">Audouin 1826: 91</bibRefCitation>
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(
<emphasis id="D9CBEAEEFF98FFA4E04D7F32F17CFCF8" box="[536,619,797,821]" italics="true" pageId="3" pageNumber="254">lap. cal.</emphasis>
).
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<taxonomicName id="2CBF4D7FFF98FFA4E2C27F13F248FC99" ID-CoL="84LMQ" authority="Edwardsii." authorityName="Edwardsii." box="[151,351,828,852]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="D9CBEAEEFF98FFA4E2C27F13F248FC99" box="[151,351,828,852]" italics="true" pageId="3" pageNumber="254">Athanas Edwardsii.</emphasis>
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<treatmentCitation id="6A1E10EDFF98FFA4E32D7F14F128FC99" author="Audouin, V." box="[376,575,827,852]" page="274" pageId="3" pageNumber="254" year="1827">
<bibRefCitation id="8F2E4B0DFF98FFA4E32D7F14F128FC99" author="Audouin, V." box="[376,575,827,852]" pageId="3" pageNumber="254" pagination="249 - 290" refId="ref11819" refString="Audouin, V. (1827) Description de l'Egypte, ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise, publie par les ordres de sa Majeste l'Empereur Napoleon le Grand. Vol. 22. Edition 2. Imprimerie Imperiale, Paris, 42 pp. [pp. 249 - 290]" type="book chapter" year="1827">Audouin 1827: 274</bibRefCitation>
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.
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<treatmentCitationGroup id="CBAF11D2FF98FFA4E2C27F75F271FC03" pageId="3" pageNumber="254">
<taxonomicName id="2CBF4D7FFF98FFA4E2C27F75F248FCBF" ID-CoL="84LB5" authority="Edwardsii." authorityName="Edwardsii." box="[151,351,858,882]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="D9CBEAEEFF98FFA4E2C27F75F248FCBF" box="[151,351,858,882]" italics="true" pageId="3" pageNumber="254">Alpheus Edwardsii.</emphasis>
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<treatmentCitation id="6A1E10EDFF98FFA4E32D7F75F156FCBE" author="Coutiere, H." box="[376,577,858,883]" page="249" pageId="3" pageNumber="254" year="1898">
<bibRefCitation id="8F2E4B0DFF98FFA4E32D7F75F156FCBE" author="Coutiere, H." box="[376,577,858,883]" pageId="3" pageNumber="254" pagination="131 - 134" refId="ref12776" refString="Coutiere, H. (1898) Note sur quelques formes nouvelles d'Alpheides voisines de A. Bouvieri A. M. - Edwards [Crust.]. Bulletin de la Societe Entomologique de France, 1898, 131 - 134. https: // doi. org / 10.5962 / bhl. part. 16847" type="journal article" year="1898">Coutière 1898: 249</bibRefCitation>
</treatmentCitation>
, figs. 4, 5;
<treatmentCitation id="6A1E10EDFF98FFA4E0EC7F75F045FCBE" author="Nobili, G." box="[697,850,858,883]" page="2" pageId="3" pageNumber="254" year="1901">
<bibRefCitation id="8F2E4B0DFF98FFA4E0EC7F75F045FCBE" author="Nobili, G." box="[697,850,858,883]" pageId="3" pageNumber="254" pagination="1 - 21" refId="ref16019" refString="Nobili, G. (1901) Decapodi e Stomatopodi Eritrei del Museo Zoologico dell'Universita di Napoli. Annuario del Museo Zoologico della R. Universita di Napoli, 1, 1 - 21." type="journal article" year="1901">Nobili 1901: 2</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF98FFA4E10A7F74F788FCBE" author="Banner, A. H. &amp; Banner, D. M." box="[863,1183,859,883]" page="1141" pageId="3" pageNumber="254" year="1973">
<bibRefCitation id="8F2E4B0DFF98FFA4E10A7F74F788FCBE" author="Banner, A. H. &amp; Banner, D. M." box="[863,1183,859,883]" pageId="3" pageNumber="254" pagination="1141 - 1146" refId="ref11995" refString="Banner, A. H. &amp; Banner, D. M. (1973 a) The establishment of a neotype for Alpheus edwardsi (Audouin). Bulletin du Museum national d'Histoire naturelle, 3 eme serie, Zoologie, 88, 1141 - 1146." type="journal article" year="1973">Banner &amp; Banner 1973a: 1141</bibRefCitation>
</treatmentCitation>
, fig. 1;
<treatmentCitation id="6A1E10EDFF98FFA4E6A47F75F227FC5C" author="Kazmi, M. A. &amp; Kazmi, Q. B." page="154" pageId="3" pageNumber="254" year="1979">
<bibRefCitation id="8F2E4B0DFF98FFA4E6A47F75F227FC5C" author="Kazmi, M. A. &amp; Kazmi, Q. B." pageId="3" pageNumber="254" pagination="151 - 157" refId="ref14783" refString="Kazmi, M. A. &amp; Kazmi, Q. B. (1979) A check-list of marine caridean prawns of Pakistan. Biologia, 25, 151 - 157." type="journal article" year="1979">Kazmi &amp; Kazmi 1979: 154</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF98FFA4E36C7F55F15BFC5C" author="Banner, D. M. &amp; Banner, A. H." box="[313,588,889,913]" page="22" pageId="3" pageNumber="254" year="1981">
<bibRefCitation id="8F2E4B0DFF98FFA4E36C7F55F15BFC5C" author="Banner, D. M. &amp; Banner, A. H." box="[313,588,889,913]" pageId="3" pageNumber="254" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 22</bibRefCitation>
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, 92;
<treatmentCitation id="6A1E10EDFF98FFA4E0297F55F08DFC5C" author="Banner, D. M. &amp; Banner, A. H." box="[636,922,889,913]" page="270" pageId="3" pageNumber="254" year="1982">
<bibRefCitation id="8F2E4B0DFF98FFA4E0297F55F08DFC5C" author="Banner, D. M. &amp; Banner, A. H." box="[636,922,889,913]" pageId="3" pageNumber="254" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982: 270</bibRefCitation>
</treatmentCitation>
, fig. 83;
<treatmentCitation id="6A1E10EDFF98FFA4E1A67F57F633FC5C" author="Lewinsohn, C. &amp; Galil, B." box="[1011,1316,888,913]" page="208" pageId="3" pageNumber="254" year="1982">
<bibRefCitation id="8F2E4B0DFF98FFA4E1A67F57F633FC5C" author="Lewinsohn, C. &amp; Galil, B." box="[1011,1316,888,913]" pageId="3" pageNumber="254" pagination="207 - 210" refId="ref15201" refString="Lewinsohn, C. &amp; Galil, B. (1982) Notes on species of Alpheus (Crustacea Decapoda) from the Mediterranean coast of Israel. Quaderni del Laboratorio di Tecnologia della Pesca, Ancona, 3, 207 - 210." type="journal article" year="1982">Lewinsohn &amp; Galil 1982: 208</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF98FFA4E7797F57F226FC7D" author="Afzal, M. &amp; Javed, A. &amp; Barkati, S." page="338" pageId="3" pageNumber="254" year="1986">
<bibRefCitation id="8F2E4B0DFF98FFA4E7797F57F226FC7D" author="Afzal, M. &amp; Javed, A. &amp; Barkati, S." pageId="3" pageNumber="254" pagination="337 - 340" refId="ref11048" refString="Afzal, M., Javed, A. &amp; Barkati, S. (1986) Checklist, synonymy and key to the genera and species of alpheid shrimps of Karachi coast. Pakistan Journal of Agriculture Research, 7, 337 - 340." type="journal article" year="1986">
Afzal
<emphasis id="D9CBEAEEFF98FFA4E73F7F55F681FC5C" box="[1386,1430,889,913]" italics="true" pageId="3" pageNumber="254">et al</emphasis>
. 1986: 338
</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF98FFA4E3697FB7F17EFC7D" author="De Grave, S. &amp; Ashelby, C. W." box="[316,617,919,944]" page="3" pageId="3" pageNumber="254" year="2011">
<bibRefCitation id="8F2E4B0DFF98FFA4E3697FB7F17EFC7D" author="De Grave, S. &amp; Ashelby, C. W." box="[316,617,919,944]" pageId="3" pageNumber="254" pagination="1 - 7" refId="ref13141" refString="De Grave, S. &amp; Ashelby, C. W. (2011) Notes on some shrimp species (Decapoda: Caridea) from Qatar, with an annotated checklist of marine caridean species known from the Persian Gulf. Marine Biodiversity Records, 4 (e 20), 1 - 7. https: // doi. org / 10.1017 / S 1755267211000224" type="journal article" year="2011">De Grave &amp; Ashelby 2011: 3</bibRefCitation>
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;
<treatmentCitation id="6A1E10EDFF98FFA4E0217FB8F083FC7D" author="Naderloo, R. &amp; Turkay, M." box="[628,916,919,944]" page="9" pageId="3" pageNumber="254" year="2012">
<bibRefCitation id="8F2E4B0DFF98FFA4E0217FB8F083FC7D" author="Naderloo, R. &amp; Turkay, M." box="[628,916,919,944]" pageId="3" pageNumber="254" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012: 9</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF98FFA4E1CA7FB8F7BEFC7D" author="Naderloo, R. &amp; Turkay, M. &amp; Sari, A." box="[927,1193,919,944]" page="448" pageId="3" pageNumber="254" year="2013">
<bibRefCitation id="8F2E4B0DFF98FFA4E1CA7FB8F7BEFC7D" author="Naderloo, R. &amp; Turkay, M. &amp; Sari, A." box="[927,1193,919,944]" pageId="3" pageNumber="254" pagination="445 - 462" refId="ref15763" refString="Naderloo, R., Turkay, M. &amp; Sari, A. (2013) Intertidal habitats and decapod (Crustacea) diversity of Qeshm Island, a biodiversity hotspot within the Persian Gulf. Marine Biodiversity, 43, 445 - 462. https: // doi. org / 10.1007 / s 12526 - 013 - 0174 - 3" type="journal article" year="2013">
Naderloo
<emphasis id="D9CBEAEEFF98FFA4E6507FB6F72EFC7D" box="[1029,1081,920,944]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
2013: 448
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="6A1E10EDFF98FFA4E6E17FB8F3F8FC02" author="Dehghani, A. &amp; Sari A. &amp; Naderloo, R." page="484" pageId="3" pageNumber="254" year="2019">
<bibRefCitation id="8F2E4B0DFF98FFA4E6E17FB8F3F8FC02" author="Dehghani, A. &amp; Sari A. &amp; Naderloo, R." pageId="3" pageNumber="254" pagination="479 - 504" refId="ref13421" refString="Dehghani, A., Sari A. &amp; Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
Dehghani
<emphasis id="D9CBEAEEFF98FFA4E7487FB6F646FC7D" box="[1309,1361,920,944]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
2019a: 484
</bibRefCitation>
</treatmentCitation>
, fig. 3A, B.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EB0036FCFF98FFA4E2C27FFAF098FC20" blockId="3.[151,1437,796,1067]" box="[151,911,980,1005]" pageId="3" pageNumber="254">
<treatmentCitationGroup id="CBAF11D2FF98FFA4E2C27FFAF098FC20" box="[151,911,980,1005]" pageId="3" pageNumber="254">
<taxonomicName id="2CBF4D7FFF98FFA4E2C27FFAF245FC20" ID-CoL="84W7L" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[151,338,981,1005]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" sensu="lato" species="edwardsii">
<emphasis id="D9CBEAEEFF98FFA4E2C27FFAF245FC20" box="[151,338,981,1005]" italics="true" pageId="3" pageNumber="254">Alpheus edwardsii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2F85795FF98FFA4E30D7FF9F2AAFC20" box="[344,445,980,1005]" pageId="3" pageNumber="254" sensu="lato">sensu lato</taxonomicNameLabel>
.—
<treatmentCitation id="6A1E10EDFF98FFA4E38F7FFBF01AFC20" author="Anker, A. &amp; De Grave, S." box="[474,781,980,1005]" page="352" pageId="3" pageNumber="254" year="2016">
<bibRefCitation id="8F2E4B0DFF98FFA4E38F7FFBF01AFC20" author="Anker, A. &amp; De Grave, S." box="[474,781,980,1005]" pageId="3" pageNumber="254" pagination="343 - 454" refId="ref11545" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016: 352</bibRefCitation>
</treatmentCitation>
, figs. 10, 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="EB0036FCFF98FFA4E2C27FDBF1C4FBC1" blockId="3.[151,1437,796,1067]" box="[151,723,1011,1036]" pageId="3" pageNumber="254">
(?)
<taxonomicName id="2CBF4D7FFF98FFA4E2EF7FDBF12EFBC1" ID-CoL="84LB5" authority="Audouini Coutiere 1905: 911" authorityName="Audouini Coutiere" authorityPageNumber="911" authorityYear="1905" box="[186,569,1011,1036]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="D9CBEAEEFF98FFA4E2EF7FDBF279FBC1" box="[186,366,1012,1036]" italics="true" pageId="3" pageNumber="254">Alpheus Audouini</emphasis>
<bibRefCitation id="8F2E4B0DFF98FFA4E3217FDCF12EFBC1" author="Coutiere, H." box="[372,569,1011,1036]" pageId="3" pageNumber="254" pagination="852 - 921" refId="ref12869" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The Fauna and Geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the Collections made by an Expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921." type="book chapter" year="1905">Coutière 1905: 911</bibRefCitation>
</taxonomicName>
, pl. 87, fig. 52.
</paragraph>
<paragraph id="EB0036FCFF98FFA4E2C2783CF790FBE6" blockId="3.[151,1437,796,1067]" box="[151,1159,1042,1067]" pageId="3" pageNumber="254">
<treatmentCitationGroup id="CBAF11D2FF98FFA4E2C2783CF790FBE6" box="[151,1159,1042,1067]" pageId="3" pageNumber="254">
<collectingRegion id="297BF81EFF98FFA4E2C2783CF3AAFBE7" box="[151,189,1043,1066]" country="Maldives" name="Uthuru" pageId="3" pageNumber="254">Not</collectingRegion>
<taxonomicName id="2CBF4D7FFF98FFA4E291783DF291FBE7" ID-CoL="84LB5" authority="edwardsii." authorityName="Edwardsii." box="[196,390,1042,1066]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="genus">
<emphasis id="D9CBEAEEFF98FFA4E291783DF291FBE7" box="[196,390,1042,1066]" italics="true" pageId="3" pageNumber="254">Alpheus edwardsii.</emphasis>
</taxonomicName>
<treatmentCitation id="6A1E10EDFF98FFA4E3CA783DF1B3FBE6" author="Al-Maliky, T. H. Y. &amp; Al-Khafaji, K. K. &amp; Khalaf, T. A." box="[415,676,1042,1067]" page="49" pageId="3" pageNumber="254" year="2017">
<bibRefCitation id="8F2E4B0DFF98FFA4E3CA783DF1B3FBE6" author="Al-Maliky, T. H. Y. &amp; Al-Khafaji, K. K. &amp; Khalaf, T. A." box="[415,676,1042,1067]" pageId="3" pageNumber="254" pagination="47 - 53" refId="ref11168" refString="Al-Maliky, T. H. Y., Al-Khafaji, K. K. &amp; Khalaf, T. A. (2017) New records of the snapping shrimp Alpheus edwardsii (Audouin, 1826) (Crustacea: Alpheoidea) in Basarh, Iraq. Arthropods, 6, 47 - 53." type="journal article" year="2017">
Al-Maliky
<emphasis id="D9CBEAEEFF98FFA4E05A783CF154FBE7" box="[527,579,1042,1066]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
2017: 49
</bibRefCitation>
</treatmentCitation>
, figs. 24 [=
<taxonomicName id="2CBF4D7FFF98FFA4E17F783DF76AFBE6" ID-CoL="84W8B" authority="De Haan, 1849" authorityName="De Haan" authorityYear="1849" box="[810,1149,1042,1067]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="lobidens">
<emphasis id="D9CBEAEEFF98FFA4E17F783DF0CFFBE7" box="[810,984,1042,1066]" italics="true" pageId="3" pageNumber="254">Alpheus lobidens</emphasis>
De Haan, 1849
</taxonomicName>
].
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</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF98FFA4E2C27877F012FAA2" pageId="3" pageNumber="254" type="materials_examined">
<paragraph id="EB0036FCFF98FFA4E2C27877F7BAFB76" blockId="3.[151,1437,1112,1931]" pageId="3" pageNumber="254">
<emphasis id="D9CBEAEEFF98FFA4E2C27877F26CFBBE" bold="true" box="[151,379,1112,1139]" pageId="3" pageNumber="254">Material examined</emphasis>
.
<materialsCitation id="5BD73CA1FF98FFA4E3DF7876F76EFB5A" collectingDate="2019-09-26" collectionCode="OUMNH, ZC" collectorName="S. De Grave" country="Kuwait" elevation="3" latitude="29.071926" location="Kubbar Island" longLatPrecision="1" longitude="48.48854" pageId="3" pageNumber="254" specimenCount="1" specimenCount-male="1">
<collectingCountry id="93A8766CFF98FFA4E3DF7876F2F6FBBE" box="[394,481,1113,1139]" name="Kuwait" pageId="3" pageNumber="254">Kuwait</collectingCountry>
:
<specimenCount id="FDB9FD75FF98FFA4E3B87876F12BFBBE" box="[493,572,1113,1139]" pageId="3" pageNumber="254" type="male">1 male</specimenCount>
,
<collectionCode id="8DAEAE39FF98FFA4E01C7876F1AEFBBE" box="[585,697,1113,1139]" pageId="3" pageNumber="254">OUMNH</collectionCode>
.
<collectionCode id="8DAEAE39FF98FFA4E0EE7876F1F7FBBE" box="[699,736,1113,1139]" country="Austria" name="Zoological Collection, University of Vienna" pageId="3" pageNumber="254" type="University or college">ZC</collectionCode>
. 2020.01.018,
<location id="EE606027FF98FFA4E1DD7876F73CFBBE" LSID="urn:lsid:plazi:treatment:631687EAFF98FFA4E2C27E9AF3E6F846:EE606027FF98FFA4E1DD7876F73CFBBE" box="[904,1067,1113,1139]" country="Kuwait" latitude="29.071926" longLatPrecision="1" longitude="48.48854" name="Kubbar Island" pageId="3" pageNumber="254">Kubbar Island</location>
,
<geoCoordinate id="8E8B503BFF98FFA4E66D7876F7F3FBBE" box="[1080,1252,1113,1139]" degrees="29" direction="north" minutes="04" orientation="latitude" pageId="3" pageNumber="254" precision="1" seconds="18.93" value="29.071926">29°0418.93”N</geoCoordinate>
<geoCoordinate id="8E8B503BFF98FFA4E6B97876F68FFBBE" box="[1260,1432,1113,1139]" degrees="48" direction="east" minutes="29" orientation="longitude" pageId="3" pageNumber="254" precision="1" seconds="18.74" value="48.48854">48°2918.74”E</geoCoordinate>
, depth
<quantity id="2C479B19FF98FFA4E2897852F233FB5A" box="[220,292,1149,1175]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="254" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">
<elevation id="6092D1CFFF98FFA4E2897852F233FB5A" box="[220,292,1149,1175]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="254" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">23 m</elevation>
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, sand flat with sparse rubble, under coral rocks, leg.
<collectorName id="464A532AFF98FFA4E1397852F0E0FB5A" box="[876,1015,1149,1175]" pageId="3" pageNumber="254">S. De Grave</collectorName>
,
<date id="9F01103CFF98FFA4E6577852F76EFB5A" box="[1026,1145,1149,1175]" pageId="3" pageNumber="254" value="2019-09-26">
<collectingDate id="8F45E9D4FF98FFA4E6577852F76EFB5A" box="[1026,1145,1149,1175]" pageId="3" pageNumber="254" value="2019-09-26">26.ix.2019</collectingDate>
</date>
</materialsCitation>
[fcn KUW-017];
<materialsCitation id="5BD73CA1FF98FFA4E76A7852F7BEFB76" collectingDate="2019-12-15" collectionCode="KISR" collectorName="M. Al-Kandari" country="Kuwait" location="Miscan Island" pageId="3" pageNumber="254" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="FDB9FD75FF98FFA4E76A7852F68FFB5A" box="[1343,1432,1149,1175]" pageId="3" pageNumber="254" type="male">2 males</specimenCount>
,
<specimenCount id="FDB9FD75FF98FFA4E2C2788EF3ECFB76" box="[151,251,1185,1211]" pageId="3" pageNumber="254" type="female">1 female</specimenCount>
,
<collectionCode id="8DAEAE39FF98FFA4E353788EF25EFB76" box="[262,329,1185,1211]" pageId="3" pageNumber="254">KISR</collectionCode>
,
<location id="EE606027FF98FFA4E301788EF2E2FB76" LSID="urn:lsid:plazi:treatment:631687EAFF98FFA4E2C27E9AF3E6F846:EE606027FF98FFA4E301788EF2E2FB76" box="[340,501,1185,1211]" country="Kuwait" name="Miscan Island" pageId="3" pageNumber="254">Miscan Island</location>
, sandy-rocky intertidal, leg.
<collectorName id="464A532AFF98FFA4E160788EF0CDFB76" box="[821,986,1185,1211]" pageId="3" pageNumber="254">M. Al-Kandari</collectorName>
<emphasis id="D9CBEAEEFF98FFA4E1B4788DF705FB76" box="[993,1042,1185,1211]" italics="true" pageId="3" pageNumber="254">et al</emphasis>
.,
<date id="9F01103CFF98FFA4E672788FF7BEFB76" box="[1063,1193,1184,1211]" pageId="3" pageNumber="254" value="2019-12-15">
<collectingDate id="8F45E9D4FF98FFA4E672788FF7BEFB76" box="[1063,1193,1184,1211]" pageId="3" pageNumber="254" value="2019-12-15">15.xii.2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="EB0036FCFF98FFA4E29278EBF012FAA2" blockId="3.[151,1437,1112,1931]" pageId="3" pageNumber="254">
<emphasis id="D9CBEAEEFF98FFA4E29278EBF2ABFB12" bold="true" box="[199,444,1220,1247]" pageId="3" pageNumber="254">General distribution</emphasis>
. Species complex (see Remarks below), reported from throughout the Indo-West Pacific, from the Red Sea, Arabian Gulf (
<collectingCountry id="93A8766CFF98FFA4E05978C6F12AFACE" box="[524,573,1257,1283]" name="Iran" pageId="3" pageNumber="254">Iran</collectingCountry>
,
<collectingCountry id="93A8766CFF98FFA4E01D78C6F18AFACE" box="[584,669,1257,1283]" name="Kuwait" pageId="3" pageNumber="254">Kuwait</collectingCountry>
,
<collectingCountry id="93A8766CFF98FFA4E0FC78C6F1F3FACE" box="[681,740,1257,1283]" name="United Arab Emirates" pageId="3" pageNumber="254">UAE</collectingCountry>
) and
<collectingCountry id="93A8766CFF98FFA4E17778C6F0A6FACE" box="[802,945,1257,1283]" name="South Africa" pageId="3" pageNumber="254">South Africa</collectingCountry>
across the Indian Ocean to
<collectingCountry id="93A8766CFF98FFA4E6BC78C6F63BFACE" box="[1257,1324,1257,1283]" name="Japan" pageId="3" pageNumber="254">Japan</collectingCountry>
,
<collectingCountry id="93A8766CFF98FFA4E76378C6F68AFACE" box="[1334,1437,1257,1283]" name="Australia" pageId="3" pageNumber="254">Australia</collectingCountry>
and
<collectingCountry id="93A8766CFF98FFA4E29C7922F29AFAEA" box="[201,397,1293,1319]" name="French Polynesia" pageId="3" pageNumber="254">French Polynesia</collectingCountry>
(e.g.,
<bibRefCitation id="8F2E4B0DFF98FFA4E38D7922F1E4FAEA" author="Banner, A. H. &amp; Banner, D. M." box="[472,755,1293,1319]" pageId="3" pageNumber="254" pagination="1141 - 1146" refId="ref11995" refString="Banner, A. H. &amp; Banner, D. M. (1973 a) The establishment of a neotype for Alpheus edwardsi (Audouin). Bulletin du Museum national d'Histoire naturelle, 3 eme serie, Zoologie, 88, 1141 - 1146." type="journal article" year="1973">Banner &amp; Banner 1973a</bibRefCitation>
,
<bibRefCitation id="8F2E4B0DFF98FFA4E1557922F02DFAEA" author="Banner, D. M. &amp; Banner, A. H." box="[768,826,1293,1319]" pageId="3" pageNumber="254" pagination="1 - 99" refId="ref12195" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">1981</bibRefCitation>
,
<bibRefCitation id="8F2E4B0DFF98FFA4E11C7922F093FAEA" author="Banner, D. M. &amp; Banner, A. H." box="[841,900,1293,1319]" pageId="3" pageNumber="254" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">1982</bibRefCitation>
,
<bibRefCitation id="8F2E4B0DFF98FFA4E1C47922F0DBFAEA" author="Banner, A. H. &amp; Banner, D. M." box="[913,972,1293,1319]" pageId="3" pageNumber="254" pagination="1 - 164" refId="ref12098" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the Western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E18F7922F773FAEA" author="Chace, F. A. Jr." box="[986,1124,1293,1319]" pageId="3" pageNumber="254" pagination="1 - 99" refId="ref12618" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E6277922F68EFAEA" author="Naderloo, R. &amp; Turkay, M." box="[1138,1433,1293,1319]" pageId="3" pageNumber="254" pagination="1 - 67" refId="ref15707" refString="Naderloo, R. &amp; Turkay, M. (2012) Decapod crustaceans of the littoral and shallow sublittoral Iranian coast of the Persian Gulf: faunistics, biodiversity and zoogeography. Zootaxa, 3374 (1), 1 - 67. https: // doi. org / 10.11646 / zootaxa. 3374.1.1" type="journal article" year="2012">Naderloo &amp; Türkay 2012</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E2C2791EF2B9FA86" author="Anker, A. &amp; De Grave, S." box="[151,430,1329,1355]" pageId="3" pageNumber="254" pagination="343 - 454" refId="ref11545" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E3EC791EF1A5FA86" author="Dehghani, A. &amp; Sari A. &amp; Naderloo, R." box="[441,690,1329,1355]" pageId="3" pageNumber="254" pagination="479 - 504" refId="ref13421" refString="Dehghani, A., Sari A. &amp; Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
Dehghani
<emphasis id="D9CBEAEEFF98FFA4E079791DF173FA86" box="[556,612,1329,1355]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
2019a
</bibRefCitation>
; present study); Lessepsian migrant into the eastern Mediterranean Sea (
<bibRefCitation id="8F2E4B0DFF98FFA4E29B797AF2DDFAA2" author="Forest, J. &amp; Guinot, D." box="[206,458,1364,1391]" pageId="3" pageNumber="254" pagination="4 - 16" refId="ref13850" refString="Forest, J. &amp; Guinot, D. (1958) Sur une collection de crustaces decapodes des cotes d'Israel. Bulletin of the Sea Fisheries Research Station of Israel, 15, 4 - 16." type="journal article" year="1958">Forest &amp; Guinot 1958</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E380797AF1EFFAA2" author="Holthuis, L. B. &amp; Gottlieb, E." box="[469,760,1364,1391]" pageId="3" pageNumber="254" pagination="1 - 126" refId="ref14516" refString="Holthuis, L. B. &amp; Gottlieb, E. (1958) An annotated list of the Decapod Crustacea of the Mediterranean Coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis &amp; Gottlieb 1958</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF98FFA4E2927956F684FA11" pageId="3" pageNumber="254" type="biology_ecology">
<paragraph id="EB0036FCFF98FFA4E2927956F684FA11" blockId="3.[151,1437,1112,1931]" pageId="3" pageNumber="254">
<emphasis id="D9CBEAEEFF98FFA4E2927956F231FA5E" bold="true" box="[199,294,1401,1427]" pageId="3" pageNumber="254">Ecology</emphasis>
. Variable (species complex, see below); found both in the intertidal and shallow subtidal (to at least
<quantity id="2C479B19FF98FFA4E7D57956F3A7FA7A" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="254" unit="m" value="36.0">36 m</quantity>
), rocky and mixed rocky-sandy shores, also on coral reefs and associated coral rubble habitats (reef flats); typically, under large rocks and coral rubble (e.g.
<bibRefCitation id="8F2E4B0DFF98FFA4E0CD79EEF0B7FA16" author="Banner, D. M. &amp; Banner, A. H." box="[664,928,1473,1499]" pageId="3" pageNumber="254" pagination="1 - 357" refId="ref12237" refString="Banner, D. M. &amp; Banner, A. H. (1982) The Alpheid Shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
;
<bibRefCitation id="8F2E4B0DFF98FFA4E1F979EEF7F5FA16" author="Corfield, J. L. &amp; Alexander, C. G." box="[940,1250,1472,1499]" pageId="3" pageNumber="254" pagination="675 - 687" refId="ref12711" refString="Corfield, J. L. &amp; Alexander, C. G. (1995) The distribution of two species of alpheid shrimp, Alpheus edwardsii and A. lobidens, on a tropical beach. Journal of the Marine Biological Association of the United Kingdom, 75, 675 - 687. https: // doi. org / 10.1017 / S 0025315400039096" type="journal article" year="1995">Corfield &amp; Alexander 1995</bibRefCitation>
; present study).
</paragraph>
</subSubSection>
<subSubSection id="A3A56577FF98FFA4E29279CAF3E6F846" pageId="3" pageNumber="254" type="discussion">
<paragraph id="EB0036FCFF98FFA4E29279CAF237F97E" blockId="3.[151,1437,1112,1931]" pageId="3" pageNumber="254">
<emphasis id="D9CBEAEEFF98FFA4E29279CAF223FA32" bold="true" box="[199,308,1509,1535]" pageId="3" pageNumber="254">Remarks</emphasis>
. Without a comprehensive revision of the
<taxonomicName id="2CBF4D7FFF98FFA4E17F79CAF0ADFA32" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[810,954,1509,1535]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="D9CBEAEEFF98FFA4E17F79CAF0ADFA32" box="[810,954,1509,1535]" italics="true" pageId="3" pageNumber="254">A. edwardsii</emphasis>
</taxonomicName>
species complex (cf.
<bibRefCitation id="8F2E4B0DFF98FFA4E6EE79CAF3C4F9E9" author="Anker, A. &amp; De Grave, S." pageId="3" pageNumber="254" pagination="343 - 454" refId="ref11545" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
), it is impossible to apply a correct species name to the likely more than one species present in the Arabian Gulf, at least in its more biodiverse eastern part (see also
<bibRefCitation id="8F2E4B0DFF98FFA4E1597A02F713F98A" author="Dehghani, A. &amp; Sari A. &amp; Naderloo, R." box="[780,1028,1581,1607]" pageId="3" pageNumber="254" pagination="479 - 504" refId="ref13421" refString="Dehghani, A., Sari A. &amp; Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
Dehghani
<emphasis id="D9CBEAEEFF98FFA4E12B7A01F0A2F98A" box="[894,949,1581,1607]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
2019a
</bibRefCitation>
). In addition, many literature records of
<taxonomicName id="2CBF4D7FFF98FFA4E2E27A7EF253F9A6" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[183,324,1617,1643]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="D9CBEAEEFF98FFA4E2E27A7EF253F9A6" box="[183,324,1617,1643]" italics="true" pageId="3" pageNumber="254">A. edwardsii</emphasis>
</taxonomicName>
are unclear and may refer to closely or even not closely related taxa, exemplified by the regional record of
<bibRefCitation id="8F2E4B0DFF98FFA4E3507A5AF104F942" author="Al-Maliky, T. H. Y. &amp; Al-Khafaji, K. K. &amp; Khalaf, T. A." box="[261,531,1653,1679]" pageId="3" pageNumber="254" pagination="47 - 53" refId="ref11168" refString="Al-Maliky, T. H. Y., Al-Khafaji, K. K. &amp; Khalaf, T. A. (2017) New records of the snapping shrimp Alpheus edwardsii (Audouin, 1826) (Crustacea: Alpheoidea) in Basarh, Iraq. Arthropods, 6, 47 - 53." type="journal article" year="2017">
Al-Maliky
<emphasis id="D9CBEAEEFF98FFA4E3D17A59F2AAF942" box="[388,445,1653,1679]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
(2017)
</bibRefCitation>
, who erroneously reported
<taxonomicName id="2CBF4D7FFF98FFA4E11F7A5AF0CFF942" baseAuthorityName="Audouin" baseAuthorityYear="1826" box="[842,984,1653,1679]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="D9CBEAEEFF98FFA4E11F7A5AF0CFF942" box="[842,984,1653,1679]" italics="true" pageId="3" pageNumber="254">A. edwardsii</emphasis>
</taxonomicName>
as
<taxonomicName id="2CBF4D7FFF98FFA4E6557A5AF797F942" authorityName="De Haan" authorityYear="1849" box="[1024,1152,1653,1679]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="lobidens">
<emphasis id="D9CBEAEEFF98FFA4E6557A5AF797F942" box="[1024,1152,1653,1679]" italics="true" pageId="3" pageNumber="254">A. lobidens</emphasis>
</taxonomicName>
from Shatt Al
<collectingRegion id="297BF81EFF98FFA4E7797A5AF67BF942" box="[1324,1388,1653,1679]" country="Iraq" name="Al Basrah" pageId="3" pageNumber="254">Basra</collectingRegion>
and Faw in
<collectingCountry id="93A8766CFF98FFA4E2BE7AB6F20BF97E" box="[235,284,1689,1715]" name="Iraq" pageId="3" pageNumber="254">Iraq</collectingCountry>
.
</paragraph>
<paragraph id="EB0036FCFF98FFA4E2927A92F3E6F846" blockId="3.[151,1437,1112,1931]" pageId="3" pageNumber="254">
The photographed male specimen from
<collectingCountry id="93A8766CFF98FFA4E0D97A92F1C9F91A" box="[652,734,1725,1751]" name="Kuwait" pageId="3" pageNumber="254">Kuwait</collectingCountry>
(
<figureCitation id="73842A79FF98FFA4E0BA7A92F023F91A" box="[751,820,1725,1751]" captionStart="FIGURE 2" captionStartId="4.[151,250,1702,1727]" captionTargetBox="[151,1435,181,1677]" captionTargetId="figure@4.[151,1435,181,1677]" captionTargetPageId="4" captionText="FIGURE 2. Alpheusedwardsii (Audouin, 1826) [sensu lato]:A—male from Kubbar Island, Kuwait (OUMNH.ZC.2020.01.018), dorsal view; B—same, major chela with dactylus opened. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.3909081" httpUri="https://zenodo.org/record/3909081/files/figure.png" pageId="3" pageNumber="254">Fig. 2</figureCitation>
) is characterised by the overall brown to brown-green colour, the presence of narrow longitudinal white streaks on the pleon flanks and two small dark spots on the dorsal surface of the third pleonite, as well as by the brightly coloured major chela, with conspicuous blue, yellow, olivegreen and red-orange areas on the mesial face of the palm. This colour pattern seems to correspond to that of the female from
<collectingCountry id="93A8766CFF98FFA4E37F7B62F240F8AA" box="[298,343,1869,1895]" name="Iran" pageId="3" pageNumber="254">Iran</collectingCountry>
illustrated by
<bibRefCitation id="8F2E4B0DFF98FFA4E3AE7B62F014F8AA" author="Dehghani, A. &amp; Sari A. &amp; Naderloo, R." box="[507,771,1869,1895]" pageId="3" pageNumber="254" pagination="479 - 504" refId="ref13421" refString="Dehghani, A., Sari A. &amp; Naderloo, R. (2019 a) Annotated checklist of the snapping shrimps of the genus Alpheus Fabricius, 1798 (Crustacea: Decapoda: Alpheidae) from the Persian Gulf and Gulf of Oman, Iran. Zootaxa, 4544 (4), 479 - 504. https: // doi. org / 10.11646 / zootaxa. 4544.4.2" type="journal article" year="2019">
Dehghani
<emphasis id="D9CBEAEEFF98FFA4E0257B61F1BEF8AA" box="[624,681,1869,1895]" italics="true" pageId="3" pageNumber="254">et al.</emphasis>
(2019a
</bibRefCitation>
: fig. 3B), although the Iranian specimen is much paler and greener.
</paragraph>
</subSubSection>
</treatment>
</document>