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<document id="02AC0F037F88C9F651B21D74EB0E98CE" ID-DOI="10.5281/zenodo.213946" ID-GBIF-Dataset="2a54e127-4d40-404e-ab7f-5b48f6e37e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="213946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564534622" checkinUser="plazi" docAuthor="Grisales, Diana, Wolff, Marta &amp; De, Claudio J. B." docDate="2012" docId="03BE87F85232D42AC0942259FC3BF97B" docLanguage="en" docName="zt03591p046.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fannia Robineau-Desvoidy" docType="treatment" docVersion="8" lastPageNumber="4" masterDocId="FF87FF805231D429C0032033FFBBFFA9" masterDocTitle="Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia" masterLastPageNumber="46" masterPageNumber="1" pageNumber="4" updateTime="1698320797981" updateUser="plazi">
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<mods:title id="4A930866C145D643F4D2B2187A4274C9">Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia</mods:title>
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<treatment id="03BE87F85232D42AC0942259FC3BF97B" ID-DOI="http://doi.org/10.5281/zenodo.6175037" ID-GBIF-Taxon="119557545" ID-Zenodo-Dep="6175037" LSID="urn:lsid:plazi:treatment:03BE87F85232D42AC0942259FC3BF97B" httpUri="http://treatment.plazi.org/id/03BE87F85232D42AC0942259FC3BF97B" lastPageNumber="4" pageId="3" pageNumber="4">
<subSubSection id="C30D65655232D42AC0942259FE57FD2D" box="[151,492,618,644]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BA836EE5232D42AC0942259FE57FD2D" blockId="3.[151,492,618,644]" box="[151,492,618,644]" pageId="3" pageNumber="4">
<heading id="D0E081825232D42AC0942259FE57FD2D" bold="true" box="[151,492,618,644]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<taxonomicName id="4C174D6D5232D42AC0942259FE57FD2D" ID-CoL="8YQF2" authority="Robineau-Desvoidy" authorityName="Robineau-Desvoidy" box="[151,492,618,644]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC0942259FE57FD2D" bold="true" box="[151,492,618,644]" pageId="3" pageNumber="4">
<emphasis id="B963EAFC5232D42AC0942259FF4AFD2D" bold="true" box="[151,241,618,644]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
Robineau-Desvoidy
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30D65655232D42AC0942281FED0FD4C" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BA836EE5232D42AC0942281FED0FD4C" blockId="3.[151,1436,689,742]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB0711C05232D42AC0942281FED0FD4C" pageId="3" pageNumber="4">
<taxonomicName id="4C174D6D5232D42AC0942281FD8BFD6E" ID-CoL="8YQF2" authority="Robineau-Desvoidy, 1830: 567" authorityName="Robineau-Desvoidy" authorityPageNumber="567" authorityYear="1830" box="[151,560,689,711]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC0942281FF5BFD6E" box="[151,224,690,711]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
<treatmentCitation id="0AB610FF5232D42AC0E92282FD8BFD6E" author="Robineau-Desvoidy" box="[234,560,689,711]" page="567" pageId="3" pageNumber="4" year="1830">
<bibRefCitation id="EF864B1F5232D42AC0E92282FE42FD6E" author="Robineau-Desvoidy" box="[234,505,689,711]" pageId="3" pageNumber="4" refString="Robineau-Desvoidy, A. J. B. (1830) Essai sur les Myodaires. Memories presentes par divers savans l'Academie Royale des Sciences de l'Institut de France (Sciences Mathematiques et Physiques), (2) 2, 813 pp. [6 June]." type="book" year="1830">Robineau-Desvoidy, 1830</bibRefCitation>
: 567
</treatmentCitation>
</taxonomicName>
. Type-species,
<taxonomicName id="4C174D6D5232D42AC2D22281FB35FD6E" ID-CoL="6HPXS" authority="Robineau-Desvoidy, 1830" authorityName="Robineau-Desvoidy" authorityYear="1830" box="[721,1166,689,711]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="saltatrix">
<emphasis id="B963EAFC5232D42AC2D22281FCC3FD6E" box="[721,888,689,711]" italics="true" pageId="3" pageNumber="4">Fannia saltatrix</emphasis>
<bibRefCitation id="EF864B1F5232D42AC3812282FB35FD6E" author="Robineau-Desvoidy" box="[898,1166,689,711]" pageId="3" pageNumber="4" refString="Robineau-Desvoidy, A. J. B. (1830) Essai sur les Myodaires. Memories presentes par divers savans l'Academie Royale des Sciences de l'Institut de France (Sciences Mathematiques et Physiques), (2) 2, 813 pp. [6 June]." type="book" year="1830">Robineau-Desvoidy, 1830</bibRefCitation>
</taxonomicName>
(mon.) =
<taxonomicName id="4C174D6D5232D42AC4FD2281FED3FD4C" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scalaris">
<emphasis id="B963EAFC5232D42AC4FD2281FA27FD6E" box="[1278,1436,689,711]" italics="true" pageId="3" pageNumber="4">Musca scalaris</emphasis>
<bibRefCitation id="EF864B1F5232D42AC0C422E3FED3FD4C" author="Fabricius" box="[199,360,720,742]" pageId="3" pageNumber="4" refString="Fabricius, J. C. (1794) Entomologia Systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vo l. 4. Impensis Christ. Gottl. Proft., Hafniae, 472 pp." type="book" year="1794">Fabricius, 1794</bibRefCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C30D65655232D42AC0942321FC71FB3B" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BA836EE5232D42AC0942321FC71FB3B" blockId="3.[151,1436,786,1746]" pageId="3" pageNumber="4">
<emphasis id="B963EAFC5232D42AC0942321FEAAFC82" bold="true" box="[151,273,786,811]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
sexually-dimorphic; male holoptic (except in some species,
<emphasis id="B963EAFC5232D42AC3BF2327FC5EFC82" box="[956,997,788,811]" italics="true" pageId="3" pageNumber="4">e.g.</emphasis>
the
<emphasis id="B963EAFC5232D42AC4152320FB22FC82" box="[1046,1177,787,811]" italics="true" pageId="3" pageNumber="4">petrocchiae</emphasis>
subgroup, Neotropical region),
<emphasis id="B963EAFC5232D42AC0F52305FEA6FCE7" box="[246,285,822,846]" italics="true" pageId="3" pageNumber="4">orb</emphasis>
absent (except in
<taxonomicName id="4C174D6D5232D42AC1E82304FD7AFCE7" box="[491,705,822,846]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5232D42AC1E82304FD7AFCE7" box="[491,705,822,846]" italics="true" pageId="3" pageNumber="4">Fannia canicularis</emphasis>
</taxonomicName>
, upper
<emphasis id="B963EAFC5232D42AC3142305FC85FCE7" box="[791,830,822,846]" italics="true" pageId="3" pageNumber="4">orb</emphasis>
present); female dichoptic, 2
<emphasis id="B963EAFC5232D42AC48C2305FB0DFCE7" box="[1167,1206,822,846]" italics="true" pageId="3" pageNumber="4">orb</emphasis>
, one reclinate upper and one lateroclinate lower seta, interfrontal setae absent. Arista bare or pubescent; 2
<emphasis id="B963EAFC5232D42AC4502368FB36FCDA" box="[1107,1165,859,883]" italics="true" pageId="3" pageNumber="4">kepst</emphasis>
; first pre-sutural
<emphasis id="B963EAFC5232D42AC54C2368FAD1FCDA" box="[1359,1386,859,883]" italics="true" pageId="3" pageNumber="4">dc</emphasis>
half length of second.
<emphasis id="B963EAFC5232D42AC15523B3FECCFC3E" box="[342,375,896,919]" italics="true" pageId="3" pageNumber="4">Dc</emphasis>
2:3 and 2 developed
<emphasis id="B963EAFC5232D42AC25C234CFDCEFC3F" box="[607,629,895,918]" italics="true" pageId="3" pageNumber="4">ia</emphasis>
(except in
<taxonomicName id="4C174D6D5232D42AC2F223B3FCC1FC3F" box="[753,890,894,919]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="quimbaya" status="sp. nov.">
<emphasis id="B963EAFC5232D42AC2F223B3FCC1FC3F" box="[753,890,894,919]" italics="true" pageId="3" pageNumber="4">F. quimbaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875232D42AC381234CFC60FC3F" box="[898,987,895,919]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B963EAFC5232D42AC381234CFC27FC3E" bold="true" box="[898,924,895,919]" pageId="3" pageNumber="4">sp</emphasis>
.
<emphasis id="B963EAFC5232D42AC3A9234CFC6FFC3E" bold="true" box="[938,980,895,919]" pageId="3" pageNumber="4">nov</emphasis>
.
</taxonomicNameLabel>
and in
<taxonomicName id="4C174D6D5232D42AC42E23B3FB39FC3F" box="[1069,1154,895,919]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pijao" status="sp. nov.">
<emphasis id="B963EAFC5232D42AC42E23B3FB39FC3F" box="[1069,1154,895,919]" italics="true" pageId="3" pageNumber="4">F. pijao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875232D42AC48A234CFB5AFC3F" box="[1161,1249,895,919]" pageId="3" pageNumber="4" rank="species">
<emphasis id="B963EAFC5232D42AC48A234CFB18FC3E" bold="true" box="[1161,1187,895,919]" pageId="3" pageNumber="4">sp</emphasis>
.
<emphasis id="B963EAFC5232D42AC4B2234CFB60FC3E" bold="true" box="[1201,1243,895,919]" pageId="3" pageNumber="4">nov</emphasis>
.
</taxonomicNameLabel>
, with
<emphasis id="B963EAFC5232D42AC52B234DFAF8FC3F" box="[1320,1347,894,918]" italics="true" pageId="3" pageNumber="4">dc</emphasis>
0:1 and 0:2 and 1 intra-alar); 2
<emphasis id="B963EAFC5232D42AC1AF2397FE49FC12" box="[428,498,932,955]" italics="true" pageId="3" pageNumber="4">prepm</emphasis>
, seta near spiracle two times longer; subcosta bowed; second anal vein strongly curved, imaginary extension of the anal veins meeting before wing margin; mid femur and tibia of male with modified ventral setae; hind tibia with 1
<emphasis id="B963EAFC5232D42AC26E23D8FDC0FBAA" box="[621,635,1003,1027]" italics="true" pageId="3" pageNumber="4">d</emphasis>
on apical third, on the same row as apical seta; male abdomen wider before posterior margin of abdominal segment 2; second abdominal segment in females distally wider than the third one; hypopygium of male usually with bacilliform process differentiated at base of cercal plate; aedeagus often membranous; ovipositor with two spiracles and 23 spermathecae (
<bibRefCitation id="EF864B1F5232D42AC3EF2465FB2DFBC7" author="Chillcott" box="[1004,1174,1110,1134]" pageId="3" pageNumber="4" refString="Chillcott, J. (1961) A Revision of the Neartic Species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, 92, 1 - 295." type="journal article" year="1961">Chillcott 1961</bibRefCitation>
;
<bibRefCitation id="EF864B1F5232D42AC4A52465FA2CFBC7" author="Rozkosny" box="[1190,1431,1110,1135]" pageId="3" pageNumber="4" refString="Rozkosny, R., Gregor, F. &amp; Pont, A. C. (1997) The European Fanniidae (Diptera). Acta Scientiarum Bohemicae Brno, 31, 1 - 80." type="journal article" year="1997">
Rozkošný
<emphasis id="B963EAFC5232D42AC523246BFAE7FBC7" box="[1312,1372,1110,1135]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="EF864B1F5232D42AC0942449FE11FB3B" author="Albuquerque" box="[151,426,1146,1171]" pageId="3" pageNumber="4" refString="Albuquerque, D. de O., Pamplona, D. &amp; de Carvalho, C. J. B. (1981) Contribuicao ao conhecimento dos Fannia R. D., 1830 da regiao Neotropical (Diptera, Fanniidae). Arquivo do Museu Nacional do Rio de Janeiro, 56, 9 - 34." type="journal article" year="1981">
Albuquerque
<emphasis id="B963EAFC5232D42AC133244FFEDAFB3A" box="[304,353,1147,1171]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
. 1981
</bibRefCitation>
;
<bibRefCitation id="EF864B1F5232D42AC1B52449FDC1FB3B" author="Dominguez" box="[438,634,1146,1170]" pageId="3" pageNumber="4" refString="Dominguez, M. C. (2007) Taxonomic revision of southern South America Fannia R. - D. Papeis Avulsos de Zoologia, 47, 289 - 347." type="journal article" year="2007">Domínguez 2007</bibRefCitation>
; Wendt &amp; de
<bibRefCitation id="EF864B1F5232D42AC3172449FC05FB3B" author="Carvalho" box="[788,958,1146,1170]" pageId="3" pageNumber="4" refString="Wendt, L. D. &amp; de Carvalho, C. J. B. (2009) Taxonomia de Fanniidae (Diptera) do sul do Brazil - II: Novas especies e chave de identificacao de Fannia Robineau - Desvoidy. Revista Brasileira de Entomologia, 53, 171 - 206." type="journal article" year="2009">Carvalho 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655232D42AC0C424ADFC3BF97B" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BA836EE5232D42AC0C424ADFAEAFA1B" blockId="3.[151,1436,786,1746]" pageId="3" pageNumber="4">
<emphasis id="B963EAFC5232D42AC0C424ADFEF4FB1E" bold="true" box="[199,335,1182,1207]" pageId="3" pageNumber="4">Comments.</emphasis>
<taxonomicName id="4C174D6D5232D42AC15424ACFE13FB1F" box="[343,424,1183,1206]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC15424ACFE13FB1F" box="[343,424,1183,1206]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
</taxonomicName>
is monophyletic (
<bibRefCitation id="EF864B1F5232D42AC27724ADFC56FB1F" author="Dominguez" box="[628,1005,1182,1206]" pageId="3" pageNumber="4" refString="Dominguez, M. C. &amp; Roig-Junent, S. (2008) A phylogeny of the family Fanniidae Schnabl (Insecta: Diptera: Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia. Invertebrate Systematics, 22, 563 - 587." type="journal article" year="2008">Domínguez &amp; Roig-Juñent 2008</bibRefCitation>
).
<bibRefCitation id="EF864B1F5232D42AC40324ADFB0FFB1F" author="Chillcott" box="[1024,1204,1182,1206]" pageId="3" pageNumber="4" refString="Chillcott, J. (1961) A Revision of the Neartic Species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, 92, 1 - 295." type="journal article" year="1961">Chillcott (1961)</bibRefCitation>
grouped the species of
<taxonomicName id="4C174D6D5232D42AC0B724F7FEBEFB72" box="[180,261,1220,1243]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC0B724F7FEBEFB72" box="[180,261,1220,1243]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
</taxonomicName>
in the Nearctic region based on the morphology of the male and female terminalia. Later,
<bibRefCitation id="EF864B1F5232D42AC4EC24F1FEB3FB57" pageId="3" pageNumber="4" refString="Albuquerque, D. de O., Pamplona, D. &amp; de Carvalho, C. J. B. (1981) Contribuicao ao conhecimento dos Fannia R. D., 1830 da regiao Neotropical (Diptera, Fanniidae). Arquivo do Museu Nacional do Rio de Janeiro, 56, 9 - 34." type="journal article">
Albuquerque
<emphasis id="B963EAFC5232D42AC58B24F7FF0FFB57" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
(1981)
</bibRefCitation>
recognized eight
<taxonomicName id="4C174D6D5232D42AC1DB24D4FD92FB57" box="[472,553,1255,1278]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC1DB24D4FD92FB57" box="[472,553,1255,1278]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
</taxonomicName>
species- groups in the Neotropical region (
<emphasis id="B963EAFC5232D42AC42124D5FCF9FA8A" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C174D6D5232D42AC42124D5FB18FB57" box="[1058,1187,1254,1278]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">canicularis</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC4B224D5FAB7FB57" box="[1201,1292,1254,1278]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scalaris">scalaris</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC51924D5FA23FB57" box="[1306,1432,1254,1278]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="anthracina">anthracina</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC0942538FF54FA8A" box="[151,239,1291,1315]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="grandis">grandis</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC1052538FED3FA8A" box="[262,360,1291,1315]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heydenii">heydenii</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC17C2538FDA1FA8A" box="[383,538,1291,1315]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="obscurinervis">obscurinervis</taxonomicName>
, flavipalpis, admirabilis
</emphasis>
) and three subgroups of the
<taxonomicName id="4C174D6D5232D42AC4C92538FAF3FA8A" box="[1226,1352,1291,1315]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5232D42AC4C92538FAF3FA8A" box="[1226,1352,1291,1315]" italics="true" pageId="3" pageNumber="4">canicularis</emphasis>
</taxonomicName>
group (
<emphasis id="B963EAFC5232D42AC0A3251DFDBBFAEF" box="[160,512,1326,1350]" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="4C174D6D5232D42AC0A3251DFE9AFAEF" box="[160,289,1326,1350]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">canicularis</taxonomicName>
,
<taxonomicName id="4C174D6D5232D42AC12C251CFED5FAEF" box="[303,366,1327,1350]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pusio">pusio</taxonomicName>
, petrocchiae
</emphasis>
), based on characters of little phylogenetic relevance (
<bibRefCitation id="EF864B1F5232D42AC478251DFA37FAEF" author="Albuquerque" box="[1147,1420,1326,1351]" pageId="3" pageNumber="4" refString="Albuquerque, D. de O., Pamplona, D. &amp; de Carvalho, C. J. B. (1981) Contribuicao ao conhecimento dos Fannia R. D., 1830 da regiao Neotropical (Diptera, Fanniidae). Arquivo do Museu Nacional do Rio de Janeiro, 56, 9 - 34." type="journal article" year="1981">
Albuquerque
<emphasis id="B963EAFC5232D42AC51A2503FAF6FAEF" box="[1305,1357,1326,1351]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
.1981
</bibRefCitation>
). Even though the species-groups of
<taxonomicName id="4C174D6D5232D42AC2202567FDCFFAC2" box="[547,628,1364,1387]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5232D42AC2202567FDCFFAC2" box="[547,628,1364,1387]" italics="true" pageId="3" pageNumber="4">Fannia</emphasis>
</taxonomicName>
are widely mentioned in the literature, their monophyly has been tested only in the work of
<bibRefCitation id="EF864B1F5232D42AC1782545FCB8FA27" author="Dominguez" box="[379,771,1398,1422]" pageId="3" pageNumber="4" refString="Dominguez, M. C. &amp; Roig-Junent, S. (2008) A phylogeny of the family Fanniidae Schnabl (Insecta: Diptera: Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia. Invertebrate Systematics, 22, 563 - 587." type="journal article" year="2008">Domínguez &amp; Roig-Juñent (2008)</bibRefCitation>
. In that contribution, only the
<taxonomicName id="4C174D6D5232D42AC45D2545FB06FA27" box="[1118,1213,1398,1422]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heydenii">
<emphasis id="B963EAFC5232D42AC45D2545FB06FA27" box="[1118,1213,1398,1422]" italics="true" pageId="3" pageNumber="4">heydenii</emphasis>
</taxonomicName>
and the
<taxonomicName id="4C174D6D5232D42AC5222545FA27FA27" box="[1313,1436,1398,1422]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="B963EAFC5232D42AC5222545FA27FA27" box="[1313,1436,1398,1422]" italics="true" pageId="3" pageNumber="4">anthracina</emphasis>
</taxonomicName>
groups, and the sub-group
<taxonomicName id="4C174D6D5232D42AC1C125AFFE45FA1A" box="[450,510,1436,1459]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B963EAFC5232D42AC1C125AFFE45FA1A" box="[450,510,1436,1459]" italics="true" pageId="3" pageNumber="4">pusio</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C174D6D5232D42AC24E25A8FD70FA1A" box="[589,715,1435,1459]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5232D42AC24E25A8FD70FA1A" box="[589,715,1435,1459]" italics="true" pageId="3" pageNumber="4">canicularis</emphasis>
</taxonomicName>
group were indicated as clades in the Neotropical region.
</paragraph>
<paragraph id="8BA836EE5232D42AC0C4258DFC3BF97B" blockId="3.[151,1436,786,1746]" pageId="3" pageNumber="4">
Species of the
<taxonomicName id="4C174D6D5232D42AC16F258DFE70FA7F" box="[364,459,1470,1494]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="heydenii">
<emphasis id="B963EAFC5232D42AC16F258DFE70FA7F" box="[364,459,1470,1494]" italics="true" pageId="3" pageNumber="4">heydenii</emphasis>
</taxonomicName>
group have the scape and the pedicel light brown, the hind femur with a group of long pre-apical
<emphasis id="B963EAFC5232D42AC10D25D7FE92FA52" box="[270,297,1508,1531]" italics="true" pageId="3" pageNumber="4">pv</emphasis>
and
<emphasis id="B963EAFC5232D42AC15C25D7FED1FA52" box="[351,362,1508,1531]" italics="true" pageId="3" pageNumber="4">v</emphasis>
, hind coxa setulose on the posterior margin, abdomen partly translucent yellow, sternite 1 bare, cercal plate wider than long, and bacilliform process cuneiform; females have two spermathecae (
<bibRefCitation id="EF864B1F5232D42AC50B2635FF69F9EB" author="Albuquerque" pageId="3" pageNumber="4" refString="Albuquerque, D. de O. (1957) Sobre especies de Fannia R. - D., 1830 novas ou pouco conhecidas (Diptera - Muscidae). Boletim do Museu Nacional (Nova Serie), 172, 1 - 31." type="journal article" year="1957">Albuquerque 1957</bibRefCitation>
,
<bibRefCitation id="EF864B1F5232D42AC0E32619FE46F9EB" author="Albuquerque" box="[224,509,1578,1603]" pageId="3" pageNumber="4" refString="Albuquerque, D. de O., Pamplona, D. &amp; de Carvalho, C. J. B. (1981) Contribuicao ao conhecimento dos Fannia R. D., 1830 da regiao Neotropical (Diptera, Fanniidae). Arquivo do Museu Nacional do Rio de Janeiro, 56, 9 - 34." type="journal article" year="1981">
Albuquerque
<emphasis id="B963EAFC5232D42AC17F261FFE0CF9EA" box="[380,439,1579,1603]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
1981
</bibRefCitation>
). The species of the
<taxonomicName id="4C174D6D5232D42AC2F22618FCD7F9EA" box="[753,876,1579,1603]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="B963EAFC5232D42AC2F22618FCD7F9EA" box="[753,876,1579,1603]" italics="true" pageId="3" pageNumber="4">anthracina</emphasis>
</taxonomicName>
group have tarsi with modified tarsomeres, fore basitarsus with 1 differentiated basal
<emphasis id="B963EAFC5232D42AC2492663FDEEF9CE" box="[586,597,1616,1639]" italics="true" pageId="3" pageNumber="4">v</emphasis>
, and bacilliform process spiral-shaped (de
<bibRefCitation id="EF864B1F5232D42AC44D267DFAE1F9CF" author="Carvalho" box="[1102,1370,1614,1638]" pageId="3" pageNumber="4" refString="Pont, A. C. &amp; de Carvalho, C. J. B. (1994) Neotropical Fanniidae (Diptera): A key to the Fannia anthracina Group. Entomologist´s Monthly Magazine, 130, 229 - 237." type="journal article" year="1994">Carvalho &amp; Pont 1994</bibRefCitation>
). The species of this group have restricted distribution in southern South
<collectingCountry id="F300767E5232D42AC38A2641FC56F923" box="[905,1005,1650,1674]" name="United States of America" pageId="3" pageNumber="4">America</collectingCountry>
. Species of the
<taxonomicName id="4C174D6D5232D42AC49D2647FB61F922" box="[1182,1242,1652,1675]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B963EAFC5232D42AC49D2647FB61F922" box="[1182,1242,1652,1675]" italics="true" pageId="3" pageNumber="4">pusio</emphasis>
</taxonomicName>
sub-group of the
<taxonomicName id="4C174D6D5232D42AC09426A5FEAEF907" box="[151,277,1686,1710]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5232D42AC09426A5FEAEF907" box="[151,277,1686,1710]" italics="true" pageId="3" pageNumber="4">canicularis</emphasis>
</taxonomicName>
group have parafacial with short bristles, no upper
<emphasis id="B963EAFC5232D42AC36926A5FC2AF907" box="[874,913,1686,1710]" italics="true" pageId="3" pageNumber="4">orb</emphasis>
, abdomen trimaculate and an epiphallus. The absence of the latter is considered an autapomorphy for
<taxonomicName id="4C174D6D5232D42AC3092689FCC7F97B" box="[778,892,1722,1746]" class="Insecta" family="Fanniidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Fanniidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>