159 lines
15 KiB
XML
159 lines
15 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.7501705" ID-ISSN="1211-87 8 8" ID-Zenodo-Dep="7501705" checkinTime="1672754391430" checkinUser="diego" docAuthor="Löbl, Ivan" docDate="2022" docId="03BE8788FFB1C75003236E1ADEE850B9" docLanguage="en" docName="ActaMusMoraviae.107.1-2.7-19.pdf" docOrigin="Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2)" docTitle="Pseudobironium rapax Löbl 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FF87FFF0FFB6C75802466814DF5C556E" masterDocTitle="Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="19" masterPageNumber="7" pageNumber="14" updateTime="1672761276865" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Löbl, Ivan</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Musei Moraviae, Scientiae biologicae</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>107</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1 - 2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7</mods:start>
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<mods:end>19</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.7501705</mods:identifier>
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<mods:identifier type="ISSN">1211-8788</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7501705</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7501619" ID-Zenodo-Dep="7501619" LSID="urn:lsid:plazi:treatment:03BE8788FFB1C75003236E1ADEE850B9" httpUri="http://treatment.plazi.org/id/03BE8788FFB1C75003236E1ADEE850B9" lastPageId="8" lastPageNumber="15" pageId="7" pageNumber="14">
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<subSubSection box="[357,705,1550,1575]" pageId="7" pageNumber="14" type="nomenclature">
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<paragraph blockId="7.[357,705,1550,1575]" box="[357,705,1550,1575]" pageId="7" pageNumber="14">
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<heading bold="true" box="[357,705,1550,1575]" fontSize="10" level="2" pageId="7" pageNumber="14" reason="2">
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<taxonomicName authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[357,610,1550,1574]" class="Insecta" family="Staphylinidae" genus="Pseudobironium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="14" phylum="Arthropoda" rank="species" species="rapax" status="sp. nov.">Pseudobironium rapax</taxonomicName>
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<taxonomicNameLabel box="[617,705,1551,1575]" pageId="7" pageNumber="14" rank="species">
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<emphasis bold="true" box="[617,697,1551,1575]" pageId="7" pageNumber="14">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1072,1196,1551,1575]" pageId="7" pageNumber="14" type="description">
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<paragraph blockId="7.[1072,1196,1551,1575]" box="[1072,1196,1551,1575]" pageId="7" pageNumber="14">
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(
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<figureCitation box="[1079,1146,1551,1575]" captionStart="Figs 8-13. 8-10" captionStartId="6.[332,370,1715,1734]" captionTargetBox="[355,1235,431,1586]" captionTargetId="figure-89@6.[345,1267,390,1595]" captionTargetPageId="6" captionText="Figs 8-13. 8-10 ̅ Pseudobironium rapax sp. nov.; 8, 9 ̅ Aedeagus in dorsal and lateral views, scale = 0.1 mm; 10 ̅ Internal sac in dorsal view, scale = 0.1 mm. 11-13 ̅ Scaphoxium sinuatum sp. nov.; 11 ̅ Aedeagus in dorsal view, scale = 0.1 mm; 12 ̅ Apical part of paramere, dorsal view, scale = 0.05 mm; 13 ̅ Internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7501709" httpUri="https://zenodo.org/record/7501709/files/figure.png" pageId="7" pageNumber="14">Figs 8</figureCitation>
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̅10)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="14" type="materials_examined">
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<paragraph blockId="7.[304,1251,1598,1643]" pageId="7" pageNumber="14">
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<materialsCitation collectedFrom="vegetation debris and forest floor litter accumulated along large fallen trees near river" collectingDate="1996-06-17" collectionCode="MHNG" country="Malaysia" location="Mawar Waterfall" municipality="Crocker Range" pageId="7" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
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<emphasis bold="true" box="[304,441,1598,1617]" pageId="7" pageNumber="14">Type material.</emphasis>
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<typeStatus box="[451,530,1598,1617]" pageId="7" pageNumber="14">Holotype</typeStatus>
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<specimenCount box="[541,585,1598,1617]" pageId="7" pageNumber="14" type="male">male</specimenCount>
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,
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<collectingCountry box="[598,678,1598,1617]" name="Malaysia" pageId="7" pageNumber="14" value="Malaysia">Malaysia</collectingCountry>
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,
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<collectingRegion box="[692,746,1598,1617]" country="Malaysia" name="Sabah" pageId="7" pageNumber="14">Sabah</collectingRegion>
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,
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<collectingMunicipality box="[759,895,1598,1617]" pageId="7" pageNumber="14">Crocker Range</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:03BE8788FFB1C75003236E1ADEE850B9:8EC86045FFB1C75F01CA6E2ADB7E533F" box="[908,1058,1598,1617]" country="Malaysia" municipality="Crocker Range" name="Mawar Waterfall" pageId="7" pageNumber="14" stateProvince="Sabah">Mawar Waterfall</location>
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env.
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<date box="[1113,1212,1598,1617]" pageId="7" pageNumber="14" value="1996-06-17">
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<collectingDate box="[1113,1212,1598,1617]" pageId="7" pageNumber="14" value="1996-06-17">17.VI.1996</collectingDate>
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</date>
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, 9c,
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<collectedFrom box="[304,1039,1623,1643]" pageId="7" pageNumber="14">vegetation debris and forest floor litter accumulated along large fallen trees near river</collectedFrom>
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(
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<collectionCode box="[1051,1121,1623,1642]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="14" type="Museum">MHNG</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="15" pageId="7" pageNumber="14" type="description">
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<paragraph blockId="7.[304,1251,1664,1720]" lastBlockId="8.[332,1280,393,1495]" lastPageId="8" lastPageNumber="15" pageId="7" pageNumber="14">
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<emphasis bold="true" box="[304,443,1664,1688]" pageId="7" pageNumber="14">Description.</emphasis>
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Length
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<quantity box="[537,637,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="7" pageNumber="14" unit="mm" value="1.95">1.95 mm</quantity>
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, width
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<quantity box="[722,823,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="14" unit="mm" value="1.3">1.30 mm</quantity>
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. Head and most of body dark reddishbrown, apex of abdomen lighter, yellowish; femora and tibiae roughly as body, tarsi and antennae light brown. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows:
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<date box="[953,1016,425,449]" pageId="8" pageNumber="15" value="2015-03">III 15</date>
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:
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<date box="[1027,1091,425,449]" pageId="8" pageNumber="15" value="2022-04">IV 22</date>
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:
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<date box="[1101,1157,425,449]" pageId="8" pageNumber="15" value="1926-05">V 26</date>
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:
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<date box="[1167,1232,425,449]" pageId="8" pageNumber="15" value="2021-06">VI 21</date>
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:
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<date pageId="8" pageNumber="15" value="1926-07">VII 26</date>
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:
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<date box="[376,463,457,481]" pageId="8" pageNumber="15" value="1926-08">VIII 26</date>
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:
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<date box="[477,546,457,481]" pageId="8" pageNumber="15" value="1927-09">IX 27</date>
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: X: 26:
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<date box="[643,711,457,481]" pageId="8" pageNumber="15" value="1935-11">XI 35</date>
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; antennomere XI about 3.5 times as long as wide. Pronotal punctation very fine, similar to that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 4 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation similar to pronotal. Hypomera impunctate, impressed posteriad. Mesoventrite lacking mesal ridge and lacking distinct setiferous punctures. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite lacking microsculpture; median area nearly flat, lacking impressions and tubercle, with two irregular rows of distinct admesal punctures. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, horizontal. Submesocoxal area short, about as fourth shortest interval to metacoxa. Submesocoxal line finely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Ventrite Ilacking microsculpture, with very fine and scattered punctures; following ventrites and exposed tergites with punctulate microsculpture.
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</paragraph>
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<paragraph blockId="8.[332,1280,393,1495]" pageId="8" pageNumber="15">
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<emphasis box="[332,387,905,929]" italics="true" pageId="8" pageNumber="15">Male</emphasis>
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. Aedeagus (
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<figureCitation box="[524,593,905,929]" captionStart="Figs 8-13. 8-10" captionStartId="6.[332,370,1715,1734]" captionTargetBox="[355,1235,431,1586]" captionTargetId="figure-89@6.[345,1267,390,1595]" captionTargetPageId="6" captionText="Figs 8-13. 8-10 ̅ Pseudobironium rapax sp. nov.; 8, 9 ̅ Aedeagus in dorsal and lateral views, scale = 0.1 mm; 10 ̅ Internal sac in dorsal view, scale = 0.1 mm. 11-13 ̅ Scaphoxium sinuatum sp. nov.; 11 ̅ Aedeagus in dorsal view, scale = 0.1 mm; 12 ̅ Apical part of paramere, dorsal view, scale = 0.05 mm; 13 ̅ Internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7501709" httpUri="https://zenodo.org/record/7501709/files/figure.png" pageId="8" pageNumber="15">Figs 8</figureCitation>
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̅10)
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<quantity box="[653,750,905,929]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="15" unit="mm" value="0.74">0.74 mm</quantity>
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long. Median lobe with basal bulb shorter than apical process, the latter slightly inflexed, with sinuate ventral margin (lateral view), and blunt tip; apical part of median lobe lacking distinct valves. Parameres nearly evenly wide, bent in dorsal view, straight in lateral view. Internal sac with pair of strongly sclerotized denticles and mesal sclerite, lacking flagellum and lacking apical spinose structures.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="etymology">
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<paragraph blockId="8.[332,1280,393,1495]" pageId="8" pageNumber="15">
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<emphasis bold="true" box="[332,460,1105,1129]" pageId="8" pageNumber="15">Etymology.</emphasis>
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The species epithet is a Latin noun, referring to the caninelike denticles of the internal sac.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="diagnosis">
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<paragraph blockId="8.[332,1280,393,1495]" pageId="8" pageNumber="15">
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<emphasis bold="true" box="[332,586,1176,1200]" pageId="8" pageNumber="15">Differential diagnosis.</emphasis>
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The species is as
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<taxonomicName authorityName="Löbl" authorityYear="2022" box="[796,934,1176,1200]" class="Insecta" family="Staphylinidae" genus="Pseudobironium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="irregulare">
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<emphasis box="[796,934,1176,1200]" italics="true" pageId="8" pageNumber="15">P. irregulare</emphasis>
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</taxonomicName>
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less than
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<quantity box="[1052,1116,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="15" unit="mm" value="2.0">2 mm</quantity>
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long and thus conspicuous by its small bodysize. It may be easily distinguished from
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<taxonomicName authorityName="Löbl" authorityYear="2022" box="[1110,1245,1208,1232]" class="Insecta" family="Staphylinidae" genus="Pseudobironium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="irregulare">
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<emphasis box="[1110,1245,1208,1232]" italics="true" pageId="8" pageNumber="15">P. irregulare</emphasis>
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</taxonomicName>
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by the evenly very fine elytral punctation, the narrow submesocoxal areas and ventrite I lacking microsculpture. The aedeagal characters suggest relationship with members of the
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<taxonomicName authorityName="Pic" authorityYear="1920" box="[376,523,1304,1328]" class="Insecta" family="Staphylinidae" genus="Pseudobironium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="subovatum">
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<emphasis box="[376,523,1304,1328]" italics="true" pageId="8" pageNumber="15">P. subovatum</emphasis>
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</taxonomicName>
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group (see
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<bibRefCitation author="LOBL I. & TANG L." box="[662,888,1304,1328]" journalOrPublisher="Revue suisse de zoologie" pageId="8" pageNumber="15" pagination="665" part="120" refId="ref7096" refString="LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734." title="A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae)" type="journal article" year="2013">LÖBL & TANG 2013</bibRefCitation>
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), though the shape of the admesal sclerites of the internal sac is unique. The species falls in the key species to couplet 27, to
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<taxonomicName authority="Lobl & Tang, 2013" authorityName="Lobl & Tang" authorityYear="2013" box="[361,678,1368,1392]" class="Insecta" family="Staphylinidae" genus="Pseudobironium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="schuhi">
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<emphasis box="[361,460,1368,1392]" italics="true" pageId="8" pageNumber="15">P. schuhi</emphasis>
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<bibRefCitation author="LOBL I. & TANG L." box="[468,678,1368,1392]" journalOrPublisher="Revue suisse de zoologie" pageId="8" pageNumber="15" pagination="665" part="120" refId="ref7096" refString="LOBL I. & TANG L. 2013: A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae). Revue suisse de zoologie 120: 665 ‾ 734." title="A review of the genus Pseudobironium Pic (Coleoptera: Staphylinidae: Scaphidiinae)" type="journal article" year="2013">Löbl & Tang, 2013</bibRefCitation>
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</taxonomicName>
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from which it differs notably by the parameres evenly narrow, except at their basis.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="discussion">
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<paragraph blockId="8.[332,1280,393,1495]" pageId="8" pageNumber="15">
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<emphasis bold="true" box="[332,443,1440,1464]" pageId="8" pageNumber="15">Remarks.</emphasis>
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The single available specimen has its forelegs broken, probably lost when collected.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |