treatments-xml/data/03/BE/87/03BE87A85255775AFF23F898D8766ECD.xml

<document ID-DOI="10.11646/zootaxa.5254.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7732079" ID-ZooBank="2301C5BB-AE1B-4D32-9201-E7D00DAB7901" checkinTime="1678782829124" checkinUser="plazi" docAuthor="Jokar, Milad, Kamangar, Barzan Bahrami, Ghaderi, Edris &amp; Freyhof, Jörg" docDate="2023" docId="03BE87A85255775AFF23F898D8766ECD" docLanguage="en" docName="zootaxa.5254.4.2.pdf" docOrigin="Zootaxa 5254 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5254.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyptothorax sardashtensis Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof 2023, new species" docType="treatment" docVersion="3" lastPageNumber="490" masterDocId="FF87FFD052537754FFB4FF9ADA326831" masterDocTitle="Glyptothorax sardashtensis, a new species of torrent catfish from the upper Lesser Zab drainage in Iran (Teleostei: Sisoridae)" masterLastPageNumber="492" masterPageNumber="476" pageNumber="482" updateTime="1678808524203" updateUser="tatiana" zenodo-license-document="CLOSED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Glyptothorax sardashtensis, a new species of torrent catfish from the upper Lesser Zab drainage in Iran (Teleostei: Sisoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jokar, Milad</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-3344-4540</mods:nameIdentifier>
<mods:affiliation>miladjokar 80 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3344 - 4540</mods:affiliation>
<mods:nameIdentifier type="email">miladjokar80@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kamangar, Barzan Bahrami</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ghaderi, Edris</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-9875-7735</mods:nameIdentifier>
<mods:affiliation>ed. ghaderi @ uok. ac. ir; https: // orcid. org / 0000 - 0002 - 9875 - 7735</mods:affiliation>
<mods:nameIdentifier type="email">ed.ghaderi@uok.ac.ir</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Freyhof, Jörg</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7042-3127</mods:nameIdentifier>
<mods:affiliation>Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, 10115 Berlin, Germany. joerg. freyhof @ mfn. berlin; https: // orcid. org / 0000 - 0002 - 7042 - 3127</mods:affiliation>
<mods:nameIdentifier type="email">freyhof@mfn.berlin</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5254</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>476</mods:start>
<mods:end>492</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5254.4.2</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5254.4.2</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7732079</mods:identifier>
<mods:identifier type="ZooBank">2301C5BB-AE1B-4D32-9201-E7D00DAB7901</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD" httpUri="http://treatment.plazi.org/id/03BE87A85255775AFF23F898D8766ECD" lastPageId="14" lastPageNumber="490" pageId="6" pageNumber="482">
<subSubSection box="[151,640,1794,1821]" pageId="6" pageNumber="482" type="nomenclature">
<paragraph blockId="6.[151,640,1794,1857]" box="[151,640,1794,1821]" pageId="6" pageNumber="482">
<heading bold="true" box="[151,640,1794,1821]" fontSize="11" level="1" pageId="6" pageNumber="482" reason="1">
<emphasis bold="true" box="[151,640,1794,1821]" pageId="6" pageNumber="482">
<taxonomicName authority="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof, 2023" authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[151,481,1794,1821]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="482" phylum="Chordata" rank="species" species="sardashtensis" status="new species">
<emphasis bold="true" box="[151,481,1794,1821]" italics="true" pageId="6" pageNumber="482">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[496,640,1795,1821]" pageId="6" pageNumber="482" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,277,1831,1857]" pageId="6" pageNumber="482" type="description">
<paragraph blockId="6.[151,640,1794,1857]" box="[151,277,1831,1857]" pageId="6" pageNumber="482">
(
<figureCitation box="[159,270,1831,1857]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="8.[151,250,1114,1138]" captionStartId-1="10.[151,250,1953,1977]" captionStartId-2="11.[151,250,1047,1071]" captionStartId-3="11.[151,250,1933,1957]" captionStartId-4="12.[151,250,562,586]" captionTargetBox-0="[156,1431,669,1091]" captionTargetBox-1="[155,1433,597,1930]" captionTargetBox-2="[152,1435,181,1023]" captionTargetBox-3="[152,1435,1132,1910]" captionTargetBox-4="[154,1433,181,538]" captionTargetId-0="figure-249@8.[156,1431,669,1091]" captionTargetId-1="figure-204@10.[155,1433,597,1930]" captionTargetId-2="figure-24@11.[152,1435,181,1023]" captionTargetId-3="figure-49@11.[152,1435,1132,1910]" captionTargetId-4="figure-20@12.[154,1433,181,538]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-1="FIGURE 7. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-2="FIGURE 8. Glyptothorax sardashtensis, paratypes, from left: FCFUK 302, 51 mm SL; FCFUK 303, paratype, 42 mm SL. Lesser Zab, Iran." captionText-3="FIGURE 9. Glyptothorax sardashtensis, from left: FCFUK 300, holotype, 60 mm SL; FCFUK 302, paratype, 51 mm SL; FCFUK 305, paratype, 55 mm SL; Lesser Zab, Iran." captionText-4="FIGURE 10. Comparison of the presence or absence of spots and blotches inG.sardashtensis and G. kurdistanicus. Glyptothorax sardashtensis, from left: FCFUK 306, 51 mm SL; FCFUK 321, 45 mm SL; Lesser Zab, Iran. Glyptothorax kurdistanicus, from right: FCFUK 332, 67 mm SL; FCFUK 337, 43 mm SL; Lesser Zab, Iran." figureDoi-0="http://doi.org/10.5281/zenodo.7732091" figureDoi-1="http://doi.org/10.5281/zenodo.7732093" figureDoi-2="http://doi.org/10.5281/zenodo.7732095" figureDoi-3="http://doi.org/10.5281/zenodo.7732099" figureDoi-4="http://doi.org/10.5281/zenodo.7732101" httpUri-0="https://zenodo.org/record/7732091/files/figure.png" httpUri-1="https://zenodo.org/record/7732093/files/figure.png" httpUri-2="https://zenodo.org/record/7732095/files/figure.png" httpUri-3="https://zenodo.org/record/7732099/files/figure.png" httpUri-4="https://zenodo.org/record/7732101/files/figure.png" pageId="6" pageNumber="482">Fig. 6–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="483" pageId="6" pageNumber="482" type="materials_examined">
<paragraph blockId="6.[151,1437,1903,2038]" pageId="6" pageNumber="482">
<materialsCitation collectionCode="FCFUK" country="Iran" latitude="36.2688" location="Nalas" longLatPrecision="6" longitude="45.5012" municipality="Lesser Zab" pageId="6" pageNumber="482" specimenCode="FCFUK 300" specimenCount="1" stateProvince="Azarbayjan-e Gharbi" typeStatus="holotype">
<emphasis bold="true" box="[151,269,1903,1929]" pageId="6" pageNumber="482">
<typeStatus box="[151,264,1903,1929]" pageId="6" pageNumber="482">Holotype</typeStatus>
:
</emphasis>
<specimenCode box="[280,426,1903,1929]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 300</specimenCode>
, 
<quantity box="[441,524,1903,1929]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="482" unit="mm" value="60.0">60 mm</quantity>
SL; 
<collectingCountry box="[586,635,1903,1929]" name="Iran" pageId="6" pageNumber="482">Iran</collectingCountry>
: 
<collectingRegion box="[649,909,1903,1929]" country="Iran" name="Azarbayjan-e Gharbi" pageId="6" pageNumber="482">West Azerbaijan prov.</collectingRegion>
: 
<collectingMunicipality box="[923,1051,1903,1929]" pageId="6" pageNumber="482">Lesser Zab</collectingMunicipality>
at bridge east of 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC8606552557752FB5BF8F5DF036FB8" box="[1263,1329,1903,1929]" country="Iran" latitude="36.2688" longLatPrecision="6" longitude="45.5012" municipality="Lesser Zab" name="Nalas" pageId="6" pageNumber="482" stateProvince="Azarbayjan-e Gharbi">Nalas</location>
, 
<geoCoordinate box="[1345,1436,1903,1929]" degrees="36.2688" direction="north" orientation="latitude" pageId="6" pageNumber="482" precision="5" value="36.2688">36.2688</geoCoordinate>
<geoCoordinate box="[151,246,1939,1966]" degrees="45.5012" direction="east" orientation="longitude" pageId="6" pageNumber="482" precision="5" value="45.5012">45.5012</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph blockId="6.[151,1437,1903,2038]" lastBlockId="7.[151,1437,150,2013]" lastPageId="7" lastPageNumber="483" pageId="6" pageNumber="482">
<materialsCitation box="[199,500,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 302, 1" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[199,328,1975,2001]" pageId="6" pageNumber="482">
<typeStatus box="[199,323,1975,2001]" pageId="6" pageNumber="482">Paratypes</typeStatus>
:
</emphasis>
<specimenCode box="[335,476,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 302</specimenCode>
, 
<specimenCode box="[486,500,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[513,592,1975,2002]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="6" pageNumber="482" unit="mm" value="51.0">51 mm</quantity>
SL; 
<materialsCitation box="[644,810,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 303, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[644,786,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 303</specimenCode>
, 
<specimenCode box="[796,810,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[823,901,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="482" unit="mm" value="42.0">42 mm</quantity>
SL; 
<materialsCitation box="[954,1120,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 309, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[954,1095,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 309</specimenCode>
, 
<specimenCode box="[1106,1120,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[1133,1211,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="482" unit="mm" value="41.0">41 mm</quantity>
SL; 
<materialsCitation box="[1263,1429,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 310, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1263,1405,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 310</specimenCode>
, 
<specimenCode box="[1415,1429,1975,2001]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[151,229,2011,2037]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="482" unit="mm" value="40.0">40 mm</quantity>
SL; 
<materialsCitation box="[282,446,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 311, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[282,421,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 311</specimenCode>
, 
<specimenCode box="[432,446,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[459,537,2011,2037]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="482" unit="mm" value="38.0">38 mm</quantity>
SL; 
<materialsCitation box="[589,755,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 312, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[589,731,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 312</specimenCode>
, 
<specimenCode box="[741,755,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[768,846,2011,2037]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="482" unit="mm" value="38.0">38 mm</quantity>
SL; 
<materialsCitation box="[898,1064,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482" specimenCode="FCFUK 313, 1" specimenCount="1" typeStatus="paratype">
<specimenCode box="[898,1040,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">FCFUK 313</specimenCode>
, 
<specimenCode box="[1050,1064,2011,2037]" collectionCode="FCFUK" pageId="6" pageNumber="482">1</specimenCode>
</materialsCitation>
, 
<quantity box="[1077,1155,2011,2037]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="482" unit="mm" value="37.0">37 mm</quantity>
SL; 
<materialsCitation box="[1207,1437,2011,2038]" collectionCode="FSJF" pageId="6" pageNumber="482" specimenCode="FSJF 4112, 2, 51-52" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1207,1329,2011,2037]" collectionCode="FSJF" pageId="6" pageNumber="482">FSJF 4112</specimenCode>
, 
<specimenCode box="[1339,1355,2011,2037]" collectionCode="FSJF" pageId="6" pageNumber="482">2</specimenCode>
, 
<specimenCode box="[1366,1437,2011,2038]" collectionCode="FSJF" pageId="6" pageNumber="482">51–52</specimenCode>
</materialsCitation>
<materialsCitation collectionCode="FCFUK" country="Iran" latitude="36.1369" location="Nisk-Abad" longLatPrecision="6" longitude="45.551" municipality="Lesser Zab" pageId="7" pageNumber="483" specimenCode="FCFUK 304, 1, FCFUK 305" specimenCount="1" stateProvince="Azarbayjan-e Gharbi">
mm; data same as holotype.— 
<specimenCode box="[489,634,151,177]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 304</specimenCode>
, 
<specimenCode box="[645,659,151,177]" collectionCode="FCFUK" pageId="7" pageNumber="483">1</specimenCode>
, 
<quantity box="[673,752,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="7" pageNumber="483" unit="mm" value="67.0">67 mm</quantity>
SL; 
<specimenCode box="[807,949,150,177]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 305</specimenCode>
, 
<specimenCode box="[960,974,151,177]" collectionCode="FCFUK" pageId="7" pageNumber="483">1</specimenCode>
, 
<quantity box="[988,1067,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="483" unit="mm" value="55.0">55 mm</quantity>
SL; 
<collectingCountry box="[1122,1171,151,177]" name="Iran" pageId="7" pageNumber="483">Iran</collectingCountry>
: 
<collectingRegion box="[1181,1433,151,177]" country="Iran" name="Azarbayjan-e Gharbi" pageId="7" pageNumber="483">West Azerbaijan prov.</collectingRegion>
: 
<collectingMunicipality box="[151,275,187,213]" pageId="7" pageNumber="483">Lesser Zab</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC8606552547753FE81FF21DB8268E4" box="[309,432,187,213]" country="Iran" latitude="36.1369" longLatPrecision="6" longitude="45.551" municipality="Lesser Zab" name="Nisk-Abad" pageId="7" pageNumber="483" stateProvince="Azarbayjan-e Gharbi">Nisk-Abad</location>
(now dawned in a reservoir), 
<geoCoordinate box="[769,860,187,213]" degrees="36.1369" direction="north" orientation="latitude" pageId="7" pageNumber="483" precision="5" value="36.1369">36.1369</geoCoordinate>
<geoCoordinate box="[867,962,186,213]" degrees="45.5510" direction="east" orientation="longitude" pageId="7" pageNumber="483" precision="5" value="45.551">45.5510</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
<emphasis bold="true" box="[199,532,222,249]" pageId="7" pageNumber="483">Material used for osteology:</emphasis>
<materialsCitation box="[539,811,222,249]" collectionCode="FCFUK" pageId="7" pageNumber="483" specimenCode="FCFUK 301" specimenCount="1">
<specimenCode box="[539,679,223,249]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 301</specimenCode>
, 
<quantity box="[691,769,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="483" unit="mm" value="58.0">58 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation box="[822,1095,223,249]" collectionCode="FCFUK" pageId="7" pageNumber="483" specimenCode="FCFUK 314" specimenCount="1">
<specimenCode box="[822,964,223,249]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 314</specimenCode>
, 
<quantity box="[975,1053,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="483" unit="mm" value="38.0">38 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation collectionCode="FCFUK" country="Iran" latitude="36.2688" location="Nalas" longLatPrecision="6" longitude="45.5012" municipality="Lesser Zab" pageId="7" pageNumber="483" specimenCode="FCFUK 320" specimenCount="1" stateProvince="Azarbayjan-e Gharbi">
<specimenCode box="[1106,1248,223,249]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 320</specimenCode>
, 
<quantity box="[1258,1337,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="483" unit="mm" value="31.0">31 mm</quantity>
SL; data same as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
<emphasis bold="true" box="[199,645,294,321]" pageId="7" pageNumber="483">Material used for molecular analysis:</emphasis>
<materialsCitation box="[652,1404,295,321]" collectionCode="FCFUK, OQ" pageId="7" pageNumber="483" specimenCode="FCFUK 304, OQ078869" specimenCount="1">
<specimenCode box="[652,795,295,321]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 304</specimenCode>
, 
<quantity box="[806,885,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="7" pageNumber="483" unit="mm" value="67.0">67 mm</quantity>
SL (GenBank accession number: 
<specimenCode box="[1267,1396,295,321]" collectionCode="OQ" pageId="7" pageNumber="483">OQ078869</specimenCode>
)
</materialsCitation>
.— 
<materialsCitation box="[151,1164,330,357]" collectionCode="FCFUK, OQ" country="Iran" latitude="36.2688" location="Nalas" longLatPrecision="6" longitude="45.5012" municipality="Lesser Zab" pageId="7" pageNumber="483" specimenCode="FCFUK 305, OQ078870" specimenCount="1" stateProvince="Azarbayjan-e Gharbi">
<specimenCode box="[151,293,330,357]" collectionCode="FCFUK" pageId="7" pageNumber="483">FCFUK 305</specimenCode>
, 
<quantity box="[304,383,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="483" unit="mm" value="55.0">55 mm</quantity>
SL (GenBank accession number: 
<specimenCode box="[765,895,331,357]" collectionCode="OQ" pageId="7" pageNumber="483">OQ078870</specimenCode>
); same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="483" type="diagnosis">
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
<emphasis bold="true" box="[199,324,366,392]" pageId="7" pageNumber="483">Diagnosis:</emphasis>
In the Iranian upper Lesser Zab drainage, 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[787,971,367,393]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[787,971,367,393]" italics="true" pageId="7" pageNumber="483">G. sardashtensis</emphasis>
</taxonomicName>
occurs in sympatry with 
<taxonomicName box="[1247,1429,367,393]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1247,1429,367,393]" italics="true" pageId="7" pageNumber="483">G. kurdistanicus</emphasis>
</taxonomicName>
. It is distinguished from this species by having a plain flank, without spots or blotches, with few, very small, darkbrown dots on the head, the dorsal- and adipose-fin bases, and on the caudal-peduncle (vs. flank with many, often large, black spots and blotches) (
<figureCitation box="[518,600,475,501]" captionStart="FIGURE 10" captionStartId="12.[151,250,562,586]" captionTargetBox="[154,1433,181,538]" captionTargetId="figure-20@12.[154,1433,181,538]" captionTargetPageId="12" captionText="FIGURE 10. Comparison of the presence or absence of spots and blotches inG.sardashtensis and G. kurdistanicus. Glyptothorax sardashtensis, from left: FCFUK 306, 51 mm SL; FCFUK 321, 45 mm SL; Lesser Zab, Iran. Glyptothorax kurdistanicus, from right: FCFUK 332, 67 mm SL; FCFUK 337, 43 mm SL; Lesser Zab, Iran." figureDoi="http://doi.org/10.5281/zenodo.7732101" httpUri="https://zenodo.org/record/7732101/files/figure.png" pageId="7" pageNumber="483">Fig. 10</figureCitation>
), three yellowish blotches, arranged in crescent-shaped arch in front of the dorsal-fin origin (vs. arranged in a triangle), a wide medial pit with a roundish anterior margin (vs. narrow with pointed anterior margin in juveniles smaller than 
<quantity box="[696,773,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="483" unit="mm" value="80.0">80 mm</quantity>
SL), and the caudal-peduncle depth 1.6–2.3 times its length (vs. (1.2) 1.3–1.7).
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[199,409,619,645]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="cous">
<emphasis box="[199,409,619,645]" italics="true" pageId="7" pageNumber="483">Glyptothorax cous</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[468,656,619,645]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="steindachneri">
<emphasis box="[468,656,619,645]" italics="true" pageId="7" pageNumber="483">G. steindachneri</emphasis>
</taxonomicName>
are two additional species found in the Iraqi part of the Lesser Zab drainage by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[293,534,655,681]" pageId="7" pageNumber="483" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[390,448,655,681]" italics="true" pageId="7" pageNumber="483">et al.</emphasis>
(2021)
</bibRefCitation>
. 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[546,855,655,681]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[546,855,655,681]" italics="true" pageId="7" pageNumber="483">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[1112,1199,655,681]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="cous">
<emphasis box="[1112,1199,655,681]" italics="true" pageId="7" pageNumber="483">G. cous</emphasis>
</taxonomicName>
by lacking tubercles on the head and body (vs. having many large, bony, striated, and elongated tubercles). In 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[1183,1370,691,717]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1183,1370,691,717]" italics="true" pageId="7" pageNumber="483">G. sardashtensis</emphasis>
</taxonomicName>
, only few, soft, small warts are present around the thoracic apparatus, on the edges of the gill cover, and on the skin of the dorsal surface of the unbranched pectoral-fin ray. The new species is distinguished from 
<taxonomicName baseAuthorityName="Pietschmann" baseAuthorityYear="1913" box="[1134,1319,763,789]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="steindachneri">
<emphasis box="[1134,1319,763,789]" italics="true" pageId="7" pageNumber="483">G. steindachneri</emphasis>
</taxonomicName>
by having a shorter adipose-fin (12–23% SL vs. 23–30% SL), the medial pit of the thoracic adhesive apparatus without striae (vs. with striae), and the head and body without tubercles (vs. many large, elongated, bony and striated tubercles).
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
Applying the identification key by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[602,843,871,897]" pageId="7" pageNumber="483" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[700,758,871,897]" italics="true" pageId="7" pageNumber="483">et al.</emphasis>
(2021)
</bibRefCitation>
, 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[857,1045,871,897]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[857,1045,871,897]" italics="true" pageId="7" pageNumber="483">G. sardashtensis</emphasis>
</taxonomicName>
keys out as 
<taxonomicName box="[1191,1301,871,897]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[1191,1301,871,897]" italics="true" pageId="7" pageNumber="483">G. silviae</emphasis>
</taxonomicName>
, by lacking tubercles and warts on the head and anterior body (vs. present in 
<taxonomicName baseAuthorityName="Berg" baseAuthorityYear="1918" box="[867,1031,907,933]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="armeniacus">
<emphasis box="[867,1031,907,933]" italics="true" pageId="7" pageNumber="483">G. armeniacus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Freyhof, Kaya, Abdullah &amp; Geiger" authorityYear="2021" box="[1083,1205,907,933]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="daemon">
<emphasis box="[1083,1205,907,933]" italics="true" pageId="7" pageNumber="483">G. daemon</emphasis>
</taxonomicName>
) and having few and short anteromedial striae (vs. numerous and long in 
<taxonomicName baseAuthorityName="Berg" baseAuthorityYear="1918" box="[732,897,943,969]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="armeniacus">
<emphasis box="[732,897,943,969]" italics="true" pageId="7" pageNumber="483">G. armeniacus</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Freyhof, Kaya, Abdullah &amp; Geiger" authorityYear="2021" box="[951,1074,943,969]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="daemon">
<emphasis box="[951,1074,943,969]" italics="true" pageId="7" pageNumber="483">G. daemon</emphasis>
</taxonomicName>
). Furthermore, 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[1251,1436,943,969]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1251,1436,943,969]" italics="true" pageId="7" pageNumber="483">G. sardashtensis</emphasis>
</taxonomicName>
always has a central yellowish blotch on the nuchal plate in front of the dorsal-fin origin (vs. usually absent in 
<taxonomicName baseAuthorityName="Berg" baseAuthorityYear="1918" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="armeniacus">
<emphasis italics="true" pageId="7" pageNumber="483">G. armeniacus</emphasis>
</taxonomicName>
). 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[304,611,1015,1041]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[304,611,1015,1041]" italics="true" pageId="7" pageNumber="483">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName authorityName="Freyhof, Kaya, Abdullah &amp; Geiger" authorityYear="2021" box="[941,1063,1015,1041]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="daemon">
<emphasis box="[941,1063,1015,1041]" italics="true" pageId="7" pageNumber="483">G. daemon</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Berg" baseAuthorityYear="1918" box="[1115,1280,1015,1041]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="armeniacus">
<emphasis box="[1115,1280,1015,1041]" italics="true" pageId="7" pageNumber="483">G. armeniacus</emphasis>
</taxonomicName>
by having the thoracic adhesive apparatus slightly elevated (vs. strongly elevated), extending from the isthmus to the base of the third pectoral-fin ray (vs. to the posterior limit of the pectoral-fin base).
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
Applying the key published by 
<bibRefCitation author="Mousavi-Sabet, H. &amp; Eagderi, S. &amp; Vatandoust, S. &amp; Freyhof, J." box="[548,863,1123,1149]" pageId="7" pageNumber="483" pagination="451 - 484" refId="ref10209" refString="Mousavi-Sabet, H., Eagderi, S., Vatandoust, S. &amp; Freyhof, J. (2021) Five new species of the sisorid catfish genus Glyptothorax from Iran (Teleostei: Sisoridae). Zootaxa, 5067 (4), 451 - 484. https: // doi. org / 10.11646 / zootaxa. 5067.4.1" type="journal article" year="2021">
Mousavi-Sabet 
<emphasis box="[724,780,1123,1149]" italics="true" pageId="7" pageNumber="483">et al.</emphasis>
(2021)
</bibRefCitation>
, 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[874,1058,1123,1149]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[874,1058,1123,1149]" italics="true" pageId="7" pageNumber="483">G. sardashtensis</emphasis>
</taxonomicName>
keys out as 
<taxonomicName box="[1195,1309,1123,1149]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="pallens">
<emphasis box="[1195,1309,1123,1149]" italics="true" pageId="7" pageNumber="483">G. pallens</emphasis>
</taxonomicName>
, 
<emphasis italics="true" pageId="7" pageNumber="483">
<taxonomicName box="[1323,1432,1123,1149]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="silviae">G. silviae</taxonomicName>
, 
<taxonomicName box="[151,271,1159,1185]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="alidaeii">G. alidaeii</taxonomicName>
</emphasis>
or 
<emphasis box="[306,433,1159,1185]" italics="true" pageId="7" pageNumber="483">
<taxonomicName box="[306,428,1159,1185]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="shapuri">G. shapuri</taxonomicName>
.
</emphasis>
Similar to 
<taxonomicName box="[555,669,1159,1185]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="pallens">
<emphasis box="[555,669,1159,1185]" italics="true" pageId="7" pageNumber="483">G. pallens</emphasis>
</taxonomicName>
, from the adjacent 
<collectingRegion box="[883,956,1159,1185]" country="Azerbaijan" name="Sirvan" pageId="7" pageNumber="483">Sirvan</collectingRegion>
drainage, the new species lack dark-brown or black spots and blotches on the flank. 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[627,936,1195,1221]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[627,936,1195,1221]" italics="true" pageId="7" pageNumber="483">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName box="[1196,1314,1195,1221]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="pallens">
<emphasis box="[1196,1314,1195,1221]" italics="true" pageId="7" pageNumber="483">G. pallens</emphasis>
</taxonomicName>
by having a shorter outer mandibular barbel (extending to posterior edge of gill cover, not reaching pectoral-fin origin vs. reaching to second pectoral-fin ray), few soft and small warts around the thoracic apparatus, the edges of the gill cover, and the dorsal surface the first unbranched pectoral-fin (vs. many small shallow warts on the head, body, and belly); a thoracic adhesive apparatus slightly elevated (vs. strongly elevated), the medial pit much broader, and its anterior end roundish (vs. narrow and spear-blade shaped), and a more slender caudal peduncle (caudal peduncle depth 1.6–2.3 times in its length vs. 1.1–1.3).
</paragraph>
<paragraph blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="483">
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[199,510,1447,1473]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[199,510,1447,1473]" italics="true" pageId="7" pageNumber="483">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName box="[776,984,1447,1473]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[776,984,1447,1473]" italics="true" pageId="7" pageNumber="483">G. hosseinpanahii</emphasis>
</taxonomicName>
by having few and short anteromedial striae (vs. numerous and long). 
<taxonomicName authorityName="Jokar &amp; Kamangar &amp; Ghaderi &amp; Freyhof" authorityYear="2023" box="[516,825,1483,1509]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[516,825,1483,1509]" italics="true" pageId="7" pageNumber="483">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName box="[1085,1207,1483,1509]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[1085,1207,1483,1509]" italics="true" pageId="7" pageNumber="483">G. alidaeii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[1223,1430,1483,1509]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[1223,1430,1483,1509]" italics="true" pageId="7" pageNumber="483">G. hosseinpanahii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[151,283,1519,1545]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[151,283,1519,1545]" italics="true" pageId="7" pageNumber="483">G. galaxias</emphasis>
</taxonomicName>
, 
<taxonomicName box="[299,422,1519,1545]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="shapuri">
<emphasis box="[299,422,1519,1545]" italics="true" pageId="7" pageNumber="483">G. shapuri</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[488,597,1519,1545]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="483" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[488,597,1519,1545]" italics="true" pageId="7" pageNumber="483">G. silviae</emphasis>
</taxonomicName>
by having the thoracic adhesive apparatus slightly elevated (vs. strongly elevated), the thoracic adhesive apparatus extending from the isthmus to the level of the base of the third pectoralfin ray (vs. to the posterior-most pectoral-fin base), a wide medial pit (vs. narrow) with a roundish anterior end (vs. pointed), a plain flank, with few, very small, dark-brown dots on the head, the dorsal-, and adipose-fin bases, and on the caudal-peduncle (vs. flank usually with many, rarely few, black spots and blotches, with many additional, greenish-silvery blotches in life, faded in few preserved individuals), and three yellowish blotches on the nuchal plate, arranged in crescent-shaped arch (vs. arranged in a triangle).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="486" pageId="7" pageNumber="483" type="description">
<paragraph blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1437,151,645]" lastPageId="8" lastPageNumber="484" pageId="7" pageNumber="483">
<emphasis bold="true" box="[199,345,1770,1796]" pageId="7" pageNumber="483">Description.</emphasis>
See 
<figureCitation box="[404,554,1771,1797]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="8.[151,250,1114,1138]" captionStartId-1="10.[151,250,1953,1977]" captionStartId-2="11.[151,250,1047,1071]" captionStartId-3="11.[151,250,1933,1957]" captionStartId-4="12.[151,250,562,586]" captionTargetBox-0="[156,1431,669,1091]" captionTargetBox-1="[155,1433,597,1930]" captionTargetBox-2="[152,1435,181,1023]" captionTargetBox-3="[152,1435,1132,1910]" captionTargetBox-4="[154,1433,181,538]" captionTargetId-0="figure-249@8.[156,1431,669,1091]" captionTargetId-1="figure-204@10.[155,1433,597,1930]" captionTargetId-2="figure-24@11.[152,1435,181,1023]" captionTargetId-3="figure-49@11.[152,1435,1132,1910]" captionTargetId-4="figure-20@12.[154,1433,181,538]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-1="FIGURE 7. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-2="FIGURE 8. Glyptothorax sardashtensis, paratypes, from left: FCFUK 302, 51 mm SL; FCFUK 303, paratype, 42 mm SL. Lesser Zab, Iran." captionText-3="FIGURE 9. Glyptothorax sardashtensis, from left: FCFUK 300, holotype, 60 mm SL; FCFUK 302, paratype, 51 mm SL; FCFUK 305, paratype, 55 mm SL; Lesser Zab, Iran." captionText-4="FIGURE 10. Comparison of the presence or absence of spots and blotches inG.sardashtensis and G. kurdistanicus. Glyptothorax sardashtensis, from left: FCFUK 306, 51 mm SL; FCFUK 321, 45 mm SL; Lesser Zab, Iran. Glyptothorax kurdistanicus, from right: FCFUK 332, 67 mm SL; FCFUK 337, 43 mm SL; Lesser Zab, Iran." figureDoi-0="http://doi.org/10.5281/zenodo.7732091" figureDoi-1="http://doi.org/10.5281/zenodo.7732093" figureDoi-2="http://doi.org/10.5281/zenodo.7732095" figureDoi-3="http://doi.org/10.5281/zenodo.7732099" figureDoi-4="http://doi.org/10.5281/zenodo.7732101" httpUri-0="https://zenodo.org/record/7732091/files/figure.png" httpUri-1="https://zenodo.org/record/7732093/files/figure.png" httpUri-2="https://zenodo.org/record/7732095/files/figure.png" httpUri-3="https://zenodo.org/record/7732099/files/figure.png" httpUri-4="https://zenodo.org/record/7732101/files/figure.png" pageId="7" pageNumber="483">Figures 6–10</figureCitation>
for general appearance. Morphometric data in 
<tableCitation box="[1098,1186,1771,1797]" captionStart="TABLE 3" captionStartId="9.[151,239,332,356]" captionTargetPageId="9" captionText="TABLE 3. Morphometric characteristics of G. sardashtensis (holotype FCFUK 300, paratypes FCFUK 302–305, n=4; FCFUK 309–313, n=5; FSJF 4112, n=2; non-types FCFUK 301, n=1; FCFUK 306, n=1; FCFUK 314, n=1) and G. kurdistanicus (FCFUK 330–338, n=9). Holotype, paratypes and non-types measurements are included in G. sardashtensis data." pageId="7" pageNumber="483">Table 3</tableCitation>
. Body subcylindrical. Head depressed, blunt and spade-shaped with a rounded snout margin. Maximum head width 1.6–2.3 times in head depth at eye. Dorsal head profile straight or slightly convex. Predorsal profile slightly convex, then almost straight until adipose-fin origin, sloping gently ventrally at adipose-fin base, almost straight to end of caudal peduncle. Ventral profile straight to anal-fin origin. A shallow keel between base of last dorsal-fin ray and adipose-fin origin. Nuchal plate small (
<figureCitation box="[376,444,1950,1977]" captionStart="FIGURE 5" captionStartId="6.[151,250,1508,1532]" captionTargetBox="[215,1329,694,1384]" captionTargetId="figure-248@6.[156,1431,597,1485]" captionTargetPageId="6" captionText="FIGURE 5. Premaxillary tooth plate, nuchal plate, dentary elements, and vomer plate of G. kurdistanicus (A, C, E, and G respectively), FCFUK 338, 41 mm SL, and G. sardashtensis (B, D, F, and H respectively), FCFUK 314, 38 mm SL; a and b, ventral view of premaxillary tooth plate; c and d, dorsal view of the nuchal plate; e and f, dorsal view of dentary elements; g and h, ventral view of vomer plate. Abbreviations: ppe, premaxillary proximal element; pde, premaxillary distal element; anp, anterior nuchal plate; mnp, middle nuchal plate; pnp, posterior nuchal plate; spl, spinlet; fdr, first dorsal-fin ray; vt, villiform teeth; vp, vomerine plate; ps, posterior stem." figureDoi="http://doi.org/10.5281/zenodo.7732089" httpUri="https://zenodo.org/record/7732089/files/figure.png" pageId="7" pageNumber="483">Fig. 5</figureCitation>
). Caudal-peduncle depth 1.6–2.3 times in its length. Body depth at dorsal-fin origin 1.3– 1.9 times in caudal-peduncle depth. Anus covered by tip of adpressed pelvic fin (
<figureCitation box="[1049,1157,1987,2013]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1953,1977]" captionStartId-1="11.[151,250,1047,1071]" captionTargetBox-0="[155,1433,597,1930]" captionTargetBox-1="[152,1435,181,1023]" captionTargetId-0="figure-204@10.[155,1433,597,1930]" captionTargetId-1="figure-24@11.[152,1435,181,1023]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-1="FIGURE 8. Glyptothorax sardashtensis, paratypes, from left: FCFUK 302, 51 mm SL; FCFUK 303, paratype, 42 mm SL. Lesser Zab, Iran." figureDoi-0="http://doi.org/10.5281/zenodo.7732093" figureDoi-1="http://doi.org/10.5281/zenodo.7732095" httpUri-0="https://zenodo.org/record/7732093/files/figure.png" httpUri-1="https://zenodo.org/record/7732095/files/figure.png" pageId="7" pageNumber="483">Figs. 7–8</figureCitation>
). Skin of head, back, and flanks without tubercles. Few, soft, small warts around thoracic adhesive apparatus, medial pit, edges of gill cover, and dorsal surface of first unbranched pectoral-fin ray. Snout 1.5–2.0 times longer than interorbital distance. Lateral line complete. Eye ovoid, horizontal axis longest. Dorsal-fin origin situated closer to adipose-fin origin than to tip of snout. Predorsal length 1.1–1.3 times in distance between dorsal-fin origin and adipose-fin origin. Posterior dorsalfin margin straight or slightly convex. Last unbranched dorsal-fin ray 11–15% SL; without serrations, distal 20% of its length soft, poorly ossified. First branched dorsal-fin ray longest, its length 13–18% SL. Anterior adipose-fin margin straight or slightly convex, posterior margin round. Adipose-fin origin above or very slightly in front of vertical of anal-fin origin. Caudal fin forked with rounded lobes, lower lobe slightly wider than upper lobe. Anterior anal-fin margin straight or slightly convex, posterior margin straight to slightly concave. Pelvic-fin origin below or slightly behind vertical of dorsal-fin origin. Its anterior margin slightly convex. Pelvic fin reaching anal-fin origin, its length 1.1–1.6 times longer in distance between pelvic-fin origin and anus. Distance between anus and anal-fin origin 4.7–6.8% SL. Pectoral fin reaching vertical of base of last branched dorsal-fin ray, its length 0.7–1.0 times in distance between pectoral and pelvic-fin origins. Posterior-fin margin slightly concave, anterior margin smooth, posterior margin with 9–11 serrations; distal 25% of ray soft without serrae (
<figureCitation box="[1009,1090,619,645]" captionStart="FIGURE 11" captionStartId="12.[151,250,1152,1176]" captionTargetBox="[219,1406,761,1028]" captionTargetId="figure-78@12.[153,1434,682,1128]" captionTargetPageId="12" captionText="FIGURE 11. Pectoral-fin spine of Glyptothorax sardashtensis. From left to right FCFUK 320, 31 mm SL, FCFUK 314, 38 mm SL, and FCFUK 301, 58 mm SL. Numbers indicate the number posterior serrae, pink is the ossified part." figureDoi="http://doi.org/10.5281/zenodo.7732103" httpUri="https://zenodo.org/record/7732103/files/figure.png" pageId="8" pageNumber="484">Fig. 11</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732091" ID-Zenodo-Dep="7732091" box="[151,1105,1114,1139]" httpUri="https://zenodo.org/record/7732091/files/figure.png" pageId="8" pageNumber="484" startId="8.[151,250,1114,1138]" targetBox="[156,1431,669,1091]" targetPageId="8">
<paragraph blockId="8.[151,1105,1114,1139]" box="[151,1105,1114,1139]" pageId="8" pageNumber="484">
<emphasis bold="true" box="[151,275,1114,1139]" pageId="8" pageNumber="484">FIGURE 6.</emphasis>
<taxonomicName authority=", FCFUK" authorityName="FCFUK" box="[281,654,1115,1139]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="484" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[281,560,1115,1139]" italics="true" pageId="8" pageNumber="484">Glyptothorax sardashtensis</emphasis>
, FCFUK
</taxonomicName>
300, holotype, 60 mm SL; Lesser Zab, Iran.
</paragraph>
</caption>
<paragraph blockId="8.[151,1437,1185,2039]" pageId="8" pageNumber="484">Dorsal fin with two visible unbranched rays and 5½ (n=18) branched rays. Caudal fin with ii,7+7,ii (10), ii,8+7,ii (5), i,7+7,ii (1), i,8+8,i (1), ii,9+9,ii (1) principal rays. Anal fin with one unbranched and 5½ (10), 6½ (8) branched fin rays. Pelvic fin with one unbranched and 5 (18) branched rays. Pectoral fin with one unbranched and 7 (11), 8 (7) branched fin rays.</paragraph>
<paragraph blockId="8.[151,1437,1185,2039]" pageId="8" pageNumber="484">
Thoracic adhesive apparatus with striae or narrow longitudinal pleats located in a rhombic field extending from isthmus to base of second branched pectoral-fin ray (
<figureCitation box="[817,916,1365,1391]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1953,1977]" captionStartId-1="11.[151,250,1047,1071]" captionTargetBox-0="[155,1433,597,1930]" captionTargetBox-1="[152,1435,181,1023]" captionTargetId-0="figure-204@10.[155,1433,597,1930]" captionTargetId-1="figure-24@11.[152,1435,181,1023]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Glyptothorax sardashtensis, FCFUK 300, holotype, 60 mm SL; Lesser Zab, Iran." captionText-1="FIGURE 8. Glyptothorax sardashtensis, paratypes, from left: FCFUK 302, 51 mm SL; FCFUK 303, paratype, 42 mm SL. Lesser Zab, Iran." figureDoi-0="http://doi.org/10.5281/zenodo.7732093" figureDoi-1="http://doi.org/10.5281/zenodo.7732095" httpUri-0="https://zenodo.org/record/7732093/files/figure.png" httpUri-1="https://zenodo.org/record/7732095/files/figure.png" pageId="8" pageNumber="484">Fig. 7–8</figureCitation>
). Anteromedial striae few and short (
<figureCitation box="[1338,1423,1365,1391]" captionStart="FIGURE 12" captionStartId="14.[151,250,1043,1067]" captionTargetBox="[155,1432,181,1019]" captionTargetId="figure-20@14.[155,1432,181,1019]" captionTargetPageId="14" captionText="FIGURE 12. Glyptothorax kurdistanicus, upper row, from left: FCFUK 339, 27 mm SL; FCFUK 337, 43 mm SL; FCFUK 332, 67 mm SL; G. sardashtensis, lower row, from left: FCFUK 322, 21 mm SL; FSJF 4112, 52 mm SL; FCFUK 305, 55 mm SL; Lesser Zab, Iran." figureDoi="http://doi.org/10.5281/zenodo.7732107" httpUri="https://zenodo.org/record/7732107/files/figure.png" pageId="8" pageNumber="484">Fig. 12</figureCitation>
). Medial pit wide, without striae, its anterior end roundish. Thoracic adhesive apparatus’s anterolateral edges almost straight or slightly concave, its length is 0.8–1.2 times its width. Barbels in four pairs. Maxillary barbel broad and thick, reaching almost to pectoral-fin base, to fourth or fifth dorsal-fin ray in some individuals. Inner mandibular barbel reaching to isthmus. Outer mandibular barbel reaching to posterior edge of gill cover, not reaching pectoralfin origin. Nasal barbel reaching almost to anterior eye margin. Mouth inferior, when closed, exposing a crescentshaped band of premaxillary teeth. Anterior and posterior nares closely spaced and separated only by base of nasal barbel. Anterior nasal circular, posterior triangular with two external flaps.
</paragraph>
<paragraph blockId="8.[151,1437,1185,2039]" lastBlockId="9.[151,1436,150,285]" lastPageId="9" lastPageNumber="485" pageId="8" pageNumber="484">
<emphasis bold="true" box="[199,323,1653,1679]" pageId="8" pageNumber="484">Osteology.</emphasis>
Premaxillary tooth plate crescent shaped consisting of four bony elements. One pair of proximal elements located medially and one pair of distal elements located laterally, all of which are connected by connective tissue (
<figureCitation box="[233,323,1725,1752]" captionStart="FIGURE 5" captionStartId="6.[151,250,1508,1532]" captionTargetBox="[215,1329,694,1384]" captionTargetId="figure-248@6.[156,1431,597,1485]" captionTargetPageId="6" captionText="FIGURE 5. Premaxillary tooth plate, nuchal plate, dentary elements, and vomer plate of G. kurdistanicus (A, C, E, and G respectively), FCFUK 338, 41 mm SL, and G. sardashtensis (B, D, F, and H respectively), FCFUK 314, 38 mm SL; a and b, ventral view of premaxillary tooth plate; c and d, dorsal view of the nuchal plate; e and f, dorsal view of dentary elements; g and h, ventral view of vomer plate. Abbreviations: ppe, premaxillary proximal element; pde, premaxillary distal element; anp, anterior nuchal plate; mnp, middle nuchal plate; pnp, posterior nuchal plate; spl, spinlet; fdr, first dorsal-fin ray; vt, villiform teeth; vp, vomerine plate; ps, posterior stem." figureDoi="http://doi.org/10.5281/zenodo.7732089" httpUri="https://zenodo.org/record/7732089/files/figure.png" pageId="8" pageNumber="484">Fig. 5B</figureCitation>
). Proximal bones pentagonal in shape, distal elements rod-shaped with straight anterior margins. Proximal element with straight inner margin, medially attached to opposite proximal element.Anterior and posterior margins straight, their outer side with anterolateral and posterolateral margins. Anterolateral margin of proximal element located at an angle with anterior margin of distal element. Premaxillary tooth plate ventrally with numerous villiform teeth in several rows, their tips slightly turned backward. Nuchal plate covered by thin skin. Anterior and middle elements, small triangular (
<figureCitation box="[546,637,1905,1932]" captionStart="FIGURE 5" captionStartId="6.[151,250,1508,1532]" captionTargetBox="[215,1329,694,1384]" captionTargetId="figure-248@6.[156,1431,597,1485]" captionTargetPageId="6" captionText="FIGURE 5. Premaxillary tooth plate, nuchal plate, dentary elements, and vomer plate of G. kurdistanicus (A, C, E, and G respectively), FCFUK 338, 41 mm SL, and G. sardashtensis (B, D, F, and H respectively), FCFUK 314, 38 mm SL; a and b, ventral view of premaxillary tooth plate; c and d, dorsal view of the nuchal plate; e and f, dorsal view of dentary elements; g and h, ventral view of vomer plate. Abbreviations: ppe, premaxillary proximal element; pde, premaxillary distal element; anp, anterior nuchal plate; mnp, middle nuchal plate; pnp, posterior nuchal plate; spl, spinlet; fdr, first dorsal-fin ray; vt, villiform teeth; vp, vomerine plate; ps, posterior stem." figureDoi="http://doi.org/10.5281/zenodo.7732089" httpUri="https://zenodo.org/record/7732089/files/figure.png" pageId="8" pageNumber="484">Fig. 5D</figureCitation>
). Anterolateral margins concave or slightly straight, forming a pointed tip in medial anterior. Medial anterior tip short reaching horizontally to level of rib of seventh vertebra. Tip is not turned upward. Posterior margin concave and its posterolateral tip turned back widely contacting posterior element. Posterior element expands horizontally, trapezoid in shape. Dentary consists of two halves connected by connective tissue at midline forming a single crescentic band (
<figureCitation box="[752,842,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1508,1532]" captionTargetBox="[215,1329,694,1384]" captionTargetId="figure-248@6.[156,1431,597,1485]" captionTargetPageId="6" captionText="FIGURE 5. Premaxillary tooth plate, nuchal plate, dentary elements, and vomer plate of G. kurdistanicus (A, C, E, and G respectively), FCFUK 338, 41 mm SL, and G. sardashtensis (B, D, F, and H respectively), FCFUK 314, 38 mm SL; a and b, ventral view of premaxillary tooth plate; c and d, dorsal view of the nuchal plate; e and f, dorsal view of dentary elements; g and h, ventral view of vomer plate. Abbreviations: ppe, premaxillary proximal element; pde, premaxillary distal element; anp, anterior nuchal plate; mnp, middle nuchal plate; pnp, posterior nuchal plate; spl, spinlet; fdr, first dorsal-fin ray; vt, villiform teeth; vp, vomerine plate; ps, posterior stem." figureDoi="http://doi.org/10.5281/zenodo.7732089" httpUri="https://zenodo.org/record/7732089/files/figure.png" pageId="9" pageNumber="485">Fig. 5F</figureCitation>
). Anterior portion of dentary more slender than its posterior part. Anterior portion with several rows of long, villiform teeth. Vomer edentulous, consisting of a large, triangular plate anteriorly, and a long stem posteriorly (
<figureCitation box="[759,848,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1508,1532]" captionTargetBox="[215,1329,694,1384]" captionTargetId="figure-248@6.[156,1431,597,1485]" captionTargetPageId="6" captionText="FIGURE 5. Premaxillary tooth plate, nuchal plate, dentary elements, and vomer plate of G. kurdistanicus (A, C, E, and G respectively), FCFUK 338, 41 mm SL, and G. sardashtensis (B, D, F, and H respectively), FCFUK 314, 38 mm SL; a and b, ventral view of premaxillary tooth plate; c and d, dorsal view of the nuchal plate; e and f, dorsal view of dentary elements; g and h, ventral view of vomer plate. Abbreviations: ppe, premaxillary proximal element; pde, premaxillary distal element; anp, anterior nuchal plate; mnp, middle nuchal plate; pnp, posterior nuchal plate; spl, spinlet; fdr, first dorsal-fin ray; vt, villiform teeth; vp, vomerine plate; ps, posterior stem." figureDoi="http://doi.org/10.5281/zenodo.7732089" httpUri="https://zenodo.org/record/7732089/files/figure.png" pageId="9" pageNumber="485">Fig. 5H</figureCitation>
). Vomer’s triangular plate concave in its anterolateral and posterior margins, resulting in a thin lateral process and a sharply pointed medial anterior tip.
</paragraph>
<caption pageId="9" pageNumber="485" startId="9.[151,239,332,356]" targetBox="[166,1406,480,1901]" targetIsTable="true" targetPageId="9">
<paragraph blockId="9.[151,1437,332,464]" pageId="9" pageNumber="485">
<emphasis bold="true" box="[151,267,332,356]" pageId="9" pageNumber="485">TABLE 3.</emphasis>
Morphometric characteristics of 
<taxonomicName box="[629,807,332,356]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="485" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[629,807,332,356]" italics="true" pageId="9" pageNumber="485">G. sardashtensis</emphasis>
</taxonomicName>
(holotype FCFUK 300, paratypes FCFUK 302–305, n=4; FCFUK 309–313, n=5; FSJF 4112, n=2; non-types FCFUK 301, n=1; FCFUK 306, n=1; FCFUK 314, n=1) and 
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="485" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis italics="true" pageId="9" pageNumber="485">G. kurdistanicus</emphasis>
</taxonomicName>
(FCFUK 330–338, n=9). Holotype, paratypes and non-types measurements are included in 
<taxonomicName box="[1261,1436,404,428]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="485" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1261,1436,404,428]" italics="true" pageId="9" pageNumber="485">G. sardashtensis</emphasis>
</taxonomicName>
data.
</paragraph>
</caption>
<paragraph pageId="9" pageNumber="485">
<table box="[166,1406,480,1901]" gridcols="10" gridrows="33" pageId="9" pageNumber="485">
<tr box="[166,1406,480,504]" gridrow="0" pageId="9" pageNumber="485" rowspan-0="1">
<th box="[591,690,480,504]" gridcol="1" gridrow="0" pageId="9" pageNumber="485">
<emphasis bold="true" box="[591,690,480,504]" pageId="9" pageNumber="485">Holotype</emphasis>
</th>
<th box="[727,1043,480,504]" colspan="4" colspanRight="3" gridcol="2" gridrow="0" pageId="9" pageNumber="485">
<emphasis bold="true" box="[727,993,480,504]" pageId="9" pageNumber="485">
<taxonomicName box="[727,899,480,504]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="485" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis bold="true" box="[727,899,480,504]" italics="true" pageId="9" pageNumber="485">G. sardashtensis</emphasis>
</taxonomicName>
(N = 15)
</emphasis>
</th>
<th box="[1089,1406,480,504]" colspan="4" colspanRight="3" gridcol="6" gridrow="0" pageId="9" pageNumber="485">
<emphasis bold="true" box="[1089,1344,480,504]" pageId="9" pageNumber="485">
<taxonomicName box="[1089,1262,480,504]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="485" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis bold="true" box="[1089,1262,480,504]" italics="true" pageId="9" pageNumber="485">G. kurdistanicus</emphasis>
</taxonomicName>
(N = 9)
</emphasis>
</th>
</tr>
<tr box="[166,1406,520,545]" gridrow="1" pageId="9" pageNumber="485" rowspan-1="1">
<th box="[166,552,520,545]" gridcol="0" gridrow="1" pageId="9" pageNumber="485">Characteristics</th>
<td box="[727,786,520,545]" gridcol="2" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[727,786,521,545]" pageId="9" pageNumber="485">mean</emphasis>
</td>
<td box="[817,861,520,545]" gridcol="3" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[817,859,520,544]" pageId="9" pageNumber="485">min</emphasis>
</td>
<td box="[908,954,520,545]" gridcol="4" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[908,954,521,545]" pageId="9" pageNumber="485">max</emphasis>
</td>
<td box="[999,1043,520,545]" gridcol="5" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[999,1031,520,544]" pageId="9" pageNumber="485">SD</emphasis>
</td>
<td box="[1089,1148,520,545]" gridcol="6" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[1089,1148,521,545]" pageId="9" pageNumber="485">mean</emphasis>
</td>
<td box="[1180,1224,520,545]" gridcol="7" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[1180,1222,520,544]" pageId="9" pageNumber="485">min</emphasis>
</td>
<td box="[1271,1328,520,545]" gridcol="8" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[1271,1317,521,545]" pageId="9" pageNumber="485">max</emphasis>
</td>
<td box="[1362,1406,520,545]" gridcol="9" gridrow="1" pageId="9" pageNumber="485">
<emphasis bold="true" box="[1362,1394,520,544]" pageId="9" pageNumber="485">SD</emphasis>
</td>
</tr>
<tr box="[166,1406,561,585]" gridrow="2" pageId="9" pageNumber="485">
<th box="[166,552,561,585]" gridcol="0" gridrow="2" pageId="9" pageNumber="485">Standard length</th>
<td box="[591,690,561,585]" gridcol="1" gridrow="2" pageId="9" pageNumber="485">60.3</td>
<td box="[727,786,561,585]" gridcol="2" gridrow="2" pageId="9" pageNumber="485">48.0</td>
<td box="[817,861,561,585]" gridcol="3" gridrow="2" pageId="9" pageNumber="485">36.7</td>
<td box="[908,954,561,585]" gridcol="4" gridrow="2" pageId="9" pageNumber="485">66.8</td>
<td box="[999,1043,561,585]" gridcol="5" gridrow="2" pageId="9" pageNumber="485">9.6</td>
<td box="[1089,1148,561,585]" gridcol="6" gridrow="2" pageId="9" pageNumber="485">57.6</td>
<td box="[1180,1224,561,585]" gridcol="7" gridrow="2" pageId="9" pageNumber="485">41.3</td>
<td box="[1271,1328,561,585]" gridcol="8" gridrow="2" pageId="9" pageNumber="485">107.7</td>
<td box="[1362,1406,561,585]" gridcol="9" gridrow="2" pageId="9" pageNumber="485">21.4</td>
</tr>
<tr box="[166,1406,601,625]" gridrow="3" pageId="9" pageNumber="485">
<th box="[166,1406,601,625]" colspan="10" colspanRight="9" gridcol="0" gridrow="3" pageId="9" pageNumber="485">
<emphasis bold="true" box="[166,482,601,625]" pageId="9" pageNumber="485">In percent of standard length</emphasis>
</th>
</tr>
<tr box="[166,1406,641,666]" gridrow="4" pageId="9" pageNumber="485">
<th box="[166,552,641,666]" gridcol="0" gridrow="4" pageId="9" pageNumber="485">Head length</th>
<td box="[591,690,641,666]" gridcol="1" gridrow="4" pageId="9" pageNumber="485">24.6</td>
<td box="[727,786,641,666]" gridcol="2" gridrow="4" pageId="9" pageNumber="485">25.8</td>
<td box="[817,861,641,666]" gridcol="3" gridrow="4" pageId="9" pageNumber="485">23.6</td>
<td box="[908,954,641,666]" gridcol="4" gridrow="4" pageId="9" pageNumber="485">29.0</td>
<td box="[999,1043,641,666]" gridcol="5" gridrow="4" pageId="9" pageNumber="485">1.7</td>
<td box="[1089,1148,641,666]" gridcol="6" gridrow="4" pageId="9" pageNumber="485">26.0</td>
<td box="[1180,1224,641,666]" gridcol="7" gridrow="4" pageId="9" pageNumber="485">23.5</td>
<td box="[1271,1328,641,666]" gridcol="8" gridrow="4" pageId="9" pageNumber="485">29.5</td>
<td box="[1362,1406,641,666]" gridcol="9" gridrow="4" pageId="9" pageNumber="485">1.8</td>
</tr>
<tr box="[166,1406,682,706]" gridrow="5" pageId="9" pageNumber="485">
<th box="[166,552,682,706]" gridcol="0" gridrow="5" pageId="9" pageNumber="485">Body depth at dorsal-fin origin</th>
<td box="[591,690,682,706]" gridcol="1" gridrow="5" pageId="9" pageNumber="485">19.5</td>
<td box="[727,786,682,706]" gridcol="2" gridrow="5" pageId="9" pageNumber="485">16.5</td>
<td box="[817,861,682,706]" gridcol="3" gridrow="5" pageId="9" pageNumber="485">13.3</td>
<td box="[908,954,682,706]" gridcol="4" gridrow="5" pageId="9" pageNumber="485">20.4</td>
<td box="[999,1043,682,706]" gridcol="5" gridrow="5" pageId="9" pageNumber="485">2.2</td>
<td box="[1089,1148,682,706]" gridcol="6" gridrow="5" pageId="9" pageNumber="485">19.2</td>
<td box="[1180,1224,682,706]" gridcol="7" gridrow="5" pageId="9" pageNumber="485">18.6</td>
<td box="[1271,1328,682,706]" gridcol="8" gridrow="5" pageId="9" pageNumber="485">20.2</td>
<td box="[1362,1406,682,706]" gridcol="9" gridrow="5" pageId="9" pageNumber="485">0.6</td>
</tr>
<tr box="[166,1406,722,746]" gridrow="6" pageId="9" pageNumber="485">
<th box="[166,552,722,746]" gridcol="0" gridrow="6" pageId="9" pageNumber="485">Predorsal length</th>
<td box="[591,690,722,746]" gridcol="1" gridrow="6" pageId="9" pageNumber="485">37.2</td>
<td box="[727,786,722,746]" gridcol="2" gridrow="6" pageId="9" pageNumber="485">38.4</td>
<td box="[817,861,722,746]" gridcol="3" gridrow="6" pageId="9" pageNumber="485">36.2</td>
<td box="[908,954,722,746]" gridcol="4" gridrow="6" pageId="9" pageNumber="485">41.7</td>
<td box="[999,1043,722,746]" gridcol="5" gridrow="6" pageId="9" pageNumber="485">1.7</td>
<td box="[1089,1148,722,746]" gridcol="6" gridrow="6" pageId="9" pageNumber="485">36.7</td>
<td box="[1180,1224,722,746]" gridcol="7" gridrow="6" pageId="9" pageNumber="485">32.2</td>
<td box="[1271,1328,722,746]" gridcol="8" gridrow="6" pageId="9" pageNumber="485">38.9</td>
<td box="[1362,1406,722,746]" gridcol="9" gridrow="6" pageId="9" pageNumber="485">1.9</td>
</tr>
<tr box="[166,1406,762,786]" gridrow="7" pageId="9" pageNumber="485">
<th box="[166,552,762,786]" gridcol="0" gridrow="7" pageId="9" pageNumber="485">Postdorsal length</th>
<td box="[591,690,762,786]" gridcol="1" gridrow="7" pageId="9" pageNumber="485">52.2</td>
<td box="[727,786,762,786]" gridcol="2" gridrow="7" pageId="9" pageNumber="485">52.3</td>
<td box="[817,861,762,786]" gridcol="3" gridrow="7" pageId="9" pageNumber="485">47.3</td>
<td box="[908,954,762,786]" gridcol="4" gridrow="7" pageId="9" pageNumber="485">58.7</td>
<td box="[999,1043,762,786]" gridcol="5" gridrow="7" pageId="9" pageNumber="485">4.0</td>
<td box="[1089,1148,762,786]" gridcol="6" gridrow="7" pageId="9" pageNumber="485">54.0</td>
<td box="[1180,1224,762,786]" gridcol="7" gridrow="7" pageId="9" pageNumber="485">48.2</td>
<td box="[1271,1328,762,786]" gridcol="8" gridrow="7" pageId="9" pageNumber="485">58.6</td>
<td box="[1362,1406,762,786]" gridcol="9" gridrow="7" pageId="9" pageNumber="485">3.8</td>
</tr>
<tr box="[166,1406,803,827]" gridrow="8" pageId="9" pageNumber="485">
<th box="[166,552,803,827]" gridcol="0" gridrow="8" pageId="9" pageNumber="485">Preanal length</th>
<td box="[591,690,803,827]" gridcol="1" gridrow="8" pageId="9" pageNumber="485">69.7</td>
<td box="[727,786,803,827]" gridcol="2" gridrow="8" pageId="9" pageNumber="485">68.1</td>
<td box="[817,861,803,827]" gridcol="3" gridrow="8" pageId="9" pageNumber="485">64.0</td>
<td box="[908,954,803,827]" gridcol="4" gridrow="8" pageId="9" pageNumber="485">73.1</td>
<td box="[999,1043,803,827]" gridcol="5" gridrow="8" pageId="9" pageNumber="485">3.1</td>
<td box="[1089,1148,803,827]" gridcol="6" gridrow="8" pageId="9" pageNumber="485">73.1</td>
<td box="[1180,1224,803,827]" gridcol="7" gridrow="8" pageId="9" pageNumber="485">67.0</td>
<td box="[1271,1328,803,827]" gridcol="8" gridrow="8" pageId="9" pageNumber="485">78.1</td>
<td box="[1362,1406,803,827]" gridcol="9" gridrow="8" pageId="9" pageNumber="485">4.3</td>
</tr>
<tr box="[166,1406,843,867]" gridrow="9" pageId="9" pageNumber="485">
<th box="[166,552,843,867]" gridcol="0" gridrow="9" pageId="9" pageNumber="485">Prepelvic length</th>
<td box="[591,690,843,867]" gridcol="1" gridrow="9" pageId="9" pageNumber="485">52.0</td>
<td box="[727,786,843,867]" gridcol="2" gridrow="9" pageId="9" pageNumber="485">51.3</td>
<td box="[817,861,843,867]" gridcol="3" gridrow="9" pageId="9" pageNumber="485">49.7</td>
<td box="[908,954,843,867]" gridcol="4" gridrow="9" pageId="9" pageNumber="485">52.5</td>
<td box="[999,1043,843,867]" gridcol="5" gridrow="9" pageId="9" pageNumber="485">0.8</td>
<td box="[1089,1148,843,867]" gridcol="6" gridrow="9" pageId="9" pageNumber="485">55.4</td>
<td box="[1180,1224,843,867]" gridcol="7" gridrow="9" pageId="9" pageNumber="485">50.9</td>
<td box="[1271,1328,843,867]" gridcol="8" gridrow="9" pageId="9" pageNumber="485">61.4</td>
<td box="[1362,1406,843,867]" gridcol="9" gridrow="9" pageId="9" pageNumber="485">3.9</td>
</tr>
<tr box="[166,1406,883,943]" gridrow="10" pageId="9" pageNumber="485">
<th box="[166,552,883,943]" gridcol="0" gridrow="10" pageId="9" pageNumber="485">Distance between pectoral and pelvic- fin origins</th>
<td box="[591,690,883,943]" gridcol="1" gridrow="10" pageId="9" pageNumber="485">31.4</td>
<td box="[727,786,883,943]" gridcol="2" gridrow="10" pageId="9" pageNumber="485">30.9</td>
<td box="[817,861,883,943]" gridcol="3" gridrow="10" pageId="9" pageNumber="485">28.6</td>
<td box="[908,954,883,943]" gridcol="4" gridrow="10" pageId="9" pageNumber="485">34.5</td>
<td box="[999,1043,883,943]" gridcol="5" gridrow="10" pageId="9" pageNumber="485">1.8</td>
<td box="[1089,1148,883,943]" gridcol="6" gridrow="10" pageId="9" pageNumber="485">32.4</td>
<td box="[1180,1224,883,943]" gridcol="7" gridrow="10" pageId="9" pageNumber="485">29.3</td>
<td box="[1271,1328,883,943]" gridcol="8" gridrow="10" pageId="9" pageNumber="485">37.9</td>
<td box="[1362,1406,883,943]" gridcol="9" gridrow="10" pageId="9" pageNumber="485">2.5</td>
</tr>
<tr box="[166,1406,959,1019]" gridrow="11" pageId="9" pageNumber="485">
<th box="[166,552,959,1019]" gridcol="0" gridrow="11" pageId="9" pageNumber="485">Distance between pelvic and anal-fin origins</th>
<td box="[591,690,959,1019]" gridcol="1" gridrow="11" pageId="9" pageNumber="485">18.1</td>
<td box="[727,786,959,1019]" gridcol="2" gridrow="11" pageId="9" pageNumber="485">17.5</td>
<td box="[817,861,959,1019]" gridcol="3" gridrow="11" pageId="9" pageNumber="485">13.9</td>
<td box="[908,954,959,1019]" gridcol="4" gridrow="11" pageId="9" pageNumber="485">21.5</td>
<td box="[999,1043,959,1019]" gridcol="5" gridrow="11" pageId="9" pageNumber="485">2.2</td>
<td box="[1089,1148,959,1019]" gridcol="6" gridrow="11" pageId="9" pageNumber="485">20.3</td>
<td box="[1180,1224,959,1019]" gridcol="7" gridrow="11" pageId="9" pageNumber="485">18.7</td>
<td box="[1271,1328,959,1019]" gridcol="8" gridrow="11" pageId="9" pageNumber="485">21.2</td>
<td box="[1362,1406,959,1019]" gridcol="9" gridrow="11" pageId="9" pageNumber="485">0.9</td>
</tr>
<tr box="[166,1406,1035,1060]" gridrow="12" pageId="9" pageNumber="485">
<th box="[166,552,1035,1060]" gridcol="0" gridrow="12" pageId="9" pageNumber="485">Depth of caudal peduncle</th>
<td box="[591,690,1035,1060]" gridcol="1" gridrow="12" pageId="9" pageNumber="485">11.9</td>
<td box="[727,786,1035,1060]" gridcol="2" gridrow="12" pageId="9" pageNumber="485">10.2</td>
<td box="[817,861,1035,1060]" gridcol="3" gridrow="12" pageId="9" pageNumber="485">8.5</td>
<td box="[908,954,1035,1060]" gridcol="4" gridrow="12" pageId="9" pageNumber="485">12.6</td>
<td box="[999,1043,1035,1060]" gridcol="5" gridrow="12" pageId="9" pageNumber="485">1.4</td>
<td box="[1089,1148,1035,1060]" gridcol="6" gridrow="12" pageId="9" pageNumber="485">11.4</td>
<td box="[1180,1224,1035,1060]" gridcol="7" gridrow="12" pageId="9" pageNumber="485">9.8</td>
<td box="[1271,1328,1035,1060]" gridcol="8" gridrow="12" pageId="9" pageNumber="485">13.4</td>
<td box="[1362,1406,1035,1060]" gridcol="9" gridrow="12" pageId="9" pageNumber="485">1.4</td>
</tr>
<tr box="[166,1406,1076,1100]" gridrow="13" pageId="9" pageNumber="485">
<th box="[166,552,1076,1100]" gridcol="0" gridrow="13" pageId="9" pageNumber="485">Length of caudal peduncle</th>
<td box="[591,690,1076,1100]" gridcol="1" gridrow="13" pageId="9" pageNumber="485">20.9</td>
<td box="[727,786,1076,1100]" gridcol="2" gridrow="13" pageId="9" pageNumber="485">19.5</td>
<td box="[817,861,1076,1100]" gridcol="3" gridrow="13" pageId="9" pageNumber="485">17.2</td>
<td box="[908,954,1076,1100]" gridcol="4" gridrow="13" pageId="9" pageNumber="485">23.3</td>
<td box="[999,1043,1076,1100]" gridcol="5" gridrow="13" pageId="9" pageNumber="485">1.6</td>
<td box="[1089,1148,1076,1100]" gridcol="6" gridrow="13" pageId="9" pageNumber="485">17.2</td>
<td box="[1180,1224,1076,1100]" gridcol="7" gridrow="13" pageId="9" pageNumber="485">15.1</td>
<td box="[1271,1328,1076,1100]" gridcol="8" gridrow="13" pageId="9" pageNumber="485">20.2</td>
<td box="[1362,1406,1076,1100]" gridcol="9" gridrow="13" pageId="9" pageNumber="485">1.9</td>
</tr>
<tr box="[166,1406,1116,1140]" gridrow="14" pageId="9" pageNumber="485">
<th box="[166,552,1116,1140]" gridcol="0" gridrow="14" pageId="9" pageNumber="485">Dorsal-fin depth</th>
<td box="[591,690,1116,1140]" gridcol="1" gridrow="14" pageId="9" pageNumber="485">13.7</td>
<td box="[727,786,1116,1140]" gridcol="2" gridrow="14" pageId="9" pageNumber="485">16.2</td>
<td box="[817,861,1116,1140]" gridcol="3" gridrow="14" pageId="9" pageNumber="485">13.2</td>
<td box="[908,954,1116,1140]" gridcol="4" gridrow="14" pageId="9" pageNumber="485">18.4</td>
<td box="[999,1043,1116,1140]" gridcol="5" gridrow="14" pageId="9" pageNumber="485">1.6</td>
<td box="[1089,1148,1116,1140]" gridcol="6" gridrow="14" pageId="9" pageNumber="485">14.8</td>
<td box="[1180,1224,1116,1140]" gridcol="7" gridrow="14" pageId="9" pageNumber="485">11.9</td>
<td box="[1271,1328,1116,1140]" gridcol="8" gridrow="14" pageId="9" pageNumber="485">16.4</td>
<td box="[1362,1406,1116,1140]" gridcol="9" gridrow="14" pageId="9" pageNumber="485">1.2</td>
</tr>
<tr box="[166,1406,1156,1180]" gridrow="15" pageId="9" pageNumber="485">
<th box="[166,552,1156,1180]" gridcol="0" gridrow="15" pageId="9" pageNumber="485">Adipose-fin length</th>
<td box="[591,690,1156,1180]" gridcol="1" gridrow="15" pageId="9" pageNumber="485">19.4</td>
<td box="[727,786,1156,1180]" gridcol="2" gridrow="15" pageId="9" pageNumber="485">17.3</td>
<td box="[817,861,1156,1180]" gridcol="3" gridrow="15" pageId="9" pageNumber="485">12.4</td>
<td box="[908,954,1156,1180]" gridcol="4" gridrow="15" pageId="9" pageNumber="485">22.6</td>
<td box="[999,1043,1156,1180]" gridcol="5" gridrow="15" pageId="9" pageNumber="485">3.6</td>
<td box="[1089,1148,1156,1180]" gridcol="6" gridrow="15" pageId="9" pageNumber="485">15.6</td>
<td box="[1180,1224,1156,1180]" gridcol="7" gridrow="15" pageId="9" pageNumber="485">11.4</td>
<td box="[1271,1328,1156,1180]" gridcol="8" gridrow="15" pageId="9" pageNumber="485">22.0</td>
<td box="[1362,1406,1156,1180]" gridcol="9" gridrow="15" pageId="9" pageNumber="485">4.6</td>
</tr>
<tr box="[166,1406,1196,1257]" gridrow="16" pageId="9" pageNumber="485">
<th box="[166,552,1196,1257]" gridcol="0" gridrow="16" pageId="9" pageNumber="485">Distance between last dorsal ray and adipose origin</th>
<td box="[591,690,1196,1257]" gridcol="1" gridrow="16" pageId="9" pageNumber="485">15.9</td>
<td box="[727,786,1196,1257]" gridcol="2" gridrow="16" pageId="9" pageNumber="485">19.5</td>
<td box="[817,861,1196,1257]" gridcol="3" gridrow="16" pageId="9" pageNumber="485">15.9</td>
<td box="[908,954,1196,1257]" gridcol="4" gridrow="16" pageId="9" pageNumber="485">21.0</td>
<td box="[999,1043,1196,1257]" gridcol="5" gridrow="16" pageId="9" pageNumber="485">1.4</td>
<td box="[1089,1148,1196,1257]" gridcol="6" gridrow="16" pageId="9" pageNumber="485">23.5</td>
<td box="[1180,1224,1196,1257]" gridcol="7" gridrow="16" pageId="9" pageNumber="485">19.7</td>
<td box="[1271,1328,1196,1257]" gridcol="8" gridrow="16" pageId="9" pageNumber="485">27.8</td>
<td box="[1362,1406,1196,1257]" gridcol="9" gridrow="16" pageId="9" pageNumber="485">2.6</td>
</tr>
<tr box="[166,1406,1273,1297]" gridrow="17" pageId="9" pageNumber="485">
<th box="[166,552,1273,1297]" gridcol="0" gridrow="17" pageId="9" pageNumber="485">Anal-fin base length</th>
<td box="[591,690,1273,1297]" gridcol="1" gridrow="17" pageId="9" pageNumber="485">11.5</td>
<td box="[727,786,1273,1297]" gridcol="2" gridrow="17" pageId="9" pageNumber="485">11.2</td>
<td box="[817,861,1273,1297]" gridcol="3" gridrow="17" pageId="9" pageNumber="485">9.6</td>
<td box="[908,954,1273,1297]" gridcol="4" gridrow="17" pageId="9" pageNumber="485">12.5</td>
<td box="[999,1043,1273,1297]" gridcol="5" gridrow="17" pageId="9" pageNumber="485">0.9</td>
<td box="[1089,1148,1273,1297]" gridcol="6" gridrow="17" pageId="9" pageNumber="485">11.4</td>
<td box="[1180,1224,1273,1297]" gridcol="7" gridrow="17" pageId="9" pageNumber="485">9.9</td>
<td box="[1271,1328,1273,1297]" gridcol="8" gridrow="17" pageId="9" pageNumber="485">13.3</td>
<td box="[1362,1406,1273,1297]" gridcol="9" gridrow="17" pageId="9" pageNumber="485">1.1</td>
</tr>
<tr box="[166,1406,1313,1337]" gridrow="18" pageId="9" pageNumber="485">
<th box="[166,552,1313,1337]" gridcol="0" gridrow="18" pageId="9" pageNumber="485">Pectoral-fin length</th>
<td box="[591,690,1313,1337]" gridcol="1" gridrow="18" pageId="9" pageNumber="485">23.6</td>
<td box="[727,786,1313,1337]" gridcol="2" gridrow="18" pageId="9" pageNumber="485">24.0</td>
<td box="[817,861,1313,1337]" gridcol="3" gridrow="18" pageId="9" pageNumber="485">20.8</td>
<td box="[908,954,1313,1337]" gridcol="4" gridrow="18" pageId="9" pageNumber="485">27.0</td>
<td box="[999,1043,1313,1337]" gridcol="5" gridrow="18" pageId="9" pageNumber="485">1.4</td>
<td box="[1089,1148,1313,1337]" gridcol="6" gridrow="18" pageId="9" pageNumber="485">22.4</td>
<td box="[1180,1224,1313,1337]" gridcol="7" gridrow="18" pageId="9" pageNumber="485">19.1</td>
<td box="[1271,1328,1313,1337]" gridcol="8" gridrow="18" pageId="9" pageNumber="485">25.1</td>
<td box="[1362,1406,1313,1337]" gridcol="9" gridrow="18" pageId="9" pageNumber="485">1.8</td>
</tr>
<tr box="[166,1406,1353,1377]" gridrow="19" pageId="9" pageNumber="485">
<th box="[166,552,1353,1377]" gridcol="0" gridrow="19" pageId="9" pageNumber="485">Pelvic-fin length</th>
<td box="[591,690,1353,1377]" gridcol="1" gridrow="19" pageId="9" pageNumber="485">14.6</td>
<td box="[727,786,1353,1377]" gridcol="2" gridrow="19" pageId="9" pageNumber="485">17.0</td>
<td box="[817,861,1353,1377]" gridcol="3" gridrow="19" pageId="9" pageNumber="485">14.6</td>
<td box="[908,954,1353,1377]" gridcol="4" gridrow="19" pageId="9" pageNumber="485">18.7</td>
<td box="[999,1043,1353,1377]" gridcol="5" gridrow="19" pageId="9" pageNumber="485">1.2</td>
<td box="[1089,1148,1353,1377]" gridcol="6" gridrow="19" pageId="9" pageNumber="485">16.3</td>
<td box="[1180,1224,1353,1377]" gridcol="7" gridrow="19" pageId="9" pageNumber="485">14.5</td>
<td box="[1271,1328,1353,1377]" gridcol="8" gridrow="19" pageId="9" pageNumber="485">18.6</td>
<td box="[1362,1406,1353,1377]" gridcol="9" gridrow="19" pageId="9" pageNumber="485">1.4</td>
</tr>
<tr box="[166,1406,1393,1417]" gridrow="20" pageId="9" pageNumber="485">
<th box="[166,1406,1393,1417]" colspan="10" colspanRight="9" gridcol="0" gridrow="20" pageId="9" pageNumber="485">
<emphasis bold="true" box="[166,437,1393,1417]" pageId="9" pageNumber="485">In percent of head length</emphasis>
</th>
</tr>
<tr box="[166,1406,1434,1458]" gridrow="21" pageId="9" pageNumber="485">
<th box="[166,552,1434,1458]" gridcol="0" gridrow="21" pageId="9" pageNumber="485">Head depth at eye</th>
<td box="[591,690,1434,1458]" gridcol="1" gridrow="21" pageId="9" pageNumber="485">51.8</td>
<td box="[727,786,1434,1458]" gridcol="2" gridrow="21" pageId="9" pageNumber="485">40.4</td>
<td box="[817,861,1434,1458]" gridcol="3" gridrow="21" pageId="9" pageNumber="485">37</td>
<td box="[908,954,1434,1458]" gridcol="4" gridrow="21" pageId="9" pageNumber="485">52</td>
<td box="[999,1043,1434,1458]" gridcol="5" gridrow="21" pageId="9" pageNumber="485">3.8</td>
<td box="[1089,1148,1434,1458]" gridcol="6" gridrow="21" pageId="9" pageNumber="485">41.6</td>
<td box="[1180,1224,1434,1458]" gridcol="7" gridrow="21" pageId="9" pageNumber="485">40</td>
<td box="[1271,1328,1434,1458]" gridcol="8" gridrow="21" pageId="9" pageNumber="485">45</td>
<td box="[1362,1406,1434,1458]" gridcol="9" gridrow="21" pageId="9" pageNumber="485">2.3</td>
</tr>
<tr box="[166,1406,1474,1498]" gridrow="22" pageId="9" pageNumber="485">
<th box="[166,552,1474,1498]" gridcol="0" gridrow="22" pageId="9" pageNumber="485">Snout length</th>
<td box="[591,690,1474,1498]" gridcol="1" gridrow="22" pageId="9" pageNumber="485">56.7</td>
<td box="[727,786,1474,1498]" gridcol="2" gridrow="22" pageId="9" pageNumber="485">53.3</td>
<td box="[817,861,1474,1498]" gridcol="3" gridrow="22" pageId="9" pageNumber="485">48</td>
<td box="[908,954,1474,1498]" gridcol="4" gridrow="22" pageId="9" pageNumber="485">59</td>
<td box="[999,1043,1474,1498]" gridcol="5" gridrow="22" pageId="9" pageNumber="485">2.8</td>
<td box="[1089,1148,1474,1498]" gridcol="6" gridrow="22" pageId="9" pageNumber="485">52.6</td>
<td box="[1180,1224,1474,1498]" gridcol="7" gridrow="22" pageId="9" pageNumber="485">46</td>
<td box="[1271,1328,1474,1498]" gridcol="8" gridrow="22" pageId="9" pageNumber="485">58</td>
<td box="[1362,1406,1474,1498]" gridcol="9" gridrow="22" pageId="9" pageNumber="485">3.8</td>
</tr>
<tr box="[166,1406,1514,1539]" gridrow="23" pageId="9" pageNumber="485">
<th box="[166,552,1514,1539]" gridcol="0" gridrow="23" pageId="9" pageNumber="485">Eye diameter</th>
<td box="[591,690,1514,1539]" gridcol="1" gridrow="23" pageId="9" pageNumber="485">12.8</td>
<td box="[727,786,1514,1539]" gridcol="2" gridrow="23" pageId="9" pageNumber="485">10.3</td>
<td box="[817,861,1514,1539]" gridcol="3" gridrow="23" pageId="9" pageNumber="485">8</td>
<td box="[908,954,1514,1539]" gridcol="4" gridrow="23" pageId="9" pageNumber="485">13</td>
<td box="[999,1043,1514,1539]" gridcol="5" gridrow="23" pageId="9" pageNumber="485">1.5</td>
<td box="[1089,1148,1514,1539]" gridcol="6" gridrow="23" pageId="9" pageNumber="485">7.8</td>
<td box="[1180,1224,1514,1539]" gridcol="7" gridrow="23" pageId="9" pageNumber="485">5</td>
<td box="[1271,1328,1514,1539]" gridcol="8" gridrow="23" pageId="9" pageNumber="485">12</td>
<td box="[1362,1406,1514,1539]" gridcol="9" gridrow="23" pageId="9" pageNumber="485">2.2</td>
</tr>
<tr box="[166,1406,1554,1579]" gridrow="24" pageId="9" pageNumber="485">
<th box="[166,552,1554,1579]" gridcol="0" gridrow="24" pageId="9" pageNumber="485">Postorbital distance</th>
<td box="[591,690,1554,1579]" gridcol="1" gridrow="24" pageId="9" pageNumber="485">35.0</td>
<td box="[727,786,1554,1579]" gridcol="2" gridrow="24" pageId="9" pageNumber="485">39.4</td>
<td box="[817,861,1554,1579]" gridcol="3" gridrow="24" pageId="9" pageNumber="485">35</td>
<td box="[908,954,1554,1579]" gridcol="4" gridrow="24" pageId="9" pageNumber="485">46</td>
<td box="[999,1043,1554,1579]" gridcol="5" gridrow="24" pageId="9" pageNumber="485">3.4</td>
<td box="[1089,1148,1554,1579]" gridcol="6" gridrow="24" pageId="9" pageNumber="485">44.5</td>
<td box="[1180,1224,1554,1579]" gridcol="7" gridrow="24" pageId="9" pageNumber="485">38</td>
<td box="[1271,1328,1554,1579]" gridcol="8" gridrow="24" pageId="9" pageNumber="485">50</td>
<td box="[1362,1406,1554,1579]" gridcol="9" gridrow="24" pageId="9" pageNumber="485">4.4</td>
</tr>
<tr box="[166,1406,1595,1619]" gridrow="25" pageId="9" pageNumber="485">
<th box="[166,552,1595,1619]" gridcol="0" gridrow="25" pageId="9" pageNumber="485">Maximum head width</th>
<td box="[591,690,1595,1619]" gridcol="1" gridrow="25" pageId="9" pageNumber="485">81.8</td>
<td box="[727,786,1595,1619]" gridcol="2" gridrow="25" pageId="9" pageNumber="485">80.4</td>
<td box="[817,861,1595,1619]" gridcol="3" gridrow="25" pageId="9" pageNumber="485">72</td>
<td box="[908,954,1595,1619]" gridcol="4" gridrow="25" pageId="9" pageNumber="485">88</td>
<td box="[999,1043,1595,1619]" gridcol="5" gridrow="25" pageId="9" pageNumber="485">4.7</td>
<td box="[1089,1148,1595,1619]" gridcol="6" gridrow="25" pageId="9" pageNumber="485">88.3</td>
<td box="[1180,1224,1595,1619]" gridcol="7" gridrow="25" pageId="9" pageNumber="485">82</td>
<td box="[1271,1328,1595,1619]" gridcol="8" gridrow="25" pageId="9" pageNumber="485">94</td>
<td box="[1362,1406,1595,1619]" gridcol="9" gridrow="25" pageId="9" pageNumber="485">4.5</td>
</tr>
<tr box="[166,1406,1635,1659]" gridrow="26" pageId="9" pageNumber="485">
<th box="[166,552,1635,1659]" gridcol="0" gridrow="26" pageId="9" pageNumber="485">Interorbital distance</th>
<td box="[591,690,1635,1659]" gridcol="1" gridrow="26" pageId="9" pageNumber="485">31.0</td>
<td box="[727,786,1635,1659]" gridcol="2" gridrow="26" pageId="9" pageNumber="485">30.9</td>
<td box="[817,861,1635,1659]" gridcol="3" gridrow="26" pageId="9" pageNumber="485">26</td>
<td box="[908,954,1635,1659]" gridcol="4" gridrow="26" pageId="9" pageNumber="485">35</td>
<td box="[999,1043,1635,1659]" gridcol="5" gridrow="26" pageId="9" pageNumber="485">2.7</td>
<td box="[1089,1148,1635,1659]" gridcol="6" gridrow="26" pageId="9" pageNumber="485">33.7</td>
<td box="[1180,1224,1635,1659]" gridcol="7" gridrow="26" pageId="9" pageNumber="485">25</td>
<td box="[1271,1328,1635,1659]" gridcol="8" gridrow="26" pageId="9" pageNumber="485">39</td>
<td box="[1362,1406,1635,1659]" gridcol="9" gridrow="26" pageId="9" pageNumber="485">5.6</td>
</tr>
<tr box="[166,1406,1675,1699]" gridrow="27" pageId="9" pageNumber="485">
<th box="[166,552,1675,1699]" gridcol="0" gridrow="27" pageId="9" pageNumber="485">Length of inner mandibular barbel</th>
<td box="[591,690,1675,1699]" gridcol="1" gridrow="27" pageId="9" pageNumber="485">25.8</td>
<td box="[727,786,1675,1699]" gridcol="2" gridrow="27" pageId="9" pageNumber="485">25.9</td>
<td box="[817,861,1675,1699]" gridcol="3" gridrow="27" pageId="9" pageNumber="485">22</td>
<td box="[908,954,1675,1699]" gridcol="4" gridrow="27" pageId="9" pageNumber="485">31</td>
<td box="[999,1043,1675,1699]" gridcol="5" gridrow="27" pageId="9" pageNumber="485">2.3</td>
<td box="[1089,1148,1675,1699]" gridcol="6" gridrow="27" pageId="9" pageNumber="485">30.7</td>
<td box="[1180,1224,1675,1699]" gridcol="7" gridrow="27" pageId="9" pageNumber="485">27</td>
<td box="[1271,1328,1675,1699]" gridcol="8" gridrow="27" pageId="9" pageNumber="485">34</td>
<td box="[1362,1406,1675,1699]" gridcol="9" gridrow="27" pageId="9" pageNumber="485">2.7</td>
</tr>
<tr box="[166,1406,1715,1740]" gridrow="28" pageId="9" pageNumber="485">
<th box="[166,552,1715,1740]" gridcol="0" gridrow="28" pageId="9" pageNumber="485">Length of outer mandibular barbel</th>
<td box="[591,690,1715,1740]" gridcol="1" gridrow="28" pageId="9" pageNumber="485">52.9</td>
<td box="[727,786,1715,1740]" gridcol="2" gridrow="28" pageId="9" pageNumber="485">43.3</td>
<td box="[817,861,1715,1740]" gridcol="3" gridrow="28" pageId="9" pageNumber="485">33</td>
<td box="[908,954,1715,1740]" gridcol="4" gridrow="28" pageId="9" pageNumber="485">53</td>
<td box="[999,1043,1715,1740]" gridcol="5" gridrow="28" pageId="9" pageNumber="485">6.0</td>
<td box="[1089,1148,1715,1740]" gridcol="6" gridrow="28" pageId="9" pageNumber="485">51.3</td>
<td box="[1180,1224,1715,1740]" gridcol="7" gridrow="28" pageId="9" pageNumber="485">45</td>
<td box="[1271,1328,1715,1740]" gridcol="8" gridrow="28" pageId="9" pageNumber="485">59</td>
<td box="[1362,1406,1715,1740]" gridcol="9" gridrow="28" pageId="9" pageNumber="485">6.7</td>
</tr>
<tr box="[166,1406,1756,1780]" gridrow="29" pageId="9" pageNumber="485">
<th box="[166,552,1756,1780]" gridcol="0" gridrow="29" pageId="9" pageNumber="485">Length of maxillary barbel</th>
<td box="[591,690,1756,1780]" gridcol="1" gridrow="29" pageId="9" pageNumber="485">94.9</td>
<td box="[727,786,1756,1780]" gridcol="2" gridrow="29" pageId="9" pageNumber="485">71.3</td>
<td box="[817,861,1756,1780]" gridcol="3" gridrow="29" pageId="9" pageNumber="485">53</td>
<td box="[908,954,1756,1780]" gridcol="4" gridrow="29" pageId="9" pageNumber="485">95</td>
<td box="[999,1043,1756,1780]" gridcol="5" gridrow="29" pageId="9" pageNumber="485">13.5</td>
<td box="[1089,1148,1756,1780]" gridcol="6" gridrow="29" pageId="9" pageNumber="485">76.0</td>
<td box="[1180,1224,1756,1780]" gridcol="7" gridrow="29" pageId="9" pageNumber="485">58</td>
<td box="[1271,1328,1756,1780]" gridcol="8" gridrow="29" pageId="9" pageNumber="485">91</td>
<td box="[1362,1406,1756,1780]" gridcol="9" gridrow="29" pageId="9" pageNumber="485">8.9</td>
</tr>
<tr box="[166,1406,1796,1820]" gridrow="30" pageId="9" pageNumber="485">
<th box="[166,552,1796,1820]" gridcol="0" gridrow="30" pageId="9" pageNumber="485">Length of nasal barbel</th>
<td box="[591,690,1796,1820]" gridcol="1" gridrow="30" pageId="9" pageNumber="485">33.7</td>
<td box="[727,786,1796,1820]" gridcol="2" gridrow="30" pageId="9" pageNumber="485">30.9</td>
<td box="[817,861,1796,1820]" gridcol="3" gridrow="30" pageId="9" pageNumber="485">26</td>
<td box="[908,954,1796,1820]" gridcol="4" gridrow="30" pageId="9" pageNumber="485">39</td>
<td box="[999,1043,1796,1820]" gridcol="5" gridrow="30" pageId="9" pageNumber="485">3.5</td>
<td box="[1089,1148,1796,1820]" gridcol="6" gridrow="30" pageId="9" pageNumber="485">31.9</td>
<td box="[1180,1224,1796,1820]" gridcol="7" gridrow="30" pageId="9" pageNumber="485">28</td>
<td box="[1271,1328,1796,1820]" gridcol="8" gridrow="30" pageId="9" pageNumber="485">42</td>
<td box="[1362,1406,1796,1820]" gridcol="9" gridrow="30" pageId="9" pageNumber="485">5.7</td>
</tr>
<tr box="[166,1406,1836,1860]" gridrow="31" pageId="9" pageNumber="485">
<th box="[166,552,1836,1860]" gridcol="0" gridrow="31" pageId="9" pageNumber="485">Thoracic apparatus length</th>
<td box="[591,690,1836,1860]" gridcol="1" gridrow="31" pageId="9" pageNumber="485">50.7</td>
<td box="[727,786,1836,1860]" gridcol="2" gridrow="31" pageId="9" pageNumber="485">52.3</td>
<td box="[817,861,1836,1860]" gridcol="3" gridrow="31" pageId="9" pageNumber="485">45</td>
<td box="[908,954,1836,1860]" gridcol="4" gridrow="31" pageId="9" pageNumber="485">60</td>
<td box="[999,1043,1836,1860]" gridcol="5" gridrow="31" pageId="9" pageNumber="485">4.6</td>
<td box="[1089,1148,1836,1860]" gridcol="6" gridrow="31" pageId="9" pageNumber="485">45.3</td>
<td box="[1180,1224,1836,1860]" gridcol="7" gridrow="31" pageId="9" pageNumber="485">39</td>
<td box="[1271,1328,1836,1860]" gridcol="8" gridrow="31" pageId="9" pageNumber="485">52</td>
<td box="[1362,1406,1836,1860]" gridcol="9" gridrow="31" pageId="9" pageNumber="485">4.0</td>
</tr>
<tr box="[166,1406,1876,1901]" gridrow="32" pageId="9" pageNumber="485">
<th box="[166,552,1876,1901]" gridcol="0" gridrow="32" pageId="9" pageNumber="485">Thoracic apparatus width</th>
<td box="[591,690,1876,1901]" gridcol="1" gridrow="32" pageId="9" pageNumber="485">59.6</td>
<td box="[727,786,1876,1901]" gridcol="2" gridrow="32" pageId="9" pageNumber="485">53.0</td>
<td box="[817,861,1876,1901]" gridcol="3" gridrow="32" pageId="9" pageNumber="485">41</td>
<td box="[908,954,1876,1901]" gridcol="4" gridrow="32" pageId="9" pageNumber="485">60</td>
<td box="[999,1043,1876,1901]" gridcol="5" gridrow="32" pageId="9" pageNumber="485">5.7</td>
<td box="[1089,1148,1876,1901]" gridcol="6" gridrow="32" pageId="9" pageNumber="485">50.2</td>
<td box="[1180,1224,1876,1901]" gridcol="7" gridrow="32" pageId="9" pageNumber="485">43</td>
<td box="[1271,1328,1876,1901]" gridcol="8" gridrow="32" pageId="9" pageNumber="485">55</td>
<td box="[1362,1406,1876,1901]" gridcol="9" gridrow="32" pageId="9" pageNumber="485">4.1</td>
</tr>
</table>
</paragraph>
<paragraph blockId="9.[151,1436,1948,2011]" lastBlockId="10.[151,1437,151,573]" lastPageId="10" lastPageNumber="486" pageId="9" pageNumber="485">
<emphasis bold="true" box="[199,335,1948,1974]" pageId="9" pageNumber="485">Coloration.</emphasis>
In 70% ethanol: background colour of head and body brown, ventral surface yellowish-brown, thoracic adhesive apparatus yellowish to pale-grey. Body immaculate, some individuals with few, minute dots, darker than background coloration around head, dorsal-fin base, adipose-fin base, and caudal-peduncle.An upper and lower yellowish blotch at posteriormost caudal peduncle. Three pale-grey blotches, organised in a crescent-shaped arch in front of dorsal-fin origin, fused in some individuals. A yellow blotch at and on back behind posteriormost dorsal-fin base, faded in most individuals. Dorsal fin with a brown base and last unbranched dorsal-fin ray brown. Dorsal-fin membranes hyaline or yellowish, rays brown or yellowish, usually brown in middle of ray and yellowish at tip. Adipose fin with a large, elongate, yellowish blotch at origin and a small yellowish blotch at posterior margin. Caudal fin dark-brown, with a blackish base, followed by a yellowish band, usually reduced into two yellowish blotches, and yellowish caudal-fin tips. All other fins yellowish with a brown base, a central, wide, brown band, often reduced to brown fin-rays. Maxillary and nasal barbels grey or blackish dorsally, pale-grey ventrally and velum pale-grey or beige. Mandibular barbels beige or pale-grey.
</paragraph>
<paragraph blockId="10.[151,1437,151,573]" pageId="10" pageNumber="486">In life: colour pattern identical to preserved individuals, background colour of head and body olive-green. Pattern on body and fins yellow.</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732093" ID-Zenodo-Dep="7732093" box="[151,1105,1953,1977]" httpUri="https://zenodo.org/record/7732093/files/figure.png" pageId="10" pageNumber="486" startId="10.[151,250,1953,1977]" targetBox="[155,1433,597,1930]" targetPageId="10">
<paragraph blockId="10.[151,1105,1953,1977]" box="[151,1105,1953,1977]" pageId="10" pageNumber="486">
<emphasis bold="true" box="[151,275,1953,1977]" pageId="10" pageNumber="486">FIGURE 7.</emphasis>
<taxonomicName authority=", FCFUK" authorityName="FCFUK" box="[281,654,1953,1977]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="486" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[281,560,1953,1977]" italics="true" pageId="10" pageNumber="486">Glyptothorax sardashtensis</emphasis>
, FCFUK
</taxonomicName>
300, holotype, 60 mm SL; Lesser Zab, Iran.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732095" ID-Zenodo-Dep="7732095" httpUri="https://zenodo.org/record/7732095/files/figure.png" pageId="11" pageNumber="487" startId="11.[151,250,1047,1071]" targetBox="[152,1435,181,1023]" targetPageId="11">
<paragraph blockId="11.[151,1437,1047,1107]" pageId="11" pageNumber="487">
<emphasis bold="true" box="[151,277,1047,1071]" pageId="11" pageNumber="487">FIGURE 8.</emphasis>
<taxonomicName box="[286,567,1047,1071]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="487" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[286,567,1047,1071]" italics="true" pageId="11" pageNumber="487">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
, paratypes, from left: FCFUK 302, 51 mm SL; FCFUK 303, paratype, 42 mm SL. Lesser Zab, Iran.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732099" ID-Zenodo-Dep="7732099" httpUri="https://zenodo.org/record/7732099/files/figure.png" pageId="11" pageNumber="487" startId="11.[151,250,1933,1957]" targetBox="[152,1435,1132,1910]" targetPageId="11">
<paragraph blockId="11.[151,1436,1933,1994]" pageId="11" pageNumber="487">
<emphasis bold="true" box="[151,278,1933,1957]" pageId="11" pageNumber="487">FIGURE 9.</emphasis>
<taxonomicName box="[287,568,1933,1957]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="487" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[287,568,1933,1957]" italics="true" pageId="11" pageNumber="487">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
, from left: FCFUK 300, holotype, 60 mm SL; FCFUK 302, paratype, 51 mm SL; FCFUK 305, paratype, 55 mm SL; Lesser Zab, Iran.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732101" ID-Zenodo-Dep="7732101" httpUri="https://zenodo.org/record/7732101/files/figure.png" pageId="12" pageNumber="488" startId="12.[151,250,562,586]" targetBox="[154,1433,181,538]" targetPageId="12">
<paragraph blockId="12.[151,1436,562,658]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[151,286,562,586]" pageId="12" pageNumber="488">FIGURE 10.</emphasis>
Comparison of the presence or absence of spots and blotches in 
<taxonomicName box="[915,1081,562,586]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[915,1081,562,586]" italics="true" pageId="12" pageNumber="488">G. sardashtensis</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1126,1291,562,586]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1126,1291,562,586]" italics="true" pageId="12" pageNumber="488">G. kurdistanicus</emphasis>
</taxonomicName>
. 
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis italics="true" pageId="12" pageNumber="488">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
, from left: FCFUK 306, 51 mm SL; FCFUK 321, 45 mm SL; Lesser Zab, Iran. 
<taxonomicName box="[1098,1375,598,622]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1098,1375,598,622]" italics="true" pageId="12" pageNumber="488">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, from right: FCFUK 332, 67 mm SL; FCFUK 337, 43 mm SL; Lesser Zab, Iran.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732103" ID-Zenodo-Dep="7732103" httpUri="https://zenodo.org/record/7732103/files/figure.png" pageId="12" pageNumber="488" startId="12.[151,250,1152,1176]" targetBox="[219,1406,761,1028]" targetPageId="12">
<paragraph blockId="12.[151,1436,1152,1212]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[151,286,1152,1176]" pageId="12" pageNumber="488">FIGURE 11.</emphasis>
Pectoral-fin spine of 
<taxonomicName box="[503,781,1152,1176]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[503,781,1152,1176]" italics="true" pageId="12" pageNumber="488">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
. From left to right FCFUK 320, 31 mm SL, FCFUK 314, 38 mm SL, and FCFUK 301, 58 mm SL. Numbers indicate the number posterior serrae, pink is the ossified part.
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="488" type="distribution">
<paragraph blockId="12.[151,1437,1258,2010]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[199,353,1258,1284]" pageId="12" pageNumber="488">Distribution.</emphasis>
Upper reaches of the Lesser Zab drainage. So far it was found at two places only. Following the construction of the Sardasht Dam and the impound of its reservoir in 2017, one place became submerged. In 
<date pageId="12" pageNumber="488" value="2017-06">June 2017</date>
, 
<taxonomicName box="[221,406,1330,1357]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[221,406,1330,1357]" italics="true" pageId="12" pageNumber="488">G. sardashtensis</emphasis>
</taxonomicName>
could only be found near Nalas village.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="488" type="etymology">
<paragraph blockId="12.[151,1437,1258,2010]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[199,328,1367,1393]" pageId="12" pageNumber="488">Etymology</emphasis>
. Named for the Sardasht County, where the species is found. This is also in the memorial of the victims of the chemical attack in the area on 
<date box="[653,805,1403,1429]" pageId="12" pageNumber="488" value="1987-06-28">28 June 1987</date>
. An adjective.
</paragraph>
<paragraph blockId="12.[151,1437,1258,2010]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[199,298,1439,1465]" pageId="12" pageNumber="488">Habitat.</emphasis>
<taxonomicName box="[311,624,1439,1465]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[311,624,1439,1465]" italics="true" pageId="12" pageNumber="488">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
was found in habitats with fast water velocity, shallow waters, and predominantly pebbles to cobbles substrate.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="490" pageId="12" pageNumber="488" type="discussion">
<paragraph blockId="12.[151,1437,1258,2010]" pageId="12" pageNumber="488">
<emphasis bold="true" box="[199,315,1512,1538]" pageId="12" pageNumber="488">Remarks.</emphasis>
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[326,632,1512,1538]" pageId="12" pageNumber="488" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[486,546,1512,1538]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
treat 
<taxonomicName box="[704,829,1512,1538]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[704,829,1512,1538]" italics="true" pageId="12" pageNumber="488">G. alidaeii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[847,1056,1512,1538]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[847,1056,1512,1538]" italics="true" pageId="12" pageNumber="488">G. hosseinpanahii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[1074,1207,1512,1538]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[1074,1207,1512,1538]" italics="true" pageId="12" pageNumber="488">G. galaxias</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1277,1402,1512,1538]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="shapuri">
<emphasis box="[1277,1402,1512,1538]" italics="true" pageId="12" pageNumber="488">G. shapuri</emphasis>
</taxonomicName>
as synonyms of 
<taxonomicName box="[309,420,1548,1574]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[309,420,1548,1574]" italics="true" pageId="12" pageNumber="488">G. silviae</emphasis>
</taxonomicName>
, and propose these species to be indistinguishable by external characters. Interestingly, 
<bibRefCitation author="Eagderi, S. &amp; Esmaeli, H. R. &amp; Sayyadzadeh, G. &amp; Nasri, M." box="[151,393,1584,1610]" pageId="12" pageNumber="488" pagination="500 - 522" refId="ref9439" refString="Eagderi, S., Mouludi-saleh, A., Esmaeli, H. R., Sayyadzadeh, G. &amp; Nasri, M. (2022) Freshwater lamprey and fishes of Iran; a revised and updated annotated checklist- 2022. Turkish Journal of Zoology, 46, 500 - 522. https: // doi. org / 10.55730 / 1300 - 0179.3104" type="journal article" year="2022">
Eagderi 
<emphasis box="[248,306,1584,1610]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
, published after 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[586,888,1584,1610]" pageId="12" pageNumber="488" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[743,801,1584,1610]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
, partly with an overlapping team of co-authors, treat these species as valid. This is not the place to discuss this opinion and the morphological characters in detail. All 
<taxonomicName box="[199,442,1657,1683]" pageId="12" pageNumber="488">
<emphasis box="[199,348,1657,1683]" italics="true" pageId="12" pageNumber="488">Glyptothorax</emphasis>
species
</taxonomicName>
of the Persian Gulf basin are superficially similar; even species found in sympatry largely overlap in their morphometric characters as already demonstrated by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[1017,1255,1693,1719]" pageId="12" pageNumber="488" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[1113,1170,1693,1719]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2021)
</bibRefCitation>
. The absence of morphometric characters, as found by 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[585,882,1729,1755]" pageId="12" pageNumber="488" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[740,797,1729,1755]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
, can therefore not prove that all these species are conspecific.
</paragraph>
<paragraph blockId="12.[151,1437,1258,2010]" lastBlockId="13.[151,1437,151,1941]" lastPageId="13" lastPageNumber="489" pageId="12" pageNumber="488">
To our surprise, 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[378,666,1802,1828]" pageId="12" pageNumber="488" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[531,586,1802,1828]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
found the molecular lineages identified as 
<taxonomicName box="[1139,1266,1802,1828]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[1139,1266,1802,1828]" italics="true" pageId="12" pageNumber="488">G. galaxias</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1318,1437,1802,1828]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[1318,1437,1802,1828]" italics="true" pageId="12" pageNumber="488">G. alidaeii</emphasis>
</taxonomicName>
in sympatry, clearly counteracting their conclusion, that these two species are conspecific. Naturally, we are aware, that two mitochondrial lineages might be found in one species following introgressive hybridisation. This needs to be tested in the future. Until then, we see no other way than to treat 
<taxonomicName box="[957,1088,1911,1937]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="488" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[957,1088,1911,1937]" italics="true" pageId="12" pageNumber="488">G. galaxias</emphasis>
</taxonomicName>
as a valid species as sympatry strongly suggest also reproductive isolation in sympatry, one of the strongest indications for the fact, that two species have to be recognised. 
<bibRefCitation author="Mousavi-Sabet, H. &amp; Eagderi, S. &amp; Vatandoust, S. &amp; Freyhof, J." box="[504,825,1983,2009]" pageId="12" pageNumber="488" pagination="451 - 484" refId="ref10209" refString="Mousavi-Sabet, H., Eagderi, S., Vatandoust, S. &amp; Freyhof, J. (2021) Five new species of the sisorid catfish genus Glyptothorax from Iran (Teleostei: Sisoridae). Zootaxa, 5067 (4), 451 - 484. https: // doi. org / 10.11646 / zootaxa. 5067.4.1" type="journal article" year="2021">
Mousavi-Sabet 
<emphasis box="[682,740,1983,2009]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2021)
</bibRefCitation>
, 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[838,1137,1983,2009]" pageId="12" pageNumber="488" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[994,1052,1983,2009]" italics="true" pageId="12" pageNumber="488">et al.</emphasis>
(2022)
</bibRefCitation>
, and 
<tableCitation box="[1198,1283,1983,2009]" captionStart="TABLE 2" captionStartId="3.[151,239,1422,1446]" captionTargetPageId="3" captionText="TABLE 2. Pairwise genetic distances (percentage, mean K2P) between Glyptothorax species from the Persian Gulf basin for COI sequences used in this study." pageId="12" pageNumber="488">
Table 
<quantity box="[1269,1283,1983,2009]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="488" unit="in" value="2.0">2</quantity>
</tableCitation>
in this study, given the K2P distances between the different species. In all these tables, the K2P distance between 
<taxonomicName box="[1261,1390,151,177]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[1261,1390,151,177]" italics="true" pageId="13" pageNumber="489">G. galaxias</emphasis>
</taxonomicName>
and 
<taxonomicName box="[151,272,187,213]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[151,272,187,213]" italics="true" pageId="13" pageNumber="489">G. alidaeii</emphasis>
</taxonomicName>
is clearly below 2%, a situation that needs some additional research as it might shed some light also in the validity of the allopatric species.
</paragraph>
<paragraph blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="489">
Other than morphometric characters have not been analysed in detail by 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[1014,1307,259,285]" pageId="13" pageNumber="489" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[1169,1225,259,285]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2022)
</bibRefCitation>
and should be the objective of future studies, before 
<taxonomicName box="[614,735,295,321]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[614,735,295,321]" italics="true" pageId="13" pageNumber="489">G. alidaeii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[749,954,295,321]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[749,954,295,321]" italics="true" pageId="13" pageNumber="489">G. hosseinpanahii</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1016,1137,295,321]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="shapuri">
<emphasis box="[1016,1137,295,321]" italics="true" pageId="13" pageNumber="489">G. shapuri</emphasis>
</taxonomicName>
are accepted as synonyms of 
<taxonomicName box="[182,291,331,357]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[182,291,331,357]" italics="true" pageId="13" pageNumber="489">G. silviae</emphasis>
</taxonomicName>
. Morphometric characters seem to be rather conservative in 
<taxonomicName box="[979,1128,331,357]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="genus">Glyptothorax</taxonomicName>
from Western Asia and we also found no differences in morphometrics between 
<taxonomicName box="[742,926,367,393]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[742,926,367,393]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
and sympatric 
<taxonomicName box="[1097,1280,367,393]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1097,1280,367,393]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
(
<tableCitation box="[1294,1380,367,393]" captionStart="TABLE 3" captionStartId="9.[151,239,332,356]" captionTargetPageId="9" captionText="TABLE 3. Morphometric characteristics of G. sardashtensis (holotype FCFUK 300, paratypes FCFUK 302–305, n=4; FCFUK 309–313, n=5; FSJF 4112, n=2; non-types FCFUK 301, n=1; FCFUK 306, n=1; FCFUK 314, n=1) and G. kurdistanicus (FCFUK 330–338, n=9). Holotype, paratypes and non-types measurements are included in G. sardashtensis data." pageId="13" pageNumber="489">Table 3</tableCitation>
). We agree with 
<bibRefCitation author="Sayyadzadeh, G. &amp; Zarei, F. &amp; Esmaeili, H. R." box="[275,566,403,429]" pageId="13" pageNumber="489" pagination="884" refId="ref10450" refString="Sayyadzadeh, G., Zarei, F. &amp; Esmaeili, H. R. (2022) Glyptothorax (Teleostei: Sisoridae) from the Middle East: An integrated molecular and morphological insight into its taxonomic diversity. Diversity, 14, 884. https: // doi. org / 10.3390 / d 14100884" type="journal article" year="2022">
Sayyadzadeh 
<emphasis box="[429,485,403,429]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2022)
</bibRefCitation>
that 
<taxonomicName box="[621,741,403,429]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[621,741,403,429]" italics="true" pageId="13" pageNumber="489">G. alidaeii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[754,958,403,429]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[754,958,403,429]" italics="true" pageId="13" pageNumber="489">G. hosseinpanahii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[971,1091,403,429]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="shapuri">
<emphasis box="[971,1091,403,429]" italics="true" pageId="13" pageNumber="489">G. shapuri</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1151,1258,403,429]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[1151,1258,403,429]" italics="true" pageId="13" pageNumber="489">G. silviae</emphasis>
</taxonomicName>
form four welldifferentiated, but closely related molecular groups, which should, in the absence of morphological differences, be all treated as conspecific. This case is similar to the 
<taxonomicName baseAuthorityName="Derjavin" baseAuthorityYear="1934" box="[756,1076,475,501]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="bergianus">
<emphasis box="[756,1076,475,501]" italics="true" pageId="13" pageNumber="489">Oxynoemacheilus bergianus</emphasis>
</taxonomicName>
(
<taxonomicName box="[1092,1264,475,501]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="489" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
) species group discussed by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Geiger, M. F." box="[297,529,511,537]" pageId="13" pageNumber="489" pagination="151 - 194" refId="ref9647" refString="Freyhof, J., Kaya, C., &amp; Geiger, M. F. (2022) A practical approach to revise the Oxynoemacheilus bergianus species group (Teleostei: Nemacheilidae). Zootaxa, 5128 (2), 151 - 194. https: // doi. org / 10.11646 / zootaxa. 5128.2.1" type="journal article" year="2022">
Freyhof 
<emphasis box="[392,446,511,537]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2022)
</bibRefCitation>
, who treat several molecular clusters as conspecific and discuss such cases and the rationals behind this conclusion, in detail. In contrast, 
<bibRefCitation author="Esmaeili, H. &amp; Teimori, A. &amp; Gholami, Z. &amp; Reichenbacher, B." box="[762,1007,547,573]" pageId="13" pageNumber="489" pagination="246 - 268" refId="ref9506" refString="Esmaeili, H. R, Teimori, A., Gholami, Z. &amp; Reichenbacher, B. (2014) Two new species of the tooth-carp Aphanius (Teleostei: Cyprinodontidae) and the evolutionary history of the Iranian inland and inland-related Aphanius species. Zootaxa, 3786 (3), 246 - 268. https: // doi. org / 10.11646 / zootaxa. 3786.3.2" type="journal article" year="2014">
Esmaeili 
<emphasis box="[868,925,547,573]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2014)
</bibRefCitation>
distinguish several molecular clusters of the 
<taxonomicName baseAuthorityName="Heckel" baseAuthorityYear="1847" box="[223,440,583,609]" family="Aphaniidae" genus="Esmaeilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sophiae">
<emphasis box="[223,440,583,609]" italics="true" pageId="13" pageNumber="489">Esmaeilius sophiae</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="Hoedeman" authorityYear="1949" box="[454,587,583,609]" class="Actinopterygii" family="Aphaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="489" phylum="Chordata" rank="family">Aphaniidae</taxonomicName>
) group as different species. Esmaeili 
<emphasis box="[1007,1063,583,609]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2020) describe a new species of 
<taxonomicName box="[151,247,619,645]" class="Actinopterygii" family="Cyprinidae" genus="Rhodeus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="489" phylum="Chordata" rank="genus">
<emphasis box="[151,247,619,645]" italics="true" pageId="13" pageNumber="489">Rhodeus</emphasis>
</taxonomicName>
(Acheilgnathidae), and 
<bibRefCitation author="Khaefi, R. &amp; Esmaeili, H. R. &amp; Geiger, M. F. &amp; Eagderi., S." box="[520,742,619,645]" pageId="13" pageNumber="489" pagination="90 - 115" refId="ref10058" refString="Khaefi, R., Esmaeili, H. R., Geiger, M. F. &amp; Eagderi., S. (2017) Taxonomic review of the cryptic Barbus lacerta species group with description of a new species (Teleostei: Cyprinidae). FishTaxa, 2 (2), 90 - 115." type="journal article" year="2017">
Khaefi 
<emphasis box="[603,660,619,645]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2017)
</bibRefCitation>
distinguish different molecular clusters of the 
<taxonomicName authorityName="Heckel" authorityYear="1843" box="[1268,1436,619,645]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="lacerta">
<emphasis box="[1268,1436,619,645]" italics="true" pageId="13" pageNumber="489">Barbus lacerta</emphasis>
</taxonomicName>
(
<taxonomicName box="[159,287,655,681]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="489" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
) group as species, without morphological differences demonstrated.
</paragraph>
<paragraph blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="489">
The diagnostic value of osteological characters has not been assessed in 
<taxonomicName box="[1008,1245,691,717]" pageId="13" pageNumber="489">
<emphasis box="[1008,1157,691,717]" italics="true" pageId="13" pageNumber="489">Glyptothorax</emphasis>
species
</taxonomicName>
from the Persian Gulf basin. 
<bibRefCitation author="Gauba, R. K." box="[282,439,727,753]" pageId="13" pageNumber="489" pagination="802 - 810" refId="ref9706" refString="Gauba, R. K. (1966) Studies on the Osteology of Indian Sisorid Catfishes II. The Skull of Glyptothorax cavia. Copeia, 4, 802 - 810." type="journal article" year="1966">Gauba (1966)</bibRefCitation>
, 
<bibRefCitation author="Vishwanath, W. &amp; Darshan, A. &amp; Anganthoibi, A." box="[450,729,727,753]" pageId="13" pageNumber="489" pagination="1245 - 1250" refId="ref10508" refString="Vishwanath, W., Darshan, A. &amp; Anganthoibi, A. (2010) Osteology of some catfishes of the genus Glyptothorax (Teleostei: Siluriformes) of northeastern India. Journal of Threatened Taxa, 11, 1245 - 1250. https: // doi. org / 10.11609 / JoTT. o 1874.1245 - 50" type="journal article" year="2010">
Vishwanath 
<emphasis box="[589,645,727,753]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2010)
</bibRefCitation>
, and 
<bibRefCitation author="Jiang, W. &amp; Ng, H. H. &amp; Yang, J. &amp; Chen, X." box="[787,928,727,753]" pageId="13" pageNumber="489" pagination="363 - 389" refId="ref9979" refString="Jiang, W., Ng, H. H., Yang, J. &amp; Chen, X. (2012) A taxonomic review of the catfish identified as Glyptothorax zanaensis (Teleostei: Siluriformes: Sisoridae), with the descriptions of two new species. Zoological Journal of the Linnean Society, 165, 363 - 389. https: // doi. org / 10.1111 / j. 1096 - 3642.2011.00811. x" type="journal article" year="2012">Jiang (2012)</bibRefCitation>
studied such characters in East Asian species of the genus. Our dataset is for sure too limited to use osteological characters to distinguish 
<taxonomicName box="[1201,1387,763,789]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1201,1387,763,789]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
and more materials of more species should be studied. We compared few individuals of 
<taxonomicName box="[1106,1292,799,825]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1106,1292,799,825]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
with few 
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
and found the premaxillary tooth plate in 
<taxonomicName box="[768,952,835,861]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[768,952,835,861]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
having a pair of rod-shaped distal elements with straight anterior margins (vs. wedge-shaped, with convex anterior margins in 
<taxonomicName box="[1088,1274,871,897]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1088,1274,871,897]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
). The anterior side of the triangular-shaped nuchal plate is less stretched reaching horizontally to the level of the seventh vertebra (vs. the sixth vertebra in 
<taxonomicName box="[422,604,943,969]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[422,604,943,969]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
). Its anterior tip is not turned up (vs. turned up in 
<taxonomicName box="[1146,1329,943,969]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1146,1329,943,969]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
), causing the three blotches in the front of the dorsal fin to look like a crescent shaped arch (vs. triangular in 
<taxonomicName box="[1237,1420,979,1005]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1237,1420,979,1005]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
). The posterior margin of the vomer’s triangular plate is concave, (vs. straight in 
<taxonomicName box="[1040,1225,1015,1041]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1040,1225,1015,1041]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="13.[151,1437,151,1941]" pageId="13" pageNumber="489">
As 
<taxonomicName box="[239,427,1051,1077]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[239,427,1051,1077]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
occurs in sympatry with 
<taxonomicName box="[722,908,1051,1077]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[722,908,1051,1077]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
, it is worthwhile to discuss, which of the two species is the one described by 
<bibRefCitation author="Berg, L. S." box="[507,643,1087,1113]" pageId="13" pageNumber="489" pagination="1267 - 1270" refId="ref8988" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267 - 1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
as 
<taxonomicName box="[682,867,1087,1113]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[682,867,1087,1113]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
. 
<taxonomicName box="[882,1190,1087,1113]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[882,1190,1087,1113]" italics="true" pageId="13" pageNumber="489">Glyptothorax sardashtensis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName box="[151,335,1123,1149]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[151,335,1123,1149]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
by having a plain flank, with few, minute, dark-brown spots on the head, the dorsal-, and adiposefin bases, and on the caudal-peduncle in some individuals (vs. flank with many, large, black spots and blotches in 
<taxonomicName box="[151,335,1195,1221]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[151,335,1195,1221]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
). 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[359,592,1195,1221]" pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[454,510,1195,1221]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021)
</bibRefCitation>
show the 
<typeStatus box="[706,754,1195,1221]" pageId="13" pageNumber="489">type</typeStatus>
of 
<taxonomicName box="[791,976,1195,1221]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[791,976,1195,1221]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
(ZIN 20780) and the figures drawn from the 
<typeStatus box="[194,242,1231,1257]" pageId="13" pageNumber="489">type</typeStatus>
published by 
<bibRefCitation author="Berg, L. S." box="[406,547,1231,1257]" pageId="13" pageNumber="489" pagination="1267 - 1270" refId="ref8988" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267 - 1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
. The 
<typeStatus box="[611,659,1231,1257]" pageId="13" pageNumber="489">type</typeStatus>
has indeed few, large black spots on the flank. Not as prominent as in individuals shown by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[431,668,1267,1293]" pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[528,585,1267,1293]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021)
</bibRefCitation>
and in our study (
<figureCitation box="[877,947,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1151,1175]" captionTargetBox="[199,1391,736,1127]" captionTargetId="figure-217@4.[196,1391,736,1127]" captionTargetPageId="4" captionText="FIGURE 2. Glyptothorax kurdistanicus, FCFUK 330, 108 mm SL; upper Lesser Zab, Iran." figureDoi="http://doi.org/10.5281/zenodo.7732083" httpUri="https://zenodo.org/record/7732083/files/figure.png" pageId="13" pageNumber="489">Fig. 2</figureCitation>
), but much larger than in 
<taxonomicName box="[1242,1429,1267,1293]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[1242,1429,1267,1293]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
. In none of the individuals of 
<taxonomicName box="[480,665,1303,1329]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[480,665,1303,1329]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
, such spots on the flank are found. The caudal-peduncle depth in 
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
is 1.6–2.3 times its length (vs. 
<quantity box="[654,767,1339,1365]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.318" metricValueMin="3.302" pageId="13" pageNumber="489" unit="in" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 in</quantity>
<taxonomicName box="[774,959,1339,1365]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[774,959,1339,1365]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
). 
<bibRefCitation author="Berg, L. S." box="[982,1117,1339,1365]" pageId="13" pageNumber="489" pagination="1267 - 1270" refId="ref8988" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267 - 1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
give the depth of the caudal peduncle as 81% of its length. This corresponds to a caudal-peduncle depth 1.2 times its length, making it also likely that our identification of 
<taxonomicName box="[439,626,1411,1437]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[439,626,1411,1437]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
is correct. ZIN 20780 has a short thoracic adhesive apparatus and this character state has also been found in other fishes identified as 
<taxonomicName box="[841,1024,1447,1473]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[841,1024,1447,1473]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
by 
<bibRefCitation author="Coad, B. W." box="[1063,1195,1447,1473]" pageId="13" pageNumber="489" pagination="291 - 295" refId="ref9162" refString="Coad, B. W. (1981) Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27 (4), 291 - 295." type="journal article" year="1981">Coad (1981</bibRefCitation>
, 
<bibRefCitation author="Coad, B. W." box="[1205,1263,1447,1473]" pageId="13" pageNumber="489" pagination="1 - 16" refId="ref9198" refString="Coad, B. W. (2010) Freshwater fishes of Iraq. Sofia and Moscow (Pensoft): 275 pp., Pls. 1 - 16." type="book chapter" year="2010">2010</bibRefCitation>
) and 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021)
</bibRefCitation>
. Also this character indicates that our identifications are correct. 
<bibRefCitation author="Berg, L. S." box="[985,1120,1483,1509]" pageId="13" pageNumber="489" pagination="1267 - 1270" refId="ref8988" refString="Berg, L. S. (1931) Description of a new siluroid fish, Glyptosternum kurdistanicum, from the basin of the Tigris River. Bulletin de l'Academie des Sciences de ' Union des Republiques Sovietiques Socialistes, VII Serie, Classe des Sciences Mathematiques et Naturelles, 1931, 1267 - 1270." type="journal article" year="1931">Berg (1931)</bibRefCitation>
figure the medial pit of ZIN 20780 with a roundish anterior tip but its shape is difficult to see in the partly destroyed, thoracic adhesive apparatus of ZIN 20780 shown by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[423,657,1555,1581]" pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[518,573,1555,1581]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021)
</bibRefCitation>
. ZIN 20780 is 
<quantity box="[820,912,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="13" pageNumber="489" unit="mm" value="113.0">113 mm</quantity>
SL and other individuals of 
<taxonomicName box="[1225,1408,1555,1581]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[1225,1408,1555,1581]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
in the same size category shown in Fig. 22 by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[637,871,1591,1617]" pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[732,788,1591,1617]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021)
</bibRefCitation>
, also have a roundish anterior tip of the medial pit. This character state was not found in 
<taxonomicName box="[570,754,1627,1653]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[570,754,1627,1653]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
collected from 
<collectingCountry box="[930,975,1627,1653]" name="Iran" pageId="13" pageNumber="489">Iran</collectingCountry>
(
<figureCitation box="[990,1072,1627,1653]" captionStart="FIGURE 12" captionStartId="14.[151,250,1043,1067]" captionTargetBox="[155,1432,181,1019]" captionTargetId="figure-20@14.[155,1432,181,1019]" captionTargetPageId="14" captionText="FIGURE 12. Glyptothorax kurdistanicus, upper row, from left: FCFUK 339, 27 mm SL; FCFUK 337, 43 mm SL; FCFUK 332, 67 mm SL; G. sardashtensis, lower row, from left: FCFUK 322, 21 mm SL; FSJF 4112, 52 mm SL; FCFUK 305, 55 mm SL; Lesser Zab, Iran." figureDoi="http://doi.org/10.5281/zenodo.7732107" httpUri="https://zenodo.org/record/7732107/files/figure.png" pageId="13" pageNumber="489">Fig. 12</figureCitation>
), except for one large individual (
<quantity box="[159,255,1663,1689]" metricMagnitude="-1" metricUnit="m" metricValue="1.08" pageId="13" pageNumber="489" unit="mm" value="108.0">108 mm</quantity>
SL, 
<figureCitation box="[312,381,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,2004,2028]" captionTargetBox="[199,1389,1200,1981]" captionTargetId="figure-234@4.[199,1389,1200,1981]" captionTargetPageId="4" captionText="FIGURE 3. Glyptothorax kurdistanicus, from left: FCFUK 330, 108 mm SL; FCFUK 333, 52 mm; SL; Lesser Zab, Iran." figureDoi="http://doi.org/10.5281/zenodo.7732085" httpUri="https://zenodo.org/record/7732085/files/figure.png" pageId="13" pageNumber="489">Fig. 3</figureCitation>
). As juvenile individuals of 
<taxonomicName box="[706,891,1663,1689]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[706,891,1663,1689]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
shown by 
<bibRefCitation author="Freyhof, J. &amp; Kaya, C. &amp; Abdullah, Y. S. &amp; Geiger, M. F." box="[1018,1251,1663,1689]" pageId="13" pageNumber="489" pagination="453 - 491" refId="ref9577" refString="Freyhof, J., Kaya, C., Abdullah, Y. S. &amp; Geiger, M. F. (2021) The Glyptothorax catfishes of the Euphrates and Tigris with the description of a new species (Teleostei: Sisoridae). Zootaxa, 4969 (3), 453 - 491. https: // doi. org / 10.11646 / zootaxa. 4969.3.2" type="journal article" year="2021">
Freyhof 
<emphasis box="[1115,1173,1663,1689]" italics="true" pageId="13" pageNumber="489">et al.</emphasis>
(2021
</bibRefCitation>
: Fig. 21) have a pointed anterior tip of the medial tip, this character might be variable depending on size, pointed in small, roundish in large individuals. In 
<emphasis box="[408,599,1735,1761]" italics="true" pageId="13" pageNumber="489">
<taxonomicName box="[408,595,1735,1761]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">G. sardashtensis</taxonomicName>
,
</emphasis>
however, we found no size-dependent variability of this character. As both, 
<taxonomicName box="[151,337,1771,1797]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[151,337,1771,1797]" italics="true" pageId="13" pageNumber="489">G. sardashtensis</emphasis>
</taxonomicName>
and large 
<taxonomicName box="[457,642,1771,1797]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[457,642,1771,1797]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
have a median pit with a roundish anterior tip, this character does not allow to distinguish larger individuals (as in ZIN 20780) of both species. Also, the skin on the nuchal plate in ZIN 20780 is destroyed, and the shape of the yellowish blotch on that plate cannot be seen. As a conclusion, we remain confident to have identified the right fishes as 
<taxonomicName box="[672,856,1879,1905]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="489" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[672,856,1879,1905]" italics="true" pageId="13" pageNumber="489">G. kurdistanicus</emphasis>
</taxonomicName>
based on the very deep caudal peduncle, the colour pattern, and the shape of the thoracic adhesive apparatus.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7732107" ID-Zenodo-Dep="7732107" httpUri="https://zenodo.org/record/7732107/files/figure.png" pageId="14" pageNumber="490" startId="14.[151,250,1043,1067]" targetBox="[155,1432,181,1019]" targetPageId="14">
<paragraph blockId="14.[151,1436,1043,1139]" pageId="14" pageNumber="490">
<emphasis bold="true" box="[151,287,1043,1067]" pageId="14" pageNumber="490">FIGURE 12.</emphasis>
<taxonomicName box="[292,569,1043,1067]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="kurdistanicus">
<emphasis box="[292,569,1043,1067]" italics="true" pageId="14" pageNumber="490">Glyptothorax kurdistanicus</emphasis>
</taxonomicName>
, upper row, from left: FCFUK 339, 27 mm SL; FCFUK 337, 43 mm SL; FCFUK 332, 67 mm SL; 
<taxonomicName box="[273,442,1079,1103]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="sardashtensis">
<emphasis box="[273,442,1079,1103]" italics="true" pageId="14" pageNumber="490">G. sardashtensis</emphasis>
</taxonomicName>
, lower row, from left: FCFUK 322, 21 mm SL; FSJF 4112, 52 mm SL; FCFUK 305, 55 mm SL; Lesser Zab, Iran.
</paragraph>
</caption>
<paragraph blockId="14.[151,1436,1185,1248]" pageId="14" pageNumber="490">
<emphasis bold="true" box="[151,421,1185,1211]" pageId="14" pageNumber="490">Comparative material.</emphasis>
Materials listed by 
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Freyhof 
<emphasis box="[728,782,1185,1211]" italics="true" pageId="14" pageNumber="490">et al.</emphasis>
(2021)
</bibRefCitation>
was also used for this study. Additional comparative materials are:
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" box="[151,1413,1293,1320]" pageId="14" pageNumber="490">
<taxonomicName authority=": FSJF" authorityName="FSJF" box="[151,466,1293,1320]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="alidaeii">
<emphasis box="[151,394,1293,1319]" italics="true" pageId="14" pageNumber="490">Glyptothorax alidaeii</emphasis>
: FSJF
</taxonomicName>
4111, 2, 
<quantity box="[568,689,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" metricValueMax="7.8" metricValueMin="3.5" pageId="14" pageNumber="490" unit="mm" value="56.5" valueMax="78.0" valueMin="35.0">35–78 mm</quantity>
SL; 
<materialsCitation box="[743,1409,1293,1320]" country="Iran" latitude="33.688" location="Zirkhaki" longLatPrecision="6" longitude="47.0639" municipality="Seimare" pageId="14" pageNumber="490" specimenCount="1" stateProvince="Lorestan">
<collectingCountry box="[743,792,1294,1320]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[803,969,1294,1320]" country="Iran" name="Lorestan" pageId="14" pageNumber="490">Lorestan prov.</collectingRegion>
: 
<collectingMunicipality box="[979,1071,1293,1319]" pageId="14" pageNumber="490">Seimare</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFBE5FA97DE866D16" box="[1105,1204,1293,1319]" country="Iran" latitude="33.688" longLatPrecision="6" longitude="47.0639" municipality="Seimare" name="Zirkhaki" pageId="14" pageNumber="490" stateProvince="Lorestan">Zirkhaki</location>
, 
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<geoCoordinate box="[1314,1409,1293,1319]" degrees="47.0639" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="47.0639">47.0639</geoCoordinate>
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.
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" pageId="14" pageNumber="490">
<taxonomicName authority=": FCFUK" authorityName="FCFUK" box="[151,506,1329,1356]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="galaxias">
<emphasis box="[151,402,1329,1355]" italics="true" pageId="14" pageNumber="490">Glyptothorax galaxias</emphasis>
: FCFUK
</taxonomicName>
344, 1, 
<quantity box="[595,673,1329,1356]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="14" pageNumber="490" unit="mm" value="59.0">59 mm</quantity>
SL; 
<materialsCitation box="[726,1025,1329,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490" specimenCode="FCFUK 345, 1" specimenCount="1">
<specimenCode box="[726,867,1329,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490">FCFUK 345</specimenCode>
, 
<specimenCode box="[878,892,1330,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[905,983,1330,1356]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="490" unit="mm" value="80.0">80 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation box="[1036,1335,1329,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490" specimenCode="FCFUK 350, 1" specimenCount="1">
<specimenCode box="[1036,1178,1329,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490">FCFUK 350</specimenCode>
, 
<specimenCode box="[1188,1202,1330,1356]" collectionCode="FCFUK" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[1215,1293,1329,1356]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="490" unit="mm" value="95.0">95 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation collectionCode="FCFUK" country="Iran" latitude="33.7856" location="Kaka-Reza" longLatPrecision="6" longitude="48.2064" municipality="Kashkan" pageId="14" pageNumber="490" specimenCode="FCFUK 351, 10" specimenCount="1" stateProvince="Lorestan">
<specimenCode collectionCode="FCFUK" pageId="14" pageNumber="490">FCFUK 351</specimenCode>
, 
<specimenCode box="[255,285,1366,1392]" collectionCode="FCFUK" pageId="14" pageNumber="490">10</specimenCode>
, 
<quantity box="[298,434,1365,1392]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" metricValueMax="10.4" metricValueMin="6.2" pageId="14" pageNumber="490" unit="mm" value="83.0" valueMax="104.0" valueMin="62.0">62–104 mm</quantity>
SL; 
<collectingCountry box="[491,540,1366,1392]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[552,719,1366,1392]" country="Iran" name="Lorestan" pageId="14" pageNumber="490">Lorestan prov.</collectingRegion>
: 
<collectingMunicipality box="[730,828,1365,1391]" pageId="14" pageNumber="490">Kashkan</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFCD5FACFD9D26D5E" box="[865,992,1365,1391]" country="Iran" latitude="33.7856" longLatPrecision="6" longitude="48.2064" municipality="Kashkan" name="Kaka-Reza" pageId="14" pageNumber="490" stateProvince="Lorestan">Kaka-Reza</location>
, 
<geoCoordinate box="[1004,1095,1365,1392]" degrees="33.7856" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="33.7856">33.7856</geoCoordinate>
<geoCoordinate box="[1103,1196,1365,1391]" degrees="48.2064" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="48.2064">48.2064</geoCoordinate>
</materialsCitation>
.— 
<materialsCitation collectionCode="FSJF" country="Iran" county="Chaharmahal-va-Bakhtiari prov." latitude="32.0287" location="Beheshtabad" longLatPrecision="6" longitude="50.6265" municipality="Beheshtabad" pageId="14" pageNumber="490" specimenCode="FSJF 4112, 2" specimenCount="1">
<specimenCode box="[1228,1353,1365,1392]" collectionCode="FSJF" pageId="14" pageNumber="490">FSJF 4112</specimenCode>
, 
<specimenCode box="[1365,1381,1366,1392]" collectionCode="FSJF" pageId="14" pageNumber="490">2</specimenCode>
, 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.95" metricValueMax="5.2" metricValueMin="4.7" pageId="14" pageNumber="490" unit="mm" value="49.5" valueMax="52.0" valueMin="47.0">47– 52mm</quantity>
SL; 
<collectingCountry box="[331,380,1402,1428]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingCounty box="[394,765,1401,1428]" pageId="14" pageNumber="490">Chaharmahal-va-Bakhtiari prov.</collectingCounty>
: stream 
<collectingMunicipality box="[864,1007,1401,1427]" pageId="14" pageNumber="490">Beheshtabad</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFBACFAE3DE986DA2" box="[1048,1194,1401,1427]" country="Iran" county="Chaharmahal-va-Bakhtiari prov." latitude="32.0287" longLatPrecision="6" longitude="50.6265" municipality="Beheshtabad" name="Beheshtabad" pageId="14" pageNumber="490">Beheshtabad</location>
, 
<geoCoordinate box="[1209,1300,1402,1428]" degrees="32.0287" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="32.0287">32.0287</geoCoordinate>
<geoCoordinate box="[1310,1403,1401,1428]" degrees="50.6265" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="50.6265">50.6265</geoCoordinate>
</materialsCitation>
.— 
<materialsCitation box="[198,983,1437,1464]" collectionCode="FSJF" country="Iran" latitude="31.2587" location="Sangan" longLatPrecision="6" longitude="51.2858" municipality="Sangan" pageId="14" pageNumber="490" specimenCode="FSJF 2214, 1" specimenCount="1">
<specimenCode box="[198,324,1438,1464]" collectionCode="FSJF" pageId="14" pageNumber="490">FSJF 2214</specimenCode>
, 
<specimenCode box="[337,351,1438,1464]" collectionCode="FSJF" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[368,449,1437,1464]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="490" unit="mm" value="57.0">57 mm</quantity>
SL; 
<collectingCountry box="[508,557,1438,1464]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingMunicipality box="[570,652,1438,1464]" pageId="14" pageNumber="490">Sangan</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFD00FA04D93B6D89" box="[692,777,1438,1464]" country="Iran" latitude="31.2587" longLatPrecision="6" longitude="51.2858" municipality="Sangan" name="Sangan" pageId="14" pageNumber="490">Sangan</location>
, 
<geoCoordinate box="[791,882,1437,1464]" degrees="31.2587" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="31.2587">31.2587</geoCoordinate>
<geoCoordinate box="[891,983,1437,1464]" degrees="51.2858" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="51.2858">51.2858</geoCoordinate>
</materialsCitation>
.— 
<materialsCitation collectionCode="FSJF" country="Iran" latitude="30.8521" location="Doruhan" longLatPrecision="6" longitude="51.342" municipality="Beshar" pageId="14" pageNumber="490" specimenCode="FSJF 3243, 24" specimenCount="1">
<specimenCode box="[1016,1143,1437,1464]" collectionCode="FSJF" pageId="14" pageNumber="490">FSJF 3243</specimenCode>
, 
<specimenCode box="[1157,1187,1438,1464]" collectionCode="FSJF" pageId="14" pageNumber="490">24</specimenCode>
, 
<quantity box="[1201,1325,1438,1464]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.8" metricValueMin="3.1" pageId="14" pageNumber="490" unit="mm" value="39.5" valueMax="48.0" valueMin="31.0">31–48 mm</quantity>
SL; 
<collectingCountry box="[1383,1432,1438,1464]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingMunicipality box="[198,276,1473,1499]" pageId="14" pageNumber="490">Beshar</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFE82FA5BDBA96DEA" box="[310,411,1473,1499]" country="Iran" latitude="30.8521" longLatPrecision="6" longitude="51.342" municipality="Beshar" name="Doruhan" pageId="14" pageNumber="490">Doruhan</location>
, 
<geoCoordinate box="[422,513,1473,1500]" degrees="30.8521" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="30.8521">30.8521</geoCoordinate>
<geoCoordinate box="[520,615,1473,1500]" degrees="51.3420" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="51.342">51.3420</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" pageId="14" pageNumber="490">
<taxonomicName authority=": FSJF" authorityName="FSJF" box="[151,548,1509,1536]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="hosseinpanahii">
<emphasis box="[151,477,1509,1535]" italics="true" pageId="14" pageNumber="490">Glyptothorax hosseinpanahii</emphasis>
: FSJF
</taxonomicName>
4113, 1, 
<quantity box="[649,727,1509,1536]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="14" pageNumber="490" unit="mm" value="36.0">36 mm</quantity>
SL; 
<materialsCitation country="Iran" latitude="30.5295" location="Kheirabad" longLatPrecision="6" longitude="50.4156" municipality="Zohreh River" pageId="14" pageNumber="490" specimenCount="1" stateProvince="Khuzestan">
<collectingCountry box="[780,829,1510,1536]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[839,1024,1509,1536]" country="Iran" name="Khuzestan" pageId="14" pageNumber="490">Khuzestan prov.</collectingRegion>
: 
<collectingMunicipality box="[1034,1183,1509,1535]" pageId="14" pageNumber="490">Zohreh River</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFB0BFA7FDF056DCE" box="[1215,1335,1509,1535]" country="Iran" latitude="30.5295" longLatPrecision="6" longitude="50.4156" municipality="Zohreh River" name="Kheirabad" pageId="14" pageNumber="490" stateProvince="Khuzestan">Kheirabad</location>
, 
<geoCoordinate box="[1345,1436,1509,1536]" degrees="30.5295" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="30.5295">30.5295</geoCoordinate>
<geoCoordinate box="[198,293,1545,1572]" degrees="50.4156" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="50.4156">50.4156</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" pageId="14" pageNumber="490">
<taxonomicName authority=": FCFUK" authorityName="FCFUK" box="[151,497,1581,1608]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="pallens">
<emphasis box="[151,390,1581,1607]" italics="true" pageId="14" pageNumber="490">Glyptothorax pallens</emphasis>
: 
<materialsCitation box="[407,497,1582,1608]" collectionCode="FCFUK" country="Iran" latitude="34.6416" location="Zamkan-e Olya" longLatPrecision="6" longitude="46.2841" municipality="Zemkan" pageId="14" pageNumber="490" specimenCode="FCFUK 372, 1, FCFUK 373" specimenCount="1" stateProvince="Kermanshah">FCFUK</materialsCitation>
</taxonomicName>
371, 1, 
<quantity box="[594,675,1582,1608]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="490" unit="mm" value="48.0">48 mm</quantity>
SL; 
<specimenCode box="[732,877,1582,1608]" collectionCode="FCFUK" pageId="14" pageNumber="490">FCFUK 372</specimenCode>
, 
<specimenCode box="[889,903,1582,1608]" collectionCode="FCFUK" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[919,1000,1581,1608]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="14" pageNumber="490" unit="mm" value="59.0">59 mm</quantity>
SL; 
<specimenCode box="[1058,1202,1582,1608]" collectionCode="FCFUK" pageId="14" pageNumber="490">FCFUK 373</specimenCode>
, 
<specimenCode box="[1215,1229,1582,1608]" collectionCode="FCFUK" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[1245,1326,1581,1608]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="490" unit="mm" value="62.0">62 mm</quantity>
SL; 
<collectingCountry box="[1383,1432,1582,1608]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[198,406,1617,1644]" country="Iran" name="Kermanshah" pageId="14" pageNumber="490">Kermanshah prov.</collectingRegion>
: stream 
<collectingMunicipality box="[496,588,1617,1643]" pageId="14" pageNumber="490">Zemkan</collectingMunicipality>
north of 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFD1BF9CBD96D6E5A" box="[687,863,1617,1643]" country="Iran" latitude="34.6416" longLatPrecision="6" longitude="46.2841" municipality="Zemkan" name="Zamkan-e Olya" pageId="14" pageNumber="490" stateProvince="Kermanshah">Zamkan-e Olya</location>
, 
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<geoCoordinate box="[970,1062,1617,1643]" degrees="46.2841" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="46.2841">46.2841</geoCoordinate>
.— 
<materialsCitation collectionCode="FSJF" country="Iran" latitude="34.6452" location="Zamkan-e Olya" longLatPrecision="6" longitude="46.2856" municipality="Zemkan" pageId="14" pageNumber="490" specimenCode="FSJF 4114, 1" specimenCount="1" stateProvince="Kermanshah">
<specimenCode box="[1095,1218,1617,1644]" collectionCode="FSJF" pageId="14" pageNumber="490">FSJF 4114</specimenCode>
, 
<specimenCode box="[1227,1241,1618,1644]" collectionCode="FSJF" pageId="14" pageNumber="490">1</specimenCode>
, 
<quantity box="[1254,1332,1617,1644]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="490" unit="mm" value="65.0">65 mm</quantity>
SL; 
<collectingCountry box="[1383,1432,1618,1644]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[198,408,1653,1680]" country="Iran" name="Kermanshah" pageId="14" pageNumber="490">Kermanshah prov.</collectingRegion>
: stream 
<collectingMunicipality box="[499,591,1653,1679]" pageId="14" pageNumber="490">Zemkan</collectingMunicipality>
<quantity box="[598,655,1653,1680]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="490" unit="km" value="3.0">3 km</quantity>
north of 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFD42F9EFD99A6EBE" box="[758,936,1653,1679]" country="Iran" latitude="34.6452" longLatPrecision="6" longitude="46.2856" municipality="Zemkan" name="Zamkan-e Olya" pageId="14" pageNumber="490" stateProvince="Kermanshah">Zamkan-e Olya</location>
, 
<geoCoordinate box="[947,1038,1653,1680]" degrees="34.6452" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="34.6452">34.6452</geoCoordinate>
<geoCoordinate box="[1045,1140,1653,1680]" degrees="46.2856" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="46.2856">46.2856</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" box="[151,1405,1689,1716]" pageId="14" pageNumber="490">
<taxonomicName authority=": FSJF" authorityName="FSJF" box="[151,466,1689,1716]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="shapuri">
<emphasis box="[151,394,1689,1715]" italics="true" pageId="14" pageNumber="490">Glyptothorax shapuri</emphasis>
: FSJF
</taxonomicName>
4115, 1, 
<quantity box="[569,648,1689,1716]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="490" unit="mm" value="57.0">57 mm</quantity>
SL; 
<materialsCitation box="[702,1401,1689,1716]" country="Iran" latitude="29.7618" location="Eslamabad" longLatPrecision="6" longitude="51.5507" municipality="Shapur River" pageId="14" pageNumber="490" specimenCount="1" stateProvince="Fars">
<collectingCountry box="[702,751,1690,1716]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<collectingRegion box="[762,879,1690,1716]" country="Iran" name="Fars" pageId="14" pageNumber="490">Fars prov.</collectingRegion>
: 
<collectingMunicipality box="[889,1037,1689,1715]" pageId="14" pageNumber="490">Shapur River</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFB84F903DE9C6E82" box="[1072,1198,1689,1715]" country="Iran" latitude="29.7618" longLatPrecision="6" longitude="51.5507" municipality="Shapur River" name="Eslamabad" pageId="14" pageNumber="490" stateProvince="Fars">Eslamabad</location>
, 
<geoCoordinate box="[1208,1299,1689,1715]" degrees="29.7618" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="29.7618">29.7618</geoCoordinate>
<geoCoordinate box="[1306,1401,1689,1716]" degrees="51.5507" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="51.5507">51.5507</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph blockId="14.[151,1437,1293,1788]" pageId="14" pageNumber="490">
<taxonomicName authority=": FSJF" authorityName="FSJF" box="[151,460,1725,1752]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="490" phylum="Chordata" rank="species" species="silviae">
<emphasis box="[151,384,1725,1751]" italics="true" pageId="14" pageNumber="490">Glyptothorax silviae</emphasis>
: FSJF
</taxonomicName>
4116, 5, 
<quantity box="[574,698,1725,1752]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.5" metricValueMin="4.5" pageId="14" pageNumber="490" unit="mm" value="70.0" valueMax="95.0" valueMin="45.0">45–95 mm</quantity>
SL; 
<materialsCitation country="Iran" county="Garab-e-Lodab" latitude="30.9455" location="Kohgiluyeh-va-Boyer Ahmad prov." longLatPrecision="6" longitude="50.9042" municipality="Maroun River" pageId="14" pageNumber="490" specimenCount="1">
<collectingCountry box="[759,808,1726,1752]" name="Iran" pageId="14" pageNumber="490">Iran</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:03BE87A85255775AFF23F898D8766ECD:8EC86065525D775AFC82F927DEFD6EE9" box="[822,1231,1725,1752]" country="Iran" county="Garab-e-Lodab" latitude="30.9455" longLatPrecision="6" longitude="50.9042" municipality="Maroun River" name="Kohgiluyeh-va-Boyer Ahmad prov." pageId="14" pageNumber="490">Kohgiluyeh-va-Boyer Ahmad prov.</location>
: 
<collectingMunicipality box="[1244,1405,1725,1752]" pageId="14" pageNumber="490">Maroun River</collectingMunicipality>
at 
<collectingCounty box="[198,372,1761,1787]" pageId="14" pageNumber="490">Garab-e-Lodab</collectingCounty>
, 
<geoCoordinate box="[383,474,1761,1788]" degrees="30.9455" direction="north" orientation="latitude" pageId="14" pageNumber="490" precision="5" value="30.9455">30.9455</geoCoordinate>
<geoCoordinate box="[481,576,1761,1788]" degrees="50.9042" direction="east" orientation="longitude" pageId="14" pageNumber="490" precision="5" value="50.9042">50.9042</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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