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<document id="AF6B915F6E859F71C4E793BF56D91489" ID-DOI="10.11646/zootaxa.5419.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10798671" ID-ZooBank="E1084FA8-2920-4A64-A9A9-5BECDFE15764" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709929310234" checkinUser="plazi" docAuthor="Poorani, J., Anuradha, C., Thanigairaj, R. &amp; Mol, P. Prashina" docDate="2024" docId="03C0BB05FFB3FD1EB3CE9DE1FD51FDDC" docLanguage="en" docName="zootaxa.5419.4.3.pdf" docOrigin="Zootaxa 5419 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scymnus (Pullus) syoitii Sasaji 1971" docType="treatment" docVersion="2" lastPageNumber="543" masterDocId="FFF9C37DFFBEFD0CB359992EFFC9FFBD" masterDocTitle="Coccinellid predators of mealybugs infesting banana in South India, including a new species and a new record of Scymnus Kugelann (Coleoptera: Coccinellidae), with notes on other natural enemies" masterLastPageNumber="544" masterPageNumber="525" pageNumber="538" updateTime="1709929739591" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5AC718E48A9412984A326BE5627381B4">Coccinellid predators of mealybugs infesting banana in South India, including a new species and a new record of Scymnus Kugelann (Coleoptera: Coccinellidae), with notes on other natural enemies</mods:title>
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<mods:affiliation id="3646CC763DC59BBF26D44B8F890284C8">ICAR-National Research Centre for Banana, Thogamalai Road, Thayanur Post, Tiruchirappalli- 620102, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="27CE45E4B221EE7EA2C4977085A3182A">Anuradha, C.</mods:namePart>
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<mods:affiliation id="AF4CA9D8FEB4C49C7C495CD3B9DF1388">ICAR-National Research Centre for Banana, Thogamalai Road, Thayanur Post, Tiruchirappalli- 620102, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="0F036D633A1F77CD22EB8A30589064DE">Thanigairaj, R.</mods:namePart>
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<mods:affiliation id="46A15E99A1D6D367CD922F024656B4BD">ICAR-National Research Centre for Banana, Thogamalai Road, Thayanur Post, Tiruchirappalli- 620102, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="C1C68F587A8D666E361C8341622DDAC7">Mol, P. Prashina</mods:namePart>
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<mods:affiliation id="91C2E00EB708FD0869A5D44FD08AC8BB">ICAR-National Research Centre for Banana, Thogamalai Road, Thayanur Post, Tiruchirappalli- 620102, Tamil Nadu, India</mods:affiliation>
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<subSubSection id="C3735998FFB3FD01B3CE9DE1FD90FB55" box="[151,601,1230,1257]" pageId="13" pageNumber="538" type="nomenclature">
<paragraph id="8BD60A13FFB3FD01B3CE9DE1FD90FB55" blockId="13.[151,601,1230,1293]" box="[151,601,1230,1257]" pageId="13" pageNumber="538">
<heading id="D09EBD7FFFB3FD01B3CE9DE1FD90FB55" bold="true" box="[151,601,1230,1257]" fontSize="11" level="1" pageId="13" pageNumber="538" reason="1">
<taxonomicName id="4C697190FFB3FD01B3CE9DE1FD90FB55" authority="Sasaji, 1971" authorityName="Sasaji" authorityYear="1971" box="[151,601,1230,1257]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="538" phylum="Arthropoda" rank="species" species="syoitii" subGenus="Pullus">
<emphasis id="B91DD601FFB3FD01B3CE9DE1FD90FB55" bold="true" box="[151,601,1230,1257]" pageId="13" pageNumber="538">
<emphasis id="B91DD601FFB3FD01B3CE9DE1FECDFB54" bold="true" box="[151,260,1231,1257]" italics="true" pageId="13" pageNumber="538">Scymnus</emphasis>
(
<emphasis id="B91DD601FFB3FD01B24C9DE0FEAAFB54" bold="true" box="[277,355,1230,1257]" italics="true" pageId="13" pageNumber="538">Pullus</emphasis>
)
<emphasis id="B91DD601FFB3FD01B22C9DE1FE74FB54" bold="true" box="[373,445,1231,1257]" italics="true" pageId="13" pageNumber="538">syoitii</emphasis>
<bibRefCitation id="EFF877E2FFB3FD01B29D9DE0FD90FB55" author="Sasaji, H." box="[452,601,1230,1256]" pageId="13" pageNumber="538" refId="ref7522" refString="Sasaji, H. (1971) Fauna Japonica, Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp." type="book" year="1971">Sasaji, 1971</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C3735998FFB3FD01B3CE9DDDFEC2FAB0" box="[151,267,1267,1293]" pageId="13" pageNumber="538" type="description">
<paragraph id="8BD60A13FFB3FD01B3CE9DDDFEC2FAB0" blockId="13.[151,601,1230,1293]" box="[151,267,1267,1293]" pageId="13" pageNumber="538">
(
<figureCitation id="13521696FFB3FD01B3C69DDDFF2EFAB0" box="[159,231,1267,1293]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[244,1343,181,1919]" captionTargetId="figure-42@14.[244,1343,181,1919]" captionTargetPageId="14" captionText="FIGURE 8. Life stages of Scymnus (Pullus) syoitii Sasaji: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult." figureDoi="http://doi.org/10.5281/zenodo.10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="13" pageNumber="538">Figs 8</figureCitation>
,
<figureCitation id="13521696FFB3FD01B3AD9DDDFECAFAB0" box="[244,259,1267,1293]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3735998FFB3FD01B3CE9C19FDF7F966" pageId="13" pageNumber="538" type="reference_group">
<paragraph id="8BD60A13FFB3FD01B3CE9C19FD80FAF2" blockId="13.[151,585,1335,1359]" box="[151,585,1335,1359]" pageId="13" pageNumber="538">
<treatmentCitationGroup id="AB792D3DFFB3FD01B3CE9C19FD80FAF2" box="[151,585,1335,1359]" pageId="13" pageNumber="538">
<treatmentCitation id="0AC82C02FFB3FD01B3CE9C19FD8CFAF2" author="Sasaji, H." box="[151,581,1335,1359]" page="146" pageId="13" pageNumber="538" year="1971">
<taxonomicName id="4C697190FFB3FD01B3CE9C19FD8CFAF2" authority="Sasaji, 1971: 146" authorityName="Sasaji" authorityPageNumber="146" authorityYear="1971" box="[151,581,1335,1359]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="538" phylum="Arthropoda" rank="species" species="syoitii" subGenus="Pullus">
<emphasis id="B91DD601FFB3FD01B3CE9C19FF39FAF3" box="[151,240,1335,1358]" italics="true" pageId="13" pageNumber="538">Scymnus</emphasis>
(
<emphasis id="B91DD601FFB3FD01B3A69C19FE89FAF2" box="[255,320,1335,1359]" italics="true" pageId="13" pageNumber="538">Pullus</emphasis>
)
<emphasis id="B91DD601FFB3FD01B2179C16FE45FAF2" box="[334,396,1336,1359]" italics="true" pageId="13" pageNumber="538">syoitii</emphasis>
<bibRefCitation id="EFF877E2FFB3FD01B2CB9C19FD8CFAF2" author="Sasaji, H." box="[402,581,1335,1359]" pageId="13" pageNumber="538" refId="ref7522" refString="Sasaji, H. (1971) Fauna Japonica, Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp." type="book" year="1971">Sasaji, 1971: 146</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD60A13FFB3FD01B3CE9C53FDF7F966" blockId="13.[151,1437,1405,2008]" pageId="13" pageNumber="538">
<emphasis id="B91DD601FFB3FD01B3CE9C53FEA1FA2A" bold="true" box="[151,360,1405,1431]" pageId="13" pageNumber="538">Brief description.</emphasis>
Length:
<quantity id="4C91A7F6FFB3FD01B2979C53FDBDFA25" box="[462,628,1405,1432]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="13" pageNumber="538" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.10–1.30 mm</quantity>
; width:
<quantity id="4C91A7F6FFB3FD01B1969C53FCBCFA25" box="[719,885,1405,1432]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="13" pageNumber="538" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.70–0.90 mm</quantity>
. Form narrow and elongate oval, dorsum weakly convex with dense white pubescence; lateral margins of elytra with few long, erect setae interspersed with shorter ones. Dorsum usually pale yellow (
<figureCitation id="13521696FFB3FD01B1469CEBFD40FA62" box="[543,649,1477,1503]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[244,1343,181,1919]" captionTargetId="figure-42@14.[244,1343,181,1919]" captionTargetPageId="14" captionText="FIGURE 8. Life stages of Scymnus (Pullus) syoitii Sasaji: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult." figureDoi="http://doi.org/10.5281/zenodo.10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="13" pageNumber="538">Fig. 8f, h</figureCitation>
), rarely darker reddish brown (
<figureCitation id="13521696FFB3FD01B0B89CEBFBFAFA62" box="[993,1075,1477,1503]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[244,1343,181,1919]" captionTargetId="figure-42@14.[244,1343,181,1919]" captionTargetPageId="14" captionText="FIGURE 8. Life stages of Scymnus (Pullus) syoitii Sasaji: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult." figureDoi="http://doi.org/10.5281/zenodo.10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="13" pageNumber="538">
Fig.
<quantity id="4C91A7F6FFB3FD01B74C9CEBFBFAFA62" box="[1045,1075,1477,1503]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="13" pageNumber="538" unit="g" value="8.0">8g</quantity>
</figureCitation>
), basal part of elytra with darker infuscate area. Head (
<figureCitation id="13521696FFB3FD01B2CB9CC7FE2DF9BE" box="[402,484,1513,1539]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9a</figureCitation>
) broad with small eyes, interocular distance a little more than 3/5
<superScript id="7C1CA75BFFB3FD01B7B99CC7FB24FA4A" attach="left" box="[1248,1261,1513,1527]" fontSize="6" pageId="13" pageNumber="538">th</superScript>
of head width. Antenna (
<figureCitation id="13521696FFB3FD01B25C9F23FE9FF99A" box="[261,342,1549,1575]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9b</figureCitation>
) with 11 antennomeres. Prosternal carinae distinctly and somewhat abruptly divergent posteriorly around middle (
<figureCitation id="13521696FFB3FD01B2089F1FFE6FF9F6" box="[337,422,1585,1611]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9c</figureCitation>
). Abdominal postcoxal line complete, strongly curved, anteriorly punctate, smooth and impunctate near posterior margin (
<figureCitation id="13521696FFB3FD01B1449F7BFD45F9D2" box="[541,652,1621,1647]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9d, e</figureCitation>
). Male genitalia (
<figureCitation id="13521696FFB3FD01B00A9F7BFC74F9D2" box="[851,957,1621,1647]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9f–h</figureCitation>
) with penis having a distinct capsule (
<figureCitation id="13521696FFB3FD01B6369F7BFF7FF92E" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9h</figureCitation>
), penis guide elongate triangular in ventral view (
<figureCitation id="13521696FFB3FD01B1BA9F57FCFCF92E" box="[739,821,1657,1683]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">
Fig.
<quantity id="4C91A7F6FFB3FD01B04E9F57FCFCF92E" box="[791,821,1657,1683]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="13" pageNumber="538" unit="g" value="9.0">9g</quantity>
</figureCitation>
), parameres and penis guide subequal in length, apices of parameres with few short hairs (
<figureCitation id="13521696FFB3FD01B1439FB3FD4AF90A" box="[538,643,1693,1719]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9f, g</figureCitation>
). Female genitalia (
<figureCitation id="13521696FFB3FD01B0079FB3FC61F90A" box="[862,936,1693,1719]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9i</figureCitation>
) and spermatheca (
<figureCitation id="13521696FFB3FD01B7D99FB3FB02F90A" box="[1152,1227,1693,1719]" captionStart="FIGURE 9" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[233,1351,181,1907]" captionTargetId="figure-18@15.[233,1354,181,1919]" captionTargetPageId="15" captionText="FIGURE 9. Scymnus (Pullus) syoitii Sasaji: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="13" pageNumber="538">Fig. 9j</figureCitation>
) as illustrated. See
<bibRefCitation id="EFF877E2FFB3FD01B3CE9FEFFEE2F966" author="Sasaji, H." box="[151,299,1729,1755]" pageId="13" pageNumber="538" refId="ref7522" refString="Sasaji, H. (1971) Fauna Japonica, Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp." type="book" year="1971">Sasaji (1971)</bibRefCitation>
for detailed description.
</paragraph>
</subSubSection>
<subSubSection id="C3735998FFB3FD01B39E9FC8FAA3F89E" pageId="13" pageNumber="538" type="description">
<paragraph id="8BD60A13FFB3FD01B39E9FC8FAA3F89E" blockId="13.[151,1437,1405,2008]" pageId="13" pageNumber="538">
<emphasis id="B91DD601FFB3FD01B39E9FC8FE44F8BD" bold="true" box="[199,397,1766,1792]" italics="true" pageId="13" pageNumber="538">Immature stages.</emphasis>
Larva (
<figureCitation id="13521696FFB3FD01B2BD9FCBFD99F942" box="[484,592,1765,1791]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[244,1343,181,1919]" captionTargetId="figure-42@14.[244,1343,181,1919]" captionTargetPageId="14" captionText="FIGURE 8. Life stages of Scymnus (Pullus) syoitii Sasaji: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult." figureDoi="http://doi.org/10.5281/zenodo.10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="13" pageNumber="538">Fig. 8a–d</figureCitation>
) creamy white with white waxy filaments, always found in association with banana root mealybug colonies. Pupa (
<figureCitation id="13521696FFB3FD01B1139E27FD7CF89E" box="[586,693,1801,1827]" captionStart="FIGURE 8" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[244,1343,181,1919]" captionTargetId="figure-42@14.[244,1343,181,1919]" captionTargetPageId="14" captionText="FIGURE 8. Life stages of Scymnus (Pullus) syoitii Sasaji: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult." figureDoi="http://doi.org/10.5281/zenodo.10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="13" pageNumber="538">Fig. 8d, e</figureCitation>
) elongate, creamy white, somewhat bereft of waxy filaments.
</paragraph>
</subSubSection>
<subSubSection id="C3735998FFB3FD01B39E9E03FCE8F80E" pageId="13" pageNumber="538" type="materials_examined">
<paragraph id="8BD60A13FFB3FD01B39E9E03FCE8F80E" blockId="13.[151,1437,1405,2008]" pageId="13" pageNumber="538">
<emphasis id="B91DD601FFB3FD01B39E9E03FE60F8F5" bold="true" box="[199,425,1837,1864]" italics="true" pageId="13" pageNumber="538">Material examined.</emphasis>
India:
<collectingRegion id="49ADC4F1FFB3FD01B2A39E03FD4DF8FA" box="[506,644,1837,1863]" country="India" name="Tamil Nadu" pageId="13" pageNumber="538">Tamil Nadu</collectingRegion>
: Mullikarumbur,
<geoCoordinate id="EE5D6CD4FFB3FD01B0119E03FC3DF8FA" box="[840,1012,1837,1863]" degrees="10" direction="north" minutes="49" orientation="latitude" pageId="13" pageNumber="538" precision="1" seconds="34.39" value="10.82622">N10°49’34.39”</geoCoordinate>
<geoCoordinate id="EE5D6CD4FFB3FD01B0A59E03FB7FF8FA" box="[1020,1206,1837,1863]" degrees="078" direction="east" minutes="36" orientation="longitude" pageId="13" pageNumber="538" precision="1" seconds="41.69" value="78.61158">E078°36’41.69”</geoCoordinate>
, Ex. Root mealybug complex on banana, 18.ix.21, 22.xii.21,
<collectionCode id="ED7892D6FFB3FD01B1079E7CFDB8F8D1" box="[606,625,1874,1900]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="538">R</collectionCode>
. Thanigairaj (
<collectionCode id="ED7892D6FFB3FD01B0499E7FFC66F8D6" box="[784,943,1873,1899]" pageId="13" pageNumber="538">ICAR-NRCB</collectionCode>
: 9);
<collectingCountry id="F37E4A83FFB3FD01B0B89E7CFBFCF8D1" box="[993,1077,1874,1900]" name="India" pageId="13" pageNumber="538">INDIA</collectingCountry>
: I.I.S.C., Bangalore,
<collectingRegion id="49ADC4F1FFB3FD01B6789E7FFA5DF8D6" box="[1313,1428,1873,1899]" country="India" name="Karnataka" pageId="13" pageNumber="538">Karnataka</collectingRegion>
/
<date id="FFD72CD3FFB3FD01B3CE9E5BFF3EF832" box="[151,247,1909,1935]" pageId="13" pageNumber="538" value="1992-07">vii.1992</date>
, Sp. no. 44/ex
<taxonomicName id="4C697190FFB3FD01B2C79E5BFD40F82D" authority="IIE" authorityName="IIE" box="[414,649,1909,1936]" class="Insecta" family="Pseudococcidae" genus="Planococcus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="538" phylum="Arthropoda" rank="species" species="citri">Planococcus citri IIE</taxonomicName>
22763/
<taxonomicName id="4C697190FFB3FD01B1869E5BFC8CF832" authorityName="Kugelann" authorityYear="1794" box="[735,837,1909,1935]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="538" phylum="Arthropoda" rank="genus">Scymnus</taxonomicName>
(P.) sp., det.
<collectionCode id="ED7892D6FFB3FD01B0819E58FC22F82D" box="[984,1003,1910,1936]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="538">R</collectionCode>
.G. Booth, 1993/Pres. by Int. Inst. Ent. B.M. 1993 (
<collectionCode id="ED7892D6FFB3FD01B2479EB7FEBDF80E" box="[286,372,1945,1971]" pageId="13" pageNumber="538">BMNH</collectionCode>
: 2M (genitalia dissected), 1 unsexed).
</paragraph>
</subSubSection>
<subSubSection id="C3735998FFB3FD1EB39E9E93FD51FDDC" lastPageId="18" lastPageNumber="543" pageId="13" pageNumber="538" type="distribution">
<paragraph id="8BD60A13FFB3FD01B39E9E93FC6EF86A" blockId="13.[151,1437,1405,2008]" box="[199,935,1981,2008]" pageId="13" pageNumber="538">
<emphasis id="B91DD601FFB3FD01B39E9E93FE93F865" bold="true" box="[199,346,1981,2008]" italics="true" pageId="13" pageNumber="538">Distribution.</emphasis>
<collectingCountry id="F37E4A83FFB3FD01B2389E90FE6DF865" box="[353,420,1982,2008]" name="Japan" pageId="13" pageNumber="538">Japan</collectingCountry>
;
<collectingCountry id="F37E4A83FFB3FD01B2E99E93FE23F86A" box="[432,490,1981,2007]" name="India" pageId="13" pageNumber="538">India</collectingCountry>
(
<collectingRegion id="49ADC4F1FFB3FD01B2A19E93FDB9F86A" box="[504,624,1981,2007]" country="India" name="Karnataka" pageId="13" pageNumber="538">Karnataka</collectingRegion>
;
<collectingRegion id="49ADC4F1FFB3FD01B1229E93FCC8F86A" box="[635,769,1981,2007]" country="India" name="Tamil Nadu" pageId="13" pageNumber="538">Tamil Nadu</collectingRegion>
) (new record).
</paragraph>
<caption id="DF165A9BFFB0FD02B3CE9EB9FD66F86E" ID-DOI="http://doi.org/10.5281/zenodo.10798693" ID-Zenodo-Dep="10798693" httpUri="https://zenodo.org/record/10798693/files/figure.png" pageId="14" pageNumber="539" startId="14.[151,250,1943,1967]" targetBox="[244,1343,181,1919]" targetPageId="14" targetType="figure">
<paragraph id="8BD60A13FFB0FD02B3CE9EB9FD66F86E" blockId="14.[151,1436,1943,2003]" pageId="14" pageNumber="539">
<emphasis id="B91DD601FFB0FD02B3CE9EB9FEDBF812" bold="true" box="[151,274,1943,1967]" pageId="14" pageNumber="539">FIGURE 8.</emphasis>
Life stages of
<taxonomicName id="4C697190FFB0FD02B2FE9EB6FD28F812" authority="Sasaji" authorityName="Sasaji" authorityYear="1971" box="[423,737,1943,1967]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="539" phylum="Arthropoda" rank="species" species="syoitii" subGenus="Pullus">
<emphasis id="B91DD601FFB0FD02B2FE9EB6FDC9F812" box="[423,512,1944,1967]" italics="true" pageId="14" pageNumber="539">Scymnus</emphasis>
(
<emphasis id="B91DD601FFB0FD02B1579EB9FD86F812" box="[526,591,1943,1967]" italics="true" pageId="14" pageNumber="539">Pullus</emphasis>
)
<emphasis id="B91DD601FFB0FD02B1049EB6FD52F812" box="[605,667,1944,1967]" italics="true" pageId="14" pageNumber="539">syoitii</emphasis>
Sasaji
</taxonomicName>
: a, c. larva feeding on banana root mealybug; b. larva on banana root; d. mature larvae and pupa in soil; e. pupa; f–h. adult.
</paragraph>
</caption>
<caption id="DF165A9BFFB1FD03B3CE9EB9FECFF84A" ID-DOI="http://doi.org/10.5281/zenodo.10798695" ID-Zenodo-Dep="10798695" httpUri="https://zenodo.org/record/10798695/files/figure.png" pageId="15" pageNumber="540" startId="15.[151,250,1943,1967]" targetBox="[233,1351,181,1907]" targetPageId="15" targetType="figure">
<paragraph id="8BD60A13FFB1FD03B3CE9EB9FECFF84A" blockId="15.[151,1437,1943,2039]" pageId="15" pageNumber="540">
<emphasis id="B91DD601FFB1FD03B3CE9EB9FED1F812" bold="true" box="[151,280,1943,1967]" pageId="15" pageNumber="540">FIGURE 9.</emphasis>
<taxonomicName id="4C697190FFB1FD03B27A9EB6FDA6F812" authority="Sasaji" authorityName="Sasaji" authorityYear="1971" box="[291,623,1943,1967]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="540" phylum="Arthropoda" rank="species" species="syoitii" subGenus="Pullus">
<emphasis id="B91DD601FFB1FD03B27A9EB6FEB5F812" box="[291,380,1944,1967]" italics="true" pageId="15" pageNumber="540">Scymnus</emphasis>
(
<emphasis id="B91DD601FFB1FD03B2C99EB9FE18F812" box="[400,465,1943,1967]" italics="true" pageId="15" pageNumber="540">Pullus</emphasis>
)
<emphasis id="B91DD601FFB1FD03B2BD9EB6FDEBF812" box="[484,546,1944,1967]" italics="true" pageId="15" pageNumber="540">syoitii</emphasis>
Sasaji
</taxonomicName>
: a. head; b. antenna; c. prosternal carinae; d. abdomen, male; e. abdominal postcoxal line; f–h. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis; i. female genitalia; j. spermatheca (enlarged).
</paragraph>
</caption>
<caption id="DF165A9BFFAEFD1CB3CE9E2CFC47F8DF" ID-DOI="http://doi.org/10.5281/zenodo.10798699" ID-Zenodo-Dep="10798699" httpUri="https://zenodo.org/record/10798699/files/figure.png" pageId="16" pageNumber="541" startId="16.[151,250,1794,1818]" targetBox="[151,1435,181,1769]" targetPageId="16" targetType="figure">
<paragraph id="8BD60A13FFAEFD1CB3CE9E2CFC47F8DF" blockId="16.[151,1437,1794,1891]" pageId="16" pageNumber="541">
<emphasis id="B91DD601FFAEFD1CB3CE9E2CFEE8F8A7" bold="true" box="[151,289,1794,1818]" pageId="16" pageNumber="541">FIGURE 10.</emphasis>
Parasitoids of banana fruit mealybugs: a.
<taxonomicName id="4C697190FFAEFD1CB18A9E2CFBC6F8A7" authority="(Subba Rao)" baseAuthorityName="Subba Rao" baseAuthorityYear="1967" box="[723,1039,1794,1818]" class="Insecta" family="Encyrtidae" genus="Anagyrus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B91DD601FFAEFD1CB18A9E2CFC4DF8A7" box="[723,900,1794,1818]" italics="true" pageId="16" pageNumber="541">Anagyrus indicus</emphasis>
(Subba Rao)
</taxonomicName>
; b.
<taxonomicName id="4C697190FFAEFD1CB76C9E2CFEC3F883" authority="Shafee, Alam &amp; Agarwal" authorityName="Shafee, Alam &amp; Agarwal" authorityYear="1975" class="Insecta" family="Encyrtidae" genus="Gyranusoidea" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="indica">
<emphasis id="B91DD601FFAEFD1CB76C9E2CFAC2F8A7" box="[1077,1291,1794,1818]" italics="true" pageId="16" pageNumber="541">Gyranusoidea indica</emphasis>
Shafee, Alam &amp; Agarwal
</taxonomicName>
; c.
<taxonomicName id="4C697190FFAEFD1CB2779E08FE7CF883" box="[302,437,1830,1854]" class="Insecta" family="Encyrtidae" genus="Anagyrus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAEFD1CB2779E08FE46F880" box="[302,399,1830,1853]" italics="true" pageId="16" pageNumber="541">Anagyrus</emphasis>
sp.
</taxonomicName>
; d.
<taxonomicName id="4C697190FFAEFD1CB2829E08FDA9F883" box="[475,608,1830,1854]" class="Insecta" family="Encyrtidae" genus="Anagyrus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAEFD1CB2829E08FDF5F880" box="[475,572,1830,1853]" italics="true" pageId="16" pageNumber="541">Anagyrus</emphasis>
sp.
</taxonomicName>
emerging from mummified mealybug; e.
<taxonomicName id="4C697190FFAEFD1CB7559E08FB0BF883" box="[1036,1218,1830,1854]" class="Insecta" family="Encyrtidae" genus="Gyranusoidea" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAEFD1CB7559E08FB55F883" box="[1036,1180,1830,1854]" italics="true" pageId="16" pageNumber="541">Gyranusoidea</emphasis>
sp.
</taxonomicName>
; f.
<taxonomicName id="4C697190FFAEFD1CB7BA9E08FEC1F8DF" authority="(Cameron)" baseAuthorityName="Cameron" baseAuthorityYear="1886" class="Insecta" family="Encyrtidae" genus="Blepyrus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B91DD601FFAEFD1CB7BA9E08FA54F883" box="[1251,1437,1830,1854]" italics="true" pageId="16" pageNumber="541">Blepyrus insularis</emphasis>
(Cameron)
</taxonomicName>
; g.
<taxonomicName id="4C697190FFAEFD1CB2759E64FE19F8DF" box="[300,464,1866,1890]" class="Insecta" family="Signiphoridae" genus="Chartocerus" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAEFD1CB2759E64FE62F8DF" box="[300,427,1866,1890]" italics="true" pageId="16" pageNumber="541">Chartocerus</emphasis>
sp.
</taxonomicName>
; h.
<taxonomicName id="4C697190FFAEFD1CB2AD9E64FC43F8DF" authority="(Girault)" baseAuthorityName="Girault" box="[500,906,1866,1890]" class="Insecta" family="Pteromalidae" genus="Promuscidea" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="541" phylum="Arthropoda" rank="species" species="unfasciativentris">
<emphasis id="B91DD601FFAEFD1CB2AD9E64FCE1F8DF" box="[500,808,1866,1890]" italics="true" pageId="16" pageNumber="541">Promuscidea unfasciativentris</emphasis>
(Girault)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF165A9BFFAFFD1DB3CE9EB6FB63F845" ID-DOI="http://doi.org/10.5281/zenodo.10798701" ID-Zenodo-Dep="10798701" httpUri="https://zenodo.org/record/10798701/files/figure.png" pageId="17" pageNumber="542" startId="17.[151,250,1944,1968]" targetBox="[184,1404,181,1920]" targetPageId="17" targetType="figure">
<paragraph id="8BD60A13FFAFFD1DB3CE9EB6FB63F845" blockId="17.[151,1436,1944,2041]" pageId="17" pageNumber="542">
<emphasis id="B91DD601FFAFFD1DB3CE9EB6FED4F80D" bold="true" box="[151,285,1944,1968]" pageId="17" pageNumber="542">FIGURE 11.</emphasis>
Parasitoids of banana mealybugs: a.
<taxonomicName id="4C697190FFAFFD1DB1D69EB7FC57F80D" authority="Compere" authorityName="Compere" authorityYear="1937" box="[655,926,1944,1968]" class="Insecta" family="Encyrtidae" genus="Aenasius" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="advena">
<emphasis id="B91DD601FFAFFD1DB1D69EB7FCF0F80D" box="[655,825,1944,1968]" italics="true" pageId="17" pageNumber="542">Aenasius advena</emphasis>
Compere
</taxonomicName>
; b.
<taxonomicName id="4C697190FFAFFD1DB0999EB7FB3CF80D" authority="(Girault)" baseAuthorityName="Girault" baseAuthorityYear="1915" box="[960,1269,1944,1968]" class="Insecta" family="Encyrtidae" genus="Aenasius" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="arizonensis">
<emphasis id="B91DD601FFAFFD1DB0999EB7FB5DF80D" box="[960,1172,1945,1968]" italics="true" pageId="17" pageNumber="542">Aenasius arizonensis</emphasis>
(Girault)
</taxonomicName>
; c–h. parasitoids of Jack Beardsley mealybug: c.
<taxonomicName id="4C697190FFAFFD1DB2839E93FDA9F869" box="[474,608,1981,2004]" class="Insecta" family="Encyrtidae" genus="Anagyrus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAFFD1DB2839E93FDF2F869" box="[474,571,1981,2004]" italics="true" pageId="17" pageNumber="542">Anagyrus</emphasis>
sp.
</taxonomicName>
, female; d.
<taxonomicName id="4C697190FFAFFD1DB18F9E93FC94F869" box="[726,861,1981,2004]" class="Insecta" family="Encyrtidae" genus="Anagyrus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAFFD1DB18F9E93FCFEF869" box="[726,823,1981,2004]" italics="true" pageId="17" pageNumber="542">Anagyrus</emphasis>
sp.
</taxonomicName>
, male; e. mealybug mummies with parasitoid emergence holes; f. mummified Jack Beardsley mealybug; g.
<taxonomicName id="4C697190FFAFFD1DB1CC9ECEFCF0F845" box="[661,825,2016,2040]" class="Insecta" family="Signiphoridae" genus="Chartocerus" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAFFD1DB1CC9ECEFCDDF845" box="[661,788,2016,2040]" italics="true" pageId="17" pageNumber="542">Chartocerus</emphasis>
sp.
</taxonomicName>
, its hyperparasitoid; h.
<taxonomicName id="4C697190FFAFFD1DB77F9ECEFB63F845" box="[1062,1194,2016,2040]" class="Insecta" family="Platygastridae" genus="Allotropa" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="542" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFAFFD1DB77F9ECEFB4FF845" box="[1062,1158,2016,2040]" italics="true" pageId="17" pageNumber="542">Allotropa</emphasis>
sp.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BD60A13FFACFD1EB39E99B9FC2CFF68" blockId="18.[151,1437,150,609]" pageId="18" pageNumber="543">
<emphasis id="B91DD601FFACFD1EB39E99B9FE26FF0C" bold="true" box="[199,495,150,177]" italics="true" pageId="18" pageNumber="543">Prey / associated habitat.</emphasis>
Predatory on banana root mealybug complex (comprising
<taxonomicName id="4C697190FFACFD1EB7C199B9FF36FF68" authority="Williams" authorityName="Williams" authorityYear="1969" class="Insecta" family="Pseudococcidae" genus="Geococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="B91DD601FFACFD1EB7C199B9FADAFF0C" box="[1176,1299,151,177]" italics="true" pageId="18" pageNumber="543">Geococcus</emphasis>
?
<emphasis id="B91DD601FFACFD1EB67099B9FA54FF0C" box="[1321,1437,151,177]" italics="true" pageId="18" pageNumber="543">johorensis</emphasis>
Williams
</taxonomicName>
,
<taxonomicName id="4C697190FFACFD1EB2529995FE64FF68" box="[267,429,187,213]" class="Insecta" family="Pseudococcidae" genus="Geococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFACFD1EB2529995FE4FFF68" box="[267,390,187,213]" italics="true" pageId="18" pageNumber="543">Geococcus</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C697190FFACFD1EB2BA9995FDB7FF68" box="[483,638,187,213]" class="Insecta" family="Pseudococcidae" genus="Rhizoecus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFACFD1EB2BA9995FD9CFF68" box="[483,597,187,213]" italics="true" pageId="18" pageNumber="543">Rhizoecus</emphasis>
sp.
</taxonomicName>
);
<taxonomicName id="4C697190FFACFD1EB1CA9995FC91FF69" box="[659,856,187,213]" class="Insecta" family="Pseudococcidae" genus="Planococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="citri">
<emphasis id="B91DD601FFACFD1EB1CA9995FC91FF69" box="[659,856,187,213]" italics="true" pageId="18" pageNumber="543">Planococcus citri</emphasis>
</taxonomicName>
(label data).
</paragraph>
<paragraph id="8BD60A13FFACFD1EB39E99F1FD51FDDC" blockId="18.[151,1437,150,609]" pageId="18" pageNumber="543">
<emphasis id="B91DD601FFACFD1EB39E99F1FEC6FF44" bold="true" box="[199,271,223,249]" italics="true" pageId="18" pageNumber="543">Notes.</emphasis>
<bibRefCitation id="EFF877E2FFACFD1EB24F99F1FE63FF44" author="Sasaji, H." box="[278,426,223,249]" pageId="18" pageNumber="543" refId="ref7522" refString="Sasaji, H. (1971) Fauna Japonica, Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp." type="book" year="1971">Sasaji (1971)</bibRefCitation>
described it from Kyushu,
<collectingCountry id="F37E4A83FFACFD1EB18299F1FCD2FF44" box="[731,795,223,249]" name="Japan" pageId="18" pageNumber="543">Japan</collectingCountry>
[see the
<typeStatus id="54D2B4B1FFACFD1EB02799F1FC29FF44" box="[894,992,223,249]" pageId="18" pageNumber="543" type="holotype">holotype</typeStatus>
photo and details in Kyushu University Museum Database [Hiroyuki Sasaji Collection, Catalog no. SH005533, URL: http://db.museum.kyushu-u.ac.jp/ entomology/?wb=det&amp;cl=31&amp;it=5533]. It appears to be a unique and characteristic species of
<taxonomicName id="4C697190FFACFD1EB7889809FAFAFEFC" authorityName="Kugelann" authorityYear="1794" box="[1233,1331,295,321]" class="Insecta" family="Coccinellidae" genus="Scymnus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="genus">
<emphasis id="B91DD601FFACFD1EB7889809FAFAFEFC" box="[1233,1331,295,321]" italics="true" pageId="18" pageNumber="543">Scymnus</emphasis>
</taxonomicName>
in South
<collectingCountry id="F37E4A83FFACFD1EB3CE9865FF18FED8" box="[151,209,331,357]" name="India" pageId="18" pageNumber="543">India</collectingCountry>
with a small, distinctly narrow and elongate oval form, head with a wide frons and small, broadly separated eyes, and prosternal carinae that are distinctly divergent posteriorly and the male genitalia are also diagnostic. The specimens collected from
<collectingRegion id="49ADC4F1FFACFD1EB2AB98BDFDB3FE10" box="[498,634,403,429]" country="India" name="Tamil Nadu" pageId="18" pageNumber="543">Tamil Nadu</collectingRegion>
, fully match
<bibRefCitation id="EFF877E2FFACFD1EB05698BDFC7EFE10" author="Sasaji, H." box="[783,951,403,429]" pageId="18" pageNumber="543" refId="ref7522" refString="Sasaji, H. (1971) Fauna Japonica, Coccinellidae (Insecta: Coleoptera). Academic Press of Japan, Tokyo, 340 pp." type="book" year="1971">Sasaji’s (1971)</bibRefCitation>
description and the male genitalia also are almost identical to the illustrations given by Sasaji. The larvae were found to be predatory on banana root mealybug complex (comprising a mixture of
<emphasis id="B91DD601FFACFD1EB14B98F5FCD3FE48" box="[530,794,475,501]" italics="true" pageId="18" pageNumber="543">
<taxonomicName id="4C697190FFACFD1EB14B98F5FCDFFE48" authorityName="Williams" authorityYear="1969" box="[530,790,475,501]" class="Insecta" family="Pseudococcidae" genus="Geococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="johorensis">Geococcus?johorensis</taxonomicName>
,
</emphasis>
<taxonomicName id="4C697190FFACFD1EB07998F5FC76FE48" box="[800,959,475,501]" class="Insecta" family="Pseudococcidae" genus="Geococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFACFD1EB07998F5FC52FE48" box="[800,923,475,501]" italics="true" pageId="18" pageNumber="543">Geococcus</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C697190FFACFD1EB0A898F5FB43FE48" box="[1009,1162,475,501]" class="Insecta" family="Pseudococcidae" genus="Rhizoecus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91DD601FFACFD1EB0A898F5FBAAFE48" box="[1009,1123,475,501]" italics="true" pageId="18" pageNumber="543">Rhizoecus</emphasis>
sp.
</taxonomicName>
). Conspecific specimens collected in association with
<taxonomicName id="4C697190FFACFD1EB2B598D1FD7CFDA5" box="[492,693,511,537]" class="Insecta" family="Pseudococcidae" genus="Planococcus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="species" species="citri">
<emphasis id="B91DD601FFACFD1EB2B598D1FD7CFDA5" box="[492,693,511,537]" italics="true" pageId="18" pageNumber="543">Planococcus citri</emphasis>
</taxonomicName>
from Bangalore,
<collectingRegion id="49ADC4F1FFACFD1EB0DE98D1FC34FDA4" box="[903,1021,511,537]" country="India" name="Karnataka" pageId="18" pageNumber="543">Karnataka</collectingRegion>
, by the Commonwealth Institute of Biological Control-Indian Station (Bangalore) in the collections of BMNH were also examined. It is a new addition to the fauna of
<taxonomicName id="4C697190FFACFD1EB2189B69FE65FDDC" authorityName="Poorani &amp; Anuradha &amp; Thanigairaj &amp; Mol" authorityYear="2024" box="[321,428,583,609]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="543" phylum="Arthropoda" rank="tribe" tribe="Scymnini">Scymnini</taxonomicName>
of the Indian region.
</paragraph>
</subSubSection>
</treatment>
</document>
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