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<document id="013C5AC2F52E7B625EEB6457ED0C5BC0" ID-CLB-Dataset="29018" ID-DOI="10.11646/zootaxa.4459.2.8" ID-GBIF-Dataset="f36314b8-a9a6-4bd2-8704-8923013c22b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458709" ID-ZooBank="7270DEAE-A08F-4015-8FE6-6E0D0DBD9C84" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289409860" checkinUser="plazi" docAuthor="Kаtаev, Boris M. &amp; Wrаse, Dаvid W." docDate="2018" docId="03952C247445FFCAFF2CF96AFA99DD8B" docLanguage="en" docName="zootaxa.4459.2.8.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Indocarterus Kаtаev &amp; Wrаse 2018, gen. n." docType="treatment" docVersion="7" lastPageNumber="355" masterDocId="FFAC545C7441FFCFFFBBFFD4FF8FD92B" masterDocTitle="Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India" masterLastPageNumber="368" masterPageNumber="350" pageNumber="354" updateTime="1698578541753" updateUser="ExternalLinkService">
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<mods:title id="68E0ECA41879574CE4BB48CE793B80DA">Notes on Ditomina, with description of two new genera and a new species (Coleoptera, Carabidae) from the Middle East and India</mods:title>
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<mods:namePart id="5356E794D132F18A39BCDA642B842D79">Kаtаev, Boris M.</mods:namePart>
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<mods:namePart id="DF87A6CFD6B91EFD10017AC58A753932">Wrаse, Dаvid W.</mods:namePart>
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<mods:date id="3621E35E45D3449248F196E2C0631510">2018</mods:date>
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<mods:number id="17F6A77B3C8B5E06A50E62520158B580">2018-08-15</mods:number>
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<treatment id="03952C247445FFCAFF2CF96AFA99DD8B" ID-DOI="http://doi.org/10.5281/zenodo.5980139" ID-GBIF-Taxon="148692762" ID-Zenodo-Dep="5980139" LSID="urn:lsid:plazi:treatment:03952C247445FFCAFF2CF96AFA99DD8B" httpUri="http://treatment.plazi.org/id/03952C247445FFCAFF2CF96AFA99DD8B" lastPageId="5" lastPageNumber="355" pageId="4" pageNumber="354">
<subSubSection id="C326CEB97445FFCBFF2CF96AFE03DFF3" box="[151,396,1726,1752]" pageId="4" pageNumber="354" type="nomenclature">
<paragraph id="8B839D327445FFCBFF2CF96AFE03DFF3" blockId="4.[151,396,1726,1752]" box="[151,396,1726,1752]" pageId="4" pageNumber="354">
<heading id="D0CB2A5E7445FFCBFF2CF96AFE03DFF3" bold="true" box="[151,396,1726,1752]" fontSize="11" level="1" pageId="4" pageNumber="354" reason="1">
<emphasis id="B94841207445FFCBFF2CF96AFE03DFF3" bold="true" box="[151,396,1726,1752]" pageId="4" pageNumber="354">
<taxonomicName id="4C3CE6B17445FFCBFF2CF96AFEBCDFF3" ID-CoL="8MKP8" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[151,307,1726,1752]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94841207445FFCBFF2CF96AFEBCDFF3" bold="true" box="[151,307,1726,1752]" italics="true" pageId="4" pageNumber="354">Indocarterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7445FFCBFE81F96AFE03DFF3" box="[314,396,1726,1752]" pageId="4" pageNumber="354" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C326CEB97445FFCBFF2CF8D2FDFFDE30" box="[151,624,1797,1819]" pageId="4" pageNumber="354" type="materials_examined">
<paragraph id="8B839D327445FFCBFF2CF8D2FDFFDE30" blockId="4.[151,624,1797,1819]" box="[151,624,1797,1819]" pageId="4" pageNumber="354">
Type species
<taxonomicName id="4C3CE6B17445FFCBFEA4F8D1FDBFDE30" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[287,560,1797,1819]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
<emphasis id="B94841207445FFCBFEA4F8D1FDBFDE30" box="[287,560,1797,1819]" italics="true" pageId="4" pageNumber="354">Indocarterus inexspectatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7445FFCBFD8DF8D2FDFFDE30" box="[566,624,1798,1819]" pageId="4" pageNumber="354" rank="species">
<emphasis id="B94841207445FFCBFD8DF8D2FDFFDE30" bold="true" box="[566,624,1798,1819]" pageId="4" pageNumber="354">sp. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C326CEB97445FFCBFF2CF893FACFDEDB" pageId="4" pageNumber="354" type="diagnosis">
<paragraph id="8B839D327445FFCBFF2CF893FACFDEDB" blockId="4.[151,1437,1863,2033]" pageId="4" pageNumber="354">
<emphasis id="B94841207445FFCBFF2CF893FE85DE4B" bold="true" box="[151,266,1863,1888]" pageId="4" pageNumber="354">Diagnosis</emphasis>
. Most similar in combination of characters (see key) to the genus
<taxonomicName id="4C3CE6B17445FFCBFB84F89DFAE9DE4B" authority="Stichel, 1923" authorityName="Stichel" authorityYear="1923" box="[1087,1382,1863,1888]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="genus">
<emphasis id="B94841207445FFCBFB84F89DFB34DE4B" box="[1087,1211,1865,1888]" italics="true" pageId="4" pageNumber="354">Eocarterus</emphasis>
<bibRefCitation id="EFADE0C37445FFCBFB72F893FAE9DE4B" box="[1225,1382,1863,1888]" pageId="4" pageNumber="354" refId="ref14177" refString="Stichel, W. (1923) Zur Phylogenesis eines geologisch jungen Formenkreises der Kafer, der Ditominen (Carab. Harpal.). Zeitschrift fur wissenschaftliche Insektenbiologie, 18, 41 - 50 + 81 - 100 + 145 - 162 + 209 - 242." type="journal article">Stichel, 1923</bibRefCitation>
</taxonomicName>
, but distinguished by having head relatively wider, mentum without median tooth, labrum more deeply concave anteriorly, pronotum not cordate, its sides largely rounded, with very short and shallow sinuation just before basal angles. From the species of subgenus
<emphasis id="B94841207445FFCBFD86F861FD5EDEE7" box="[573,721,1973,1996]" italics="true" pageId="4" pageNumber="354">Baeticocarus</emphasis>
Jeanne, 1971, the member of the new genus additionally differs in complete basal elytral border (
<taxonomicName id="4C3CE6B17445FFCBFDB1F80DFD09DEDB" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[522,646,2009,2032]" class="Insecta" family="Carabidae" genus="Eocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B94841207445FFCBFDB1F80DFD09DEDB" box="[522,646,2009,2032]" italics="true" pageId="4" pageNumber="354">Eocarterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7445FFCBFD36F80DFD47DEDB" box="[653,712,2009,2032]" pageId="4" pageNumber="354" sensu="stricto">s. str.</taxonomicNameLabel>
has incomplete basal border as in
<taxonomicName id="4C3CE6B17445FFCBFBF1F80CFB54DEDA" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[1098,1243,2008,2033]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="354" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94841207445FFCBFBF1F80CFB54DEDA" box="[1098,1243,2008,2033]" italics="true" pageId="4" pageNumber="354">Indocarterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7445FFCBFB5AF80CFAA0DEDA" box="[1249,1327,2008,2033]" pageId="4" pageNumber="354" rank="genus">
<emphasis id="B94841207445FFCBFB5AF80CFAA0DEDA" bold="true" box="[1249,1327,2008,2033]" pageId="4" pageNumber="354">gen. n.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C326CEB97444FFCAFF7CFF43FDF3DAAB" pageId="5" pageNumber="355" type="description">
<paragraph id="8B839D327444FFCAFF7CFF43FBB1D984" blockId="5.[151,1437,151,1184]" box="[199,1086,151,176]" pageId="5" pageNumber="355">
<emphasis id="B94841207444FFCAFF7CFF43FEDED99B" bold="true" box="[199,337,151,176]" pageId="5" pageNumber="355">Description</emphasis>
. Dorsum (
<figureCitation id="130781B77444FFCAFE73FF43FDA3D99B" box="[456,556,151,176]" captionStart="FIGURES 5, 6" captionStartId="6.[151,264,1392,1415]" captionTargetBox="[192,1402,451,1356]" captionTargetId="figure@6.[163,1427,432,1382]" captionTargetPageId="6" captionText="FIGURES 5, 6. Indocarterus inexspectatus sp. n., habitus. 5, male holotype; 6, female paratype. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/1458713/files/figure.png" pageId="5" pageNumber="355">Figs 5, 6</figureCitation>
) and venter rather densely punctate and setose.
</paragraph>
<paragraph id="8B839D327444FFCAFF7CFF68FE69D8FB" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
Head large, slightly narrower than pronotum, constricted behind eyes, with only one long seta near inner margin of eyes on each side. Lateral margin of frons (
<figureCitation id="130781B77444FFCAFCA4FF0BFCE7D9D3" box="[799,872,223,248]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="5" pageNumber="355">Fig. 7</figureCitation>
) above insertion of antennae angularly rounded anteriorly. Labrum deeply concave anteriorly. Clypeus along anterior margin without convex smooth bead, rather deeply arcuately emarginate, and membranous base of labrum exposed. Genae wide. Mentum (
<figureCitation id="130781B77444FFCAFB5AFEF3FABAD86B" box="[1249,1333,295,320]" captionStart="FIGURES 10–17" captionStartId="8.[151,264,1206,1229]" captionTargetBox="[170,1428,620,1169]" captionTargetId="figure@8.[151,1436,589,1185]" captionTargetPageId="8" captionText="FIGURES 10–17. Indocarterus inexspectatus sp. n. (paratypes). 10, labium; 11, apex of median lobe; 12, 13, median lobe; 14, left paramere, inner surface; 15, left paramere, inner surface; 16, stylus, 17, right laterotergite and stylus. 10, 17, ventral view; 11, 13, dorsal view; 12, 14–16, lateral view. Scale bars: A = 1.0 mm (Fig. 10), B = 0.5 mm (Figs 11–16)." httpUri="https://zenodo.org/record/1458717/files/figure.png" pageId="5" pageNumber="355">Fig. 10</figureCitation>
) without median tooth, separated from submentum by complete transverse suture. Ligular sclerite not widened at apex, without ventral plate, and with several rather long, visible in ventral view, dorsal setae in addition to two ventral distal setae. Paraglossae rather narrow, glabrous at margins. Terminal maxillary and labial palpomeres more or less cylindrical, narrowed distally.
</paragraph>
<paragraph id="8B839D327444FFCAFF7CFE09FD6DDB33" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
Pronotum (
<figureCitation id="130781B77444FFCAFEF3FE08FE01D8DF" box="[328,398,476,501]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="5" pageNumber="355">Fig. 8</figureCitation>
) not cordate, its sides not serrate, largely rounded, with very short and shallow prebasal sinuation sharply delimited anteriorly (see remarks).
</paragraph>
<paragraph id="8B839D327444FFCAFF7CFDF0FB5ADB4B" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">Mesothorax and extreme base of elytra narrowed forming a very short neck-like constriction with scutellum above. Elytra not serrate at sides, basal border complete, strongly bent and raised near scutellum.</paragraph>
<paragraph id="8B839D327444FFCAFF7CFDB8FD9ADB83" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">Three penultimate abdominal sternites, in addition to short and dense pubescence, only with two obligatory long setae along posterior margin.</paragraph>
<paragraph id="8B839D327444FFCAFF7CFD61FDF3DAAB" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
Metacoxa without posteromedial setigerous pore. Profemur not denticulate along inner margin. Metatrochanter short, about a third as long as metafemur, rounded at apex. Protibia without long, finger-like acute apical process, with three or four preapical spines on outer margin, densely setose on ventral surface. Tarsi densely setose dorsally. Protarsus and mesotarsus of male not dilated and without ventral adhesive vestiture. Protarsomeres 1, 2 and 3 (sometimes also 4) each of both sexes with a short acute spine in middle of ventro-apical margin (
<figureCitation id="130781B77444FFCAFAF8FC90FA01DA77" box="[1347,1422,836,861]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="5" pageNumber="355">Fig. 9</figureCitation>
). Metatarsomere 1 scarcely shorter than 2+3.
</paragraph>
</subSubSection>
<subSubSection id="C326CEB97444FFCAFF7CFC58FECEDD3B" pageId="5" pageNumber="355" type="etymology">
<paragraph id="8B839D327444FFCAFF7CFC58FEC5DAEC" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
<emphasis id="B94841207444FFCAFF7CFC58FEC9DA8E" bold="true" box="[199,326,908,933]" pageId="5" pageNumber="355">Etymology</emphasis>
. The genus name is a combination of the name of the country
<collectingCountry id="F32BDDA27444FFCAFBA2FC58FBDBDA8E" box="[1049,1108,908,933]" name="India" pageId="5" pageNumber="355">
<emphasis id="B94841207444FFCAFBA2FC58FBDBDA8E" box="[1049,1108,908,933]" italics="true" pageId="5" pageNumber="355">India</emphasis>
</collectingCountry>
and the name of the carabid genus
<taxonomicName id="4C3CE6B17444FFCAFF64FC64FECCDAEC" authorityName="Dejean" authorityYear="1830" box="[223,323,944,967]" class="Insecta" family="Carabidae" genus="Carterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="B94841207444FFCAFF64FC64FECCDAEC" box="[223,323,944,967]" italics="true" pageId="5" pageNumber="355">Carterus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B839D327444FFCAFF7CFC00FECEDD3B" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
<emphasis id="B94841207444FFCAFF7CFC00FDA2DAC6" bold="true" box="[199,557,980,1005]" pageId="5" pageNumber="355">Composition and distribution</emphasis>
. The genus includes only one species,
<taxonomicName id="4C3CE6B17444FFCAFC4DFC01FB27DAC7" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[1014,1192,981,1004]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="species" species="inexspectatus" status="sp. nov.">
<emphasis id="B94841207444FFCAFC4DFC01FB27DAC7" box="[1014,1192,981,1004]" italics="true" pageId="5" pageNumber="355">I. inexspectatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7444FFCAFB09FC00FB7BDAC6" box="[1202,1268,980,1005]" pageId="5" pageNumber="355" rank="species">
<emphasis id="B94841207444FFCAFB09FC00FB7BDAC6" bold="true" box="[1202,1268,980,1005]" pageId="5" pageNumber="355">sp. n.</emphasis>
</taxonomicNameLabel>
, from western India (
<figureCitation id="130781B77444FFCAFF5BFC23FEBBDD3B" box="[224,308,1015,1040]" captionStart="FIGURE 24" captionStartId="11.[151,250,1105,1127]" captionTargetBox="[152,1436,195,1083]" captionTargetId="figure@11.[151,1436,195,1083]" captionTargetPageId="11" captionText="FIGURE 24. Geographical range of the subtribe Ditomina." httpUri="https://zenodo.org/record/1458721/files/figure.png" pageId="5" pageNumber="355">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C326CEB97444FFCAFF7CFBC8FA99DD8B" pageId="5" pageNumber="355" type="discussion">
<paragraph id="8B839D327444FFCAFF7CFBC8FA99DD8B" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="355">
<emphasis id="B94841207444FFCAFF7CFBC8FEBBDD1E" bold="true" box="[199,308,1052,1077]" pageId="5" pageNumber="355">Remarks</emphasis>
. It is possible that the short and shallow prebasal sinuation of pronotal sides (
<figureCitation id="130781B77444FFCAFB01FBC8FB70DD1F" box="[1210,1279,1052,1077]" captionStart="FIGURES 7–9" captionStartId="7.[151,264,1867,1890]" captionTargetBox="[234,1316,203,1823]" captionTargetId="figure@7.[151,1436,193,1846]" captionTargetPageId="7" captionText="FIGURES 7–9. Indocarterus inexspectatus sp. n. (paratypes: 7, 8 female; 9, male). 7, anterior part of head; 8, pronotum; 9, protarsus and apical part of protibia. 7, 8, dorsal view; 9, lateral view. Scales bars = 1.0 mm (A, C: Figs 7, 9; B: Fig. 8)." httpUri="https://zenodo.org/record/1458715/files/figure.png" pageId="5" pageNumber="355">Fig. 8</figureCitation>
) is actually a lateral part of pronotal base as in the members of the subgenus
<taxonomicName id="4C3CE6B17444FFCAFC34FBEBFB4EDD73" authority="Stephens, 1825" authorityName="Stephens" authorityYear="1825" box="[911,1217,1087,1112]" class="Insecta" family="Carabidae" genus="Philochtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="B94841207444FFCAFC34FBEBFB88DD73" box="[911,1031,1087,1112]" italics="true" pageId="5" pageNumber="355">Philochtus</emphasis>
Stephens, 1825
</taxonomicName>
(genus
<taxonomicName id="4C3CE6B17444FFCAFA98FBEBFECDDD57" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Bembidion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="B94841207444FFCAFA98FBEBFA12DD73" box="[1315,1437,1087,1112]" italics="true" pageId="5" pageNumber="355">Bembidion</emphasis>
Latreille, 1802
</taxonomicName>
). In this case the pronotal sides of
<taxonomicName id="4C3CE6B17444FFCAFD6EFBB0FCE9DD56" authority="Kаtаev &amp; Wrаse, 2018" authorityName="Kаtаev &amp; Wrаse" authorityYear="2018" box="[725,870,1124,1149]" class="Insecta" family="Carabidae" genus="Indocarterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="355" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B94841207444FFCAFD6EFBB0FCE9DD56" box="[725,870,1124,1149]" italics="true" pageId="5" pageNumber="355">Indocarterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BFC5B7444FFCAFCD4FBB0FC4FDD56" box="[879,960,1124,1149]" pageId="5" pageNumber="355" rank="genus">
<emphasis id="B94841207444FFCAFCD4FBB0FC4FDD56" bold="true" box="[879,960,1124,1149]" pageId="5" pageNumber="355">gen. n.</emphasis>
</taxonomicNameLabel>
are rounded to basal angles, but the base inside them is sinuate and the basal pronotal angles are correspondingly shifted from the base anteriad.
</paragraph>
</subSubSection>
</treatment>
</document>
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