150 lines
19 KiB
XML
150 lines
19 KiB
XML
<document id="48A3B9DBCFB0545E79E49B4A87BAF898" ID-CLB-Dataset="29018" ID-DOI="10.11646/zootaxa.4459.2.8" ID-GBIF-Dataset="f36314b8-a9a6-4bd2-8704-8923013c22b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458709" ID-ZooBank="7270DEAE-A08F-4015-8FE6-6E0D0DBD9C84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289409860" checkinUser="plazi" docAuthor="Kаtаev, Boris M. & Wrаse, Dаvid W." docDate="2018" docId="03952C24744AFFC2FF2CFB63FA12D9D3" docLanguage="en" docName="zootaxa.4459.2.8.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Proditomus mirus Schauberger 1934" docType="treatment" docVersion="7" lastPageNumber="363" masterDocId="FFAC545C7441FFCFFFBBFFD4FF8FD92B" masterDocTitle="Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India" masterLastPageNumber="368" masterPageNumber="350" pageNumber="361" updateTime="1698578541753" updateUser="ExternalLinkService">
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<mods:mods id="B340EDC808B44408137636C818B692FF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0D3ACEF14A16F405A576F465C4CED4E2">
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<mods:title id="89AA8B84C56A084F5CE5D6A69FF5D339">Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India</mods:title>
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<mods:name id="F0E0977299319BFD3A6470BFEA296863" type="personal">
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<mods:namePart id="E34107DBDF5754A7F965E50EA2ADB525">Kаtаev, Boris M.</mods:namePart>
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<mods:name id="46F6CEF8FADA91238A6EBFAA53ACF784" type="personal">
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<mods:roleTerm id="4871DC1E20304553496A394D5E3A365D">Author</mods:roleTerm>
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<mods:namePart id="E3C34E01F9B60727C7CD00BFB008C383">Wrаse, Dаvid W.</mods:namePart>
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<mods:typeOfResource id="1B998A026D442586CBD12065C72CE061">text</mods:typeOfResource>
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<mods:title id="C0B6833CB1C854F46758F823CF513F1A">Zootaxa</mods:title>
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<mods:part id="40D559C4D8A2A6FA93ED97EF7853078E">
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<mods:date id="DC502A06D7A8391688C72836AF4D2299">2018</mods:date>
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<mods:number id="BA4FCBB2544C6831C2216026E32D6C17">2018-08-15</mods:number>
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<mods:number id="A250174FC3C8ACE0B1B32A0A21B4E2D0">4459</mods:number>
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<mods:identifier id="34B9154C9E3DB2100DA1C321140731EC" type="DOI">10.11646/zootaxa.4459.2.8</mods:identifier>
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<treatment id="03952C24744AFFC2FF2CFB63FA12D9D3" ID-DOI="http://doi.org/10.5281/zenodo.6494758" ID-GBIF-Taxon="148692761" ID-Zenodo-Dep="6494758" LSID="urn:lsid:plazi:treatment:03952C24744AFFC2FF2CFB63FA12D9D3" httpUri="http://treatment.plazi.org/id/03952C24744AFFC2FF2CFB63FA12D9D3" lastPageId="13" lastPageNumber="363" pageId="11" pageNumber="361">
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<subSubSection id="C326CEB9744AFFC4FF2CFB63FEA8DDD8" pageId="11" pageNumber="361" type="nomenclature">
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<paragraph id="8B839D32744AFFC4FF2CFB63FDD1DDFA" blockId="11.[151,606,1207,1267]" box="[151,606,1207,1233]" pageId="11" pageNumber="361">
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<heading id="D0CB2A5E744AFFC4FF2CFB63FDD1DDFA" bold="true" box="[151,606,1207,1233]" fontSize="11" level="1" pageId="11" pageNumber="361" reason="1">
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<taxonomicName id="4C3CE6B1744AFFC4FF2CFB63FDD1DDFA" ID-CoL="4MQD6" authority="Schauberger, 1934" authorityName="Schauberger" authorityYear="1934" box="[151,606,1207,1233]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="mirus">
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<emphasis id="B9484120744AFFC4FF2CFB63FDD1DDFA" bold="true" box="[151,606,1207,1233]" pageId="11" pageNumber="361">
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<emphasis id="B9484120744AFFC4FF2CFB63FEE0DDFA" bold="true" box="[151,367,1207,1233]" italics="true" pageId="11" pageNumber="361">Proditomus mirus</emphasis>
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Schauberger, 1934
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B839D32744AFFC4FF2CFB0EFEA8DDD8" blockId="11.[151,606,1207,1267]" box="[151,295,1242,1267]" pageId="11" pageNumber="361">
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(
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<figureCitation id="130781B7744AFFC4FF24FB0EFE91DDD8" box="[159,286,1242,1267]" captionStart="FIGURES 25–29" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 25–29. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="11" pageNumber="361">Figs 25–29</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB9744AFFC4FF2CFAF6FDC0DC13" box="[151,591,1314,1336]" pageId="11" pageNumber="361" type="reference_group">
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<paragraph id="8B839D32744AFFC4FF2CFAF6FDC0DC13" blockId="11.[151,591,1314,1336]" box="[151,591,1314,1336]" pageId="11" pageNumber="361">
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<treatmentCitationGroup id="AB2CBA1C744AFFC4FF2CFAF6FDC0DC13" box="[151,591,1314,1336]" pageId="11" pageNumber="361">
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<taxonomicName id="4C3CE6B1744AFFC4FF2CFAF6FDC0DC13" ID-CoL="4MQD6" authority="Schauberger, 1934 b: 233" authorityName="Schauberger" authorityPageNumber="233" authorityYear="1934" box="[151,591,1314,1336]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="mirus">
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<emphasis id="B9484120744AFFC4FF2CFAF6FEC3DC13" box="[151,332,1314,1336]" italics="true" pageId="11" pageNumber="361">Proditomus mirus</emphasis>
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<treatmentCitation id="0A9DBB23744AFFC4FEE9FAF6FDC0DC13" author="" box="[338,591,1314,1336]" page="233" pageId="11" pageNumber="361">
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<bibRefCitation id="EFADE0C3744AFFC4FEE9FAF6FD90DC13" box="[338,543,1314,1336]" pageId="11" pageNumber="361" refId="ref13722" refString="Schauberger, E. (1934 b) Zur Kenntnis der palaarktischen Harpalinen (Col.). Stylops, 3, 232 - 240. https: // doi. org / 10.1111 / j. 1365 - 3113.1934. tb 01528. x" type="journal article">Schauberger, 1934b</bibRefCitation>
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: 233
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB9744AFFC4FF2CFAB0FBAEDF19" pageId="11" pageNumber="361" type="materials_examined">
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<paragraph id="8B839D32744AFFC4FF2CFAB0FBAEDF19" blockId="11.[151,1437,1380,2018]" pageId="11" pageNumber="361">
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<emphasis id="B9484120744AFFC4FF2CFAB0FE34DC56" bold="true" box="[151,443,1380,1405]" pageId="11" pageNumber="361">Type material examined</emphasis>
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. Holotype: ♀, labeled “Turkestan” [handwritten], “Type”, “
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<taxonomicName id="4C3CE6B1744AFFC4FBDDFAB0FF45DC8A" authority="Schaub., Loc." authorityName="Schaub., Loc." class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="mirus">Proditomus mirus Schaub., Loc.</taxonomicName>
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class.” [last two labels handwritten by Schauberger, the first one on red label], “Coll. Dr. E. Schauberger “[printed on white label], “
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<taxonomicName id="4C3CE6B1744AFFC4FE63FA79FC80DCED" authority="Schaub." authorityName="Schauberger" authorityYear="1934" box="[472,783,1453,1478]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="mirus">Proditomus mirus Schaub.</taxonomicName>
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[handwritten by Schauberger], “
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<taxonomicName id="4C3CE6B1744AFFC4FB2AFA79FAB4DCED" authority="Schaub." authorityName="Schauberger" authorityYear="1934" box="[1169,1339,1453,1478]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="361" phylum="Arthropoda" rank="species" species="mirus">mirus Schaub.</taxonomicName>
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” [partly handwritten, partly printed on white label], “Type” [handwritten by Schauberger on red label] (OÖLL). The specimen is in very poor condition with a part of appendages completely or partly missing. Mouthparts and female genitalia were already dissected out and glued to a separate card by Schauberger.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB9744AFFC3FF7CF9E9FD05D814" lastPageId="12" lastPageNumber="362" pageId="11" pageNumber="361" type="description">
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<paragraph id="8B839D32744AFFC3FF7CF9E9FD05D814" blockId="11.[151,1437,1380,2018]" lastBlockId="12.[151,1437,151,464]" lastPageId="12" lastPageNumber="362" pageId="11" pageNumber="361">
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<emphasis id="B9484120744AFFC4FF7CF9E9FEDDDF7D" bold="true" box="[199,338,1597,1622]" pageId="11" pageNumber="361">Description</emphasis>
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(female). With characteristics of the genus. Habitus as in
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<figureCitation id="130781B7744AFFC4FC61F9E9FBA3DF7E" box="[986,1068,1597,1622]" captionStart="FIGURES 25–29" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 25–29. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="11" pageNumber="361">Fig. 28</figureCitation>
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. Body robust and convex, length 14.0 mm from apex of mandibles to apex of elytra, and
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<quantity id="4CC430D7744AFFC4FCA6F9B5FC0CDF52" box="[797,899,1633,1657]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="11" pageNumber="361" unit="mm" value="13.2">13.2 mm</quantity>
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from apex of clypeus to apex of elytra. Black, including palpi, antennae and legs, only apices palpi paler. Head large (HWmax/PWmax = 1.08; HWmin/PWmax = 0.97), with small, very moderately convex eyes (HWmax/HWmin = 1.11), irregularly and coarsely punctate; distance between punctures longer than their diameter. Pronotum (PWmax/PL = 1.50; PWmax/PWmin = 1.76) very coarsely and sparsely punctate in anterio-medial portion and more densely and finely laterally and basally. Elytra oval (EL/EW = 1.42; EL/PL = 2.55; EW/PWmax = 1.20), widest at middle. Humerus rather widely rounded. Elytral striae thin, superficial, punctate, coarsely in basal half and finely apically, not reaching elytral base. Parascutellar striae long, basal pore puncture very small. Elytral epipleurae impunctate, distinctly bordered along inner margin. Intervals throughout flat, very finely and sparsely punctate; punctation uniform. Pro-, meso- and metathorax coarsely punctate, setation moderately long. Metepisternum (
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<figureCitation id="130781B7744AFFC4FC76F871FB90DE96" box="[973,1055,1957,1982]" captionStart="FIGURES 25–29" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 25–29. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="11" pageNumber="361">Fig. 29</figureCitation>
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) much longer than wide, strongly narrowed posteriorly. Hind wings fairly long but not folded (see remarks below). Аbdominal sternites finely and densely punctate, covered with short setae. Three penultimate abdominal sternites each with two long setae along posterior margin. Last visible sternite (in female) not modificated, with distal margin medially not enlarged and not plate-like. Female genitalia (
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<figureCitation id="130781B7744DFFC3FE67FF0BFDEED9D3" box="[476,609,223,248]" captionStart="FIGURES 25–29" captionStartId="12.[151,264,1946,1969]" captionTargetBox="[189,1404,537,1897]" captionTargetId="figure@12.[151,1436,517,1925]" captionTargetPageId="12" captionText="FIGURES 25–29. Proditomus mirus Schauberger, 1934 (holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27)." httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="12" pageNumber="362">Figs 26, 27</figureCitation>
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): laterotergite with numerous short setae on outer margin apically and at apex; basal and apical stylomeres wide and short, apical one rounded at apex, covered with large setigerous pore on inner margin and with numerous short setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="C326CEB9744DFFC3FF7CFE98FD53D8FB" pageId="12" pageNumber="362" type="distribution">
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<paragraph id="8B839D32744DFFC3FF7CFE98FD53D8FB" blockId="12.[151,1437,151,464]" pageId="12" pageNumber="362">
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<emphasis id="B9484120744DFFC3FF7CFE98FED6D84E" bold="true" box="[199,345,332,357]" pageId="12" pageNumber="362">Distribution</emphasis>
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. The species is only known from the
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<typeStatus id="54872390744DFFC3FD45FE98FCEFD84E" box="[766,864,332,357]" pageId="12" pageNumber="362" type="holotype">holotype</typeStatus>
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described from “Turkestan” without more detailed data. Since the territory of
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<collectingCountry id="F32BDDA2744DFFC3FE5EFEBBFD0DD8A3" box="[485,642,367,392]" name="Turkmenistan" pageId="12" pageNumber="362">Turkmenistan</collectingCountry>
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,
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<collectingCountry id="F32BDDA2744DFFC3FD2EFEBBFC99D8A3" box="[661,790,367,392]" name="Uzbekistan" pageId="12" pageNumber="362">Uzbekistan</collectingCountry>
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,
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<collectingCountry id="F32BDDA2744DFFC3FC9CFEBBFC17D8A3" box="[807,920,367,392]" name="Tajikistan" pageId="12" pageNumber="362">Tajikistan</collectingCountry>
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,
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<collectingCountry id="F32BDDA2744DFFC3FC11FEBBFBA2D8A3" box="[938,1069,367,392]" name="Kazakhstan" pageId="12" pageNumber="362">Kazakhstan</collectingCountry>
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and
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<collectingCountry id="F32BDDA2744DFFC3FBCAFEA4FB7DD8A3" box="[1137,1266,368,392]" name="Kyrgyzstan" pageId="12" pageNumber="362">Kyrgyzstan</collectingCountry>
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was carefully explored during last century, the range of this remarkable species may be restricted to northern
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<collectingCountry id="F32BDDA2744DFFC3FB5DFE41FA9CD887" box="[1254,1299,405,428]" name="Iran" pageId="12" pageNumber="362">Iran</collectingCountry>
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or northern
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<collectingCountry id="F32BDDA2744DFFC3FF2CFE63FEAFD8FB" box="[151,288,439,464]" name="Afghanistan" pageId="12" pageNumber="362">Аfghanistan</collectingCountry>
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as well as to western
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<collectingRegion id="49F853D0744DFFC3FDADFE63FDF7D8FB" box="[534,632,439,464]" country="China" name="Xinjiang Uygur" pageId="12" pageNumber="362">Xinjiang</collectingRegion>
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(
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<collectingCountry id="F32BDDA2744DFFC3FD3CFE63FD5FD8FB" box="[647,720,439,464]" name="China" pageId="12" pageNumber="362">China</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF43CDBA744DFFC3FF2CF84EFD82DEC5" httpUri="https://zenodo.org/record/1458723/files/figure.png" pageId="12" pageNumber="362" startId="12.[151,264,1946,1969]" targetBox="[189,1404,537,1897]" targetPageId="12">
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<paragraph id="8B839D32744DFFC3FF2CF84EFD82DEC5" blockId="12.[151,1436,1946,2030]" pageId="12" pageNumber="362">
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<emphasis id="B9484120744DFFC3FF2CF84EFEDCDE9B" bold="true" box="[151,339,1946,1969]" pageId="12" pageNumber="362">FIGURES 25–29.</emphasis>
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<taxonomicName id="4C3CE6B1744DFFC3FEE3F84FFD41DE9A" authority="Schauberger, 1934" authorityName="Schauberger" authorityYear="1934" box="[344,718,1947,1969]" class="Insecta" family="Carabidae" genus="Proditomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="362" phylum="Arthropoda" rank="species" species="mirus">
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<emphasis id="B9484120744DFFC3FEE3F84FFD82DE9A" box="[344,525,1947,1969]" italics="true" pageId="12" pageNumber="362">Proditomus mirus</emphasis>
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Schauberger, 1934
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</taxonomicName>
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(holotype). 25, labium; 26, right laterotergite and stylus; 27, stylus; 28, habitus; 29, left metepisternum. 25, 26, 29, ventral view; 27, lateral view; 28, dorsal view. Scale bars = 1.0 mm (A: Fig. 25, 29; C: Fig. 28), 0.5 mm (B: Figs 26, 27).
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</paragraph>
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</caption>
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<subSubSection id="C326CEB9744CFFC2FF7CFF43FA12D9D3" pageId="13" pageNumber="363" type="discussion">
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<paragraph id="8B839D32744CFFC2FF7CFF43FA12D9D3" blockId="13.[151,1437,151,248]" pageId="13" pageNumber="363">
|
||
<emphasis id="B9484120744CFFC2FF7CFF43FEB4D99B" bold="true" box="[199,315,151,176]" pageId="13" pageNumber="363">Remarks.</emphasis>
|
||
The only known specimen of this species is in very bad condition, so we do not dare to re-prepare it for investigation of the hind wings, fearing it will fall into pieces, so we can only cite the statement of
|
||
<bibRefCitation id="EFADE0C3744CFFC2FAAAFF68FF60D9D3" pageId="13" pageNumber="363" refId="ref13722" refString="Schauberger, E. (1934 b) Zur Kenntnis der palaarktischen Harpalinen (Col.). Stylops, 3, 232 - 240. https: // doi. org / 10.1111 / j. 1365 - 3113.1934. tb 01528. x" type="journal article">Schauberger (1934b)</bibRefCitation>
|
||
concerning this character. Аs the wings are not folded at apex, the species is seemingly incapable of flight.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |